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<document id="CB8CC6DAD272D17D1C7ED0A3D95A546A" ID-DOI="10.5281/zenodo.4665621" ID-ISSN="1638-9395" ID-Zenodo-Dep="4665621" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617712844033" checkinUser="felipe" docAuthor="Venczel, Márton &amp; Stiuc, Emanoil" docDate="2008" docId="03D08790FFC0FFEF5473A385FB71540B" docLanguage="en" docName="Geodiversitas.30.4.731-763.pdf" docOrigin="Geodiversitas 30 (4)" docStyle="DocumentStyle:9DCE1246056655C2F9CBE9822E831E4E.3:Geodiversitas.2000-2008.journal_article.type3" docStyleId="9DCE1246056655C2F9CBE9822E831E4E" docStyleName="Geodiversitas.2000-2008.journal_article.type3" docStyleVersion="3" docTitle="Carpathotriton Venczel 2008" docType="treatment" docVersion="3" lastPageNumber="738" masterDocId="FFE9FFE8FFC5FFE85703A736FFA55105" masterDocTitle="Late middle Miocene amphibians and squamate reptiles from Taut ,, Romania" masterLastPageNumber="763" masterPageNumber="731" pageNumber="736" updateTime="1698901057390" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="F66BC4792FCED090EB8CEBE42920E9E6">Late middle Miocene amphibians and squamate reptiles from Taut ,, Romania</mods:title>
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<mods:name id="99F3FD5AA2A0A5E31F4E9EA9C88088E9" type="personal">
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<mods:namePart id="A289047CDA59E29B0427CBA7C0D3CE55">Venczel, Márton</mods:namePart>
<mods:affiliation id="5505321EF62B4CC01C7D47083DFCE788">Tării Crişurilor Museum, B-dul Dacia 1 - 3, RO- 410464 Oradea (Romania) mvenczel @ rdslink. ro</mods:affiliation>
<mods:nameIdentifier id="08A2ED5BAEA8543A3F17B1FA93E3FB32" type="email">mvenczel@rdslink.ro</mods:nameIdentifier>
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<mods:namePart id="A6BDF6881B459D77E03D92F2AD13D6AD">Stiuc, Emanoil</mods:namePart>
<mods:affiliation id="514A7ACDFA327A0D6CCAEECF9955828D">Speleological Institute “ Emil Racovit, ă ”, Calea 13 Septembrie nr. 13, RO- 050711 Bucureşti (Romania) stiucaemil @ yahoo. com.</mods:affiliation>
<mods:nameIdentifier id="D3FACAD209296CC66C7F1183565E2AC8" type="email">stiucaemil@yahoo.com</mods:nameIdentifier>
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<mods:title id="79909236A861A6F2B71E6272262B2466">Geodiversitas</mods:title>
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<mods:date id="B07FD69F2C8BCE824B1617C08DE298C9">2008</mods:date>
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<treatment id="03D08790FFC0FFEF5473A385FB71540B" LSID="urn:lsid:plazi:treatment:03D08790FFC0FFEF5473A385FB71540B" httpUri="http://treatment.plazi.org/id/03D08790FFC0FFEF5473A385FB71540B" lastPageId="7" lastPageNumber="738" pageId="5" pageNumber="736">
<subSubSection id="C363650DFFC0FFED5473A385FC5855EA" pageId="5" pageNumber="736" type="nomenclature">
<paragraph id="8BC63686FFC0FFED5473A385FC5855EA" blockId="5.[880,1082,1203,1263]" pageId="5" pageNumber="736">
<heading id="D08E81EAFFC0FFED5473A385FC5855EA" centered="true" fontSize="11" level="2" pageId="5" pageNumber="736" reason="2">
<taxonomicName id="4C794D05FFC0FFED5473A385FBB055CB" authorityName="Venczel" authorityYear="2008" box="[880,1045,1203,1230]" class="Amphibia" family="Salamandridae" genus="Carpathotriton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="5" pageNumber="736" phylum="Chordata" rank="genus">
<emphasis id="B90DEA94FFC0FFED5473A385FBB055CB" bold="true" box="[880,1045,1203,1230]" italics="true" pageId="5" pageNumber="736">Carpathotriton</emphasis>
</taxonomicName>
