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<document id="4E33125347B12BD2EF36AE819E4CAEE1" ID-DOI="10.5281/zenodo.4665621" ID-ISSN="1638-9395" ID-Zenodo-Dep="4665621" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617712844033" checkinUser="felipe" docAuthor="Venczel, Márton &amp; Stiuc, Emanoil" docDate="2008" docId="03D08790FFD7FFFB5649A264FD345390" docLanguage="en" docName="Geodiversitas.30.4.731-763.pdf" docOrigin="Geodiversitas 30 (4)" docStyle="DocumentStyle:9DCE1246056655C2F9CBE9822E831E4E.3:Geodiversitas.2000-2008.journal_article.type3" docStyleId="9DCE1246056655C2F9CBE9822E831E4E" docStyleName="Geodiversitas.2000-2008.journal_article.type3" docStyleVersion="3" docTitle="Hyla arborea" docType="treatment" docVersion="3" lastPageNumber="750" masterDocId="FFE9FFE8FFC5FFE85703A736FFA55105" masterDocTitle="Late middle Miocene amphibians and squamate reptiles from Taut ,, Romania" masterLastPageNumber="763" masterPageNumber="731" pageNumber="749" updateTime="1698901057390" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="337A82DE5192F09A0B2D57511C804AF8">Late middle Miocene amphibians and squamate reptiles from Taut ,, Romania</mods:title>
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<mods:name id="14BD94C4A9B2FD3FA46804BE5900FF56" type="personal">
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<mods:namePart id="1DC485100636C994D4ED0F0648F838E4">Venczel, Márton</mods:namePart>
<mods:affiliation id="7637C9110FC28F094F8D3784C6C63C3F">Tării Crişurilor Museum, B-dul Dacia 1 - 3, RO- 410464 Oradea (Romania) mvenczel @ rdslink. ro</mods:affiliation>
<mods:nameIdentifier id="FC0E6ADC9D3FA4434F0DF4E956645585" type="email">mvenczel@rdslink.ro</mods:nameIdentifier>
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</mods:role>
<mods:namePart id="F72F4B1D18DD6EE6A3FA32586EA8B5FD">Stiuc, Emanoil</mods:namePart>
<mods:affiliation id="5311D693776F7EC4D3EB13D1F85147A6">Speleological Institute “ Emil Racovit, ă ”, Calea 13 Septembrie nr. 13, RO- 050711 Bucureşti (Romania) stiucaemil @ yahoo. com.</mods:affiliation>
<mods:nameIdentifier id="813612BF87DD2283CF589EAA560282FE" type="email">stiucaemil@yahoo.com</mods:nameIdentifier>
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<mods:date id="90033D1F6FDCAFF6A554FD55233639FB">2008</mods:date>
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<treatment id="03D08790FFD7FFFB5649A264FD345390" LSID="urn:lsid:plazi:treatment:03D08790FFD7FFFB5649A264FD345390" httpUri="http://treatment.plazi.org/id/03D08790FFD7FFFB5649A264FD345390" lastPageId="19" lastPageNumber="750" pageId="18" pageNumber="749">
<subSubSection id="C363650DFFD7FFFA5649A264FE4D5488" pageId="18" pageNumber="749" type="nomenclature">
<paragraph id="8BC63686FFD7FFFA5649A264FE4D5488" blockId="18.[330,505,1362,1421]" pageId="18" pageNumber="749">
<heading id="D08E81EAFFD7FFFA5649A264FE4D5488" centered="true" fontSize="11" level="2" pageId="18" pageNumber="749" reason="2">
<taxonomicName id="4C794D05FFD7FFFA5649A264FE5C5468" ID-CoL="3NCKV" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[330,505,1362,1389]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="18" pageNumber="749" phylum="Chordata" rank="species" species="arborea">
<emphasis id="B90DEA94FFD7FFFA5649A264FEDA5468" bold="true" box="[330,383,1362,1389]" italics="true" pageId="18" pageNumber="749">Hyla</emphasis>
cf.
<emphasis id="B90DEA94FFD7FFFA56A5A264FE5C5468" bold="true" box="[422,505,1362,1389]" italics="true" pageId="18" pageNumber="749">arborea</emphasis>
</taxonomicName>
(
<figureCitation id="13422A03FFD7FFFA5667A244FE085488" box="[356,429,1394,1421]" captionStart="FIG" captionStartId="18.[714,725,1008,1025]" captionTargetBox="[738,1228,254,963]" captionTargetId="figure-21@18.[715,1245,233,977]" captionTargetPageId="18" captionText="FIG. 11. — A-C, Hyla cf.arborea; A, ilium (ISER Tt-0411/1);B, same (ISER Tt-0411/2); C, prearticular (ISER Tt-0410);D-G, Pelophylax sp.; D, presacral vertebra (ISER Tt-0421/1); E, humerus (ISER Tt- 0422/1); F, ilium (ISER Tt-0423/1); G, same (ISER Tt-0423/2);A, B, G, left lateral views; C, D, dorsal views; E, ventral view; F, right lateral view. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4665655" httpUri="https://zenodo.org/record/4665655/files/figure.png" pageId="18" pageNumber="749">Fig. 11</figureCitation>
A-C)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C363650DFFD7FFFA5794A29AFE1454DB" pageId="18" pageNumber="749" type="materials_examined">
<paragraph id="8BC63686FFD7FFFA5794A29AFE1454DB" blockId="18.[149,685,1451,1502]" pageId="18" pageNumber="749">
MATERIAL EXAMINED. — One prearticular (
<collectionCode id="ED68AE43FFD7FFFA5518A29DFDEB54C6" box="[539,590,1451,1475]" country="Romania" httpUri="http://grbio.org/cool/911h-bwmh" name="Institutul Speologie Emil G. Racovita" pageId="18" pageNumber="749">ISER</collectionCode>
Tt-0410), five ilia (
<collectionCode id="ED68AE43FFD7FFFA57E9A2F0FE8554DB" box="[234,288,1478,1502]" country="Romania" httpUri="http://grbio.org/cool/911h-bwmh" name="Institutul Speologie Emil G. Racovita" pageId="18" pageNumber="749">ISER</collectionCode>
Tt-0411/1-5).
</paragraph>
</subSubSection>
<subSubSection id="C363650DFFD7FFFA5794A127FBDD5488" pageId="18" pageNumber="749" type="description">
<paragraph id="8BC63686FFD7FFFA5794A127FE8D572C" blockId="18.[150,685,1553,1706]" box="[151,296,1553,1579]" pageId="18" pageNumber="749">DESCRIPTION</paragraph>
<paragraph id="8BC63686FFD7FFFA5794A106FEB6574F" blockId="18.[150,685,1553,1706]" box="[151,275,1584,1610]" pageId="18" pageNumber="749">
<emphasis id="B90DEA94FFD7FFFA5794A106FEB6574F" box="[151,275,1584,1610]" italics="true" pageId="18" pageNumber="749">Prearticular</emphasis>
</paragraph>
<paragraph id="8BC63686FFD7FFFA5794A166FD0857AF" blockId="18.[150,685,1553,1706]" pageId="18" pageNumber="749">The bone is faintly S-shaped with both the anterior and posterior margins broken off. The coronoid process is elongated, relatively narrow and provided with</paragraph>
<paragraph id="8BC63686FFD7FFFA5796A199FD3657CF" blockId="18.[149,659,1711,1738]" box="[149,659,1711,1738]" pageId="18" pageNumber="749">
small tubercles on its medial margin (
<figureCitation id="13422A03FFD7FFFA5527A199FD2257CF" box="[548,647,1711,1738]" captionStart="FIG" captionStartId="18.[714,725,1008,1025]" captionTargetBox="[738,1228,254,963]" captionTargetId="figure-21@18.[715,1245,233,977]" captionTargetPageId="18" captionText="FIG. 11. — A-C, Hyla cf.arborea; A, ilium (ISER Tt-0411/1);B, same (ISER Tt-0411/2); C, prearticular (ISER Tt-0410);D-G, Pelophylax sp.; D, presacral vertebra (ISER Tt-0421/1); E, humerus (ISER Tt- 0422/1); F, ilium (ISER Tt-0423/1); G, same (ISER Tt-0423/2);A, B, G, left lateral views; C, D, dorsal views; E, ventral view; F, right lateral view. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4665655" httpUri="https://zenodo.org/record/4665655/files/figure.png" pageId="18" pageNumber="749">Fig. 11C</figureCitation>
).
</paragraph>
<paragraph id="8BC63686FFD7FFFA55E4A742FB6D5180" blockId="18.[743,1224,115,136]" box="[743,1224,115,136]" pageId="18" pageNumber="749">
Late middle Miocene amphibians and reptiles from Taut,,
<collectingCountry id="F36E7616FFD7FFFA5384A745FB6D5180" box="[1159,1224,115,133]" name="Romania" pageId="18" pageNumber="749">Romania</collectingCountry>
</paragraph>
<paragraph id="8BC63686FFD7FFFA55C9A382FCA155CB" blockId="18.[712,1248,1204,1421]" box="[714,772,1204,1230]" pageId="18" pageNumber="749">
<emphasis id="B90DEA94FFD7FFFA55C9A382FCA155CB" box="[714,772,1204,1230]" italics="true" pageId="18" pageNumber="749">Ilium</emphasis>
</paragraph>
<paragraph id="8BC63686FFD7FFFA55C9A3E2FBDD5488" blockId="18.[712,1248,1204,1421]" pageId="18" pageNumber="749">
The supraacetabular region of the ilium is distinctly smaller than the preacetabular region (
<figureCitation id="13422A03FFD7FFFA5355A3C5FB74540B" box="[1110,1233,1267,1294]" captionStart="FIG" captionStartId="18.[714,725,1008,1025]" captionTargetBox="[738,1228,254,963]" captionTargetId="figure-21@18.[715,1245,233,977]" captionTargetPageId="18" captionText="FIG. 11. — A-C, Hyla cf.arborea; A, ilium (ISER Tt-0411/1);B, same (ISER Tt-0411/2); C, prearticular (ISER Tt-0410);D-G, Pelophylax sp.; D, presacral vertebra (ISER Tt-0421/1); E, humerus (ISER Tt- 0422/1); F, ilium (ISER Tt-0423/1); G, same (ISER Tt-0423/2);A, B, G, left lateral views; C, D, dorsal views; E, ventral view; F, right lateral view. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4665655" httpUri="https://zenodo.org/record/4665655/files/figure.png" pageId="18" pageNumber="749">Fig. 11A, B</figureCitation>
). The tuber superior, of roughly oval shape, is prominent and projects laterally. The iliac shaft is curved ventrally and strongly compressed mediolaterally; the iliac crest is reduced to a faint ridge.
</paragraph>
</subSubSection>
<subSubSection id="C363650DFFD7FFFB55C9A284FD345390" lastPageId="19" lastPageNumber="750" pageId="18" pageNumber="749" type="discussion">
<paragraph id="8BC63686FFD7FFFA55C9A284FC8B54CF" blockId="18.[712,1249,1458,1706]" box="[714,814,1458,1484]" pageId="18" pageNumber="749">REMARKS</paragraph>
<paragraph id="8BC63686FFD7FFFA55C9A2E7FB4457AF" blockId="18.[712,1249,1458,1706]" pageId="18" pageNumber="749">
The assignment of the above remains to
<taxonomicName id="4C794D05FFD7FFFA5368A2E7FCA4570E" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="18" pageNumber="749" phylum="Chordata" rank="species" species="arborea">
<emphasis id="B90DEA94FFD7FFFA5368A2E7FB3854EE" box="[1131,1181,1489,1515]" italics="true" pageId="18" pageNumber="749">Hyla</emphasis>
cf.
<emphasis id="B90DEA94FFD7FFFA53C2A2E4FCA4570E" italics="true" pageId="18" pageNumber="749">arborea</emphasis>
</taxonomicName>
is based on their close resemblance with those of Recent species. On the other hand,
<taxonomicName id="4C794D05FFD7FFFA536DA127FC27574E" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="18" pageNumber="749" phylum="Chordata" rank="species" species="arborea">
<emphasis id="B90DEA94FFD7FFFA536DA127FB45572F" box="[1134,1248,1552,1578]" italics="true" pageId="18" pageNumber="749">H. arborea</emphasis>
(Linnaeus, 1758)
</taxonomicName>
may be easily distinguished from
<taxonomicName id="4C794D05FFD7FFFA55CAA167FBAA576E" authority="Boettger, 1874" authorityName="Boettger" authorityYear="1874" box="[713,1039,1616,1643]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="18" pageNumber="749" phylum="Chordata" rank="species" species="meridionalis">
<emphasis id="B90DEA94FFD7FFFA55CAA167FCCF576F" box="[713,874,1616,1642]" italics="true" pageId="18" pageNumber="749">H. meridionalis</emphasis>
Boettger, 1874
</taxonomicName>
by the morphology of the ilium, in the latter the iliac
<emphasis id="B90DEA94FFD7FFFA534FA159FB7A578C" box="[1100,1247,1647,1673]" italics="true" pageId="18" pageNumber="749">tuber superius</emphasis>
having a rounded shape (
<bibRefCitation id="EFE84B77FFD7FFFA54E6A1B9FB2D57AF" author="HOLMAN J. A." box="[997,1160,1679,1706]" pageId="18" pageNumber="749" pagination="12 - 14" refId="ref16945" refString="HOLMAN J. A. 1992. - Hyla meridionalis from the late Pleistocene (last interglacial age: Ipswichian) of Britain. British Herpetological Society Bulletin 41: 12 - 14." type="journal article" year="1992">Holman 1992</bibRefCitation>
). More-
</paragraph>
<paragraph id="8BC63686FFD7FFFA55C9A199FB7A57CF" blockId="18.[714,1247,1711,1738]" box="[714,1247,1711,1738]" pageId="18" pageNumber="749">
over, the spread of
<taxonomicName id="4C794D05FFD7FFFA548CA186FB9557CC" authorityName="Boettger" authorityYear="1874" box="[911,1072,1711,1737]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="18" pageNumber="749" phylum="Chordata" rank="species" species="meridionalis">
<emphasis id="B90DEA94FFD7FFFA548CA186FB9557CC" box="[911,1072,1711,1737]" italics="true" pageId="18" pageNumber="749">H. meridionalis</emphasis>
</taxonomicName>
in southwestern
</paragraph>
<paragraph id="8BC63686FFD6FFFB57ADA745FEFC5181" blockId="19.[174,345,115,133]" box="[174,345,115,133]" pageId="19" pageNumber="750">Venczel M. &amp; Ştiucă E.</paragraph>
<paragraph id="8BC63686FFD6FFFB5794A7C8FEBF503D" blockId="19.[149,685,254,566]" pageId="19" pageNumber="750">
Europe may have happened rather recently (
<bibRefCitation id="EFE84B77FFD6FFFB5556A7C8FEA8503D" author="RECUERO E. &amp; IRAOLA A. &amp; RUBIO X. &amp; MACHORDOM A. &amp; PARIS M." pageId="19" pageNumber="750" pagination="1207 - 1219" refId="ref17684" refString="RECUERO E., IRAOLA A., RUBIO X., MACHORDOM A. &amp; GARCIA- PARIS M. 2007. - Mitochondrial differentiation and biogeography of Hyla meridionalis (Anura: Hylidae): an unusual phylogeographical pattern. Journal of Biogeography 34: 1207 - 1219." type="journal article" year="2007">
Recuero
<emphasis id="B90DEA94FFD6FFFB5796A628FF645032" box="[149,193,285,311]" italics="true" pageId="19" pageNumber="750">et al</emphasis>
. 2007
</bibRefCitation>
).
