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<document id="065E8C0326F1EEA0A8DA3C3DD269EC3B" ID-DOI="10.11646/phytotaxa.235.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13631327" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725310362704" checkinUser="felipe" docAuthor="Mongkolsuk, Pachara, Meesim, Sanya, Poengsungnoen, Vasun, Buaruang, Kawinnat, Schumm, Felix &amp; Kalb, Klaus" docDate="2015" docId="03EF7631FFDB6A46FF584B9EFD99FB51" docLanguage="en" docName="phytotaxa.235.1.1.pdf" docOrigin="Phytotaxa 235 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.235.1.1" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.4:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="4" docTitle="Leucodermia arsenei Kalb 2015, comb. nov." docType="treatment" docVersion="2" lastPageNumber="450" masterDocId="FFD60E49FFFF6A62FFCF4960FF8EFF81" masterDocTitle="The lichen family Physciaceae in Thailand-II. Contributions to the genus Heterodermia sensu lato" masterLastPageNumber="450" masterPageNumber="448" pageNumber="448" updateTime="1725311109013" updateUser="ExternalLinkService">
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<mods:title id="3C51758962B40644BD7923901135F0FD">The lichen family Physciaceae in Thailand-II. Contributions to the genus Heterodermia sensu lato</mods:title>
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<mods:namePart id="E29C4317CE256BECD4F6EAA2CB8EAAD9">Mongkolsuk, Pachara</mods:namePart>
<mods:affiliation id="E6F4CEDB263A842A32423F93B3791FEC">Lichen Research Unit, Biology Department, Faculty of Science, Ramkhamhaeng University, Ramkhamhaeng 24 road, Bangkok, 10240 Thailand.</mods:affiliation>
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<mods:namePart id="EC377D48E6184EAA49A72BECEF5997CB">Meesim, Sanya</mods:namePart>
<mods:affiliation id="3D65694B4A6D323545B1711FD5D29C0C">Lichen Research Unit, Biology Department, Faculty of Science, Ramkhamhaeng University, Ramkhamhaeng 24 road, Bangkok, 10240 Thailand.</mods:affiliation>
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<mods:namePart id="B274B99F3987153570D4DEA8DCDA85B0">Poengsungnoen, Vasun</mods:namePart>
<mods:affiliation id="03708825EB6B2C74E090306EB6EB3852">Lichen Research Unit, Biology Department, Faculty of Science, Ramkhamhaeng University, Ramkhamhaeng 24 road, Bangkok, 10240 Thailand.</mods:affiliation>
</mods:name>
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<mods:namePart id="736AA48B4E4560FF402090BBB73836F1">Buaruang, Kawinnat</mods:namePart>
<mods:affiliation id="73451C2E8C18241C5CE519E3C7974193">Lichen Research Unit, Biology Department, Faculty of Science, Ramkhamhaeng University, Ramkhamhaeng 24 road, Bangkok, 10240 Thailand.</mods:affiliation>
</mods:name>
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<mods:namePart id="FED3DC159ECA6920AF7BDEAFD541CCFE">Schumm, Felix</mods:namePart>
<mods:affiliation id="A6D0C68DFCA4F66FC26FA63731D64FE7">Mozartstrasse 9, D- 73117 Wangen, Germany.</mods:affiliation>
</mods:name>
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<mods:namePart id="2C4B67A6572505432CF0EC251069E1B5">Kalb, Klaus</mods:namePart>
<mods:affiliation id="B25E224040ED6AAB72F9F9C6D9BC4AE1">Lichen Research Unit, Biology Department, Faculty of Science, Ramkhamhaeng University, Ramkhamhaeng 24 road, Bangkok, 10240 Thailand. &amp; Lichenologisches Institut Neumarkt, Im Tal 12, D- 92318 Neumarkt, Germany. Email: klaus. kalb @ arcor. de &amp; Lichen Research Unit, Biology Department, Faculty of Science, Ramkhamhaeng University, Ramkhamhaeng 24 road, Bangkok, 10240 Thailand. &amp; University of Regensburg, Institute of Plant Sciences, Universitätsstraβe 31, D- 39040 Regensburg, Germany Corresponding author-email: pm-tamrapap @ ru. ac. th &amp; Lichen Research Unit, Biology Department, Faculty of Science, Ramkhamhaeng University, Ramkhamhaeng 24 road, Bangkok, 10240 Thailand.</mods:affiliation>
<mods:nameIdentifier id="23E3999BDF90C02A5A14961BF5C69951" type="email">klaus.kalb@arcor.de</mods:nameIdentifier>
