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<document id="1B6D416AD0CB07B0921531740ECE10ED" ID-DOI="10.1649/0010-065X-67.2.97" ID-Zenodo-Dep="5368835" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1630261439700" checkinUser="carolina" docAuthor="Otto, Robert L." docDate="2013" docId="03D087AAA007FFE6FDD151B7FE0E857C" docLanguage="en" docName="0010-065X-67.2.97.pdf" docOrigin="The Coleopterists Bulletin 67 (2)" docSource="http://dx.doi.org/10.1649/0010-065x-67.2.97" docStyle="DocumentStyle:A0A70CFE30726088A421CCB226FF86C7.4:TheColeopBull.2010-2016.journal_article.1cover" docStyleId="A0A70CFE30726088A421CCB226FF86C7" docStyleName="TheColeopBull.2010-2016.journal_article.1cover" docStyleVersion="4" docTitle="Onichodon rugicollis" docType="treatment" docVersion="3" lastPageNumber="106" masterDocId="FFE9FFD2A000FFEFFD275368FFDD837B" masterDocTitle="Eucnemid Larvae of the Nearctic Region. Part III: Mature Larval Descriptions for Three Species ofOnichodonNewman, 1838 (Coleoptera: Eucnemidae: Macraulacinae: Macraulacini), with Notes on Their Biology" masterLastPageNumber="106" masterPageNumber="97" pageNumber="97" updateTime="1699201950542" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="35EA1C5FF34FBB32C356F2B997A714D6">Eucnemid Larvae of the Nearctic Region. Part III: Mature Larval Descriptions for Three Species ofOnichodonNewman, 1838 (Coleoptera: Eucnemidae: Macraulacinae: Macraulacini), with Notes on Their Biology</mods:title>
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<mods:name id="BE78A32EB1555A1B954E1C9EB7C3C947" type="personal">
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<mods:namePart id="786D1CA66E502CAD9004F716F46DA3F6">Otto, Robert L.</mods:namePart>
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<mods:title id="E2DD80786F06EF15EEBAE3662E6911BA">The Coleopterists Bulletin</mods:title>
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<mods:part id="7FE0020CD7762845F836B34661551CC1">
<mods:date id="FB5ACE1714AEC994F986B700AE6C8EE7">2013</mods:date>
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<mods:number id="1F6D88B0E6F8B58149FBBBF8926D83EF">2013-06-30</mods:number>
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<mods:number id="74F430D77E3DE2F7E181676405501E7A">67</mods:number>
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<mods:identifier id="BC0BBAB88A7B7A3B4D6AE75050504B15" type="Zenodo-Dep">5368835</mods:identifier>
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<treatment id="03D087AAA007FFE6FDD151B7FE0E857C" LSID="urn:lsid:plazi:treatment:03D087AAA007FFE6FDD151B7FE0E857C" httpUri="http://treatment.plazi.org/id/03D087AAA007FFE6FDD151B7FE0E857C" lastPageId="9" pageId="7">
<subSubSection id="C3636537A007FFE8FDD151B7FE3C8051" pageId="7" type="nomenclature">
<paragraph id="8BC636BCA007FFE8FDD151B7FE3C8051" blockId="7.[246,586,735,811]" pageId="7">
<taxonomicName id="4C794D3FA007FFE8FDD151B7FD97818E" authority="(Fall, 1928)" baseAuthorityName="Fall" baseAuthorityYear="1928" box="[246,586,735,757]" class="Insecta" family="Eucnemidae" genus="Onichodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="rugicollis">
<emphasis id="B90DEAAEA007FFE8FDD151B7FD97818E" bold="true" box="[246,586,735,757]" pageId="7">
<emphasis id="B90DEAAEA007FFE8FDD151B7FE1A818E" bold="true" box="[246,455,735,757]" italics="true" pageId="7">Onichodon rugicollis</emphasis>
(Fall, 1928)
</emphasis>
</taxonomicName>
fifth instar (
