192 lines
24 KiB
XML
192 lines
24 KiB
XML
<document id="8B26C445AAC2B6BEAC27833804FD558C" ID-DOI="10.5281/zenodo.4665621" ID-ISSN="1638-9395" ID-Zenodo-Dep="4665621" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617712844033" checkinUser="felipe" docAuthor="Venczel, Márton & Stiuc, Emanoil" docDate="2008" docId="03D08790FFDDFFF15412A303FC195534" docLanguage="en" docName="Geodiversitas.30.4.731-763.pdf" docOrigin="Geodiversitas 30 (4)" docStyle="DocumentStyle:9DCE1246056655C2F9CBE9822E831E4E.3:Geodiversitas.2000-2008.journal_article.type3" docStyleId="9DCE1246056655C2F9CBE9822E831E4E" docStyleName="Geodiversitas.2000-2008.journal_article.type3" docStyleVersion="3" docTitle="Coronella miocaenica Venczel 1998" docType="treatment" docVersion="3" lastPageNumber="756" masterDocId="FFE9FFE8FFC5FFE85703A736FFA55105" masterDocTitle="Late middle Miocene amphibians and squamate reptiles from Taut ,, Romania" masterLastPageNumber="763" masterPageNumber="731" pageNumber="755" updateTime="1698901057390" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:titleInfo id="BC5CDAAC70492994A71D804C2AFCD799">
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<mods:title id="635998FCD973F1D0744470FDAA5F84ED">Late middle Miocene amphibians and squamate reptiles from Taut ,, Romania</mods:title>
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</mods:titleInfo>
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<mods:name id="E8CE1526D3F85FF7A4C9208C72C941E8" type="personal">
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<mods:roleTerm id="CF9276C09178326DCE9E6AA9B907BF7A">Author</mods:roleTerm>
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<mods:namePart id="177BC04D068E76052E583593C62FF1C8">Venczel, Márton</mods:namePart>
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<mods:affiliation id="F5938C248BA03DC19991D99A1ECBC938">Tării ‚ Crişurilor Museum, B-dul Dacia 1 - 3, RO- 410464 Oradea (Romania) mvenczel @ rdslink. ro</mods:affiliation>
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<mods:nameIdentifier id="047699E276D675A0C9B045DCBAE6FF8D" type="email">mvenczel@rdslink.ro</mods:nameIdentifier>
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<mods:roleTerm id="697ED2D6210AC95641D38B8EECD597F6">Author</mods:roleTerm>
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<mods:namePart id="BFFC744013E9A9153A7B2355E36BA31F">Stiuc, Emanoil</mods:namePart>
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<mods:affiliation id="25AB92539AD8DE59439D323C530CE0B3">Speleological Institute “ Emil Racovit, ă ”, Calea 13 Septembrie nr. 13, RO- 050711 Bucureşti (Romania) stiucaemil @ yahoo. com.</mods:affiliation>
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<mods:nameIdentifier id="28D222EDAA4E4CC2B843698158DCF5FC" type="email">stiucaemil@yahoo.com</mods:nameIdentifier>
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<mods:title id="41866378C62EF8C782DEC024F2A598EA">Geodiversitas</mods:title>
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<mods:date id="EB0DC1CED13BD71E7F5D8F2A0B9A0998">2008</mods:date>
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<mods:number id="81673F77F5EC8AC7C0C821CF27D024C7">30</mods:number>
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<mods:number id="6C5AEFEA0A4E2C2E493AF382C6C32D11">4</mods:number>