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<mods:title id="3283552217603B57D0478B1B99866924">Phytotaxa</mods:title>
</mods:titleInfo>
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<mods:date id="F12DA93C79F861BD030C2145118EA2EC">2015</mods:date>
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<mods:number id="B3BA47CF761006E4983DA894DE85980D">2015-11-20</mods:number>
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<mods:number id="852AB3132DCFDCC0043C56E01A5F30FD">235</mods:number>
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<mods:url id="7BB278AE9D10F6F5445E05311A9CED7C">http://dx.doi.org/10.11646/phytotaxa.235.1.1</mods:url>
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<mods:classification id="C81D2874D91B90EEA85F46AF174C61A4">journal article</mods:classification>
<mods:identifier id="F7BEDF9C809FC54CC25F0C48C73E293E" type="DOI">10.11646/phytotaxa.235.1.1</mods:identifier>
<mods:identifier id="A6E5A861E862B4A0097B9030CCE14148" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="FDB5FA712A8BDA010A18F1815F557D23" type="Zenodo-Dep">13631327</mods:identifier>
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<treatment id="03EF7631FFDB6A46FF584B9EFD99FB51" ID-DOI="http://doi.org/10.5281/zenodo.13631363" ID-Zenodo-Dep="13631363" LSID="urn:lsid:plazi:treatment:03EF7631FFDB6A46FF584B9EFD99FB51" httpUri="http://treatment.plazi.org/id/03EF7631FFDB6A46FF584B9EFD99FB51" pageId="36">
<subSubSection id="C35C94ACFFDB6A46FF584B9EFE12FCBA" pageId="36" type="nomenclature">
<paragraph id="8BF9C727FFDB6A46FF584B9EFD33FC97" blockId="36.[151,701,766,827]" box="[151,701,766,791]" pageId="36">
<heading id="D0B1704BFFDB6A46FF584B9EFD33FC97" bold="true" box="[151,701,766,791]" fontSize="10" level="2" pageId="36" reason="2">
<emphasis id="B9321B35FFDB6A46FF584B9EFD33FC97" bold="true" box="[151,701,766,791]" pageId="36">
<taxonomicName id="4C46BCA4FFDB6A46FF584B9EFDB9FC96" ID-CoL="724Y8" authority="(Kurok.) Kalb" authorityName="Kalb" authorityYear="2015" baseAuthorityName="Kurok." box="[151,567,766,791]" class="Lecanoromycetes" family="Physciaceae" genus="Leucodermia" kingdom="Fungi" order="Caliciales" pageId="36" phylum="Ascomycota" rank="species" species="arsenei" status="comb. nov.">
<emphasis id="B9321B35FFDB6A46FF584B9EFE08FC96" bold="true" box="[151,390,766,791]" italics="true" pageId="36">Leucodermia arsenei</emphasis>
(Kurok.) Kalb
</taxonomicName>
,
<emphasis id="B9321B35FFDB6A46FD8D4B9EFD33FC97" bold="true" box="[578,701,766,791]" italics="true" pageId="36">
<taxonomicNameLabel id="A201A64EFFDB6A46FD8D4B9EFD33FC97" box="[578,701,766,791]" pageId="36" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF9C727FFDB6A46FF584A42FE12FCBA" blockId="36.[151,701,766,827]" box="[151,412,802,827]" pageId="36">Mycobank MB 813830</paragraph>
</subSubSection>
<subSubSection id="C35C94ACFFDB6A46FF584A0AFB3DFC56" pageId="36" type="reference_group">
<paragraph id="8BF9C727FFDB6A46FF584A0AFB03FC1D" blockId="36.[151,1437,874,1232]" pageId="36">
<materialsCitation id="3B2ECD7AFFDB6A46FF584A0AFBDCFCFE" box="[151,1106,874,895]" country="Mexico" location="Puebla" pageId="36" specimenCount="1" stateProvince="Puebla">
Basionym:
<taxonomicName id="4C46BCA4FFDB6A46FECB4A0AFDC9FCFE" ID-CoL="66KJH" authority="Kurok., Beih." authorityName="Kurok., Beih." box="[260,583,874,895]" class="Lecanoromycetes" family="Physciaceae" genus="Anaptychia" kingdom="Fungi" order="Caliciales" pageId="36" phylum="Ascomycota" rank="species" species="arsenei">
<emphasis id="B9321B35FFDB6A46FECB4A0AFE30FCFE" box="[260,446,874,895]" italics="true" pageId="36">Anaptychia arsenei</emphasis>
Kurok., Beih.
</taxonomicName>
<bibCitation id="5E6CCF4BFFDB6A46FD844A0AFCEFFCFE" box="[587,865,874,895]" journal="Nova Hedwigia" page="89" pageId="36" score="202.82472018338322" volume="6" year="1962">Nova Hedwigia 6: 89 (1962)</bibCitation>
. Type:—
<collectingCountry id="F35187B7FFDB6A46FC774A0AFB8AFCFE" box="[952,1028,874,895]" name="Mexico" pageId="36">Mexico</collectingCountry>
.