<figureCitation id="13422A39A007FFE8FC41507DFE048051" box="[358,473,789,811]" captionStart-0="Fig" captionStart-1="Figs" captionStartId-0="7.[731,764,1504,1523]" captionStartId-1="8.[175,215,1454,1473]" captionTargetBox-0="[717,1192,183,1481]" captionTargetBox-1="[151,1153,190,1430]" captionTargetId-0="figure-440@7.[707,1193,183,1481]" captionTargetId-1="figure-92@8.[151,1171,183,1430]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Fig. 11. Onichodon rugicollis, fifth instar, dorsal habitus." captionText-1="Figs. 1215. Onichodon rugicollis, fifth instar. 12) Head, dorsal view; 13) Head and thoracic region, dorsal view;" figureDoi-0="http://doi.org/10.5281/zenodo.5368845" figureDoi-1="http://doi.org/10.5281/zenodo.5368847" httpUri-0="https://zenodo.org/record/5368845/files/figure.png" httpUri-1="https://zenodo.org/record/5368847/files/figure.png" pageId="7">Figs. 1115</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3636537A007FFE8FDE25028FEB48785" pageId="7" type="diagnosis">
<paragraph id="8BC636BCA007FFE8FDE25028FDEE80A2" blockId="7.[173,659,832,1729]" pageId="7">
<emphasis id="B90DEAAEA007FFE8FDE25028FEF0802E" bold="true" box="[197,301,832,853]" pageId="7">Diagnosis.</emphasis>
The shape of the dorsal prothoracic scleromes, along with a widely bifurcate first lateral projection of the head capsule, and sclerotized caudal end of the ninth abdominal segment should distinguish this species from its close relatives,
<taxonomicName id="4C794D3FA007FFE8FDCE50ADFEAE80A1" baseAuthorityName="Brown" baseAuthorityYear="1940" box="[233,371,965,986]" class="Insecta" family="Eucnemidae" genus="Onichodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="canadensis">
<emphasis id="B90DEAAEA007FFE8FDCE50ADFEAE80A1" box="[233,371,965,986]" italics="true" pageId="7">O. canadensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C794D3FA007FFE8FC8E50ADFDF080A1" authorityName="Newman" authorityYear="1838" box="[425,557,965,986]" class="Insecta" family="Eucnemidae" genus="Onichodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="orchesides">
<emphasis id="B90DEAAEA007FFE8FC8E50ADFDF080A1" box="[425,557,965,986]" italics="true" pageId="7">O. orchesides</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC636BCA007FFE8FDE250B7FEB48785" blockId="7.[173,659,832,1729]" pageId="7">
<emphasis id="B90DEAAEA007FFE8FDE250B7FE78808F" bold="true" box="[197,421,991,1012]" pageId="7">Specimens Examined.</emphasis>
Five mature larvae and three larval exuviae collected at
<materialsCitation id="3B113CE1A007FFE8FCCE5092FE658702" collectorName="E. Buckley Wildlife Sanctuary &amp; March &amp; Robert L. Otto &amp; April" country="United States of America" county="Woodford County" location="Clyde" pageId="7" specimenCount="1" stateProvince="Kentucky">
<emphasis id="B90DEAAEA007FFE8FCCE5092FD508774" bold="true" box="[489,653,1018,1039]" pageId="7">
<collectingCountry id="F36E762CA007FFE8FCCE5092FDC08774" box="[489,541,1018,1039]" name="United States of America" pageId="7">USA</collectingCountry>
:
<collectingRegion id="49BDF85EA007FFE8FF0D5092FD508774" box="[554,653,1018,1039]" country="United States of America" name="Kentucky" pageId="7">Kentucky</collectingRegion>
</emphasis>
:
<collectingCounty id="62A74E30A007FFE8FD8A577CFEBD8752" box="[173,352,1044,1065]" pageId="7">Woodford County</collectingCounty>
,
<locationDeviation id="9A742F9EA007FFE8FC4A577CFD908752" box="[365,589,1044,1065]" location="Lexington" pageId="7">
<quantity id="4C819B59A007FFE8FC4A577CFE718752" box="[365,428,1044,1065]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="7" unit="km" value="40.0">40 km</quantity>
NW Lexington
</locationDeviation>
,
<location id="8EA66067A007FFE8FF7E577CFD4E8752" LSID="urn:lsid:plazi:treatment:03D087AAA007FFE6FDD151B7FE0E857C:8EA66067A007FFE8FF7E577CFD4E8752" box="[601,659,1044,1065]" country="United States of America" county="Woodford County" name="Clyde" pageId="7" stateProvince="Kentucky">Clyde</location>
<collectorName id="268C536AA007FFE8FD8A5747FE17873F" box="[173,458,1071,1092]" pageId="7">E. Buckley Wildlife Sanctuary</collectorName>
, 2425
<collectorName id="268C536AA007FFE8FF315747FD89873F" box="[534,596,1071,1092]" pageId="7">March</collectorName>
, 1995,