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<mods:identifier id="90FF38726AADDD3822A718C727AF933C" type="DOI">10.5281/zenodo.4665621</mods:identifier>
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<mods:identifier id="C957E3F2E3FE8849AB890424C9FD16C7" type="ISSN">1638-9395</mods:identifier>
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<subSubSection id="C363650DFFDDFFF05412A303FBA65575" pageId="24" pageNumber="755" type="nomenclature">
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<paragraph id="8BC63686FFDDFFF05412A303FBA65575" blockId="24.[785,1177,1077,1136]" pageId="24" pageNumber="755">
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<heading id="D08E81EAFFDDFFF05412A303FBA65575" centered="true" fontSize="11" level="2" pageId="24" pageNumber="755" reason="2">
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<taxonomicName id="4C794D05FFDDFFF05412A303FB3C5555" authority="Venczel, 1998" authorityName="Venczel" authorityYear="1998" box="[785,1177,1077,1104]" class="Reptilia" family="Colubridae" genus="Coronella" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="755" phylum="Chordata" rank="species" species="miocaenica">
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<emphasis id="B90DEA94FFDDFFF05412A303FC5F554A" bold="true" box="[785,1018,1077,1104]" italics="true" pageId="24" pageNumber="755">Coronella miocaenica</emphasis>
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<bibRefCitation id="EFE84B77FFDDFFF05302A303FB3C5555" author="VENCZEL M." box="[1025,1177,1077,1104]" pageId="24" pageNumber="755" pagination="1 - 22" refId="ref18311" refString="VENCZEL M. 1998. - Late Miocene snakes (Reptilia: Serpentes) from Polgardi (Hungary): a second contribution. Acta Zoologica Cracoviensia 41: 1 - 22." type="journal article" year="1998">Venczel, 1998</bibRefCitation>
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</taxonomicName>
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(
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<figureCitation id="13422A03FFDDFFF054ADA363FC5E5575" box="[942,1019,1109,1136]" captionStart="FIG" captionStartId="25.[151,162,610,627]" captionTargetBox="[180,1213,254,575]" captionTargetId="figure-15@25.[151,1241,250,580]" captionTargetPageId="25" captionText="FIG. 14. — Coronella miocaenica Venczel 1998, presacral vertebra (ISER Tt-0490/1): A, dorsal view; B, right lateral view; C, ventral view. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4665661" httpUri="https://zenodo.org/record/4665661/files/figure.png" pageId="24" pageNumber="755">Fig. 14</figureCitation>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C363650DFFDDFFF055C9A3B8FC6D55C4" pageId="24" pageNumber="755" type="materials_examined">
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<paragraph id="8BC63686FFDDFFF055C9A3B8FC6D55C4" blockId="24.[714,1247,1166,1217]" pageId="24" pageNumber="755">
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MATERIAL EXAMINED. — Four fragmentary trunk vertebrae (
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<collectionCode id="ED68AE43FFDDFFF05402A39FFC9255C4" box="[769,823,1193,1217]" country="Romania" httpUri="http://grbio.org/cool/911h-bwmh" name="Institutul Speologie Emil G. Racovita" pageId="24" pageNumber="755">ISER</collectionCode>
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Tt-0490/1-4).