<collectingRegion id="498209C5FFDB6A46FBC24A0AFBDCFCFE" box="[1037,1106,874,895]" country="Mexico" name="Puebla" pageId="36">Puebla</collectingRegion>
</materialsCitation>
:
<materialsCitation id="3B2ECD7AFFDB6A46FB954A0AFB87FC1D" country="Estonia" elevation="2400" location="Estonia" pageId="36" specimenCount="1" typeStatus="holotype">
Esperanza,
<quantity id="4CBE6AC2FFDB6A46FB064A0AFA9DFCFE" box="[1225,1299,874,895]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="36" unit="m" value="2400.0">
<elevation id="006B2014FFDB6A46FB064A0AFA9DFCFE" box="[1225,1299,874,895]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="36" unit="m" value="2400.0">2400 m</elevation>
</quantity>
, 1907,
<emphasis id="B9321B35FFDB6A46FA964A0AFF79FC1D" italics="true" pageId="36">Arsène 8089</emphasis>
(
<typeStatus id="54FD7985FFDB6A46FECC4AE7FED6FC1D" box="[259,344,903,924]" pageId="36" type="holotype">holotype</typeStatus>
US, n.v.).
<treatmentCitation id="0AE7E136FFDB6A46FE704AE7FB87FC1D" author="Trass" box="[447,1033,903,924]" journal="Folia Cryptog. Estonia" page="7" pageId="36" score="0.06320979317596026" volume="29" year="1992">
<taxonomicName id="4C46BCA4FFDB6A46FE704AE7FC33FC1D" ID-CoL="3L95H" authority="(Kurok.) Trass, Folia Cryptog." authorityName="Trass, Folia Cryptog." baseAuthorityName="Kurok." box="[447,957,903,924]" class="Lecanoromycetes" family="Physciaceae" genus="Heterodermia" kingdom="Fungi" order="Caliciales" pageId="36" phylum="Ascomycota" rank="species" species="arsenei">
<emphasis id="B9321B35FFDB6A46FE704AE7FD1FFC1D" box="[447,657,903,924]" italics="true" pageId="36">Heterodermia arsenei</emphasis>
(Kurok.) Trass, Folia Cryptog.
</taxonomicName>
<collectingCountry id="F35187B7FFDB6A46FC0E4AE7FB87FC1D" box="[961,1033,903,924]" name="Estonia" pageId="36">Estonia</collectingCountry>
</treatmentCitation>
</materialsCitation>
29: 7 (1992).
</paragraph>
<paragraph id="8BF9C727FFDB6A46FF584AC5FB3DFC56" blockId="36.[151,1437,874,1232]" pageId="36">Chemistry: Cortex K+ yellow, C, KC, P+ yellow; medulla K+ yellow then red, C, P+ yellow-orange; containing atranorin (major), zeorin (major), salazinic acid (major), consalazinic acid (major), 16β-acetoxyhopane-6α,22-diol (minor).</paragraph>
</subSubSection>
<subSubSection id="C35C94ACFFDB6A46FF584ABFFD99FB51" pageId="36" type="discussion">
<paragraph id="8BF9C727FFDB6A46FF584ABFFC57FBE4" blockId="36.[151,1437,874,1232]" pageId="36">
<emphasis id="B9321B35FFDB6A46FF584ABFFE8AFC79" bold="true" box="[151,260,991,1016]" pageId="36">Remarks</emphasis>
:
<emphasis id="B9321B35FFDB6A46FEC34ABFFEA6FC79" bold="true" box="[268,296,991,1016]" pageId="36"></emphasis>
Although we have not seen the
<typeStatus id="54FD7985FFDB6A46FD424A80FD33FC79" box="[653,701,992,1016]" pageId="36">type</typeStatus>
material, we have investigated several collections from
<collectingRegion id="498209C5FFDB6A46FAF44ABFFA16FC79" box="[1339,1432,991,1016]" country="Mexico" name="Chiapas" pageId="36">Chiapas</collectingRegion>
, SE of
<collectingRegion id="498209C5FFDB6A46FF2A4D64FEB9FB9C" box="[229,311,1028,1053]" country="Mexico" name="Puebla" pageId="36">Puebla</collectingRegion>
which matched the description given by
<bibRefCitation id="EFD7BAD6FFDB6A46FCF44D64FB99FB9C" author="Kurokawa, S." box="[827,1047,1028,1053]" pageId="36" pagination="1 - 115" refId="ref53593" refString="Kurokawa, S. (1962) A monograph of the genus Anaptychia. Beihefte zur Nova Hedwigia 6: 1 - 115, 9 plates." type="book chapter" year="1962">Kurokawa (1962)</bibRefCitation>
. The species was placed in ser.
<emphasis id="B9321B35FFDB6A46FF584D47FEAEFBC1" box="[151,288,1063,1088]" italics="true" pageId="36">Podocarpae</emphasis>
by
<bibRefCitation id="EFD7BAD6FFDB6A46FE854D47FD9DFBC1" author="Kurokawa, S." box="[330,531,1063,1088]" pageId="36" pagination="1 - 115" refId="ref53593" refString="Kurokawa, S. (1962) A monograph of the genus Anaptychia. Beihefte zur Nova Hedwigia 6: 1 - 115, 9 plates." type="book chapter" year="1962">Kurokawa (1962)</bibRefCitation>
, but it seems more closely related to
<emphasis id="B9321B35FFDB6A46FC7B4D47FB79FBC1" box="[948,1271,1063,1088]" italics="true" pageId="36">
<taxonomicName id="4C46BCA4FFDB6A46FC7B4D47FB7DFBC1" box="[948,1267,1063,1088]" class="Lecanoromycetes" family="Physciaceae" genus="Leucodermia" kingdom="Fungi" order="Caliciales" pageId="36" phylum="Ascomycota" rank="species" species="appalachensis">Leucodermia appalachensis</taxonomicName>
.
</emphasis>
It differs from
<taxonomicName id="4C46BCA4FFDB6A46FF584D2DFEDAFBE4" box="[151,340,1100,1125]" class="Lecanoromycetes" family="Physciaceae" genus="Leucodermia" kingdom="Fungi" order="Caliciales" pageId="36" phylum="Ascomycota" rank="species" species="appalachensis">
<emphasis id="B9321B35FFDB6A46FF584D2DFEDAFBE4" box="[151,340,1100,1125]" italics="true" pageId="36">L. appalachensis</emphasis>
</taxonomicName>
in having broader lobes and by containing salazinic acid.
</paragraph>
<paragraph id="8BF9C727FFDB6A46FF094D0FFD99FB51" blockId="36.[151,1437,874,1232]" pageId="36">
<emphasis id="B9321B35FFDB6A46FF094D0FFE36FB09" bold="true" box="[198,440,1135,1160]" pageId="36">Specimen examined</emphasis>
:—
<collectingCountry id="F35187B7FFDB6A46FE164D0FFDC0FB09" box="[473,590,1135,1160]" name="Mexico" pageId="36">MEXICO</collectingCountry>
.
<collectingRegion id="498209C5FFDB6A46FD934D0FFD35FB09" box="[604,699,1135,1160]" country="Mexico" name="Chiapas" pageId="36">Chiapas</collectingRegion>
: ca.
<quantity id="4CBE6AC2FFDB6A46FD3A4D0FFCCEFB09" box="[757,832,1135,1160]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="36" unit="km" value="25.0">25 km</quantity>
NW of Comitan de Dominguez, in a dark and dry forest, dominated by
<taxonomicName id="4C46BCA4FFDB6A46FE494DF4FDABFB2A" box="[390,549,1172,1195]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="36" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B9321B35FFDB6A46FE494DF4FE6AFB2A" box="[390,484,1172,1195]" italics="true" pageId="36">Quercus</emphasis>
spec.
</taxonomicName>
and
<taxonomicName id="4C46BCA4FFDB6A46FD934DF5FD51FB2A" box="[604,735,1172,1196]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="36" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B9321B35FFDB6A46FD934DF5FD12FB2D" box="[604,668,1173,1196]" italics="true" pageId="36">Pinus</emphasis>
spec.
</taxonomicName>
,
<quantity id="4CBE6AC2FFDB6A46FD254DF4FCCCFB2A" box="[746,834,1172,1197]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="36" unit="m" value="2150.0">2150 m</quantity>
,
<geoCoordinate id="EE72A1E0FFDB6A46FC814DF4FC37FB2A" box="[846,953,1172,1197]" degrees="16" direction="north" minutes="27" orientation="latitude" pageId="36" precision="925" value="16.45">16°27 N</geoCoordinate>
,
<geoCoordinate id="EE72A1E0FFDB6A46FC0A4DF4FBBAFB2A" box="[965,1076,1172,1197]" degrees="92" direction="west" minutes="19" orientation="longitude" pageId="36" precision="925" value="-92.316666">92°19 W</geoCoordinate>
,
<date id="FFF8E1E7FFDB6A46FB8F4DF4FB73FB2C" box="[1088,1277,1172,1197]" pageId="36" value="1979-01-25">25 January 1979</date>
,
<emphasis id="B9321B35FFDB6A46FAC74DF5FF51FB51" italics="true" pageId="36">K. Kalb &amp; G. Plöbst</emphasis>
(herb. Kalb 39870, 39872).
</paragraph>
</subSubSection>
</treatment>
</document>