sp. (
<figureCitation id="13422A03FFC0FFED54B6A3E2FC5155EA" box="[949,1012,1236,1263]" captionStart="FIG" captionStartId="6.[151,162,1407,1424]" captionTargetBox="[193,1207,256,1375]" captionTargetId="figure-21@6.[155,1245,218,1395]" captionTargetPageId="6" captionText="FIG. 3. — Carpathotriton sp.: A, frontal (ISER Tt-0330/1); B, B, squamosal (ISER Tt-0331); C, angulo-prearticular (ISER Tt-0333/1); D, dentary (ISER Tt-0332/1); E, E, presacral vertebrae (ISER Tt-0335/1); F, F, same (ISER Tt-0335/2); G, G, humerus (ISER Tt- 0336/1); H, femur (ISER Tt-0338/1); I, same (ISER Tt-0338/2); A, E, G, dorsal views; C, E, F, G, left lateral views; B, right lateral view; B, D, medial views; F, H, I, ventral views. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4665630" httpUri="https://zenodo.org/record/4665630/files/figure.png" pageId="5" pageNumber="736">Fig. 3</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C363650DFFC0FFED55C9A23BFC3354C1" pageId="5" pageNumber="736" type="materials_examined">
<paragraph id="8BC63686FFC0FFED55C9A23BFC3354C1" blockId="5.[712,1249,1293,1476]" pageId="5" pageNumber="736">
MATERIAL EXAMINED. — One frontal (
<collectionCode id="ED68AE43FFC0FFED5340A23BFBDC5420" box="[1091,1145,1293,1317]" country="Romania" httpUri="http://grbio.org/cool/911h-bwmh" name="Institutul Speologie Emil G. Racovita" pageId="5" pageNumber="736">ISER</collectionCode>
Tt-0330), one squamosal (
<collectionCode id="ED68AE43FFC0FFED5461A211FC32543A" box="[866,919,1319,1343]" country="Romania" httpUri="http://grbio.org/cool/911h-bwmh" name="Institutul Speologie Emil G. Racovita" pageId="5" pageNumber="736">ISER</collectionCode>
Tt-0331), six angulo-prearticulars (
<collectionCode id="ED68AE43FFC0FFED55CCA274FCA0545F" box="[719,773,1346,1370]" country="Romania" httpUri="http://grbio.org/cool/911h-bwmh" name="Institutul Speologie Emil G. Racovita" pageId="5" pageNumber="736">ISER</collectionCode>
Tt-0333/1-6), two dentaries (
<collectionCode id="ED68AE43FFC0FFED5334A274FBC8545F" box="[1079,1133,1346,1370]" country="Romania" httpUri="http://grbio.org/cool/911h-bwmh" name="Institutul Speologie Emil G. Racovita" pageId="5" pageNumber="736">ISER</collectionCode>
Tt-0332/1 and 2), six atlases (
<collectionCode id="ED68AE43FFC0FFED5493A26AFC625471" box="[912,967,1372,1396]" country="Romania" httpUri="http://grbio.org/cool/911h-bwmh" name="Institutul Speologie Emil G. Racovita" pageId="5" pageNumber="736">ISER</collectionCode>
Tt-0334/1-6), 35 presacral vertebrae (
<collectionCode id="ED68AE43FFC0FFED543BA241FCCA548A" box="[824,879,1399,1423]" country="Romania" httpUri="http://grbio.org/cool/911h-bwmh" name="Institutul Speologie Emil G. Racovita" pageId="5" pageNumber="736">ISER</collectionCode>
Tt-0335/1-35), four humeri (
<collectionCode id="ED68AE43FFC0FFED53A9A241FB44548A" box="[1194,1249,1399,1423]" country="Romania" httpUri="http://grbio.org/cool/911h-bwmh" name="Institutul Speologie Emil G. Racovita" pageId="5" pageNumber="736">ISER</collectionCode>
Tt-0336/1-4), one ischium (
<collectionCode id="ED68AE43FFC0FFED54D0A2A7FBA254AC" box="[979,1031,1425,1449]" country="Romania" httpUri="http://grbio.org/cool/911h-bwmh" name="Institutul Speologie Emil G. Racovita" pageId="5" pageNumber="736">ISER</collectionCode>
Tt-0337), three femurs (
<collectionCode id="ED68AE43FFC0FFED55CCA29AFCA054C1" box="[719,773,1452,1476]" country="Romania" httpUri="http://grbio.org/cool/911h-bwmh" name="Institutul Speologie Emil G. Racovita" pageId="5" pageNumber="736">ISER</collectionCode>
Tt-0338/1-3).