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</paragraph>
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</subSubSection>
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<subSubSection id="C363650DFFDDFFF155C9A3C2FE6B52F4" lastPageId="25" lastPageNumber="756" pageId="24" pageNumber="755" type="description">
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<paragraph id="8BC63686FFDDFFF055C9A3C2FCFE5409" blockId="24.[712,1248,1268,1706]" box="[714,859,1268,1294]" pageId="24" pageNumber="755">DESCRIPTION</paragraph>
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<paragraph id="8BC63686FFDDFFF055C9A225FB7A57AF" blockId="24.[712,1248,1268,1706]" pageId="24" pageNumber="755">
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All the vertebrae are of minute size. The centrum length of the largest specimen is
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<quantity id="4C819B63FFDDFFF05332A205FB3C544B" box="[1073,1177,1331,1358]" metricMagnitude="-3" metricUnit="m" metricValue="3.45" pageId="24" pageNumber="755" unit="mm" value="3.45">3.45 mm</quantity>
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, while the centrum width is
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<quantity id="4C819B63FFDDFFF054BBA264FBB85468" box="[952,1053,1362,1389]" metricMagnitude="-3" metricUnit="m" metricValue="2.38" pageId="24" pageNumber="755" unit="mm" value="2.38">2.38 mm</quantity>
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(centrum length/ centrum width ratio = 1.45). The centrum is elongated; the neural arch is moderately vaulted and provided with a longer than high neural spine. The dorsal margin of the latter structure is not thickened and both the anterior and posterior margins are overhanging. The posterior margin of the neural arch lacks epizygapophyseal spines. The zygosphene lacks a medial lobe, or it is provided with a rather indistinct medial and two small lateral lobes. The anterior portion of the haemal keel is rather promi-
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</paragraph>
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<paragraph id="8BC63686FFDDFFF055C9A199FB4557CF" blockId="24.[714,1248,1711,1738]" box="[714,1248,1711,1738]" pageId="24" pageNumber="755">nent and thin, while the posterior part diminishes in</paragraph>
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<paragraph id="8BC63686FFDCFFF157ADA745FEFC5181" blockId="25.[174,345,115,133]" box="[174,345,115,133]" pageId="25" pageNumber="756">Venczel M. & Ştiucă E.</paragraph>
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<caption id="DF06660EFFDCFFF15794A554FEF3538C" ID-DOI="http://doi.org/10.5281/zenodo.4665661" ID-Zenodo-Dep="4665661" httpUri="https://zenodo.org/record/4665661/files/figure.png" pageId="25" pageNumber="756" startId="25.[151,162,610,627]" targetBox="[180,1213,254,575]" targetPageId="25">
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<paragraph id="8BC63686FFDCFFF15794A554FEF3538C" blockId="25.[151,1247,609,649]" pageId="25" pageNumber="756">
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FIG. 14. —
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<taxonomicName id="4C794D05FFDCFFF157F7A557FD845376" authority="Venczel 1998" authorityName="Venczel" authorityYear="1998" box="[244,545,609,627]" class="Reptilia" family="Colubridae" genus="Coronella" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="756" phylum="Chordata" rank="species" species="miocaenica">
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||
<emphasis id="B90DEA94FFDCFFF157F7A557FE0C5377" box="[244,425,609,626]" italics="true" pageId="25" pageNumber="756">Coronella miocaenica</emphasis>
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||
<bibRefCitation id="EFE84B77FFDCFFF156ACA554FD845376" author="VENCZEL M." box="[431,545,610,627]" pageId="25" pageNumber="756" pagination="1 - 22" refId="ref18311" refString="VENCZEL M. 1998. - Late Miocene snakes (Reptilia: Serpentes) from Polgardi (Hungary): a second contribution. Acta Zoologica Cracoviensia 41: 1 - 22." type="journal article" year="1998">Venczel 1998</bibRefCitation>
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||
</taxonomicName>
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||
, presacral vertebra (ISER Tt-0490/1):
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||
<emphasis id="B90DEA94FFDCFFF1545DA557FCCE5377" bold="true" box="[862,875,609,626]" pageId="25" pageNumber="756">A</emphasis>
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||
, dorsal view;
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||
<emphasis id="B90DEA94FFDCFFF154E3A557FC485377" bold="true" box="[992,1005,609,626]" pageId="25" pageNumber="756">B</emphasis>
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||
, right lateral view;
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||
<emphasis id="B90DEA94FFDCFFF1538DA557FB395377" bold="true" box="[1166,1180,609,626]" pageId="25" pageNumber="756">C</emphasis>
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||
, ventral view. Scale bar: 2 mm.