<collectorName id="268C536AA007FFE8FD8A5721FE938725" box="[173,334,1097,1118]" pageId="7">Robert L. Otto</collectorName>
, reared on 27
<collectorName id="268C536AA007FFE8FCD55721FDF68725" box="[498,555,1097,1118]" pageId="7">April</collectorName>
, 1995, in rotten wood (3 exuviae)
</materialsCitation>
;
<materialsCitation id="3B113CE1A007FFE8FCEF570CFD5287D5" collectingDate="1960-05-03" collectorName="Ingham County, M. S. U. &amp; Sanford Woodlot &amp; C. F. Gibbons" country="United States of America" county="Ingham County" location="Ingham County" pageId="7" specimenCount="3" specimenCount-larva="3" stateProvince="Michigan">
<emphasis id="B90DEAAEA007FFE8FCEF570CFDED8702" bold="true" box="[456,560,1124,1145]" pageId="7">
<collectingRegion id="49BDF85EA007FFE8FCEF570CFDED8702" box="[456,560,1124,1145]" country="United States of America" name="Michigan" pageId="7">Michigan</collectingRegion>
</emphasis>
:
<collectorName id="268C536AA007FFE8FF62570CFE9587EF" pageId="7">
<collectingCounty id="62A74E30A007FFE8FF62570CFF2887EF" pageId="7">Ingham County</collectingCounty>
, M.S.U.
</collectorName>
<collectorName id="268C536AA007FFE8FC735717FE2087EF" box="[340,509,1151,1172]" pageId="7">Sanford Woodlot</collectorName>
,
<date id="FFC7107CA007FFE8FF2C5717FD5287EF" box="[523,655,1151,1172]" pageId="7" value="1960-05-03">
<collectingDate id="EF83E994A007FFE8FF2C5717FD5287EF" box="[523,655,1151,1172]" pageId="7" value="1960-05-03">May 3, 1960</collectingDate>
</date>
,
<collectorName id="268C536AA007FFE8FD8A57F1FE9B87D5" box="[173,326,1177,1198]" pageId="7">C.F. Gibbons</collectorName>
, in rotten wood (
<specimenCount id="9D7FFD35A007FFE8FF0457F1FD5A87D5" box="[547,647,1177,1198]" pageId="7" type="larva">3 larvae</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="3B113CE1A007FFE8FD8A57DCFEB88785" collectorName="Pine Hollow &amp; May &amp; Kelly M. Zivicki" country="United States of America" county="Sauk County" latitude="43.373886" location="Sauk County" longLatPrecision="19" longitude="-89.905556" pageId="7" specimenCount="2" specimenCount-larva="2" stateProvince="Wisconsin">
<emphasis id="B90DEAAEA007FFE8FD8A57DCFECC87B2" bold="true" box="[173,273,1204,1225]" pageId="7">
<collectingRegion id="49BDF85EA007FFE8FD8A57DCFECC87B2" box="[173,273,1204,1225]" country="United States of America" name="Wisconsin" pageId="7">Wisconsin</collectingRegion>
</emphasis>
:
<collectingCounty id="62A74E30A007FFE8FC3A57DCFE4B87B2" box="[285,406,1204,1225]" pageId="7">Sauk County</collectingCounty>
,
<collectorName id="268C536AA007FFE8FCB857DCFDC987B2" box="[415,532,1204,1225]" pageId="7">Pine Hollow</collectorName>
,
<geoCoordinate id="EE4D507BA007FFE8FF3A57DCFD5187B2" box="[541,652,1203,1225]" degrees="43" direction="north" minutes="22" orientation="latitude" pageId="7" precision="15" seconds="26" value="43.373886">43°2226″N</geoCoordinate>
/
<geoCoordinate id="EE4D507BA007FFE8FD8A57A6FEF58798" box="[173,296,1230,1252]" degrees="89" direction="west" minutes="54" orientation="longitude" pageId="7" precision="15" seconds="20" value="-89.905556">89°5420″W</geoCoordinate>
, 7
<collectorName id="268C536AA007FFE8FC6C57A6FEA58798" box="[331,376,1230,1251]" pageId="7">May</collectorName>
, 2007,
<collectorName id="268C536AA007FFE8FCE157A6FDAE8798" box="[454,627,1230,1251]" pageId="7">Kelly M. Zivicki</collectorName>
, in dead log (
<specimenCount id="9D7FFD35A007FFE8FC295781FE838785" box="[270,350,1257,1278]" pageId="7" type="larva">2 larvae</specimenCount>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3636537A007FFE6FDE2566BFD8E8041" lastPageId="9" pageId="7" type="description">
<paragraph id="8BC636BCA007FFE8FDE2566BFB7485BA" blockId="7.[173,659,832,1729]" lastBlockId="7.[707,1194,1602,1729]" pageId="7">
<emphasis id="B90DEAAEA007FFE8FDE2566BFE9C8663" bold="true" box="[197,321,1283,1304]" pageId="7">Description.</emphasis>
Length 30.034.0 mm, width 3.04.0 mm. Orthosomatic.