</paragraph>
</subSubSection>
<subSubSection id="C363650DFFC0FFEF55C9A2C7FCE353D1" lastPageId="7" lastPageNumber="738" pageId="5" pageNumber="736" type="description">
<paragraph id="8BC63686FFC0FFED55C9A2C7FCFE570C" blockId="5.[713,1247,1521,1706]" box="[714,859,1521,1547]" pageId="5" pageNumber="736">DESCRIPTION</paragraph>
<paragraph id="8BC63686FFC0FFED55C9A126FCB3572F" blockId="5.[713,1247,1521,1706]" box="[714,790,1552,1578]" pageId="5" pageNumber="736">
<emphasis id="B90DEA94FFC0FFED55C9A126FCB3572F" box="[714,790,1552,1578]" italics="true" pageId="5" pageNumber="736">Frontal</emphasis>
</paragraph>
<paragraph id="8BC63686FFC0FFED55C9A106FB7B57AF" blockId="5.[713,1247,1521,1706]" pageId="5" pageNumber="736">The dorsal surface is smooth except some sculpture near the supraorbital margin. The anterior portion of the dorsal surface exhibits a groove bordered by two ridges suggesting a contact with the dorsal proc-</paragraph>
<paragraph id="8BC63686FFC0FFED55C9A199FB7A57CF" blockId="5.[714,1247,1711,1738]" box="[714,1247,1711,1738]" pageId="5" pageNumber="736">
ess of the premaxillary (
<figureCitation id="13422A03FFC0FFED54BCA199FBB457CF" box="[959,1041,1711,1738]" captionStart="FIG" captionStartId="6.[151,162,1407,1424]" captionTargetBox="[193,1207,256,1375]" captionTargetId="figure-21@6.[155,1245,218,1395]" captionTargetPageId="6" captionText="FIG. 3. — Carpathotriton sp.: A, frontal (ISER Tt-0330/1); B, B, squamosal (ISER Tt-0331); C, angulo-prearticular (ISER Tt-0333/1); D, dentary (ISER Tt-0332/1); E, E, presacral vertebrae (ISER Tt-0335/1); F, F, same (ISER Tt-0335/2); G, G, humerus (ISER Tt- 0336/1); H, femur (ISER Tt-0338/1); I, same (ISER Tt-0338/2); A, E, G, dorsal views; C, E, F, G, left lateral views; B, right lateral view; B, D, medial views; F, H, I, ventral views. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4665630" httpUri="https://zenodo.org/record/4665630/files/figure.png" pageId="5" pageNumber="736">Fig. 3A</figureCitation>
). The posterolateral
</paragraph>
<paragraph id="8BC63686FFC3FFEE55E4A742FB6D5180" blockId="6.[743,1224,115,136]" box="[743,1224,115,136]" pageId="6" pageNumber="737">
Late middle Miocene amphibians and reptiles from Taut,,
<collectingCountry id="F36E7616FFC3FFEE5384A745FB6D5180" box="[1159,1224,115,133]" name="Romania" pageId="6" pageNumber="737">Romania</collectingCountry>
</paragraph>
<caption id="DF06660EFFC3FFEE5794A249FD2E54D1" ID-DOI="http://doi.org/10.5281/zenodo.4665630" ID-Zenodo-Dep="4665630" httpUri="https://zenodo.org/record/4665630/files/figure.png" pageId="6" pageNumber="737" startId="6.[151,162,1407,1424]" targetBox="[193,1207,256,1375]" targetPageId="6">
<paragraph id="8BC63686FFC3FFEE5794A249FD2E54D1" blockId="6.[151,1249,1407,1492]" pageId="6" pageNumber="737">
FIG. 3. —
<taxonomicName id="4C794D05FFC3FFEE57E9A249FEC75495" authorityName="Venczel" authorityYear="2008" box="[234,354,1407,1424]" class="Amphibia" family="Salamandridae" genus="Carpathotriton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="6" pageNumber="737" phylum="Chordata" rank="genus">
<emphasis id="B90DEA94FFC3FFEE57E9A249FEC75495" box="[234,354,1407,1424]" italics="true" pageId="6" pageNumber="737">Carpathotriton</emphasis>
</taxonomicName>
sp.:
<emphasis id="B90DEA94FFC3FFEE568EA249FE3F5495" bold="true" box="[397,410,1407,1424]" pageId="6" pageNumber="737">A</emphasis>
, frontal (ISER Tt-0330/1);
<emphasis id="B90DEA94FFC3FFEE5575A249FD265495" bold="true" box="[630,643,1407,1424]" pageId="6" pageNumber="737">B</emphasis>
,
<emphasis id="B90DEA94FFC3FFEE558DA249FD055495" bold="true" box="[654,672,1407,1424]" pageId="6" pageNumber="737">B</emphasis>
, squamosal (ISER Tt-0331);
<emphasis id="B90DEA94FFC3FFEE5491A249FC055495" bold="true" box="[914,928,1407,1424]" pageId="6" pageNumber="737">C</emphasis>