</paragraph>
<paragraph id="8BC63686FFD6FFFB57ADA60BFE5F5333" blockId="19.[149,685,254,566]" pageId="19" pageNumber="750">
The oldest fossils referable with some doubts to
<taxonomicName id="4C794D05FFD6FFFB5794A66BFF4E507D" authorityName="Gray" authorityYear="1825" box="[151,235,349,376]" class="Amphibia" family="Hylidae" kingdom="Animalia" order="Anura" pageId="19" pageNumber="750" phylum="Chordata" rank="family">Hylidae</taxonomicName>
are known from the Maastrichtian of
<collectingCountry id="F36E7616FFD6FFFB5576A66BFD08507D" box="[629,685,349,376]" name="India" pageId="19" pageNumber="750">India</collectingCountry>
(
<bibRefCitation id="EFE84B77FFD6FFFB579FA64AFEDF5092" author="PRASAD G. V. R. &amp; RAGE J. - C." box="[156,378,380,407]" pageId="19" pageNumber="750" pagination="95 - 107" refId="ref17352" refString="PRASAD G. V. R. &amp; RAGE J. - C. 1995. - Amphibians and squamates from the Maastrichtian of Naskal, India. Cretaceous Research 16: 95 - 107." type="journal article" year="1995">Prasad &amp; Rage 1995</bibRefCitation>
,
<bibRefCitation id="EFE84B77FFD6FFFB5687A64AFE655092" author="PRASAD G. V. R. &amp; RAGE J. - C." box="[388,448,380,407]" pageId="19" pageNumber="750" pagination="99 - 116" refId="ref17388" refString="PRASAD G. V. R. &amp; RAGE J. - C. 2004. - Fossil frogs (Amphibia: Anura) from the Upper Cretaceous intertrappean beds of Naskal, Andhra Pradesh, India. Revue de Paleobiologie 23: 99 - 116." type="journal article" year="2004">2004</bibRefCitation>
), while the first record of the genus
<taxonomicName id="4C794D05FFD6FFFB5620A6AAFEF050B3" authorityName="Laurenti" authorityYear="1768" box="[291,341,412,438]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="19" pageNumber="750" phylum="Chordata" rank="genus">
<emphasis id="B90DEA94FFD6FFFB5620A6AAFEF050B3" box="[291,341,412,438]" italics="true" pageId="19" pageNumber="750">Hyla</emphasis>
</taxonomicName>
in Europe is from the late early Miocene (MN 4) of
<collectingCountry id="F36E7616FFD6FFFB5671A68AFE1A50D2" box="[370,447,444,471]" name="Austria" pageId="19" pageNumber="750">Austria</collectingCountry>
(
<bibRefCitation id="EFE84B77FFD6FFFB56C8A68AFDC850D2" author="SANCHIZ B." box="[459,621,444,471]" pageId="19" pageNumber="750" pagination="13 - 29" refId="ref17825" refString="SANCHIZ B. 1998 a. - Vertebrates from the early Miocene lignite deposits of the opencast mine Oberdorf (Western Styrian Basin, Austria). 2. Amphibia. Annalen des Naturhistorischen Museums in Wien 99 A: 13 - 29." type="journal article" year="1998">Sanchiz 1998a</bibRefCitation>
). Several fossils from the late middle Miocene (MN 7) of Mátraszőlős,
<collectingCountry id="F36E7616FFD6FFFB5641A6CDFE065313" box="[322,419,507,534]" name="Hungary" pageId="19" pageNumber="750">Hungary</collectingCountry>
were already assigned to
<taxonomicName id="4C794D05FFD6FFFB5794A52AFE8C5330" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[151,297,539,566]" class="Amphibia" family="Hylidae" genus="Hyla" kingdom="Animalia" order="Anura" pageId="19" pageNumber="750" phylum="Chordata" rank="species" species="arborea">
<emphasis id="B90DEA94FFD6FFFB5794A52AFF115330" box="[151,180,540,565]" italics="true" pageId="19" pageNumber="750">H.</emphasis>
cf.
<emphasis id="B90DEA94FFD6FFFB57D8A52DFE8C5330" box="[219,297,539,565]" italics="true" pageId="19" pageNumber="750">arborea</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFE84B77FFD6FFFB5652A52DFE535333" author="VENCZEL M." box="[337,502,539,566]" pageId="19" pageNumber="750" pagination="151 - 174" refId="ref18344" refString="VENCZEL M. 2004. - Middle Miocene anurans from the Carpathian Basin. Palaeontographica abt. A. 271: 151 - 174." type="journal article" year="2004">Venczel (2004)</bibRefCitation>
.
</paragraph>
<paragraph id="8BC63686FFD6FFFB57B1A54CFD345390" blockId="19.[178,657,634,692]" box="[178,657,634,661]" pageId="19" pageNumber="750">
Family
<taxonomicName id="4C794D05FFD6FFFB5601A54CFD345390" authority="Rafinesque-Schmaltz, 1814" authorityName="Rafinesque-Schmaltz" authorityYear="1814" box="[258,657,634,661]" class="Amphibia" family="Ranidae" kingdom="Animalia" order="Anura" pageId="19" pageNumber="750" phylum="Chordata" rank="family">RANIDAE Rafinesque-Schmaltz, 1814</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>