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</paragraph>
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||
</caption>
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||
<paragraph id="8BC63686FFDCFFF15794A5EFFE6B52F4" blockId="25.[150,684,729,1009]" pageId="25" pageNumber="756">height and becomes spatulate-shaped. Tiny paired tubercles are present on the subcotylar lips. The subcentral ridges are weakly defined.The prezygapophyseal articular facets are of oval shape, while the prezygapophyseal processes are extremely short and with obtuse extremities. The paradiapophyses are evidently differentiated into diapophyses and parapophyses. The latter structures are noticeably longer than the diapophyses.</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C363650DFFDCFFF15794A320FC195534" pageId="25" pageNumber="756" type="discussion">
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||
<paragraph id="8BC63686FFDCFFF15794A320FF5E552B" blockId="25.[149,686,1046,1706]" box="[151,251,1046,1072]" pageId="25" pageNumber="756">REMARKS</paragraph>
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||
<paragraph id="8BC63686FFDCFFF15794A303FD27544B" blockId="25.[149,686,1046,1706]" pageId="25" pageNumber="756">
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The vertebral morphology of the above fossils closely resembles that of
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<taxonomicName id="4C794D05FFDCFFF1564AA363FDBA556A" authorityName="Venczel" authorityYear="1998" box="[329,543,1109,1135]" class="Reptilia" family="Colubridae" genus="Coronella" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="756" phylum="Chordata" rank="species" species="miocaenica">
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<emphasis id="B90DEA94FFDCFFF1564AA363FDBA556A" box="[329,543,1109,1135]" italics="true" pageId="25" pageNumber="756">Coronella miocaenica</emphasis>
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||
</taxonomicName>
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, known from the late Miocene (MN 13) of Polgárdi 4,
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<collectingCountry id="F36E7616FFDCFFF15549A343FD0F5595" box="[586,682,1141,1168]" name="Hungary" pageId="25" pageNumber="756">Hungary</collectingCountry>
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||
(see
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||
<bibRefCitation id="EFE84B77FFDCFFF157C6A3A2FEFD55AA" author="VENCZEL M." box="[197,344,1172,1199]" pageId="25" pageNumber="756" pagination="1 - 22" refId="ref18311" refString="VENCZEL M. 1998. - Late Miocene snakes (Reptilia: Serpentes) from Polgardi (Hungary): a second contribution. Acta Zoologica Cracoviensia 41: 1 - 22." type="journal article" year="1998">Venczel 1998</bibRefCitation>
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||
: fig. 6a-e). The centrum length/ centrum width ratio approaches also the intraspecific variation of this species (
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||
<quantity id="4C819B63FFDCFFF1568BA3E2FDA255EA" box="[392,519,1236,1263]" metricMagnitude="-2" metricUnit="m" metricValue="3.0987999999999998" metricValueMax="3.6068" metricValueMin="2.5908" pageId="25" pageNumber="756" unit="in" value="1.22" valueMax="1.42" valueMin="1.02">1.02-1.42 in</quantity>
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||
specimens coming from Polgárdi 4U). The neural arch in the extant
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<taxonomicName id="4C794D05FFDCFFF15796A225FE1A542B" authority="Laurenti, 1768" authorityName="Laurenti" authorityYear="1768" box="[149,447,1299,1326]" class="Reptilia" family="Colubridae" genus="Coronella" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="756" phylum="Chordata" rank="species" species="austriaca">
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||
<emphasis id="B90DEA94FFDCFFF15796A225FEB75428" box="[149,274,1299,1325]" italics="true" pageId="25" pageNumber="756">C. austriaca</emphasis>
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Laurenti, 1768
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</taxonomicName>
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is more flattened and provided with a somewhat lower neural spine.