, angulo-prearticular (ISER Tt-0333/1);
<emphasis id="B90DEA94FFC3FFEE579BA2A0FF0054A2" bold="true" box="[152,165,1430,1447]" pageId="6" pageNumber="737">D</emphasis>
, dentary (ISER Tt-0332/1);
<emphasis id="B90DEA94FFC3FFEE5693A2A0FE3954A2" bold="true" box="[400,412,1430,1447]" pageId="6" pageNumber="737">E</emphasis>
,
<emphasis id="B90DEA94FFC3FFEE56ABA2A0FE1C54A2" bold="true" box="[424,441,1430,1447]" pageId="6" pageNumber="737">E</emphasis>
, presacral vertebrae (ISER Tt-0335/1);
<emphasis id="B90DEA94FFC3FFEE540BA2A0FCB654A2" bold="true" box="[776,787,1430,1447]" pageId="6" pageNumber="737">F</emphasis>
,
<emphasis id="B90DEA94FFC3FFEE541CA2A0FC8A54A2" bold="true" box="[799,815,1430,1447]" pageId="6" pageNumber="737">F</emphasis>
, same (ISER Tt-0335/2);
<emphasis id="B90DEA94FFC3FFEE530AA2A0FBB254A2" bold="true" box="[1033,1047,1430,1447]" pageId="6" pageNumber="737">G</emphasis>
,
<emphasis id="B90DEA94FFC3FFEE5320A2A0FB9354A2" bold="true" box="[1059,1078,1430,1447]" pageId="6" pageNumber="737">G</emphasis>
, humerus (ISER Tt- 0336/1);
<emphasis id="B90DEA94FFC3FFEE57E3A29AFF4B54B8" bold="true" box="[224,238,1452,1469]" pageId="6" pageNumber="737">H</emphasis>
, femur (ISER Tt-0338/1);
<emphasis id="B90DEA94FFC3FFEE56C3A29AFE6354B8" bold="true" box="[448,454,1452,1469]" pageId="6" pageNumber="737">I</emphasis>
, same (ISER Tt-0338/2);
<emphasis id="B90DEA94FFC3FFEE5597A29AFD0454B8" bold="true" box="[660,673,1452,1469]" pageId="6" pageNumber="737">A</emphasis>
,
<emphasis id="B90DEA94FFC3FFEE55A8A29AFD1254B8" bold="true" box="[683,695,1452,1469]" pageId="6" pageNumber="737">E</emphasis>
,
<emphasis id="B90DEA94FFC3FFEE55C2A29AFD7154B8" bold="true" box="[705,724,1452,1469]" pageId="6" pageNumber="737">G</emphasis>
, dorsal views;
<emphasis id="B90DEA94FFC3FFEE544CA29AFCF854B8" bold="true" box="[847,861,1452,1469]" pageId="6" pageNumber="737">C</emphasis>
,
<emphasis id="B90DEA94FFC3FFEE5465A29AFCD254B8" bold="true" box="[870,887,1452,1469]" pageId="6" pageNumber="737">E</emphasis>
,
<emphasis id="B90DEA94FFC3FFEE5482A29AFC2954B8" bold="true" box="[897,908,1452,1469]" pageId="6" pageNumber="737">F</emphasis>
,
<emphasis id="B90DEA94FFC3FFEE5495A29AFC0154B8" bold="true" box="[918,932,1452,1469]" pageId="6" pageNumber="737">G</emphasis>
, left lateral views;
<emphasis id="B90DEA94FFC3FFEE533FA29AFBEC54B8" bold="true" box="[1084,1097,1452,1469]" pageId="6" pageNumber="737">B</emphasis>
, right lateral view;
<emphasis id="B90DEA94FFC3FFEE5794A2F5FF0C54D1" bold="true" box="[151,169,1475,1492]" pageId="6" pageNumber="737">B</emphasis>
,
<emphasis id="B90DEA94FFC3FFEE57B7A2F5FF6754D1" bold="true" box="[180,194,1475,1492]" pageId="6" pageNumber="737">D</emphasis>
, medial views;
<emphasis id="B90DEA94FFC3FFEE5641A2F5FEF754D1" bold="true" box="[322,338,1475,1492]" pageId="6" pageNumber="737">F</emphasis>
,
<emphasis id="B90DEA94FFC3FFEE565EA2F5FECE54D1" bold="true" box="[349,363,1475,1492]" pageId="6" pageNumber="737">H</emphasis>
,
<emphasis id="B90DEA94FFC3FFEE5676A2F5FEDE54D1" bold="true" box="[373,379,1475,1492]" pageId="6" pageNumber="737">I</emphasis>
, ventral views. Scale bar: 2 mm.
</paragraph>
</caption>
<paragraph id="8BC63686FFC3FFEE5794A127FD0957AF" blockId="6.[151,685,1553,1706]" pageId="6" pageNumber="737">process of the frontal is broken off but we assume that it might have been rather long and formed a complete fronto-squamosal arch. A low crest runs from the medial margin of the posterolateral process toward the posteromedial margin of the frontal</paragraph>
<paragraph id="8BC63686FFC3FFEE5794A199FC9C572F" blockId="6.[151,578,1711,1738]" box="[151,825,1552,1738]" lastBlockId="6.[714,1248,1552,1706]" pageId="6" pageNumber="737">
delimiting a less elevated posterior area.