<emphasis id="B90DEAAEA007FFE8FCE45676FE258648" bold="true" box="[451,504,1310,1331]" pageId="7">Body</emphasis>
(
<figureCitation id="13422A39A007FFE8FF2F5676FD938648" box="[520,590,1310,1331]" captionStart="Fig" captionStartId="7.[731,764,1504,1523]" captionTargetBox="[717,1192,183,1481]" captionTargetId="figure-440@7.[707,1193,183,1481]" captionTargetPageId="7" captionText="Fig. 11. Onichodon rugicollis, fifth instar, dorsal habitus." figureDoi="http://doi.org/10.5281/zenodo.5368845" httpUri="https://zenodo.org/record/5368845/files/figure.png" pageId="7">Fig. 11</figureCitation>
): Subcylindrical, sides parallel, yellow with head, prothoracic sclerome patches, and caudal end of 9
<superScript id="7C0C9BF4A007FFE8FDFC5600FF37860D" attach="left" box="[219,234,1384,1398]" fontSize="6" pageId="7">th</superScript>
abdominal segment dark brown. Setae reduced or absent. Pair of small legs reduced to dome-like structures present near posterolateral areas of each thoracic segment. Dorsal and ventral microtrichial patches slightly darker than their surrounding areas.
<emphasis id="B90DEAAEA007FFE8FCBB569AFE0F857C" bold="true" box="[412,466,1522,1543]" pageId="7">Head</emphasis>
(
<figureCitation id="13422A39A007FFE8FCC5569AFDF0857C" box="[482,557,1522,1543]" captionStart="Figs" captionStartId="8.[175,215,1454,1473]" captionTargetBox="[151,1153,190,1430]" captionTargetId="figure-92@8.[151,1171,183,1430]" captionTargetPageId="8" captionText="Figs. 1215. Onichodon rugicollis, fifth instar. 12) Head, dorsal view; 13) Head and thoracic region, dorsal view;" figureDoi="http://doi.org/10.5281/zenodo.5368847" httpUri="https://zenodo.org/record/5368847/files/figure.png" pageId="7">Fig. 12</figureCitation>
): Strongly flattened, prognathous, and inserted into prothorax. Dorsal and ventral cephalic discs oblonged, subtrapezoidal and simple. Anterior portion of the head capsule heavily sclerotized. Posterior areas of head capsule unsclerotized. Each lateral side of head capsule with 6 projections. First lateral projections slightly elongate, very widely bifurcate, directed anteriorally. Second through 6
<superScript id="7C0C9BF4A007FFE8FFE8553FFD03851E" attach="left" box="[719,734,1623,1637]" fontSize="6" pageId="7">th</superScript>
lateral projections directed anterolaterally. Second lateral projection short and narrowly separated from 3
<superScript id="7C0C9BF4A007FFE8FE5E55E6FC5485E7" attach="none" box="[889,905,1678,1692]" fontSize="6" pageId="7">rd</superScript>
. Antennae minute, arising between 5
<superScript id="7C0C9BF4A007FFE8FE0055CFFCE885CE" attach="left" box="[807,821,1703,1717]" fontSize="6" pageId="7">th</superScript>
and 6
<superScript id="7C0C9BF4A007FFE8FE5255CFFC5E85CE" attach="left" box="[885,899,1703,1717]" fontSize="6" pageId="7">th</superScript>
lateral projections. Scape not
</paragraph>
<caption id="DF066634A007FFE8FFFC5688FCDB8570" ID-DOI="http://doi.org/10.5281/zenodo.5368845" ID-Zenodo-Dep="5368845" httpUri="https://zenodo.org/record/5368845/files/figure.png" pageId="7" startId="7.[731,764,1504,1523]" targetBox="[717,1192,183,1481]" targetPageId="7">