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</paragraph>
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<paragraph id="8BC63686FFDCFFF157ADA264FD0957AF" blockId="25.[149,686,1046,1706]" pageId="25" pageNumber="756">
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The genus
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<taxonomicName id="4C794D05FFDCFFF1561CA264FE265469" authorityName="Laurenti" authorityYear="1768" box="[287,387,1362,1388]" class="Reptilia" family="Colubridae" genus="Coronella" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="756" phylum="Chordata" rank="genus">
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<emphasis id="B90DEA94FFDCFFF1561CA264FE265469" box="[287,387,1362,1388]" italics="true" pageId="25" pageNumber="756">Coronella</emphasis>
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</taxonomicName>
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is considered closely related to Old World rat snakes (
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<bibRefCitation id="EFE84B77FFDCFFF156B0A244FF7154A8" author="DOWLING H. G. & DUELLMAN W. E." pageId="25" pageNumber="756" refId="ref16162" refString="DOWLING H. G. & DUELLMAN W. E. 1978. - Systematic Herpetology: A Synopsis of Families and Higher Categories. HISS Publications, New York, 240 p." type="book" year="1978">Dowling & Duellman 1978</bibRefCitation>
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||
;
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||
<bibRefCitation id="EFE84B77FFDCFFF157DCA2A4FE3854A8" author="UTIGER U. & HELFENBERGER N. & SCHATTI B. & SCHMIDT C. & RUF M. & ZISWILER V." box="[223,413,1426,1453]" pageId="25" pageNumber="756" pagination="105 - 124" refId="ref18225" refString="UTIGER U., HELFENBERGER N., SCHATTI B., SCHMIDT C., RUF M. & ZISWILER V. 2002. - Molecular systematics and phylogeny of Old and New World ratsnakes, Elaphe Auct., and related genera (Reptilia, Squamata, Colubridae). Russian Journal of Herpetology 9: 105 - 124." type="journal article" year="2002">
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||
Utiger
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||
<emphasis id="B90DEA94FFDCFFF1562BA2A5FEF954A9" box="[296,348,1426,1452]" italics="true" pageId="25" pageNumber="756">et al.</emphasis>
|
||
2002
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||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE84B77FFDCFFF156A4A2A4FDF854A8" author="NAGY Z. T. & LAWSON R. & JOGER U. & WINK M." box="[423,605,1426,1453]" pageId="25" pageNumber="756" pagination="223 - 233" refId="ref17300" refString="NAGY Z. T., LAWSON R., JOGER U. & WINK M. 2004. - Molecular systematics of racers, whipsnakes and relatives (Reptilia: Colubridae) using mitochondrial and nuclear markers. Journal of Zoological Systematics and Evolutionary Research 42: 223 - 233." type="journal article" year="2004">
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Nagy
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||
<emphasis id="B90DEA94FFDCFFF156E4A2A5FDBF54A9" box="[487,538,1426,1452]" italics="true" pageId="25" pageNumber="756">et al.</emphasis>
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2004
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</bibRefCitation>
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||
). There is some evidences that the ancestry of the above groups could have differentiated in tropical Asia during late Eocene and afterward dispersed among others to the western Palearctic during Oligocene times (
|
||
<bibRefCitation id="EFE84B77FFDCFFF157E3A106FDA5574E" author="BURBRINK F. T. & LAWSON R." box="[224,512,1584,1611]" pageId="25" pageNumber="756" pagination="173 - 189" refId="ref16104" refString="BURBRINK F. T. & LAWSON R. 2007. - How and when did Old World rat snakes disperse into the New World? Molecular Phylogenetics and Evolution 43: 173 - 189." type="journal article" year="2007">Burbrink & Lawson 2007</bibRefCitation>
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||
). As a result the genus