<emphasis id="B90DEA94FFC3FFEE55C9A126FC9C572F" box="[714,825,1552,1578]" italics="true" pageId="6" pageNumber="737">Squamosal</emphasis>
</paragraph>
<paragraph id="8BC63686FFC3FFEE55C9A106FB4557AF" blockId="6.[714,1248,1552,1706]" pageId="6" pageNumber="737">
The ventral ramus is relatively long, while the anterior ramus is somewhat shorter than the posterior one (
<figureCitation id="13422A03FFC3FFEE5401A146FCF3578E" box="[770,854,1648,1675]" captionStart="FIG" captionStartId="6.[151,162,1407,1424]" captionTargetBox="[193,1207,256,1375]" captionTargetId="figure-21@6.[155,1245,218,1395]" captionTargetPageId="6" captionText="FIG. 3. — Carpathotriton sp.: A, frontal (ISER Tt-0330/1); B, B, squamosal (ISER Tt-0331); C, angulo-prearticular (ISER Tt-0333/1); D, dentary (ISER Tt-0332/1); E, E, presacral vertebrae (ISER Tt-0335/1); F, F, same (ISER Tt-0335/2); G, G, humerus (ISER Tt- 0336/1); H, femur (ISER Tt-0338/1); I, same (ISER Tt-0338/2); A, E, G, dorsal views; C, E, F, G, left lateral views; B, right lateral view; B, D, medial views; F, H, I, ventral views. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4665630" httpUri="https://zenodo.org/record/4665630/files/figure.png" pageId="6" pageNumber="737">Fig. 3B</figureCitation>
, B). The anterior ramus is widened distally for the articulation with the posterolateral
</paragraph>
<paragraph id="8BC63686FFC3FFEE55C9A199FC1057CF" blockId="6.[714,949,1711,1738]" box="[714,949,1711,1738]" pageId="6" pageNumber="737">process of the frontal.</paragraph>
<paragraph id="8BC63686FFC2FFEF57ADA745FEFC5181" blockId="7.[174,345,115,133]" box="[174,345,115,133]" pageId="7" pageNumber="738">Venczel M. &amp; Ştiucă E.</paragraph>
<caption id="DF06660EFFC2FFEF5794A5B1FD2153C0" ID-DOI="http://doi.org/10.5281/zenodo.4665634" ID-Zenodo-Dep="4665634" httpUri="https://zenodo.org/record/4665634/files/figure.png" pageId="7" pageNumber="738" startId="7.[151,161,647,664]" targetBox="[158,675,255,597]" targetPageId="7">
<paragraph id="8BC63686FFC2FFEF5794A5B1FD2153C0" blockId="7.[150,686,646,709]" pageId="7" pageNumber="738">
FIG. 4.—
<taxonomicName id="4C794D05FFC2FFEF57E2A5B0FE905392" authorityName="Bell" authorityYear="1839" box="[225,309,646,663]" class="Amphibia" family="Salamandridae" genus="Lissotriton" kingdom="Animalia" order="Caudata" pageId="7" pageNumber="738" phylum="Chordata" rank="genus">
<emphasis id="B90DEA94FFC2FFEF57E2A5B0FE905392" box="[225,309,646,663]" italics="true" pageId="7" pageNumber="738">Lissotriton</emphasis>
</taxonomicName>
sp.:
<emphasis id="B90DEA94FFC2FFEF5658A5B0FECD5392" bold="true" box="[347,360,646,663]" pageId="7" pageNumber="738">A</emphasis>
, frontal (ISER Tt-0350);
<emphasis id="B90DEA94FFC2FFEF5527A5B0FD945392" bold="true" box="[548,561,646,663]" pageId="7" pageNumber="738">B</emphasis>
, atlas (ISER Tt- 0351);
<emphasis id="B90DEA94FFC2FFEF57CFA5ABFF5253AB" bold="true" box="[204,247,669,687]" pageId="7" pageNumber="738">C -C</emphasis>
, presacral vertebra (ISER Tt-0352/1);
<emphasis id="B90DEA94FFC2FFEF5518A5ABFD8253AB" bold="true" box="[539,551,669,686]" pageId="7" pageNumber="738">A</emphasis>
,
<emphasis id="B90DEA94FFC2FFEF552CA5ABFD9953AB" bold="true" box="[559,572,669,686]" pageId="7" pageNumber="738">C</emphasis>
, dorsal views;
<emphasis id="B90DEA94FFC2FFEF5795A582FF0653C0" bold="true" box="[150,163,692,709]" pageId="7" pageNumber="738">B</emphasis>
,
<emphasis id="B90DEA94FFC2FFEF57ACA582FF6753C0" bold="true" box="[175,194,692,709]" pageId="7" pageNumber="738">C</emphasis>
, ventral views;
<emphasis id="B90DEA94FFC2FFEF5640A582FEFF53C0" bold="true" box="[323,346,692,709]" pageId="7" pageNumber="738">C</emphasis>
, right lateral view. Scale bar: 2 mm.
</paragraph>
</caption>
<paragraph id="8BC63686FFC2FFEF5794A42EFEC75237" blockId="7.[149,685,792,914]" box="[151,354,792,818]" pageId="7" pageNumber="738">
<emphasis id="B90DEA94FFC2FFEF5794A42EFEC75237" box="[151,354,792,818]" italics="true" pageId="7" pageNumber="738">Angulo-prearticular</emphasis>
</paragraph>
<paragraph id="8BC63686FFC2FFEF5794A40EFD355297" blockId="7.[149,685,792,914]" pageId="7" pageNumber="738">
The bone is elongated and provided with a moderately low coronoid process; the lateral flange is low and borders the Meckels groove lingually (
<figureCitation id="13422A03FFC2FFEF552CA441FD265297" box="[559,643,887,914]" captionStart="FIG" captionStartId="6.[151,162,1407,1424]" captionTargetBox="[193,1207,256,1375]" captionTargetId="figure-21@6.[155,1245,218,1395]" captionTargetPageId="6" captionText="FIG. 3. — Carpathotriton sp.: A, frontal (ISER Tt-0330/1); B, B, squamosal (ISER Tt-0331); C, angulo-prearticular (ISER Tt-0333/1); D, dentary (ISER Tt-0332/1); E, E, presacral vertebrae (ISER Tt-0335/1); F, F, same (ISER Tt-0335/2); G, G, humerus (ISER Tt- 0336/1); H, femur (ISER Tt-0338/1); I, same (ISER Tt-0338/2); A, E, G, dorsal views; C, E, F, G, left lateral views; B, right lateral view; B, D, medial views; F, H, I, ventral views. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4665630" httpUri="https://zenodo.org/record/4665630/files/figure.png" pageId="7" pageNumber="738">Fig. 3C</figureCitation>
).
</paragraph>
<paragraph id="8BC63686FFC2FFEF5794A481FF4952D5" blockId="7.[149,686,951,1168]" box="[151,236,951,976]" pageId="7" pageNumber="738">
<emphasis id="B90DEA94FFC2FFEF5794A481FF4952D5" box="[151,236,951,976]" italics="true" pageId="7" pageNumber="738">Dentary</emphasis>
</paragraph>
<paragraph id="8BC63686FFC2FFEF5794A4E0FEA15595" blockId="7.[149,686,951,1168]" pageId="7" pageNumber="738">
The only specimen (ISER Tt-0332/1) is rather fragmentary with the anterior and posterior portions broken off. The dental row is extremely short when compared to the toothless posterior ramus, a feature that is unparalleled in other salamandrids (
<figureCitation id="13422A03FFC2FFEF579CA343FF535595" box="[159,246,1141,1168]" captionStart="FIG" captionStartId="6.[151,162,1407,1424]" captionTargetBox="[193,1207,256,1375]" captionTargetId="figure-21@6.[155,1245,218,1395]" captionTargetPageId="6" captionText="FIG. 3. — Carpathotriton sp.: A, frontal (ISER Tt-0330/1); B, B, squamosal (ISER Tt-0331); C, angulo-prearticular (ISER Tt-0333/1); D, dentary (ISER Tt-0332/1); E, E, presacral vertebrae (ISER Tt-0335/1); F, F, same (ISER Tt-0335/2); G, G, humerus (ISER Tt- 0336/1); H, femur (ISER Tt-0338/1); I, same (ISER Tt-0338/2); A, E, G, dorsal views; C, E, F, G, left lateral views; B, right lateral view; B, D, medial views; F, H, I, ventral views. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4665630" httpUri="https://zenodo.org/record/4665630/files/figure.png" pageId="7" pageNumber="738">Fig. 3D</figureCitation>
).
</paragraph>
<paragraph id="8BC63686FFC2FFEF5794A382FF6F55CB" blockId="7.[149,685,1204,1358]" box="[151,202,1204,1230]" pageId="7" pageNumber="738">
<emphasis id="B90DEA94FFC2FFEF5794A382FF6F55CB" box="[151,202,1204,1230]" italics="true" pageId="7" pageNumber="738">Atlas</emphasis>
</paragraph>
<paragraph id="8BC63686FFC2FFEF5794A3E2FDA1544B" blockId="7.[149,685,1204,1358]" pageId="7" pageNumber="738">In anterior view, the articular surfaces are faintly convex and circular in shape. The odontoid process is relatively wide and slightly concave dorsally; the atlantal cotyle is circular in shape.</paragraph>
<paragraph id="8BC63686FFC2FFEF5794A244FEF35489" blockId="7.[149,686,1394,1706]" box="[151,342,1394,1420]" pageId="7" pageNumber="738">
<emphasis id="B90DEA94FFC2FFEF5794A244FEF35489" box="[151,342,1394,1420]" italics="true" pageId="7" pageNumber="738">Presacral vertebrae</emphasis>
</paragraph>
<paragraph id="8BC63686FFC2FFEF5794A2A4FD0B57AF" blockId="7.[149,686,1394,1706]" pageId="7" pageNumber="738">
All the vertebrae are opisthocoelous and provided with spinal nerve foramina. The neural arch is vaulted and provided with an extremely high neural spine (in most specimens it is completely broken off) (
<figureCitation id="13422A03FFC2FFEF57CCA127FEAA5729" box="[207,271,1553,1580]" captionStart="FIG" captionStartId="6.[151,162,1407,1424]" captionTargetBox="[193,1207,256,1375]" captionTargetId="figure-21@6.[155,1245,218,1395]" captionTargetPageId="6" captionText="FIG. 3. — Carpathotriton sp.: A, frontal (ISER Tt-0330/1); B, B, squamosal (ISER Tt-0331); C, angulo-prearticular (ISER Tt-0333/1); D, dentary (ISER Tt-0332/1); E, E, presacral vertebrae (ISER Tt-0335/1); F, F, same (ISER Tt-0335/2); G, G, humerus (ISER Tt- 0336/1); H, femur (ISER Tt-0338/1); I, same (ISER Tt-0338/2); A, E, G, dorsal views; C, E, F, G, left lateral views; B, right lateral view; B, D, medial views; F, H, I, ventral views. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4665630" httpUri="https://zenodo.org/record/4665630/files/figure.png" pageId="7" pageNumber="738">Fig. 3</figureCitation>
E-F). The subcentral lamina projects ventrolaterally and is connected to the lower rib bearer (= parapophysys); the subcentral foramina are of variable size. The transverse processes are widely spaced while the rib articulating surface is
</paragraph>
<paragraph id="8BC63686FFC2FFEF5794A199FC8F5012" blockId="7.[151,464,1711,1738]" box="[151,810,254,1738]" lastBlockId="7.[712,1248,254,471]" pageId="7" pageNumber="738">
relatively small and rounded.
<emphasis id="B90DEA94FFC2FFEF55C9A7C8FC8F5012" box="[714,810,254,279]" italics="true" pageId="7" pageNumber="738">Humerus</emphasis>
</paragraph>
<paragraph id="8BC63686FFC2FFEF55C9A62BFBE450D2" blockId="7.[712,1248,254,471]" pageId="7" pageNumber="738">
The head of the humerus is moderately widened and connected to a well-defined ventral humeral crest. The dorsal humeral crest is weakly defined forming a triangular bony lamina.The humeral shaft is elongated and slender with the distal extremity moderately widened (
<figureCitation id="13422A03FFC2FFEF54B7A68AFBAF50D2" box="[948,1034,444,471]" captionStart="FIG" captionStartId="6.[151,162,1407,1424]" captionTargetBox="[193,1207,256,1375]" captionTargetId="figure-21@6.[155,1245,218,1395]" captionTargetPageId="6" captionText="FIG. 3. — Carpathotriton sp.: A, frontal (ISER Tt-0330/1); B, B, squamosal (ISER Tt-0331); C, angulo-prearticular (ISER Tt-0333/1); D, dentary (ISER Tt-0332/1); E, E, presacral vertebrae (ISER Tt-0335/1); F, F, same (ISER Tt-0335/2); G, G, humerus (ISER Tt- 0336/1); H, femur (ISER Tt-0338/1); I, same (ISER Tt-0338/2); A, E, G, dorsal views; C, E, F, G, left lateral views; B, right lateral view; B, D, medial views; F, H, I, ventral views. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4665630" httpUri="https://zenodo.org/record/4665630/files/figure.png" pageId="7" pageNumber="738">Fig. 3G</figureCitation>
, G).
</paragraph>
<paragraph id="8BC63686FFC2FFEF55C9A6CAFCA85310" blockId="7.[713,1247,508,724]" box="[714,781,508,533]" pageId="7" pageNumber="738">
<emphasis id="B90DEA94FFC2FFEF55C9A6CAFCA85310" box="[714,781,508,533]" italics="true" pageId="7" pageNumber="738">Femur</emphasis>
</paragraph>
<paragraph id="8BC63686FFC2FFEF55C9A52DFCE353D1" blockId="7.[713,1247,508,724]" pageId="7" pageNumber="738">
In lateral view the bone is slightly sigmoid shaped with a moderately widened femoral head, a rather shallow ventral depression and a relatively short trochanter (
<figureCitation id="13422A03FFC2FFEF544BA54CFC3B5390" box="[840,926,634,661]" captionStart="FIG" captionStartId="6.[151,162,1407,1424]" captionTargetBox="[193,1207,256,1375]" captionTargetId="figure-21@6.[155,1245,218,1395]" captionTargetPageId="6" captionText="FIG. 3. — Carpathotriton sp.: A, frontal (ISER Tt-0330/1); B, B, squamosal (ISER Tt-0331); C, angulo-prearticular (ISER Tt-0333/1); D, dentary (ISER Tt-0332/1); E, E, presacral vertebrae (ISER Tt-0335/1); F, F, same (ISER Tt-0335/2); G, G, humerus (ISER Tt- 0336/1); H, femur (ISER Tt-0338/1); I, same (ISER Tt-0338/2); A, E, G, dorsal views; C, E, F, G, left lateral views; B, right lateral view; B, D, medial views; F, H, I, ventral views. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4665630" httpUri="https://zenodo.org/record/4665630/files/figure.png" pageId="7" pageNumber="738">Fig. 3H</figureCitation>
). The distal part of the femoral shaft is moderately widened and there is a shallow distal fossa.
</paragraph>
</subSubSection>
<subSubSection id="C363650DFFC2FFEF55C9A5CFFB71540B" pageId="7" pageNumber="738" type="discussion">
<paragraph id="8BC63686FFC2FFEF55C9A5CFFC8B5214" blockId="7.[712,1249,761,1294]" box="[714,814,761,787]" pageId="7" pageNumber="738">REMARKS</paragraph>
<paragraph id="8BC63686FFC2FFEF55C9A42EFB71540B" blockId="7.[712,1249,761,1294]" pageId="7" pageNumber="738">
For the first time,
<taxonomicName id="4C794D05FFC2FFEF5478A42EFBAB5237" authorityName="Venczel" authorityYear="2008" box="[891,1038,792,818]" class="Amphibia" family="Salamandridae" genus="Carpathotriton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="7" pageNumber="738" phylum="Chordata" rank="genus">
<emphasis id="B90DEA94FFC2FFEF5478A42EFBAB5237" box="[891,1038,792,818]" italics="true" pageId="7" pageNumber="738">Carpathotriton</emphasis>
</taxonomicName>
was found in the late middle Miocene (MN 7) of Mátraszőlős,
<collectingCountry id="F36E7616FFC2FFEF5365A40EFB605256" box="[1126,1221,824,851]" name="Hungary" pageId="7" pageNumber="738">Hungary</collectingCountry>
; it was then identified erroneously as
<taxonomicName id="4C794D05FFC2FFEF532FA461FCEF5297" authority="von Meyer, 1859" authorityName="von Meyer" authorityYear="1859" class="Amphibia" family="Salamandridae" genus="Archaeotriton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="7" pageNumber="738" phylum="Chordata" rank="genus">
<emphasis id="B90DEA94FFC2FFEF532FA461FB165274" box="[1068,1203,855,881]" italics="true" pageId="7" pageNumber="738">Archaeotriton</emphasis>
von Meyer, 1859
</taxonomicName>
(
<bibRefCitation id="EFE84B77FFC2FFEF5456A441FC4F5297" author="GAL E. &amp; HIR J. &amp; KESSLER E. &amp; KOKAI J. &amp; MESZAROS L. &amp; VENCZEL M." box="[853,1002,887,914]" pageId="7" pageNumber="738" pagination="39 - 75" refId="ref16535" refString="GAL E., HIR J., KESSLER E., KOKAI J., MESZAROS L. &amp; VENCZEL M. 2000. - Kozepso-miocen osmaradvanyok a Matraszolos, Rakoczi-kapolna alatti utbevagasbol. II. A Matrszolos 2. Lelohely [Middle Miocene fossils from the section of the road at the Rakoczi Chapel, Matraszolos. II. LocalityMatraszolos 2. Lelohely]. FoliaHistoricoNaturalia Musei Matraensis 24: 39 - 75 (in Hungarian)." type="journal article" year="2000">
Gál
<emphasis id="B90DEA94FFC2FFEF5483A44EFC0B5294" box="[896,942,887,913]" italics="true" pageId="7" pageNumber="738">et al.</emphasis>
2000
</bibRefCitation>
).The basis of this assignment was its extremely high neural spines of presacral vertebrae. However, a more detailed morphological study and the phylogenetic analysis indicates that it might have belonged to a more advanced stock of salamandrids reminiscent of some recent Asiatic newts (e.g.,
<taxonomicName id="4C794D05FFC2FFEF5406A300FC495555" authority="Tschudi, 1838" authorityName="Tschudi" authorityYear="1838" box="[773,1004,1077,1104]" class="Amphibia" family="Salamandridae" genus="Cynops" kingdom="Animalia" order="Caudata" pageId="7" pageNumber="738" phylum="Chordata" rank="genus">
<emphasis id="B90DEA94FFC2FFEF5406A300FCE95555" box="[773,844,1078,1104]" italics="true" pageId="7" pageNumber="738">Cynops</emphasis>
Tschudi, 1838
</taxonomicName>
,
<taxonomicName id="4C794D05FFC2FFEF54F4A303FCA15575" authority="Boulenger, 1878" authorityName="Boulenger" authorityYear="1878" class="Amphibia" family="Salamandridae" genus="Pachytriton" kingdom="Animalia" order="Caudata" pageId="7" pageNumber="738" phylum="Chordata" rank="genus">
<emphasis id="B90DEA94FFC2FFEF54F4A303FBCD554A" box="[1015,1128,1077,1103]" italics="true" pageId="7" pageNumber="738">Pachytriton</emphasis>
Boulenger, 1878
</taxonomicName>
,
<taxonomicName id="4C794D05FFC2FFEF5413A360FB9A5575" authority="Chang, 1935" authorityName="Chang" authorityYear="1935" box="[784,1087,1109,1136]" class="Amphibia" family="Salamandridae" genus="Paramesotriton" kingdom="Animalia" order="Caudata" pageId="7" pageNumber="738" phylum="Chordata" rank="genus">
<emphasis id="B90DEA94FFC2FFEF5413A360FC02556A" box="[784,935,1110,1135]" italics="true" pageId="7" pageNumber="738">Paramesotriton</emphasis>
Chang, 1935
</taxonomicName>
). Furthermore, some features (e.g.,the extremely high neural spines of the trunk vertebrae,limb bones only slightly widened distally, long toothless posterior ramus of dentary) indicate that it might have had a long aquatic phase and a peculiar feeding mechanism (
<bibRefCitation id="EFE84B77FFC2FFEF5334A3C5FB6D540B" author="VENCZEL M." box="[1079,1224,1267,1294]" pageId="7" pageNumber="738" pagination="41 - 59" refId="ref18421" refString="VENCZEL M. 2008. - A new salamandrid amphibian from the Middle Miocene of Hungary and its phylogenetic relationships. Journal of Systematic Palaeontology 6: 41 - 59." type="journal article" year="2008">Venczel 2008</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>