<paragraph id="8BC636BCA007FFE8FFFC5688FCDB8570" blockId="7.[707,1193,1504,1547]" pageId="7">
<emphasis id="B90DEAAEA007FFE8FFFC5688FCFF8688" bold="true" box="[731,802,1504,1523]" pageId="7">Fig. 11.</emphasis>
<taxonomicName id="4C794D3FA007FFE8FE1F5688FC2A8689" baseAuthorityName="Fall" baseAuthorityYear="1928" box="[824,1015,1504,1522]" class="Insecta" family="Eucnemidae" genus="Onichodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="rugicollis">
<emphasis id="B90DEAAEA007FFE8FE1F5688FC2A8689" box="[824,1015,1504,1522]" italics="true" pageId="7">Onichodon rugicollis</emphasis>
</taxonomicName>
, fifth instar, dorsal habitus.
</paragraph>
</caption>
<caption id="DF066634A008FFE7FD8856C6FB4986BA" ID-DOI="http://doi.org/10.5281/zenodo.5368847" ID-Zenodo-Dep="5368847" box="[175,1172,1453,1473]" httpUri="https://zenodo.org/record/5368847/files/figure.png" pageId="8" startId="8.[175,215,1454,1473]" targetBox="[151,1153,190,1430]" targetPageId="8">
<paragraph id="8BC636BCA008FFE7FD8856C6FB4986BA" blockId="8.[151,1172,1453,1497]" box="[175,1172,1453,1473]" pageId="8">
<emphasis id="B90DEAAEA008FFE7FD8856C6FEC286BA" bold="true" box="[175,287,1453,1473]" pageId="8">Figs. 1215.</emphasis>
<taxonomicName id="4C794D3FA008FFE7FC1256C6FE3486BB" baseAuthorityName="Fall" baseAuthorityYear="1928" box="[309,489,1454,1472]" class="Insecta" family="Eucnemidae" genus="Onichodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="rugicollis">
<emphasis id="B90DEAAEA008FFE7FC1256C6FE3486BB" box="[309,489,1454,1472]" italics="true" pageId="8">Onichodon rugicollis</emphasis>
</taxonomicName>
, fifth instar.
<emphasis id="B90DEAAEA008FFE7FF7E56C6FDAA86BA" bold="true" box="[601,631,1454,1473]" pageId="8">12)</emphasis>
Head, dorsal view;
<emphasis id="B90DEAAEA008FFE7FE0356C6FC9F86BA" bold="true" box="[804,834,1454,1473]" pageId="8">13)</emphasis>
Head and thoracic region, dorsal view;
</paragraph>
</caption>
<paragraph id="8BC636BCA008FFE7FDB056AEFC6C86A2" blockId="8.[151,1172,1453,1497]" box="[151,945,1478,1497]" pageId="8">
<emphasis id="B90DEAAEA008FFE7FDB056AEFF6B86A2" bold="true" box="[151,182,1478,1497]" pageId="8">14)</emphasis>
Head and thoracic region, ventral view;
<emphasis id="B90DEAAEA008FFE7FF3D56AEFDE586A2" bold="true" box="[538,568,1478,1497]" pageId="8">15)</emphasis>
Abdominal segments VII-IX, ventral view.
</paragraph>
<paragraph id="8BC636BCA008FFE6FDB05565FD8E8041" blockId="8.[151,638,1549,1729]" lastBlockId="9.[173,660,194,1543]" lastPageId="9" pageId="8">
visible. Pedicel elongate. Sensorum and flagellum subequal in length. Sensory papillae indistinct. Mandibles minute, resting in the mesal acumination of the head capsule between 6
<superScript id="7C0C9BF4A008FFE7FF60553FFD88851E" attach="left" box="[583,597,1623,1637]" fontSize="6" pageId="8">th</superScript>
lateral projections. Each mandible heavily sclerotized, distinct, oval, longer than wide with 2 outwardly projecting teeth. Maxillary palpi extremely small, 3-segmented. Ligula, mala, lacinia, and galea not visible. Hypostomal rods absent.
<emphasis id="B90DEAAEA008FFE7FFD8552AFCB7852C" bold="true" box="[767,874,1602,1623]" pageId="8">Prothorax</emphasis>
(
<figureCitation id="13422A39A008FFE7FE5A552AFBDF852C" box="[893,1026,1602,1623]" captionStart="Figs" captionStartId="8.[175,215,1454,1473]" captionTargetBox="[151,1153,190,1430]" captionTargetId="figure-92@8.[151,1171,183,1430]" captionTargetPageId="8" captionText="Figs. 1215. Onichodon rugicollis, fifth instar. 12) Head, dorsal view; 13) Head and thoracic region, dorsal view;" figureDoi="http://doi.org/10.5281/zenodo.5368847" httpUri="https://zenodo.org/record/5368847/files/figure.png" pageId="8">Figs. 13, 14</figureCitation>
): Subequal to subsequent 2 thoracic segments. Tergum with pair of square scleromes, caudally scalloped, irregularly shaped. Subtriangular microtrichial patch between scleromes. Tergum and sternum without areoles. Sternum with triangular scleromes and more rounded, slightly oblonged and smaller microthrichial patch.
<emphasis id="B90DEAAEA009FFE6FCA8539FFD518277" bold="true" box="[399,652,247,268]" pageId="9">Meso- and metathorax</emphasis>
: Each terga and sterna with kidney-shaped microtrichial patch of short triangular extension coming from mediocaudal region directed towards the posterior end. Surfaces of terga and sterna with longitudinal plicae and carinae just caudad of each microtrichial patch and extending towards posterior end. Each terga with pair of elongate oval-shaped areoles near posterior end. Oblong areole present near posterior end of metathoracic sterna. Mesothorax without spiracles.
<emphasis id="B90DEAAEA009FFE6FF015169FD51816D" bold="true" box="[550,652,513,534]" pageId="9">Abdomen</emphasis>
: Segments I-IX subequal in length and width. Terga and sterna I-VIII with microtrichial patches that successively change from oval on segment I to subtriangular on segment VIII. Terga and sterna I-VIII with oval areole near posterior end, each successively narrower. Tergum IX without microtrichial patch, sparsely punctate near the caudal end; sternum (
<figureCitation id="13422A39A009FFE6FCA451BDFE178191" box="[387,458,725,746]" captionStart="Figs" captionStartId="8.[175,215,1454,1473]" captionTargetBox="[151,1153,190,1430]" captionTargetId="figure-92@8.[151,1171,183,1430]" captionTargetPageId="8" captionText="Figs. 1215. Onichodon rugicollis, fifth instar. 12) Head, dorsal view; 13) Head and thoracic region, dorsal view;" figureDoi="http://doi.org/10.5281/zenodo.5368847" httpUri="https://zenodo.org/record/5368847/files/figure.png" pageId="9">Fig. 15</figureCitation>
) heavily sclerotized at caudal half with prominent, circular ring of asperities around anal region. Urogomphi absent on segment IX. Spiracles annular-biforous.
</paragraph>
</subSubSection>
<subSubSection id="C3636537A009FFE6FDE25028FD93808F" pageId="9" type="distribution">
<paragraph id="8BC636BCA009FFE6FDE25028FD93808F" blockId="9.[173,660,194,1543]" pageId="9">
<emphasis id="B90DEAAEA009FFE6FDE25028FE9A802E" bold="true" box="[197,327,832,853]" pageId="9">Distribution.</emphasis>
<taxonomicName id="4C794D3FA009FFE6FC725028FDF6802E" baseAuthorityName="Fall" baseAuthorityYear="1928" box="[341,555,832,853]" class="Insecta" family="Eucnemidae" genus="Onichodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="rugicollis">
<emphasis id="B90DEAAEA009FFE6FC725028FDF6802E" box="[341,555,832,853]" italics="true" pageId="9">Onichodon rugicollis</emphasis>
</taxonomicName>
is known from
<emphasis id="B90DEAAEA009FFE6FDC35032FEC08014" bold="true" box="[228,285,858,879]" pageId="9">
<collectingCountry id="F36E762CA009FFE6FDC35032FEC58014" box="[228,280,858,879]" name="United States of America" pageId="9">USA</collectingCountry>
:
</emphasis>
<collectingRegion id="49BDF85EA009FFE6FC015032FEA28014" box="[294,383,858,879]" country="United States of America" name="Alabama" pageId="9">Alabama</collectingRegion>
,
<collectingRegion id="49BDF85EA009FFE6FCAC5032FE0F8014" box="[395,466,858,879]" country="United States of America" name="Florida" pageId="9">Florida</collectingRegion>
,
<collectingRegion id="49BDF85EA009FFE6FCF85032FDF88014" box="[479,549,858,879]" country="United States of America" name="Illinois" pageId="9">Illinois</collectingRegion>
,
<collectingRegion id="49BDF85EA009FFE6FF165032FD528014" box="[561,655,858,879]" country="United States of America" name="Kentucky" pageId="9">Kentucky</collectingRegion>
,
<collectingRegion id="49BDF85EA009FFE6FD8A501DFED580F1" box="[173,264,885,906]" country="United States of America" name="Louisiana" pageId="9">Louisiana</collectingRegion>
,
<collectingRegion id="49BDF85EA009FFE6FC36501DFEA780F1" box="[273,378,885,906]" country="United States of America" name="Mississippi" pageId="9">Mississippi</collectingRegion>
,
<collectingRegion id="49BDF85EA009FFE6FCA5501DFDD180F1" box="[386,524,885,906]" country="United States of America" name="North Carolina" pageId="9">North Carolina</collectingRegion>
,
<collectingRegion id="49BDF85EA009FFE6FF32501DFD4D80F1" box="[533,656,885,906]" country="United States of America" name="Pennsylvania" pageId="9">Pennsylvania</collectingRegion>
,
<collectingRegion id="49BDF85EA009FFE6FD8A50E7FF3580DF" box="[173,232,911,932]" country="United States of America" name="Texas" pageId="9">Texas</collectingRegion>
, and
<collectingRegion id="49BDF85EA009FFE6FC3850E7FE5880DF" box="[287,389,911,932]" country="United States of America" name="Wisconsin" pageId="9">Wisconsin</collectingRegion>
(
<bibRefCitation id="EFE84B4DA009FFE6FCB450E7FDC880DF" author="Muona &amp; Revision of the Nearctic Eucnemidae" box="[403,533,911,932]" pageId="9" pagination="1 - 106" refId="ref5259" refString="Muona, J. 2000. A Revision of the Nearctic Eucnemidae. Acta Zoologica Fennica. 212: 1 - 106." type="journal article" year="2000">Muona 2000</bibRefCitation>
;
<bibRefCitation id="EFE84B4DA009FFE6FF0650E7FD5480DF" author="Otto" box="[545,649,911,932]" pageId="9" pagination="92 - 93" refId="ref5395" refString="Otto, R. L. 2010. New records for seven rare Nearctic species of false click beetles (Coleoptera: Eucnemidae). The Coleopterists Bulletin 64 (1): 92 - 93." type="journal article" year="2010">Otto 2010</bibRefCitation>
). It is recorded here as a
<emphasis id="B90DEAAEA009FFE6FC9450C2FDB680C4" bold="true" box="[435,619,938,959]" pageId="9">new state record</emphasis>
for
<collectingRegion id="49BDF85EA009FFE6FD8A50ACFED080A2" box="[173,269,964,985]" country="United States of America" name="Michigan" pageId="9">Michigan</collectingRegion>
. All specimens used in this study came from
<collectingRegion id="49BDF85EA009FFE6FDC450B7FE9F808F" box="[227,322,991,1012]" country="United States of America" name="Kentucky" pageId="9">Kentucky</collectingRegion>
,
<collectingRegion id="49BDF85EA009FFE6FC6A50B7FE71808F" box="[333,428,991,1012]" country="United States of America" name="Michigan" pageId="9">Michigan</collectingRegion>
, and
<collectingRegion id="49BDF85EA009FFE6FCC650B7FD94808F" box="[481,585,991,1012]" country="United States of America" name="Wisconsin" pageId="9">Wisconsin</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C3636537A009FFE6FDE25092FE0E857C" pageId="9" type="biology_ecology">
<paragraph id="8BC636BCA009FFE6FDE25092FE0E857C" blockId="9.[173,660,194,1543]" pageId="9">
<emphasis id="B90DEAAEA009FFE6FDE25092FECB8774" bold="true" box="[197,278,1018,1039]" pageId="9">Biology.</emphasis>
<taxonomicName id="4C794D3FA009FFE6FC035092FE328774" baseAuthorityName="Fall" baseAuthorityYear="1928" box="[292,495,1018,1039]" class="Insecta" family="Eucnemidae" genus="Onichodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="rugicollis">
<emphasis id="B90DEAAEA009FFE6FC035092FE328774" box="[292,495,1018,1039]" italics="true" pageId="9">Onichodon rugicollis</emphasis>
</taxonomicName>
is an uncommon and widespread species, largely distributed in the southern
<collectingCountry id="F36E762CA009FFE6FC225747FE5A873F" box="[261,391,1071,1092]" name="United States of America" pageId="9">United States</collectingCountry>
. In
<collectingRegion id="49BDF85EA009FFE6FC8C5747FDCC873F" box="[427,529,1071,1092]" country="United States of America" name="Wisconsin" pageId="9">Wisconsin</collectingRegion>
,
<taxonomicName id="4C794D3FA009FFE6FF3C5747FD4E873F" baseAuthorityName="Fall" baseAuthorityYear="1928" box="[539,659,1071,1092]" class="Insecta" family="Eucnemidae" genus="Onichodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="rugicollis">
<emphasis id="B90DEAAEA009FFE6FF3C5747FD4E873F" box="[539,659,1071,1092]" italics="true" pageId="9">O. rugicollis</emphasis>
</taxonomicName>
was found in a variety of forest systems. Adults and larvae were taken in northern hardwood swamp, northern mesic forest, oak barrens, oak opening, southern dry forest, and southern mesic forest. Two adults were reared from damp oak wood collected in oak opening woodland in southwestern
<collectingRegion id="49BDF85EA009FFE6FC145781FE448785" box="[307,409,1257,1278]" country="United States of America" name="Wisconsin" pageId="9">Wisconsin</collectingRegion>
. One adult was observed emerging from a dead, standing tree trunk. Based on label information, many adults were taken in a flight intercept trap, Lindgren funnel traps, and purple prism traps deployed throughout the southeastern two-thirds of
<collectingRegion id="49BDF85EA009FFE6FCAC5606FE2886F8" box="[395,501,1390,1411]" country="United States of America" name="Wisconsin" pageId="9">Wisconsin</collectingRegion>
.
<bibRefCitation id="EFE84B4DA009FFE6FF255606FD4E86F8" author="Muona &amp; Revision of the Nearctic Eucnemidae" box="[514,659,1390,1411]" pageId="9" pagination="1 - 106" refId="ref5259" refString="Muona, J. 2000. A Revision of the Nearctic Eucnemidae. Acta Zoologica Fennica. 212: 1 - 106." type="journal article" year="2000">Muona (2000)</bibRefCitation>
wrote that the species was taken at blacklight traps.
<bibRefCitation id="EFE84B4DA009FFE6FDCC56CBFE8386C3" author="Otto" box="[235,350,1443,1464]" pageId="9" pagination="92 - 93" refId="ref5395" refString="Otto, R. L. 2010. New records for seven rare Nearctic species of false click beetles (Coleoptera: Eucnemidae). The Coleopterists Bulletin 64 (1): 92 - 93." type="journal article" year="2010">Otto (2010)</bibRefCitation>
wrote that several adults were reared from larvae in a rotten log in
<collectingRegion id="49BDF85EA009FFE6FF1456D5FD4E86A9" box="[563,659,1469,1490]" country="United States of America" name="Kentucky" pageId="9">Kentucky</collectingRegion>
and adults were taken on dead trees and fungus in northwestern
<collectingRegion id="49BDF85EA009FFE6FC6A569AFE12857C" box="[333,463,1522,1543]" country="United States of America" name="Pennsylvania" pageId="9">Pennsylvania</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>