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<taxonomicName id="4C794D05FFDCFFF157DFA166FEE5576F" authorityName="Laurenti" authorityYear="1768" box="[220,320,1616,1642]" class="Reptilia" family="Colubridae" genus="Coronella" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="756" phylum="Chordata" rank="genus">
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<emphasis id="B90DEA94FFDCFFF157DFA166FEE5576F" box="[220,320,1616,1642]" italics="true" pageId="25" pageNumber="756">Coronella</emphasis>
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</taxonomicName>
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could have reached the European continent at an earlier date than considered before (
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<bibRefCitation id="EFE84B77FFDCFFF1579FA1B9FE8E57AF" author="VENCZEL M." box="[156,299,1679,1706]" pageId="25" pageNumber="756" pagination="1 - 22" refId="ref18311" refString="VENCZEL M. 1998. - Late Miocene snakes (Reptilia: Serpentes) from Polgardi (Hungary): a second contribution. Acta Zoologica Cracoviensia 41: 1 - 22." type="journal article" year="1998">Venczel 1998</bibRefCitation>
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||
). Unfortunately, remains of
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<taxonomicName id="4C794D05FFDCFFF15545A1B9FD0057AC" authorityName="Laurenti" authorityYear="1768" box="[582,677,1679,1705]" class="Reptilia" family="Colubridae" genus="Coronella" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="756" phylum="Chordata" rank="genus">
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<emphasis id="B90DEA94FFDCFFF15545A1B9FD0057AC" box="[582,677,1679,1705]" italics="true" pageId="25" pageNumber="756">Coronella</emphasis>
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||
</taxonomicName>
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,
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</paragraph>
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<paragraph id="8BC63686FFDCFFF15794A199FC195534" blockId="25.[151,684,1711,1738]" lastBlockId="25.[712,1247,729,1073]" pageId="25" pageNumber="756">
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except those from a number of Quaternary localities (
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<bibRefCitation id="EFE84B77FFDCFFF155CCA5EFFCCE53F1" author="SZYNDLAR Z." box="[719,875,729,756]" pageId="25" pageNumber="756" pagination="103 - 126" refId="ref18079" refString="SZYNDLAR Z. 1991. - A review of Neogene and Quaternary snakes of Central and Eastern Europe. Part I: Scolecophidia, Boidae, Colubrinae. Estudios Geologicos 47: 103 - 126." type="journal article" year="1991">Szyndlar 1991</bibRefCitation>
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), were rarely reported from Europe. Nevertheless, small sized colubrids, reminiscent of
|
||
<taxonomicName id="4C794D05FFDCFFF155C9A42EFC8B5237" authorityName="Laurenti" authorityYear="1768" box="[714,814,792,818]" class="Reptilia" family="Colubridae" genus="Coronella" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="756" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90DEA94FFDCFFF155C9A42EFC8B5237" box="[714,814,792,818]" italics="true" pageId="25" pageNumber="756">Coronella</emphasis>
|
||
</taxonomicName>
|
||
, were described from a number of European Tertiary localities but referred to other genera (e.g.,
|
||
<taxonomicName id="4C794D05FFDCFFF15409A461FB815276" authority="Holman, 1977" authorityName="Holman" authorityYear="1977" box="[778,1060,855,883]" class="Reptilia" family="Colubridae" genus="Texasophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="756" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90DEA94FFDCFFF15409A461FCD25274" box="[778,887,855,881]" italics="true" pageId="25" pageNumber="756">Texasophis</emphasis>
|
||
Holman, 1977
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C794D05FFDCFFF1535DA461FCC95297" authority="Szyndlar, 1985" authorityName="Szyndlar" authorityYear="1985" class="Reptilia" family="Colubridae" genus="Hispanophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="756" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90DEA94FFDCFFF1535DA461FB7A5274" box="[1118,1247,855,881]" italics="true" pageId="25" pageNumber="756">Hispanophis</emphasis>
|
||
<bibRefCitation id="EFE84B77FFDCFFF155C9A441FCC95297" author="SZYNDLAR Z." box="[714,876,887,914]" pageId="25" pageNumber="756" pagination="447 - 465" refId="ref18010" refString="SZYNDLAR Z. 1985. - Ophidian fauna (Reptilia, Serpentes) from the uppermost Miocene of Algora (Spain). Estudios geologicos 41: 447 - 465." type="journal article" year="1985">Szyndlar, 1985</bibRefCitation>
|
||
</taxonomicName>
|
||
) (
|
||
<bibRefCitation id="EFE84B77FFDCFFF15481A441FBDD5297" author="RAGE J. - C. & HOLMAN J. A." box="[898,1144,887,914]" pageId="25" pageNumber="756" pagination="89 - 104" refId="ref17559" refString="RAGE J. - C. & HOLMAN J. A. 1984. - Des serpents (Reptilia, Squamata) de type nord-americain dans le Miocene francais. Evolution parallele ou dispersion? Geobios 17: 89 - 104." type="journal article" year="1984">Rage & Holman 1984</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE84B77FFDCFFF15381A441FCA052B7" author="SZYNDLAR Z." pageId="25" pageNumber="756" pagination="447 - 465" refId="ref18010" refString="SZYNDLAR Z. 1985. - Ophidian fauna (Reptilia, Serpentes) from the uppermost Miocene of Algora (Spain). Estudios geologicos 41: 447 - 465." type="journal article" year="1985">Szyndlar 1985</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE84B77FFDCFFF15412A4A1FCE952B7" author="SZYNDLAR Z." box="[785,844,919,946]" pageId="25" pageNumber="756" refId="ref18041" refString="SZYNDLAR Z. 1987. - Snakes from the lower Miocene" type="book" year="1987">1987</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE84B77FFDCFFF1545BA4A1FC3752B7" author="SZYNDLAR Z." box="[856,914,919,946]" pageId="25" pageNumber="756" pagination="103 - 126" refId="ref18079" refString="SZYNDLAR Z. 1991. - A review of Neogene and Quaternary snakes of Central and Eastern Europe. Part I: Scolecophidia, Boidae, Colubrinae. Estudios Geologicos 47: 103 - 126." type="journal article" year="1991">1991</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE84B77FFDCFFF1549CA4A1FC7F52B7" author="SZYNDLAR Z." box="[927,986,919,946]" pageId="25" pageNumber="756" pagination="24 - 37" refId="ref18115" refString="SZYNDLAR Z. 1994. - Oligocene snakes of southern Germany. Journal of Vertebrate Paleontology 14: 24 - 37." type="journal article" year="1994">1994</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE84B77FFDCFFF154E5A4A1FB2E52B7" author="GAL E. & HIR J. & KESSLER E. & KOKAY J. & MESZAROS L. & VENCZEL M." box="[998,1163,919,946]" pageId="25" pageNumber="756" pagination="33 - 78" refId="ref16464" refString="GAL E., HIR J., KESSLER E., KOKAY J., MESZAROS L. & VENCZEL M. 1999. - Kozepso-miocen osmaradvanyok, a Matraszolos, Rakoczi-kapolna alatti utbevagasbol I. A Matraszolos 1. Lelohely [= Middle Miocene fossils from the Rakoczi Chapel sections in Matraszolos]. Folia Historico Naturalia Musei Matraensis 23: 33 - 78 (in Hungarian)." type="journal article" year="1999">
|
||
Gál
|
||
<emphasis id="B90DEA94FFDCFFF15316A4AEFBEC52B4" box="[1045,1097,919,945]" italics="true" pageId="25" pageNumber="756">et al.</emphasis>
|
||
1999
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE84B77FFDCFFF15394A4A1FCA252D7" author="IVANOV M." pageId="25" pageNumber="756" pagination="559 - 588" refId="ref17084" refString="IVANOV M. 2000. - Snakes of the lower / middle Miocene transition at Vieux-Collonges (Rhone, France), with comments on the colonization of Western Europe by colubroids. Geodiversitas 22 (4): 559 - 588." type="journal article" year="2000">Ivanov 2000</bibRefCitation>
|
||
). Conceivably, a detailed revision of all these remains should indicate (at least in some cases) a putatively early presence of the genus
|
||
<taxonomicName id="4C794D05FFDCFFF1535CA4C0FB665515" authorityName="Laurenti" authorityYear="1768" box="[1119,1219,1014,1040]" class="Reptilia" family="Colubridae" genus="Coronella" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="756" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90DEA94FFDCFFF1535CA4C0FB665515" box="[1119,1219,1014,1040]" italics="true" pageId="25" pageNumber="756">Coronella</emphasis>
|
||
</taxonomicName>
|
||
in the European Tertiary.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |