treatments-xml/data/03/B5/68/03B5686E4F74FFC5DFCEEDD2FA7A85AC.xml

<document id="877BFCE8AE6EC0DBFB78C834FC8E89CF" ID-CLB-Dataset="27001" ID-DOI="10.11646/zootaxa.4586.3.7" ID-GBIF-Dataset="7e67aad1-4b75-4cdd-b9a9-e2950ff82bcb" ID-ISSN="1175-5326" ID-Zenodo-Dep="2647308" ID-ZooBank="A45BD48D-446F-4819-AF73-40FC4409C9F5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1555665207516" checkinUser="plazi" docAuthor="Dow, Rory A. &amp; Ngiam, In. W. J." docDate="2019" docId="03B5686E4F74FFC5DFCEEDD2FA7A85AC" docLanguage="en" docName="zootaxa.4586.3.7.pdf" docOrigin="Zootaxa 4586 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Podolestes parvus Dow &amp; Ngiam 2019, sp. nov." docType="treatment" docVersion="7" lastPageNumber="514" masterDocId="FF8C10164F76FFCCDF59E97CFFE1847C" masterDocTitle="A new species of Podolestes Selys, 1862 from peat swamp forest in Sarawak (Odonata: Zygoptera: Argiolestidae)" masterLastPageNumber="516" masterPageNumber="505" pageNumber="507" updateTime="1698723294620" updateUser="ExternalLinkService">
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<mods:title id="3E1C417D85E07291DBC6343CD371F1E0">A new species of Podolestes Selys, 1862 from peat swamp forest in Sarawak (Odonata: Zygoptera: Argiolestidae)</mods:title>
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<mods:namePart id="F00FE98304363EBCA13E18F95F2FB2A3">Dow, Rory A.</mods:namePart>
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<mods:namePart id="D64DC812FB8765A460D696E98B95D512">Ngiam, In. W. J.</mods:namePart>
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<mods:date id="E1060B418F5FBA94F7E5C22E31834327">2019</mods:date>
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<subSubSection id="C3068AF34F74FFCEDFCEEDD2FE3580B4" box="[151,468,1198,1224]" pageId="2" pageNumber="507" type="nomenclature">
<paragraph id="8BA3D9784F74FFCEDFCEEDD2FE3580B4" blockId="2.[151,1162,1198,1257]" box="[151,468,1198,1224]" pageId="2" pageNumber="507">
<heading id="D0EB6E144F74FFCEDFCEEDD2FE3580B4" bold="true" box="[151,468,1198,1224]" fontSize="11" level="1" pageId="2" pageNumber="507" reason="1">
<emphasis id="B968056A4F74FFCEDFCEEDD2FE3580B4" bold="true" box="[151,468,1198,1224]" pageId="2" pageNumber="507">
<taxonomicName id="4C1CA2FB4F74FFCEDFCEEDD2FE8F80B4" authority="Dow &amp; Ngiam, 2019" authorityName="Dow &amp; Ngiam" authorityYear="2019" box="[151,366,1198,1224]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="507" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
<emphasis id="B968056A4F74FFCEDFCEEDD2FE8F80B4" bold="true" box="[151,366,1198,1224]" italics="true" pageId="2" pageNumber="507">Podolestes parvus</emphasis>
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<taxonomicNameLabel id="A25BB8114F74FFCEDE2FEDD2FE3580B4" box="[374,468,1198,1224]" pageId="2" pageNumber="507" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3068AF34F74FFCEDFCEEDACFBAE8153" pageId="2" pageNumber="507" type="description">
<paragraph id="8BA3D9784F74FFCEDFCEEDACFB6B8095" blockId="2.[151,1162,1198,1257]" box="[151,1162,1232,1257]" pageId="2" pageNumber="507">
(
<figureCitation id="1327C5FD4F74FFCEDFC6EDACFEE28095" box="[159,259,1232,1257]" captionStart="FIGURES 1–4" captionStartId="1.[151,264,1826,1849]" captionTargetBox="[151,1435,878,1805]" captionTargetId="figure@1.[151,1435,878,1805]" captionTargetPageId="1" captionText="FIGURES 1–4. Podolestes species males, head. P. atomarius (KAL12_MEG28): (1) dorsal; (3) frontal. P. parvus sp. nov. holotype: (2) dorsal; (4) dorsal-frontal." httpUri="https://zenodo.org/record/2647310/files/figure.png" pageId="2" pageNumber="507">Figs 2, 4</figureCitation>
, 
<figureCitation id="1327C5FD4F74FFCEDE56EDACFEDA8095" box="[271,315,1232,1257]" captionStart="FIGURES 5–8" captionStartId="2.[151,264,1069,1092]" captionTargetBox="[151,1435,193,1049]" captionTargetId="figure@2.[151,1435,193,1049]" captionTargetPageId="2" captionText="FIGURES 5–8. Podolestes species females, head. P. atomarius (KAL12_MEG38): (5) dorsal; (7) frontal. P. parvus sp. nov. paratype (SAR15_ARG4): (6) dorsal; (8) frontal." httpUri="https://zenodo.org/record/2647312/files/figure.png" pageId="2" pageNumber="507">6, 8</figureCitation>
, 
<figureCitation id="1327C5FD4F74FFCEDE1EEDADFE6F8095" box="[327,398,1233,1257]" captionStart="FIGURES 9–12" captionStartId="3.[151,264,1454,1477]" captionTargetBox="[151,1435,193,1434]" captionTargetId="figure@3.[151,1435,193,1434]" captionTargetPageId="3" captionText="FIGURES 9–12. Podolestes species, prothorax dorsal view. P. atomarius: (9) male (KAL12_MEG28); (10) female (KAL12_MEG38). P. parvus sp. nov. (11) holotype; (12) female paratype (SAR15_ARG4)." httpUri="https://zenodo.org/record/2647314/files/figure.png" pageId="2" pageNumber="507">11, 12</figureCitation>
, 
<figureCitation id="1327C5FD4F74FFCEDEC3EDACFE038095" box="[410,482,1232,1257]" captionStart="FIGURES 13–16" captionStartId="4.[151,264,1790,1813]" captionTargetBox="[151,1435,628,1769]" captionTargetId="figure@4.[151,1435,628,1769]" captionTargetPageId="4" captionText="FIGURES 13–16. Podolestes species, prothorax lateral view. P. atomarius: (13) male (KAL12_MEG28); (14) female (KAL12_MEG38). P. parvus sp. nov. (15) holotype; (16) female paratype (SAR15_ARG4)." httpUri="https://zenodo.org/record/2647316/files/figure.png" pageId="2" pageNumber="507">15, 16</figureCitation>
, 
<figureCitation id="1327C5FD4F74FFCEDEB7EDADFDD78095" box="[494,566,1233,1257]" captionStart="FIGURES 17–20" captionStartId="5.[151,264,1357,1380]" captionTargetBox="[151,1435,481,1337]" captionTargetId="figure@5.[151,1435,481,1337]" captionTargetPageId="5" captionText="FIGURES 17–20. Podolestes species, synthorax dorsal view. P. atomarius: (17) male (KAL12_MEG28); (18) female (KAL12_MEG38). P. parvus sp. nov. (19) holotype; (20) female paratype (SAR15_ARG4)." httpUri="https://zenodo.org/record/2647318/files/figure.png" pageId="2" pageNumber="507">19, 20</figureCitation>
, 
<figureCitation id="1327C5FD4F74FFCEDD1BEDADFD6A8095" box="[578,651,1233,1257]" captionStart="FIGURES 21–24" captionStartId="6.[151,264,1212,1235]" captionTargetBox="[151,1435,193,1192]" captionTargetId="figure@6.[151,1435,193,1192]" captionTargetPageId="6" captionText="FIGURES 21–24. Podolestes species, synthorax lateral view. P. atomarius: (21) male (KAL12_MEG28); (22) female (KAL12_MEG38). P. parvus sp. nov. (23) holotype; (24) female paratype (SAR15_ARG4)." httpUri="https://zenodo.org/record/2647320/files/figure.png" pageId="2" pageNumber="507">23, 24</figureCitation>
, 
<figureCitation id="1327C5FD4F74FFCEDDCEEDADFD548095" box="[663,693,1233,1257]" captionStart="FIGURES 25–28" captionStartId="6.[151,264,1978,2001]" captionTargetBox="[151,1435,1316,1958]" captionTargetId="figure@6.[151,1435,1316,1958]" captionTargetPageId="6" captionText="FIGURES 25–28. Pterostigma of Podolestes species. P. atomarius: (25) male Fw (KAL12_MEG28); (26) male Hw (KAL12_MEG28); (27) female Fw (KAL12_MEG38). (28) P. parvus female (SAR15_ARG4) Hw." httpUri="https://zenodo.org/record/2647322/files/figure.png" pageId="2" pageNumber="507">28</figureCitation>
, 
<figureCitation id="1327C5FD4F74FFCEDD99EDADFCE98095" box="[704,776,1233,1257]" captionStart="FIGURES 29–32" captionStartId="7.[151,264,1212,1235]" captionTargetBox="[151,1435,193,1192]" captionTargetId="figure@7.[151,1435,193,1192]" captionTargetPageId="7" captionText="FIGURES 29–32. Genital ligula of Podolestes species. P. atomarius (KAL12_MEG1): (29) ventral; (31) lateral. P. parvus sp. nov. holotype: (30) ventral; (32) lateral. All scale bars 0.2mm." httpUri="https://zenodo.org/record/2647328/files/figure.png" pageId="2" pageNumber="507">30, 32</figureCitation>
, 
<figureCitation id="1327C5FD4F74FFCEDC4DEDACFCBC8095" box="[788,861,1232,1257]" captionStart="FIGURES 33–36" captionStartId="7.[151,264,1978,2001]" captionTargetBox="[151,1435,1316,1958]" captionTargetId="figure@7.[151,1435,1316,1958]" captionTargetPageId="7" captionText="FIGURES 33–36. Podolestes species, markings of terminal abdominal segments, dorsal view. P. atomarius: (33) male (KAL12_MEG28); (34) female (KAL12_MEG38). P. parvus sp. nov. (35) holotype; (36) female paratype (SAR15_ARG4)." httpUri="https://zenodo.org/record/2647330/files/figure.png" pageId="2" pageNumber="507">35, 36</figureCitation>
, 
<figureCitation id="1327C5FD4F74FFCEDC31EDADFC518095" box="[872,944,1233,1257]" captionStart="FIGURES 37–40" captionStartId="8.[151,264,1426,1449]" captionTargetBox="[151,1435,193,1405]" captionTargetId="figure@8.[151,1435,193,1405]" captionTargetPageId="8" captionText="FIGURES 37–40. Anal appendages of Podolestes species males. P. atomarius (KAL12_MEG2): (37) dorsal; (39) lateral. P. parvus sp. nov. holotype: (38) dorsal; (40) lateral. All scale bars 0.5mm." httpUri="https://zenodo.org/record/2647332/files/figure.png" pageId="2" pageNumber="507">38, 40</figureCitation>
, 
<figureCitation id="1327C5FD4F74FFCEDCE5EDADFC3B8095" box="[956,986,1233,1257]" captionStart="FIGURE 41" captionStartId="9.[151,250,1912,1934]" captionTargetBox="[151,1435,519,1890]" captionTargetId="figure@9.[151,1435,519,1890]" captionTargetPageId="9" captionText="FIGURE 41. Distribution of Podolestes species. Yellow circles: P. atomarius; red squares: P. parvus." httpUri="https://zenodo.org/record/2647334/files/figure.png" pageId="2" pageNumber="507">41</figureCitation>
, 
<figureCitation id="1327C5FD4F74FFCEDCBFEDADFB608095" box="[998,1153,1232,1257]" captionStart="FIGURES 42–45" captionStartId="10.[151,264,962,985]" captionTargetBox="[151,1435,193,942]" captionTargetId="figure@10.[151,1435,193,942]" captionTargetPageId="10" captionText="FIGURES 42–45. Podolestes parvus sp. nov. in life. Holotype male: (42) whole animal in lateral view; (44) head and thorax in dorsal view. Female paratype (SAR11_12_MEG95): (43) whole animal in lateral view; (45) head and thorax in dorsal view. Photographs by R.W.J. Ngiam." httpUri="https://zenodo.org/record/2647336/files/figure.png" pageId="2" pageNumber="507">42, 43, 44, 45</figureCitation>
)
</paragraph>
<paragraph id="8BA3D9784F74FFCEDFCEEC65FBAE8153" blockId="2.[151,1103,1305,1327]" box="[151,1103,1305,1327]" pageId="2" pageNumber="507">
<taxonomicName id="4C1CA2FB4F74FFCEDFCEEC65FEE48153" authorityName="Selys" authorityYear="1862" box="[151,261,1305,1327]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="507" phylum="Arthropoda" rank="genus">
<emphasis id="B968056A4F74FFCEDFCEEC65FEE48153" box="[151,261,1305,1327]" italics="true" pageId="2" pageNumber="507">Podolestes</emphasis>
</taxonomicName>
new species;—Dow 
<emphasis id="B968056A4F74FFCEDE85EC66FDEE8153" box="[476,527,1305,1327]" italics="true" pageId="2" pageNumber="507">et al.</emphasis>
2015a: 6767, Image 5 (
<specimenCount id="9D1A12F14F74FFCEDC58EC65FCF78153" box="[769,790,1305,1327]" pageId="2" pageNumber="507" type="male">♂</specimenCount>
, 
<specimenCount id="9D1A12F14F74FFCEDC79EC66FCA78153" box="[800,838,1306,1327]" pageId="2" pageNumber="507" type="female">♀♀</specimenCount>
Maludam National Park).
</paragraph>
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<paragraph id="8BA3D9784F74FFCEDFCEEC27FEBB81C0" blockId="2.[151,1437,1371,2008]" pageId="2" pageNumber="507">
<materialsCitation id="3B74D3254F74FFCEDFCEEC27FEBB81C0" ID-GBIF-Occurrence="2236620144" collectingDate="2012-07-10" collectionCode="RMNH" collectorName="R. A. Dow" country="Malaysia" latitude="1.6188" location="Maludam National Park" longLatPrecision="7" longitude="111.0584" municipality="Betong Division" pageId="2" pageNumber="507" specimenCode="MEG79" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B968056A4F74FFCEDFCEEC27FEEA8108" bold="true" box="[151,267,1371,1396]" pageId="2" pageNumber="507">
<typeStatus id="54A767DA4F74FFCEDFCEEC27FEE78108" box="[151,262,1371,1396]" pageId="2" pageNumber="507" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="9D1A12F14F74FFCEDE4BEC20FEDD810F" box="[274,316,1372,1396]" pageId="2" pageNumber="507" type="male">1 ♂</specimenCount>
(SAR11_12_ 
<specimenCode id="DBBA71034F74FFCEDE8CEC20FDD58108" box="[469,564,1372,1396]" collectionCode="MEG" pageId="2" pageNumber="507">MEG79</specimenCode>
), perched low on stick over hole, peat swamp forest, 
<location id="8EC38FA34F74FFCEDBD1EC27FA798108" LSID="urn:lsid:plazi:treatment:03B5686E4F74FFC5DFCEEDD2FA7A85AC:8EC38FA34F74FFCEDBD1EC27FA798108" box="[1160,1432,1371,1396]" country="Malaysia" latitude="1.6188" longLatPrecision="7" longitude="111.0584" municipality="Betong Division" name="Maludam National Park" pageId="2" pageNumber="507">Maludam National Park</location>
, 
<collectingMunicipality id="6BC743024F74FFCEDFCEECFCFEB581E4" box="[151,340,1407,1432]" pageId="2" pageNumber="507">Betong Division</collectingMunicipality>
, 
<collectingCountry id="F30B99E84F74FFCEDE38EC03FE2581E4" box="[353,452,1407,1432]" name="Malaysia" pageId="2" pageNumber="507">Sarawak</collectingCountry>
, 
<collectingCountry id="F30B99E84F74FFCEDE8BEC03FDDD81E4" box="[466,572,1407,1432]" name="Malaysia" pageId="2" pageNumber="507">Malaysia</collectingCountry>
, 
<geoCoordinate id="EE28BFBF4F74FFCEDD10EC03FD4C81E4" box="[585,685,1407,1432]" direction="north" orientation="latitude" pageId="2" pageNumber="507" precision="5" value="1.6188">1.6188N</geoCoordinate>
, 
<geoCoordinate id="EE28BFBF4F74FFCEDDE3EC03FCD281E4" box="[698,819,1407,1432]" direction="east" orientation="longitude" pageId="2" pageNumber="507" precision="5" value="111.0584">111.0584E</geoCoordinate>
(approx. coordinates), 
<date id="FFA2FFB84F74FFCEDB64ECFCFB2581E4" box="[1085,1220,1407,1432]" pageId="2" pageNumber="507" value="2012-07-10">
<collectingDate id="EFE606504F74FFCEDB64ECFCFB2581E4" box="[1085,1220,1407,1432]" pageId="2" pageNumber="507" value="2012-07-10">10 vii 2012</collectingDate>
</date>
, leg. 
<collectorName id="26E9BCAE4F74FFCEDA5DECFCFA9881EB" box="[1284,1401,1408,1432]" pageId="2" pageNumber="507">R.A. Dow</collectorName>
, in 
<collectionCode id="ED0D41BD4F74FFCEDFCEECD8FF0A81C7" box="[151,235,1444,1467]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="2" pageNumber="507" type="Museum">RMNH</collectionCode>
(Leiden).
</materialsCitation>
</paragraph>
<paragraph id="8BA3D9784F74FFCEDF9EECBBFCC18283" blockId="2.[151,1437,1371,2008]" pageId="2" pageNumber="507">
<materialsCitation id="3B74D3254F74FFCEDF9EECBBFAC78278" ID-GBIF-Occurrence="2236620141" collectingDate="2012-07-07" collectionCode="MEG" collectorName="In" country="Malaysia" location="Maludam National Park" municipality="Betong Division" pageId="2" pageNumber="507" specimenCode="MEG95" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<emphasis id="B968056A4F74FFCEDF9EECBBFEA7819C" bold="true" box="[199,326,1479,1504]" pageId="2" pageNumber="507">
<typeStatus id="54A767DA4F74FFCEDF9EECBBFEA0819C" box="[199,321,1479,1504]" pageId="2" pageNumber="507" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectorName id="26E9BCAE4F74FFCEDE08ECB4FE8981A3" box="[337,360,1480,1503]" pageId="2" pageNumber="507">In</collectorName>
coll. Dow unless stated otherwise, all from 
<collectingCountry id="F30B99E84F74FFCEDC21ECBBFC3A819C" box="[888,987,1479,1504]" name="Malaysia" pageId="2" pageNumber="507">Sarawak</collectingCountry>
, 
<collectingCountry id="F30B99E84F74FFCEDCB2ECBBFBB7819C" box="[1003,1110,1479,1504]" name="Malaysia" pageId="2" pageNumber="507">Malaysia</collectingCountry>
: 
<specimenCount id="9D1A12F14F74FFCEDB3CECB4FB7581A3" box="[1125,1172,1480,1504]" pageId="2" pageNumber="507" type="female">1 ♀</specimenCount>
(SAR11_12_ 
<specimenCode id="DBBA71034F74FFCEDA68ECBBFA71819C" box="[1329,1424,1479,1504]" collectionCode="MEG" pageId="2" pageNumber="507">MEG95</specimenCode>
), perched by a track through peat swamp forest, 
<location id="8EC38FA34F74FFCEDDE2EC97FC308278" LSID="urn:lsid:plazi:treatment:03B5686E4F74FFC5DFCEEDD2FA7A85AC:8EC38FA34F74FFCEDDE2EC97FC308278" box="[699,977,1515,1540]" country="Malaysia" municipality="Betong Division" name="Maludam National Park" pageId="2" pageNumber="507">Maludam National Park</location>
, 
<collectingMunicipality id="6BC743024F74FFCEDC86EC90FB7F8278" box="[991,1182,1515,1540]" pageId="2" pageNumber="507">Betong Division</collectingMunicipality>
, 
<date id="FFA2FFB84F74FFCEDBF5EC90FAC78278" box="[1196,1318,1515,1540]" pageId="2" pageNumber="507" value="2012-07-07">
<collectingDate id="EFE606504F74FFCEDBF5EC90FAC78278" box="[1196,1318,1515,1540]" pageId="2" pageNumber="507" value="2012-07-07">7 vii 2012</collectingDate>
</date>
</materialsCitation>
, 
<materialsCitation id="3B74D3254F74FFCEDA6DEC97FD728230" ID-GBIF-Occurrence="2236620139" collectingDate="2015-06-30" collectionCode="RMNH" collectorName="R. A. Dow" country="Malaysia" location="Maludam National Park" municipality="Betong Division" pageId="2" pageNumber="507" specimenCode="MEG96, SAR15, ARG3" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
leg. 
<collectorName id="26E9BCAE4F74FFCEDA3EEC90FF30825B" pageId="2" pageNumber="507">R.A. Dow</collectorName>
; 
<specimenCount id="9D1A12F14F74FFCEDFB9EF6CFEEC825B" box="[224,269,1552,1576]" pageId="2" pageNumber="507" type="female">1 ♀</specimenCount>
(SAR11_12_ 
<specimenCode id="DBBA71034F74FFCEDEF0EF73FDE88254" box="[425,521,1551,1576]" collectionCode="MEG" pageId="2" pageNumber="507">MEG96</specimenCode>
; 
<collectionCode id="ED0D41BD4F74FFCEDD4EEF73FD908254" box="[535,625,1551,1576]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="2" pageNumber="507" type="Museum">RMNH</collectionCode>
.INS.506881), same data but in 
<collectionCode id="ED0D41BD4F74FFCEDCB8EF6CFBD8825B" box="[993,1081,1552,1575]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="2" pageNumber="507" type="Museum">RMNH</collectionCode>
; 
<specimenCount id="9D1A12F14F74FFCEDB1EEF6CFB94825B" box="[1095,1141,1552,1576]" pageId="2" pageNumber="507" type="female">1 ♀</specimenCount>
(
<specimenCode id="DBBA71034F74FFCEDBDEEF73FB3A8254" box="[1159,1243,1551,1576]" collectionCode="SAR" country="Malaysia" lsid="urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="2" pageNumber="507" type="Herbarium">SAR15</specimenCode>
_ 
<specimenCode id="DBBA71034F74FFCEDBB3EF73FAD78254" box="[1258,1334,1551,1576]" collectionCode="ARG" country="Malta" lsid="urn:lsid:biocol.org:col:12605" name="Argotti Botanic Garden" pageId="2" pageNumber="507" type="Herbarium">ARG3</specimenCode>
), in peat swamp forest, same national park, 
<date id="FFA2FFB84F74FFCEDD40EF48FD728230" box="[537,659,1587,1612]" pageId="2" pageNumber="507" value="2015-06-30">
<collectingDate id="EFE606504F74FFCEDD40EF48FD728230" box="[537,659,1587,1612]" pageId="2" pageNumber="507" value="2015-06-30">30 vi 2015</collectingDate>
</date>
</materialsCitation>
, 
<materialsCitation id="3B74D3254F74FFCEDDC4EF4FFAA68230" ID-GBIF-Occurrence="2236620140" box="[669,1351,1587,1612]" collectingDate="2015-06-30" collectionCode="SAR, ARG" collectorName="R. A. Dow" country="Malaysia" location="Maludam National Park" municipality="Betong Division" pageId="2" pageNumber="507" specimenCode="SAR15, ARG4" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
leg. 
<collectorName id="26E9BCAE4F74FFCEDD94EF48FCA38237" box="[717,834,1588,1612]" pageId="2" pageNumber="507">R.A. Dow</collectorName>
; 
<specimenCount id="9D1A12F14F74FFCEDC14EF48FC978237" box="[845,886,1588,1612]" pageId="2" pageNumber="507" type="female">1 ♀</specimenCount>
(
<specimenCode id="DBBA71034F74FFCEDCDDEF4FFC368230" box="[900,983,1587,1612]" collectionCode="SAR" country="Malaysia" lsid="urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="2" pageNumber="507" type="Herbarium">SAR15</specimenCode>
_ 
<specimenCode id="DBBA71034F74FFCEDCBFEF4FFBD58230" box="[998,1076,1587,1612]" collectionCode="ARG" country="Malta" lsid="urn:lsid:biocol.org:col:12605" name="Argotti Botanic Garden" pageId="2" pageNumber="507" type="Herbarium">ARG4</specimenCode>
), same location and date
</materialsCitation>
, 
<materialsCitation id="3B74D3254F74FFCEDA08EF4FFB4182E8" ID-GBIF-Occurrence="2236620145" collectingDate="2015-01-30" collectionCode="SAR, ARG" collectorName="H. Hasbi" country="Malaysia" latitude="1.4016" location="Ulu Sebuyau National Park" longLatPrecision="7" longitude="110.9325" municipality="Samarahan Division" pageId="2" pageNumber="507" specimenCode="SAR15, ARG1" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
leg. 
<collectorName id="26E9BCAE4F74FFCEDAD8EF48FF39820C" pageId="2" pageNumber="507">H. Hasbi</collectorName>
&amp; L. Ului; 
<specimenCount id="9D1A12F14F74FFCEDE3DEF24FE718213" box="[356,400,1624,1648]" pageId="2" pageNumber="507" type="male">1 ♂</specimenCount>
(
<specimenCode id="DBBA71034F74FFCEDEF8EF2BFE17820C" box="[417,502,1623,1648]" collectionCode="SAR" country="Malaysia" lsid="urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="2" pageNumber="507" type="Herbarium">SAR15</specimenCode>
_ 
<specimenCode id="DBBA71034F74FFCEDD5CEF2BFDB1820C" box="[517,592,1623,1648]" collectionCode="ARG" country="Malta" lsid="urn:lsid:biocol.org:col:12605" name="Argotti Botanic Garden" pageId="2" pageNumber="507" type="Herbarium">ARG1</specimenCode>
), on transect between 2 and 
<quantity id="4CE4749D4F74FFCEDCC0EF24FC35820C" box="[921,980,1623,1648]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="507" unit="km" value="3.0">3 km</quantity>
inside peat swamp forest, 
<location id="8EC38FA34F74FFCEDA50EF2BFED782E8" LSID="urn:lsid:plazi:treatment:03B5686E4F74FFC5DFCEEDD2FA7A85AC:8EC38FA34F74FFCEDA50EF2BFED782E8" country="Malaysia" latitude="1.4016" longLatPrecision="7" longitude="110.9325" municipality="Samarahan Division" name="Ulu Sebuyau National Park" pageId="2" pageNumber="507">Ulu Sebuyau National Park</location>
, 
<collectingMunicipality id="6BC743024F74FFCEDE1DEF07FDCD82E8" box="[324,556,1659,1684]" pageId="2" pageNumber="507">Samarahan Division</collectingMunicipality>
, 
<geoCoordinate id="EE28BFBF4F74FFCEDD63EF07FD7F82E8" box="[570,670,1659,1684]" direction="north" orientation="latitude" pageId="2" pageNumber="507" precision="5" value="1.4016">1.4016N</geoCoordinate>
, 
<geoCoordinate id="EE28BFBF4F74FFCEDDF2EF07FCC482E8" box="[683,805,1659,1684]" direction="east" orientation="longitude" pageId="2" pageNumber="507" precision="5" value="110.9325">110.9325E</geoCoordinate>
(approx. coordinates), 
<date id="FFA2FFB84F74FFCEDB76EF00FB4182E8" box="[1071,1184,1659,1684]" pageId="2" pageNumber="507" value="2015-01-30">
<collectingDate id="EFE606504F74FFCEDB76EF00FB4182E8" box="[1071,1184,1659,1684]" pageId="2" pageNumber="507" value="2015-01-30">30 i 2015</collectingDate>
</date>
</materialsCitation>
, 
<materialsCitation id="3B74D3254F74FFCEDBF7EF07FC0F82C4" ID-GBIF-Occurrence="2236620142" collectingDate="2015-01-30" collectionCode="SMSM" collectorName="R. A. Dow" country="Malaysia" latitude="1.4016" location="Ulu Sebuyau National Park" longLatPrecision="7" longitude="110.9325" municipality="Samarahan Division" pageId="2" pageNumber="507" specimenCode="SAR15, ARG2" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
leg. 
<collectorName id="26E9BCAE4F74FFCEDBB9EF00FAB482EF" box="[1248,1365,1660,1684]" pageId="2" pageNumber="507">R.A. Dow</collectorName>
, to be deposited in 
<collectionCode id="ED0D41BD4F74FFCEDE64EFDCFE7082C4" box="[317,401,1696,1720]" country="Malaysia" httpUri="http://grbio.org/cool/yhsc-qrhn" name="Sarawak Museum" pageId="2" pageNumber="507">SMSM</collectionCode>
; 
<specimenCount id="9D1A12F14F74FFCEDEF0EFDCFE3E82CB" box="[425,479,1696,1720]" pageId="2" pageNumber="507" type="male">1 ♂</specimenCount>
(
<specimenCode id="DBBA71034F74FFCEDEA0EFDCFDAA82C4" box="[505,587,1695,1720]" collectionCode="SAR" country="Malaysia" lsid="urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="2" pageNumber="507" type="Herbarium">SAR15</specimenCode>
_ 
<specimenCode id="DBBA71034F74FFCEDD03EFE3FD4982C4" box="[602,680,1695,1720]" collectionCode="ARG" country="Malta" lsid="urn:lsid:biocol.org:col:12605" name="Argotti Botanic Garden" pageId="2" pageNumber="507" type="Herbarium">ARG2</specimenCode>
), same location and date
</materialsCitation>
, 
<materialsCitation id="3B74D3254F74FFCEDB5DEFE3FD8D837C" ID-GBIF-Occurrence="2236620143" collectingDate="2018-10-15" collectionCode="SAR, ARG" collectorName="L. Southwell" country="Malaysia" county="Sri Aman Division" latitude="1.1174" location="Marau Klasau" longLatPrecision="7" longitude="111.0947" municipality="Pantu area" pageId="2" pageNumber="507" specimenCode="SAR18, ARG9, ARG10" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
leg. 
<collectorName id="26E9BCAE4F74FFCEDB19EFDCFB0182C4" box="[1088,1248,1695,1720]" pageId="2" pageNumber="507">L. Southwell</collectorName>
; 
<specimenCount id="9D1A12F14F74FFCEDBAEEFDCFACC82CB" box="[1271,1325,1696,1720]" pageId="2" pageNumber="507" type="male">1 ♂</specimenCount>
(teneral, 
<specimenCode id="DBBA71034F74FFCEDFCEEFB8FF0882A0" box="[151,233,1732,1756]" collectionCode="SAR" country="Malaysia" lsid="urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="2" pageNumber="507" type="Herbarium">SAR18</specimenCode>
_ 
<specimenCode id="DBBA71034F74FFCEDFA1EFB8FEA482A0" box="[248,325,1732,1756]" collectionCode="ARG" country="Malta" lsid="urn:lsid:biocol.org:col:12605" name="Argotti Botanic Garden" pageId="2" pageNumber="507" type="Herbarium">ARG9</specimenCode>
), 
<specimenCount id="9D1A12F14F74FFCEDE02EFB8FE6682A7" box="[347,391,1732,1756]" pageId="2" pageNumber="507" type="female">1 ♀</specimenCount>
(
<specimenCode id="DBBA71034F74FFCEDEC0EFB8FE0D82A0" box="[409,492,1732,1756]" collectionCode="SAR" country="Malaysia" lsid="urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="2" pageNumber="507" type="Herbarium">SAR18</specimenCode>
_ 
<specimenCode id="DBBA71034F74FFCEDEA2EFB8FDB682A0" box="[507,599,1732,1756]" collectionCode="ARG" country="Malta" lsid="urn:lsid:biocol.org:col:12605" name="Argotti Botanic Garden" pageId="2" pageNumber="507" type="Herbarium">ARG10</specimenCode>
), disturbed peat swamp forest near 
<location id="8EC38FA34F74FFCEDCAAEFB8FB7782A0" LSID="urn:lsid:plazi:treatment:03B5686E4F74FFC5DFCEEDD2FA7A85AC:8EC38FA34F74FFCEDCAAEFB8FB7782A0" box="[1011,1174,1731,1756]" country="Malaysia" county="Sri Aman Division" latitude="1.1174" longLatPrecision="7" longitude="111.0947" municipality="Pantu area" name="Marau Klasau" pageId="2" pageNumber="507">Marau Klasau</location>
, 
<collectingMunicipality id="6BC743024F74FFCEDBFDEFB8FAFE82A7" box="[1188,1311,1732,1755]" pageId="2" pageNumber="507">Pantu area</collectingMunicipality>
, 
<collectingCounty id="62C2A1F44F74FFCEDA74EFBFFF1B837C" pageId="2" pageNumber="507">Sri Aman Division</collectingCounty>
, 
<geoCoordinate id="EE28BFBF4F74FFCEDE5CEF94FE88837C" box="[261,361,1768,1792]" direction="north" orientation="latitude" pageId="2" pageNumber="507" precision="5" value="1.1174">1.1174N</geoCoordinate>
, 
<geoCoordinate id="EE28BFBF4F74FFCEDE2DEF94FE0F837C" box="[372,494,1768,1792]" direction="east" orientation="longitude" pageId="2" pageNumber="507" precision="5" value="111.0947">111.0947E</geoCoordinate>
, 
<date id="FFA2FFB84F74FFCEDEA0EF9BFD8D837C" box="[505,620,1767,1792]" pageId="2" pageNumber="507" value="2018-10-15">
<collectingDate id="EFE606504F74FFCEDEA0EF9BFD8D837C" box="[505,620,1767,1792]" pageId="2" pageNumber="507" value="2018-10-15">15 x 2018</collectingDate>
</date>
</materialsCitation>
, leg. R.A. Dow.
</paragraph>
<paragraph id="8BA3D9784F74FFCEDF9EEE77FEDE83F3" blockId="2.[151,1437,1371,2008]" pageId="2" pageNumber="507">
<emphasis id="B968056A4F74FFCEDF9EEE77FE678358" bold="true" box="[199,390,1803,1828]" pageId="2" pageNumber="507">Other material.</emphasis>
<materialsCitation id="3B74D3254F74FFCEDEC8EE70FEDA83F3" ID-GBIF-Occurrence="2236620138" collectingDate="2012-07-07" collectionCode="RMNH" collectorName="R. A. Dow &amp; This" country="Malaysia" location="Maludam National Park" municipality="Betong Division" pageId="2" pageNumber="507" specimenCode="MEG96" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D1A12F14F74FFCEDEC8EE70FE21835F" box="[401,448,1804,1828]" pageId="2" pageNumber="507" type="female">1 ♀</specimenCount>
(SAR11_12_ 
<specimenCode id="DBBA71034F74FFCEDD05EE77FD5D8358" box="[604,700,1803,1828]" collectionCode="MEG" pageId="2" pageNumber="507">MEG96</specimenCode>
; 
<collectionCode id="ED0D41BD4F74FFCEDD92EE77FCC78358" box="[715,806,1803,1828]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="2" pageNumber="507" type="Museum">RMNH</collectionCode>
.INS.506881), perched by a track through peat swamp forest, 
<location id="8EC38FA34F74FFCEDFBFEE53FE198334" LSID="urn:lsid:plazi:treatment:03B5686E4F74FFC5DFCEEDD2FA7A85AC:8EC38FA34F74FFCEDFBFEE53FE198334" box="[230,504,1839,1864]" country="Malaysia" municipality="Betong Division" name="Maludam National Park" pageId="2" pageNumber="507">Maludam National Park</location>
, 
<collectingMunicipality id="6BC743024F74FFCEDD5CEE4CFD208334" box="[517,705,1839,1864]" pageId="2" pageNumber="507">Betong Division</collectingMunicipality>
, 
<collectingCountry id="F30B99E84F74FFCEDD97EE53FCD08334" box="[718,817,1839,1864]" name="Malaysia" pageId="2" pageNumber="507">Sarawak</collectingCountry>
, 
<collectingCountry id="F30B99E84F74FFCEDC64EE53FC468334" box="[829,935,1839,1864]" name="Malaysia" pageId="2" pageNumber="507">Malaysia</collectingCountry>
, 
<date id="FFA2FFB84F74FFCEDCEAEE4CFBCB8334" box="[947,1066,1839,1864]" pageId="2" pageNumber="507" value="2012-07-07">
<collectingDate id="EFE606504F74FFCEDCEAEE4CFBCB8334" box="[947,1066,1839,1864]" pageId="2" pageNumber="507" value="2012-07-07">7 vii 2012</collectingDate>
</date>
, leg. 
<collectorName id="26E9BCAE4F74FFCEDB31EE4CFB3C833B" box="[1128,1245,1840,1864]" pageId="2" pageNumber="507">R.A. Dow</collectorName>
, in 
<collectionCode id="ED0D41BD4F74FFCEDA5EEE4CFABE833B" box="[1287,1375,1840,1863]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="2" pageNumber="507" type="Museum">RMNH</collectionCode>
. 
<collectorName id="26E9BCAE4F74FFCEDA32EE53FA7C8334" box="[1387,1437,1839,1864]" pageId="2" pageNumber="507">This</collectorName>
has been excluded from the 
<typeStatus id="54A767DA4F74FFCEDEBDEE28FDF58310" box="[484,532,1876,1900]" pageId="2" pageNumber="507">type</typeStatus>
series merely because it was not available for a final check on variation and measurements
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3068AF34F74FFCEDF9EEEE7FF0E83A4" pageId="2" pageNumber="507" type="etymology">
<paragraph id="8BA3D9784F74FFCEDF9EEEE7FF0E83A4" blockId="2.[151,1437,1371,2008]" pageId="2" pageNumber="507">
<emphasis id="B968056A4F74FFCEDF9EEEE7FEAF83C8" bold="true" box="[199,334,1947,1972]" pageId="2" pageNumber="507">Etymology.</emphasis>
The species is named 
<taxonomicName id="4C1CA2FB4F74FFCEDD10EEE1FD7483C8" authorityName="Dow &amp; Ngiam" authorityYear="2019" box="[585,661,1949,1972]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="507" phylum="Arthropoda" rank="species" species="parvus">
<emphasis id="B968056A4F74FFCEDD10EEE1FD7483C8" box="[585,661,1949,1972]" italics="true" pageId="2" pageNumber="507">parvus</emphasis>
</taxonomicName>
, a Latin adjective meaning small, in reference to the small size of the species.
</paragraph>
</subSubSection>
<caption id="DF6389F04F75FFCFDFCEECD2FBDA8198" httpUri="https://zenodo.org/record/2647314/files/figure.png" pageId="3" pageNumber="508" startId="3.[151,264,1454,1477]" targetBox="[151,1435,193,1434]" targetPageId="3">
<paragraph id="8BA3D9784F75FFCFDFCEECD2FBDA8198" blockId="3.[151,1436,1454,1508]" pageId="3" pageNumber="508">
<emphasis id="B968056A4F75FFCFDFCEECD2FEB181B8" bold="true" box="[151,336,1454,1477]" pageId="3" pageNumber="508">FIGURES 9–12.</emphasis>
<taxonomicName id="4C1CA2FB4F75FFCFDE06ECD3FE2C81B9" authorityName="Selys" authorityYear="1862" box="[351,461,1455,1477]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="508" phylum="Arthropoda" rank="genus">
<emphasis id="B968056A4F75FFCFDE06ECD3FE2C81B9" box="[351,461,1455,1477]" italics="true" pageId="3" pageNumber="508">Podolestes</emphasis>
</taxonomicName>
species, prothorax dorsal view. 
<taxonomicName id="4C1CA2FB4F75FFCFDC66ECCCFC2681B9" authorityName=", Lieftinck" authorityYear="1950" box="[831,967,1456,1477]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="atomarius">
<emphasis id="B968056A4F75FFCFDC66ECCCFC2681B9" box="[831,967,1456,1477]" italics="true" pageId="3" pageNumber="508">P. atomarius</emphasis>
</taxonomicName>
: (9) male (KAL12_MEG28); (10) female (KAL12_MEG38). 
<taxonomicName id="4C1CA2FB4F75FFCFDE39ECB3FE5F8198" authority="Dow &amp; Ngiam, 2019" authorityName="Dow &amp; Ngiam" authorityYear="2019" box="[352,446,1487,1508]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="508" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
<emphasis id="B968056A4F75FFCFDE39ECB3FE5F8198" box="[352,446,1487,1508]" italics="true" pageId="3" pageNumber="508">P. parvus</emphasis>
</taxonomicName>
<emphasis id="B968056A4F75FFCFDE9DECB2FDF5819F" bold="true" box="[452,532,1486,1507]" pageId="3" pageNumber="508">
<taxonomicNameLabel id="A25BB8114F75FFCFDE9DECB2FDF5819F" box="[452,532,1486,1507]" pageId="3" pageNumber="508" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(11) holotype; (12) female paratype (SAR15_ARG4).
</paragraph>
</caption>
<subSubSection id="C3068AF34F75FFC4DF9EEF6CFA7C8270" lastPageId="8" lastPageNumber="513" pageId="3" pageNumber="508" type="description">
<paragraph id="8BA3D9784F75FFCFDF9EEF6CFC358311" blockId="3.[151,1437,1552,2009]" pageId="3" pageNumber="508">
<emphasis id="B968056A4F75FFCFDF9EEF6CFDC88255" bold="true" box="[199,553,1552,1577]" pageId="3" pageNumber="508">
Description of 
<typeStatus id="54A767DA4F75FFCFDE23EF6CFE008255" box="[378,481,1552,1577]" pageId="3" pageNumber="508" type="holotype">holotype</typeStatus>
male.
</emphasis>
<emphasis id="B968056A4F75FFCFDD6BEF6CFD8E8255" box="[562,623,1552,1577]" italics="true" pageId="3" pageNumber="508">Head</emphasis>
(
<figureCitation id="1327C5FD4F75FFCFDD26EF6CFD068255" box="[639,743,1552,1577]" captionStart="FIGURES 1–4" captionStartId="1.[151,264,1826,1849]" captionTargetBox="[151,1435,878,1805]" captionTargetId="figure@1.[151,1435,878,1805]" captionTargetPageId="1" captionText="FIGURES 1–4. Podolestes species males, head. P. atomarius (KAL12_MEG28): (1) dorsal; (3) frontal. P. parvus sp. nov. holotype: (2) dorsal; (4) dorsal-frontal." httpUri="https://zenodo.org/record/2647310/files/figure.png" pageId="3" pageNumber="508">Figs 2, 4</figureCitation>
). Labium pale yellowish except hooks of labial palps, which are dark brown. Labrum pale blue, narrowly brown along free margin. Mandible bases largely bluish white. Postclypeus shining black, anteclypeus pale blue. Genae mostly pale blue adjacent to mandible bases, very dark brown adjacent to compound eye. Antenna (missing above pedicel on right) with scape black, pedicel black and dark brown with whitish ring at top. Frons and vertex mostly bronzy black, with tiny pale mark in front median ocellus, pale blue marks adjacent to compound eye at ca level of antennae; pair dull pale streaks based on outer edge of lateral ocellus, directed towards rear of antenna base, dull pale mark between ocelli, almost divided centrally, these marks together approximately crescentic. Ocelli whitish. Pale and brown band centrally on upper occiput, narrowly separated from crescentic marking on vertex, continued onto underside. Underside of head mostly black with pale markings around point of attachment of prothorax.
</paragraph>
<paragraph id="8BA3D9784F75FFC8DF9EEE04FEB38643" blockId="3.[151,1437,1552,2009]" lastBlockId="4.[151,1437,151,575]" lastPageId="4" lastPageNumber="509" pageId="3" pageNumber="508">
<emphasis id="B968056A4F75FFCFDF9EEE04FEF983ED" box="[199,280,1912,1937]" italics="true" pageId="3" pageNumber="508">Thorax</emphasis>
. Prothorax (
<figureCitation id="1327C5FD4F75FFCFDEE9EE04FDED83ED" box="[432,524,1912,1937]" captionStart="FIGURES 9–12" captionStartId="3.[151,264,1454,1477]" captionTargetBox="[151,1435,193,1434]" captionTargetId="figure@3.[151,1435,193,1434]" captionTargetPageId="3" captionText="FIGURES 9–12. Podolestes species, prothorax dorsal view. P. atomarius: (9) male (KAL12_MEG28); (10) female (KAL12_MEG38). P. parvus sp. nov. (11) holotype; (12) female paratype (SAR15_ARG4)." httpUri="https://zenodo.org/record/2647314/files/figure.png" pageId="3" pageNumber="508">Figs 11</figureCitation>
, 
<figureCitation id="1327C5FD4F75FFCFDD47EE04FDDA83ED" box="[542,571,1912,1937]" captionStart="FIGURES 13–16" captionStartId="4.[151,264,1790,1813]" captionTargetBox="[151,1435,628,1769]" captionTargetId="figure@4.[151,1435,628,1769]" captionTargetPageId="4" captionText="FIGURES 13–16. Podolestes species, prothorax lateral view. P. atomarius: (13) male (KAL12_MEG28); (14) female (KAL12_MEG38). P. parvus sp. nov. (15) holotype; (16) female paratype (SAR15_ARG4)." httpUri="https://zenodo.org/record/2647316/files/figure.png" pageId="3" pageNumber="508">15</figureCitation>
) ground colour brown, propleuron with obscure paler and darker areas. Anterior pronotal lobe brown, except at lateral extremities and centrally to rear, where pale. Posterior pronotal lobe brown with obscure darker and paler areas. Middle pronotal lobe brown with pair large pale blue lateral marks, pair smaller central marks, narrowly divided (
<figureCitation id="1327C5FD4F72FFC8DD35E9EBFD5F84CC" box="[620,702,151,176]" captionStart="FIGURES 9–12" captionStartId="3.[151,264,1454,1477]" captionTargetBox="[151,1435,193,1434]" captionTargetId="figure@3.[151,1435,193,1434]" captionTargetPageId="3" captionText="FIGURES 9–12. Podolestes species, prothorax dorsal view. P. atomarius: (9) male (KAL12_MEG28); (10) female (KAL12_MEG38). P. parvus sp. nov. (11) holotype; (12) female paratype (SAR15_ARG4)." httpUri="https://zenodo.org/record/2647314/files/figure.png" pageId="4" pageNumber="509">Fig. 11</figureCitation>
). Posterior pronotal lobe simple, collar like. Synthorax (
<figureCitation id="1327C5FD4F72FFC8DA66E9EBFA7684CC" box="[1343,1431,151,176]" captionStart="FIGURES 17–20" captionStartId="5.[151,264,1357,1380]" captionTargetBox="[151,1435,481,1337]" captionTargetId="figure@5.[151,1435,481,1337]" captionTargetPageId="5" captionText="FIGURES 17–20. Podolestes species, synthorax dorsal view. P. atomarius: (17) male (KAL12_MEG28); (18) female (KAL12_MEG38). P. parvus sp. nov. (19) holotype; (20) female paratype (SAR15_ARG4)." httpUri="https://zenodo.org/record/2647318/files/figure.png" pageId="4" pageNumber="509">Figs 19</figureCitation>
, 
<figureCitation id="1327C5FD4F72FFC8DFCEE9C0FF5584A8" box="[151,180,188,212]" captionStart="FIGURES 21–24" captionStartId="6.[151,264,1212,1235]" captionTargetBox="[151,1435,193,1192]" captionTargetId="figure@6.[151,1435,193,1192]" captionTargetPageId="6" captionText="FIGURES 21–24. Podolestes species, synthorax lateral view. P. atomarius: (21) male (KAL12_MEG28); (22) female (KAL12_MEG38). P. parvus sp. nov. (23) holotype; (24) female paratype (SAR15_ARG4)." httpUri="https://zenodo.org/record/2647320/files/figure.png" pageId="4" pageNumber="509">23</figureCitation>
) with mesepisternum brown speckled with pale blue marks, pair of larger pale blue markings adjacent to antealar triangles (
<figureCitation id="1327C5FD4F72FFC8DE37E9A3FE248484" box="[366,453,223,248]" captionStart="FIGURES 17–20" captionStartId="5.[151,264,1357,1380]" captionTargetBox="[151,1435,481,1337]" captionTargetId="figure@5.[151,1435,481,1337]" captionTargetPageId="5" captionText="FIGURES 17–20. Podolestes species, synthorax dorsal view. P. atomarius: (17) male (KAL12_MEG28); (18) female (KAL12_MEG38). P. parvus sp. nov. (19) holotype; (20) female paratype (SAR15_ARG4)." httpUri="https://zenodo.org/record/2647318/files/figure.png" pageId="4" pageNumber="509">Fig. 19</figureCitation>
). Mesepimeron similar, larger, irregular marks at either end adjacent to mesepleural suture, metepisternum pale brown with obscure bluish areas, metepimeron largely pale brown. Mesinfraepisternum marked similarly to mesepimeron, metinfraepisternum pale with darker areas. Venter of synthorax pale. Legs (left anterior missing below trochanter) with coxae pale brownish cream, with black streak on posterior coxa, tiny black marks middle coxa, trochanters similar with black marks on middle and posterior pairs. Femur pale brownish cream, two black rings, one not far below trochanter, other around two thirds length, black above joint with tibia, partially on flexor surface. Tibia mostly pale, black below joint with femur, dark central ring, darker on outer surface. Wings with arc situated slightly distal to Ax 2. 17 Px except left Hw with 15 Px. One post quadrilateral cell in all wings. R 
<subScript id="1798DB3D4F72FFC8DE67EB72FEA78660" attach="left" box="[318,326,526,540]" fontSize="6" pageId="4" pageNumber="509">4</subScript>
proximal to Sn, IR 
<subScript id="1798DB3D4F72FFC8DD46EB72FDC68660" attach="left" box="[543,551,526,540]" fontSize="6" pageId="4" pageNumber="509">3</subScript>
slightly distal to it. Pt greyish brown, almost a rhombus, covering one to two underlying cells.
</paragraph>
<caption id="DF6389F04F72FFC8DFCEEF82FBDD834F" httpUri="https://zenodo.org/record/2647316/files/figure.png" pageId="4" pageNumber="509" startId="4.[151,264,1790,1813]" targetBox="[151,1435,628,1769]" targetPageId="4">
<paragraph id="8BA3D9784F72FFC8DFCEEF82FBDD834F" blockId="4.[151,1436,1790,1843]" pageId="4" pageNumber="509">
<emphasis id="B968056A4F72FFC8DFCEEF82FEBB8368" bold="true" box="[151,346,1790,1813]" pageId="4" pageNumber="509">FIGURES 13–16.</emphasis>
<taxonomicName id="4C1CA2FB4F72FFC8DE3EEF82FE348368" authorityName="Selys" authorityYear="1862" box="[359,469,1790,1812]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="509" phylum="Arthropoda" rank="genus">
<emphasis id="B968056A4F72FFC8DE3EEF82FE348368" box="[359,469,1790,1812]" italics="true" pageId="4" pageNumber="509">Podolestes</emphasis>
</taxonomicName>
species, prothorax lateral view. 
<taxonomicName id="4C1CA2FB4F72FFC8DC67EE7CFC248369" authorityName=", Lieftinck" authorityYear="1950" box="[830,965,1792,1813]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="atomarius">
<emphasis id="B968056A4F72FFC8DC67EE7CFC248369" box="[830,965,1792,1813]" italics="true" pageId="4" pageNumber="509">P. atomarius</emphasis>
</taxonomicName>
: (13) male (KAL12_MEG28); (14) female (KAL12_MEG38). 
<taxonomicName id="4C1CA2FB4F72FFC8DE39EE62FE5F834F" authority="Dow &amp; Ngiam, 2019" authorityName="Dow &amp; Ngiam" authorityYear="2019" box="[352,446,1822,1843]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="509" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
<emphasis id="B968056A4F72FFC8DE39EE62FE5F834F" box="[352,446,1822,1843]" italics="true" pageId="4" pageNumber="509">P. parvus</emphasis>
</taxonomicName>
<emphasis id="B968056A4F72FFC8DE9DEE61FDF5834E" bold="true" box="[452,532,1821,1842]" pageId="4" pageNumber="509">
<taxonomicNameLabel id="A25BB8114F72FFC8DE9DEE61FDF5834E" box="[452,532,1821,1842]" pageId="4" pageNumber="509" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(15) holotype; (16) female paratype (SAR15_ARG4).
</paragraph>
</caption>
<paragraph id="8BA3D9784F72FFC9DF9EEE26FBA1853C" blockId="4.[151,1436,1882,2015]" lastBlockId="5.[151,1436,151,429]" lastPageId="5" lastPageNumber="510" pageId="4" pageNumber="509">
<emphasis id="B968056A4F72FFC8DF9EEE26FED0830F" box="[199,305,1882,1907]" italics="true" pageId="4" pageNumber="509">Abdomen</emphasis>
(see 
<figureCitation id="1327C5FD4F72FFC8DE32EE26FE5C830F" box="[363,445,1882,1907]" captionStart="FIGURES 42–45" captionStartId="10.[151,264,962,985]" captionTargetBox="[151,1435,193,942]" captionTargetId="figure@10.[151,1435,193,942]" captionTargetPageId="10" captionText="FIGURES 42–45. Podolestes parvus sp. nov. in life. Holotype male: (42) whole animal in lateral view; (44) head and thorax in dorsal view. Female paratype (SAR11_12_MEG95): (43) whole animal in lateral view; (45) head and thorax in dorsal view. Photographs by R.W.J. Ngiam." httpUri="https://zenodo.org/record/2647336/files/figure.png" pageId="4" pageNumber="509">Fig. 42</figureCitation>
). S1 pale brown. S2 pale brown dorsally, brownish cream laterally, pale blue dorsal mark in basal ca one quarter, almost divided centrally. S3–S8 pale brown, darker apically, with pale blue basal dorsal mark, pale laterally below this. S9 brown with pale blue dorsal mark running to posterior carina but narrowing in apical part. S10 dark brown with sub triangular pale blue dorsal mark (
<figureCitation id="1327C5FD4F72FFC8DCE8EEBAFBE583A3" box="[945,1028,1990,2015]" captionStart="FIGURES 33–36" captionStartId="7.[151,264,1978,2001]" captionTargetBox="[151,1435,1316,1958]" captionTargetId="figure@7.[151,1435,1316,1958]" captionTargetPageId="7" captionText="FIGURES 33–36. Podolestes species, markings of terminal abdominal segments, dorsal view. P. atomarius: (33) male (KAL12_MEG28); (34) female (KAL12_MEG38). P. parvus sp. nov. (35) holotype; (36) female paratype (SAR15_ARG4)." httpUri="https://zenodo.org/record/2647330/files/figure.png" pageId="4" pageNumber="509">Fig. 35</figureCitation>
), obscure pale areas to either side of this basally. Anal appendages (
<figureCitation id="1327C5FD4F73FFC9DEA2E9EBFD6584CC" box="[507,644,151,176]" captionStart="FIGURES 37–40" captionStartId="8.[151,264,1426,1449]" captionTargetBox="[151,1435,193,1405]" captionTargetId="figure@8.[151,1435,193,1405]" captionTargetPageId="8" captionText="FIGURES 37–40. Anal appendages of Podolestes species males. P. atomarius (KAL12_MEG2): (37) dorsal; (39) lateral. P. parvus sp. nov. holotype: (38) dorsal; (40) lateral. All scale bars 0.5mm." httpUri="https://zenodo.org/record/2647332/files/figure.png" pageId="5" pageNumber="510">Figs 38, 40</figureCitation>
): cerci brown, paraprocts pale brown at base, rest dark brown. Cerci longer than paraprocts, in lateral view tapering evenly from base to rounded tip, in dorsal view running almost straight from base to ca one third length where directed inwards and rearwards so that they cross one-another (
<figureCitation id="1327C5FD4F73FFC9DA36E9A3FF578560" captionStart="FIGURES 37–40" captionStartId="8.[151,264,1426,1449]" captionTargetBox="[151,1435,193,1405]" captionTargetId="figure@8.[151,1435,193,1405]" captionTargetPageId="8" captionText="FIGURES 37–40. Anal appendages of Podolestes species males. P. atomarius (KAL12_MEG2): (37) dorsal; (39) lateral. P. parvus sp. nov. holotype: (38) dorsal; (40) lateral. All scale bars 0.5mm." httpUri="https://zenodo.org/record/2647332/files/figure.png" pageId="5" pageNumber="510">Fig. 38</figureCitation>
), slightly expanded along inner margin after the turn, tapering to rounded apices. Paraprocts in lateral view contracting from broad base then expanding slightly again to subrectangular apices.
</paragraph>
<paragraph id="8BA3D9784F73FFC9DF9EE830FB9F85F4" blockId="5.[151,1436,151,429]" pageId="5" pageNumber="510">
<emphasis id="B968056A4F73FFC9DF9EE830FE828519" box="[199,355,332,357]" italics="true" pageId="5" pageNumber="510">Genital ligula</emphasis>
. Terminal segment almost square ended in ventral view, with pair of lateral terminal arms folded back and above segment, narrow at origin but then expanding considerably (
<figureCitation id="1327C5FD4F73FFC9DCB7E813FB9085F4" box="[1006,1137,367,392]" captionStart="FIGURES 29–32" captionStartId="7.[151,264,1212,1235]" captionTargetBox="[151,1435,193,1192]" captionTargetId="figure@7.[151,1435,193,1192]" captionTargetPageId="7" captionText="FIGURES 29–32. Genital ligula of Podolestes species. P. atomarius (KAL12_MEG1): (29) ventral; (31) lateral. P. parvus sp. nov. holotype: (30) ventral; (32) lateral. All scale bars 0.2mm." httpUri="https://zenodo.org/record/2647328/files/figure.png" pageId="5" pageNumber="510">Figs 30, 32</figureCitation>
).
</paragraph>
<paragraph id="8BA3D9784F73FFC9DF9EE8E9FBBB85D0" blockId="5.[151,1436,151,429]" box="[199,1114,404,429]" pageId="5" pageNumber="510">Measurements (mm). Abdomen without anal appendages 22, cerci 1.1, Hw ca 20.</paragraph>
<caption id="DF6389F04F73FFC9DFCEEC31FBDD81FE" httpUri="https://zenodo.org/record/2647318/files/figure.png" pageId="5" pageNumber="510" startId="5.[151,264,1357,1380]" targetBox="[151,1435,481,1337]" targetPageId="5">
<paragraph id="8BA3D9784F73FFC9DFCEEC31FBDD81FE" blockId="5.[151,1436,1357,1411]" pageId="5" pageNumber="510">
<emphasis id="B968056A4F73FFC9DFCEEC31FEBA811F" bold="true" box="[151,347,1357,1380]" pageId="5" pageNumber="510">FIGURES 17–20.</emphasis>
<taxonomicName id="4C1CA2FB4F73FFC9DE3EEC32FE348118" authorityName="Selys" authorityYear="1862" box="[359,469,1358,1380]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="510" phylum="Arthropoda" rank="genus">
<emphasis id="B968056A4F73FFC9DE3EEC32FE348118" box="[359,469,1358,1380]" italics="true" pageId="5" pageNumber="510">Podolestes</emphasis>
</taxonomicName>
species, synthorax dorsal view. 
<taxonomicName id="4C1CA2FB4F73FFC9DC67EC33FC248118" authorityName=", Lieftinck" authorityYear="1950" box="[830,965,1359,1380]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="510" phylum="Arthropoda" rank="species" species="atomarius">
<emphasis id="B968056A4F73FFC9DC67EC33FC248118" box="[830,965,1359,1380]" italics="true" pageId="5" pageNumber="510">P. atomarius</emphasis>
</taxonomicName>
: (17) male (KAL12_MEG28); (18) female (KAL12_MEG38). 
<taxonomicName id="4C1CA2FB4F73FFC9DE39EC12FE5F81FF" authority="Dow &amp; Ngiam, 2019" authorityName="Dow &amp; Ngiam" authorityYear="2019" box="[352,446,1390,1411]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="510" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
<emphasis id="B968056A4F73FFC9DE39EC12FE5F81FF" box="[352,446,1390,1411]" italics="true" pageId="5" pageNumber="510">P. parvus</emphasis>
</taxonomicName>
<emphasis id="B968056A4F73FFC9DE9DEC11FDF581FE" bold="true" box="[452,532,1389,1410]" pageId="5" pageNumber="510">
<taxonomicNameLabel id="A25BB8114F73FFC9DE9DEC11FDF581FE" box="[452,532,1389,1410]" pageId="5" pageNumber="510" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(19) holotype; (20) female paratype (SAR15_ARG4).
</paragraph>
</caption>
<paragraph id="8BA3D9784F73FFC9DF9EECD3FD308248" blockId="5.[151,1437,1455,2020]" pageId="5" pageNumber="510">
<emphasis id="B968056A4F73FFC9DF9EECD3FDD681B4" bold="true" box="[199,567,1455,1480]" pageId="5" pageNumber="510">
Description of female 
<typeStatus id="54A767DA4F73FFC9DE95ECD3FDD681B4" box="[460,567,1455,1480]" pageId="5" pageNumber="510" type="paratype">paratype</typeStatus>
</emphasis>
(based on SAR15_ARG4). Pedicel whitish in basal ca two-thirds, rest brown, flagella brown. Frons bronzy black, vertex bronzy black with approximately crescentic brownish cream mark (
<figureCitation id="1327C5FD4F73FFC9DA36ECAFFF47826C" captionStart="FIGURES 5–8" captionStartId="2.[151,264,1069,1092]" captionTargetBox="[151,1435,193,1049]" captionTargetId="figure@2.[151,1435,193,1049]" captionTargetPageId="2" captionText="FIGURES 5–8. Podolestes species females, head. P. atomarius (KAL12_MEG38): (5) dorsal; (7) frontal. P. parvus sp. nov. paratype (SAR15_ARG4): (6) dorsal; (8) frontal." httpUri="https://zenodo.org/record/2647312/files/figure.png" pageId="5" pageNumber="510">Fig. 6</figureCitation>
) almost surrounding the lateral ocelli with its lateral parts directed towards the antennae; this joined to subrectangular marking of same colour on occiput.
</paragraph>
<paragraph id="8BA3D9784F73FFC9DF9EEF43FAC78200" blockId="5.[151,1437,1455,2020]" pageId="5" pageNumber="510">
<emphasis id="B968056A4F73FFC9DF9EEF43FEF98224" box="[199,280,1599,1624]" italics="true" pageId="5" pageNumber="510">Thorax</emphasis>
. Markings very similar to male except in tiny details (see 
<figureCitation id="1327C5FD4F73FFC9DCEBEF43FBE88224" box="[946,1033,1599,1624]" captionStart="FIGURES 9–12" captionStartId="3.[151,264,1454,1477]" captionTargetBox="[151,1435,193,1434]" captionTargetId="figure@3.[151,1435,193,1434]" captionTargetPageId="3" captionText="FIGURES 9–12. Podolestes species, prothorax dorsal view. P. atomarius: (9) male (KAL12_MEG28); (10) female (KAL12_MEG38). P. parvus sp. nov. (11) holotype; (12) female paratype (SAR15_ARG4)." httpUri="https://zenodo.org/record/2647314/files/figure.png" pageId="5" pageNumber="510">Figs 12</figureCitation>
, 
<figureCitation id="1327C5FD4F73FFC9DB4FEF43FBD58224" box="[1046,1076,1599,1624]" captionStart="FIGURES 13–16" captionStartId="4.[151,264,1790,1813]" captionTargetBox="[151,1435,628,1769]" captionTargetId="figure@4.[151,1435,628,1769]" captionTargetPageId="4" captionText="FIGURES 13–16. Podolestes species, prothorax lateral view. P. atomarius: (13) male (KAL12_MEG28); (14) female (KAL12_MEG38). P. parvus sp. nov. (15) holotype; (16) female paratype (SAR15_ARG4)." httpUri="https://zenodo.org/record/2647316/files/figure.png" pageId="5" pageNumber="510">16</figureCitation>
, 
<figureCitation id="1327C5FD4F73FFC9DB1BEF3CFB818224" box="[1090,1120,1600,1624]" captionStart="FIGURES 17–20" captionStartId="5.[151,264,1357,1380]" captionTargetBox="[151,1435,481,1337]" captionTargetId="figure@5.[151,1435,481,1337]" captionTargetPageId="5" captionText="FIGURES 17–20. Podolestes species, synthorax dorsal view. P. atomarius: (17) male (KAL12_MEG28); (18) female (KAL12_MEG38). P. parvus sp. nov. (19) holotype; (20) female paratype (SAR15_ARG4)." httpUri="https://zenodo.org/record/2647318/files/figure.png" pageId="5" pageNumber="510">20</figureCitation>
, 
<figureCitation id="1327C5FD4F73FFC9DB34EF3CFB6C8224" box="[1133,1165,1600,1624]" captionStart="FIGURES 21–24" captionStartId="6.[151,264,1212,1235]" captionTargetBox="[151,1435,193,1192]" captionTargetId="figure@6.[151,1435,193,1192]" captionTargetPageId="6" captionText="FIGURES 21–24. Podolestes species, synthorax lateral view. P. atomarius: (21) male (KAL12_MEG28); (22) female (KAL12_MEG38). P. parvus sp. nov. (23) holotype; (24) female paratype (SAR15_ARG4)." httpUri="https://zenodo.org/record/2647320/files/figure.png" pageId="5" pageNumber="510">24</figureCitation>
). 16 Px in Fw and right Hw, 15 Px in left Hw. Pt covering one and most of another underlying cell in all wings, brown (
<figureCitation id="1327C5FD4F73FFC9DB9CEF1FFAF88200" box="[1221,1305,1635,1660]" captionStart="FIGURES 25–28" captionStartId="6.[151,264,1978,2001]" captionTargetBox="[151,1435,1316,1958]" captionTargetId="figure@6.[151,1435,1316,1958]" captionTargetPageId="6" captionText="FIGURES 25–28. Pterostigma of Podolestes species. P. atomarius: (25) male Fw (KAL12_MEG28); (26) male Hw (KAL12_MEG28); (27) female Fw (KAL12_MEG38). (28) P. parvus female (SAR15_ARG4) Hw." httpUri="https://zenodo.org/record/2647322/files/figure.png" pageId="5" pageNumber="510">Fig. 28</figureCitation>
).
</paragraph>
<paragraph id="8BA3D9784F73FFC9DF9EEFFBFDF682B8" blockId="5.[151,1437,1455,2020]" pageId="5" pageNumber="510">
<emphasis id="B968056A4F73FFC9DF9EEFFBFED082DC" box="[199,305,1671,1696]" italics="true" pageId="5" pageNumber="510">Abdomen</emphasis>
(
<figureCitation id="1327C5FD4F73FFC9DE1BEFFBFE7882DC" box="[322,409,1671,1696]" captionStart="FIGURES 33–36" captionStartId="7.[151,264,1978,2001]" captionTargetBox="[151,1435,1316,1958]" captionTargetId="figure@7.[151,1435,1316,1958]" captionTargetPageId="7" captionText="FIGURES 33–36. Podolestes species, markings of terminal abdominal segments, dorsal view. P. atomarius: (33) male (KAL12_MEG28); (34) female (KAL12_MEG38). P. parvus sp. nov. (35) holotype; (36) female paratype (SAR15_ARG4)." httpUri="https://zenodo.org/record/2647330/files/figure.png" pageId="5" pageNumber="510">Fig. 36</figureCitation>
). Cerci dark brown, sharply conical, ca same length as S10. Ovipositor mottled dark and pale brown, styles missing on left.
</paragraph>
<paragraph id="8BA3D9784F73FFC9DF9EEFACFB568294" blockId="5.[151,1437,1455,2020]" box="[199,1207,1743,1768]" pageId="5" pageNumber="510">Measurements (mm). Abdomen without anal appendages or ovipositor ca 21, Hw ca 20.5.</paragraph>
<paragraph id="8BA3D9784F73FFC9DF9EEF8FFD2E8328" blockId="5.[151,1437,1455,2020]" pageId="5" pageNumber="510">
<emphasis id="B968056A4F73FFC9DF9EEF8FFDC88370" bold="true" box="[199,553,1779,1804]" pageId="5" pageNumber="510">
Variation in male 
<typeStatus id="54A767DA4F73FFC9DEF5EF8FFDC58370" box="[428,548,1779,1804]" pageId="5" pageNumber="510" type="paratype">paratypes</typeStatus>
.
</emphasis>
Aside from small variations in markings, and size variation, there is little variation in the non-teneral males. The basal half of the scape is pale in one of the males from Ulu Sebuyau National Park and the basal two thirds in the other.
</paragraph>
<paragraph id="8BA3D9784F73FFC9DF9EEE1CFF2483E7" blockId="5.[151,1437,1455,2020]" pageId="5" pageNumber="510">
Measurements (mm). Abdomen without anal appendages 22–24, Hw 20– 
<date id="FFA2FFB84F73FFC9DB54EE23FB878304" box="[1037,1126,1887,1912]" pageId="5" pageNumber="510" value="14-05-20">20.5. 14</date>
–17 Px in Fw, 13–16 Px in Hw.
</paragraph>
<paragraph id="8BA3D9784F73FFC9DF9EEEDBFB5F8398" blockId="5.[151,1437,1455,2020]" pageId="5" pageNumber="510">
<emphasis id="B968056A4F73FFC9DF9EEEDBFDD483BC" bold="true" box="[199,565,1959,1984]" pageId="5" pageNumber="510">
Variation in female 
<typeStatus id="54A767DA4F73FFC9DEEEEEDBFDD183BC" box="[439,560,1959,1984]" pageId="5" pageNumber="510" type="paratype">paratypes</typeStatus>
.
</emphasis>
The female from Sri Aman has a slightly lower density of pale spots on the dorsum of the synthorax. Fw Pt only covering half of a second underlying cell in one 
<typeStatus id="54A767DA4F73FFC9DB0FEEB0FB5A8398" box="[1110,1211,1996,2020]" pageId="5" pageNumber="510" type="paratype">paratype</typeStatus>
.
</paragraph>
<caption id="DF6389F04F70FFCADFCEEDC0FBDD808E" httpUri="https://zenodo.org/record/2647320/files/figure.png" pageId="6" pageNumber="511" startId="6.[151,264,1212,1235]" targetBox="[151,1435,193,1192]" targetPageId="6">
<paragraph id="8BA3D9784F70FFCADFCEEDC0FBDD808E" blockId="6.[151,1436,1212,1266]" pageId="6" pageNumber="511">
<emphasis id="B968056A4F70FFCADFCEEDC0FEBB80AE" bold="true" box="[151,346,1212,1235]" pageId="6" pageNumber="511">FIGURES 21–24.</emphasis>
<taxonomicName id="4C1CA2FB4F70FFCADE3EEDC1FE3480AF" authorityName="Selys" authorityYear="1862" box="[359,469,1213,1235]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="511" phylum="Arthropoda" rank="genus">
<emphasis id="B968056A4F70FFCADE3EEDC1FE3480AF" box="[359,469,1213,1235]" italics="true" pageId="6" pageNumber="511">Podolestes</emphasis>
</taxonomicName>
species, synthorax lateral view. 
<taxonomicName id="4C1CA2FB4F70FFCADC66EDC2FC2480AF" authorityName=", Lieftinck" authorityYear="1950" box="[831,965,1214,1235]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="511" phylum="Arthropoda" rank="species" species="atomarius">
<emphasis id="B968056A4F70FFCADC66EDC2FC2480AF" box="[831,965,1214,1235]" italics="true" pageId="6" pageNumber="511">P. atomarius</emphasis>
</taxonomicName>
: (21) male (KAL12_MEG28); (22) female (KAL12_MEG38). 
<taxonomicName id="4C1CA2FB4F70FFCADE39EDA1FE5F808E" authority="Dow &amp; Ngiam, 2019" authorityName="Dow &amp; Ngiam" authorityYear="2019" box="[352,446,1245,1266]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="511" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
<emphasis id="B968056A4F70FFCADE39EDA1FE5F808E" box="[352,446,1245,1266]" italics="true" pageId="6" pageNumber="511">P. parvus</emphasis>
</taxonomicName>
<emphasis id="B968056A4F70FFCADE9DEDA0FDF5808D" bold="true" box="[452,532,1244,1265]" pageId="6" pageNumber="511">
<taxonomicNameLabel id="A25BB8114F70FFCADE9DEDA0FDF5808D" box="[452,532,1244,1265]" pageId="6" pageNumber="511" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(23) holotype; (24) female paratype (SAR15_ARG4).
</paragraph>
</caption>
<caption id="DF6389F04F70FFCADFCEEEC6FB67838C" httpUri="https://zenodo.org/record/2647322/files/figure.png" pageId="6" pageNumber="511" startId="6.[151,264,1978,2001]" targetBox="[151,1435,1316,1958]" targetPageId="6">
<paragraph id="8BA3D9784F70FFCADFCEEEC6FB67838C" blockId="6.[151,1435,1978,2032]" pageId="6" pageNumber="511">
<emphasis id="B968056A4F70FFCADFCEEEC6FEBA83AC" bold="true" box="[151,347,1978,2001]" pageId="6" pageNumber="511">FIGURES 25–28.</emphasis>
Pterostigma of 
<taxonomicName id="4C1CA2FB4F70FFCADD4AEEC7FD6183AD" authorityName="Selys" authorityYear="1862" box="[531,640,1979,2001]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="511" phylum="Arthropoda" rank="genus">
<emphasis id="B968056A4F70FFCADD4AEEC7FD6183AD" box="[531,640,1979,2001]" italics="true" pageId="6" pageNumber="511">Podolestes</emphasis>
</taxonomicName>
species. 
<taxonomicName id="4C1CA2FB4F70FFCADDB5EEC0FC9583AD" authorityName=", Lieftinck" authorityYear="1950" box="[748,884,1980,2001]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="511" phylum="Arthropoda" rank="species" species="atomarius">
<emphasis id="B968056A4F70FFCADDB5EEC0FC9583AD" box="[748,884,1980,2001]" italics="true" pageId="6" pageNumber="511">P. atomarius</emphasis>
</taxonomicName>
: (25) male Fw (KAL12_MEG28); (26) male Hw (KAL12_MEG28); (27) female Fw (KAL12_MEG38). (28) 
<taxonomicName id="4C1CA2FB4F70FFCADDA5EEA7FCBB838C" authorityName="Dow &amp; Ngiam" authorityYear="2019" box="[764,858,2011,2032]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="511" phylum="Arthropoda" rank="species" species="parvus">
<emphasis id="B968056A4F70FFCADDA5EEA7FCBB838C" box="[764,858,2011,2032]" italics="true" pageId="6" pageNumber="511">P. parvus</emphasis>
</taxonomicName>
female (SAR15_ARG4) Hw.
</paragraph>
</caption>
<caption id="DF6389F04F71FFCBDFCEEDC0FCEF808E" httpUri="https://zenodo.org/record/2647328/files/figure.png" pageId="7" pageNumber="512" startId="7.[151,264,1212,1235]" targetBox="[151,1435,193,1192]" targetPageId="7">
<paragraph id="8BA3D9784F71FFCBDFCEEDC0FCEF808E" blockId="7.[151,1436,1212,1266]" pageId="7" pageNumber="512">
<emphasis id="B968056A4F71FFCBDFCEEDC0FEB580AE" bold="true" box="[151,340,1212,1235]" pageId="7" pageNumber="512">FIGURES 29–32.</emphasis>
Genital ligula of 
<taxonomicName id="4C1CA2FB4F71FFCBDD51EDC1FD9780AF" authorityName="Selys" authorityYear="1862" box="[520,630,1213,1235]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="512" phylum="Arthropoda" rank="genus">
<emphasis id="B968056A4F71FFCBDD51EDC1FD9780AF" box="[520,630,1213,1235]" italics="true" pageId="7" pageNumber="512">Podolestes</emphasis>
</taxonomicName>
species. 
<taxonomicName id="4C1CA2FB4F71FFCBDD8BEDC2FCB280AF" authorityName=", Lieftinck" authorityYear="1950" box="[722,851,1214,1235]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="atomarius">
<emphasis id="B968056A4F71FFCBDD8BEDC2FCB280AF" box="[722,851,1214,1235]" italics="true" pageId="7" pageNumber="512">P. atomarius</emphasis>
</taxonomicName>
(KAL12_MEG1): (29) ventral; (31) lateral. 
<taxonomicName id="4C1CA2FB4F71FFCBDA40EDC2FA9680AF" authority="Dow &amp; Ngiam, 2019" authorityName="Dow &amp; Ngiam" authorityYear="2019" box="[1305,1399,1214,1235]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
<emphasis id="B968056A4F71FFCBDA40EDC2FA9680AF" box="[1305,1399,1214,1235]" italics="true" pageId="7" pageNumber="512">P. parvus</emphasis>
</taxonomicName>
<emphasis id="B968056A4F71FFCBDA24EDC1FF22808D" bold="true" pageId="7" pageNumber="512">
<taxonomicNameLabel id="A25BB8114F71FFCBDA24EDC1FF22808D" pageId="7" pageNumber="512" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
holotype: (30) ventral; (32) lateral. All scale bars 0.2mm.
</paragraph>
</caption>
<caption id="DF6389F04F71FFCBDFCEEEC6FA60838C" httpUri="https://zenodo.org/record/2647330/files/figure.png" pageId="7" pageNumber="512" startId="7.[151,264,1978,2001]" targetBox="[151,1435,1316,1958]" targetPageId="7">
<paragraph id="8BA3D9784F71FFCBDFCEEEC6FA60838C" blockId="7.[151,1436,1978,2032]" pageId="7" pageNumber="512">
<emphasis id="B968056A4F71FFCBDFCEEEC6FEBB83AC" bold="true" box="[151,346,1978,2001]" pageId="7" pageNumber="512">FIGURES 33–36.</emphasis>
<taxonomicName id="4C1CA2FB4F71FFCBDE3CEEC7FE3283AD" authorityName="Selys" authorityYear="1862" box="[357,467,1979,2001]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="512" phylum="Arthropoda" rank="genus">
<emphasis id="B968056A4F71FFCBDE3CEEC7FE3283AD" box="[357,467,1979,2001]" italics="true" pageId="7" pageNumber="512">Podolestes</emphasis>
</taxonomicName>
species, markings of terminal abdominal segments, dorsal view. 
<taxonomicName id="4C1CA2FB4F71FFCBDBC5EEC0FAC083AD" authorityName=", Lieftinck" authorityYear="1950" box="[1180,1313,1980,2001]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="atomarius">
<emphasis id="B968056A4F71FFCBDBC5EEC0FAC083AD" box="[1180,1313,1980,2001]" italics="true" pageId="7" pageNumber="512">P. atomarius</emphasis>
</taxonomicName>
: (33) male (KAL12_MEG28); (34) female (KAL12_MEG38). 
<taxonomicName id="4C1CA2FB4F71FFCBDDFCEEA7FCE2838C" authority="Dow &amp; Ngiam, 2019" authorityName="Dow &amp; Ngiam" authorityYear="2019" box="[677,771,2011,2032]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="512" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
<emphasis id="B968056A4F71FFCBDDFCEEA7FCE2838C" box="[677,771,2011,2032]" italics="true" pageId="7" pageNumber="512">P. parvus</emphasis>
</taxonomicName>
<emphasis id="B968056A4F71FFCBDC50EEA6FCBB8393" bold="true" box="[777,858,2010,2031]" pageId="7" pageNumber="512">
<taxonomicNameLabel id="A25BB8114F71FFCBDC50EEA6FCBB8393" box="[777,858,2010,2031]" pageId="7" pageNumber="512" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(35) holotype; (36) female paratype (SAR15_ARG4).
</paragraph>
</caption>
<caption id="DF6389F04F7EFFC4DFCEECEEFC9781BB" httpUri="https://zenodo.org/record/2647332/files/figure.png" pageId="8" pageNumber="513" startId="8.[151,264,1426,1449]" targetBox="[151,1435,193,1405]" targetPageId="8">
<paragraph id="8BA3D9784F7EFFC4DFCEECEEFC9781BB" blockId="8.[151,1437,1426,1479]" pageId="8" pageNumber="513">
<emphasis id="B968056A4F7EFFC4DFCEECEEFEB781D4" bold="true" box="[151,342,1426,1449]" pageId="8" pageNumber="513">FIGURES 37–40.</emphasis>
Anal appendages of 
<taxonomicName id="4C1CA2FB4F7EFFC4DD6BECEFFD4181D5" authorityName="Selys" authorityYear="1862" box="[562,672,1427,1449]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="513" phylum="Arthropoda" rank="genus">
<emphasis id="B968056A4F7EFFC4DD6BECEFFD4181D5" box="[562,672,1427,1449]" italics="true" pageId="8" pageNumber="513">Podolestes</emphasis>
</taxonomicName>
species males. 
<taxonomicName id="4C1CA2FB4F7EFFC4DC1BECE8FC2281D5" authorityName=", Lieftinck" authorityYear="1950" box="[834,963,1428,1449]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="513" phylum="Arthropoda" rank="species" species="atomarius">
<emphasis id="B968056A4F7EFFC4DC1BECE8FC2281D5" box="[834,963,1428,1449]" italics="true" pageId="8" pageNumber="513">P. atomarius</emphasis>
</taxonomicName>
(KAL12_MEG2): (37) dorsal; (39) lateral. 
<taxonomicName id="4C1CA2FB4F7EFFC4DAD0ECE8FF3D81BB" authority="Dow &amp; Ngiam, 2019" authorityName="Dow &amp; Ngiam" authorityYear="2019" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="513" phylum="Arthropoda" rank="species" species="parvus" status="sp. nov.">
<emphasis id="B968056A4F7EFFC4DAD0ECE8FF3D81BB" italics="true" pageId="8" pageNumber="513">P. parvus</emphasis>
</taxonomicName>
<emphasis id="B968056A4F7EFFC4DFBBECCEFED381BB" bold="true" box="[226,306,1458,1479]" pageId="8" pageNumber="513">
<taxonomicNameLabel id="A25BB8114F7EFFC4DFBBECCEFED381BB" box="[226,306,1458,1479]" pageId="8" pageNumber="513" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
holotype: (38) dorsal; (40) lateral. All scale bars 0.5mm.
</paragraph>
</caption>
<paragraph id="8BA3D9784F7EFFC4DF9EEC89FA7C8270" blockId="8.[151,1437,1524,2017]" box="[199,1437,1524,1549]" pageId="8" pageNumber="513">Measurements (mm). Abdomen without anal appendages 21–22, Hw 20–21. 13–15 Px in Fw, 14–16 Px in Hw.</paragraph>
</subSubSection>
<subSubSection id="C3068AF34F7EFFC5DF9EEF64FCC084A8" lastPageId="9" lastPageNumber="514" pageId="8" pageNumber="513" type="diagnosis">
<paragraph id="8BA3D9784F7EFFC5DF9EEF64FCC084A8" blockId="8.[151,1437,1524,2017]" lastBlockId="9.[151,1436,151,464]" lastPageId="9" lastPageNumber="514" pageId="8" pageNumber="513">
<emphasis id="B968056A4F7EFFC4DF9EEF64FEA0824D" bold="true" box="[199,321,1560,1585]" pageId="8" pageNumber="513">Diagnosis.</emphasis>
Both sexes are easily separated from all others known except 
<taxonomicName id="4C1CA2FB4F7EFFC4DB53EF65FB78824C" authorityName=", Lieftinck" authorityYear="1950" box="[1034,1177,1561,1584]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="513" phylum="Arthropoda" rank="species" species="atomarius">
<emphasis id="B968056A4F7EFFC4DB53EF65FB78824C" box="[1034,1177,1561,1584]" italics="true" pageId="8" pageNumber="513">P. atomarius</emphasis>
</taxonomicName>
by the combination of small size and the speckled pattern of the synthorax. It is separated from 
<taxonomicName id="4C1CA2FB4F7EFFC4DCBAEF41FB938228" authorityName=", Lieftinck" authorityYear="1950" box="[995,1138,1597,1620]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="513" phylum="Arthropoda" rank="species" species="atomarius">
<emphasis id="B968056A4F7EFFC4DCBAEF41FB938228" box="[995,1138,1597,1620]" italics="true" pageId="8" pageNumber="513">P. atomarius</emphasis>
</taxonomicName>
by the presence of a pale brown band on the occiput (compare 
<figureCitation id="1327C5FD4F7EFFC4DD62EF1CFD608205" box="[571,641,1632,1657]" captionStart="FIGURES 1–4" captionStartId="1.[151,264,1826,1849]" captionTargetBox="[151,1435,878,1805]" captionTargetId="figure@1.[151,1435,878,1805]" captionTargetPageId="1" captionText="FIGURES 1–4. Podolestes species males, head. P. atomarius (KAL12_MEG28): (1) dorsal; (3) frontal. P. parvus sp. nov. holotype: (2) dorsal; (4) dorsal-frontal." httpUri="https://zenodo.org/record/2647310/files/figure.png" pageId="8" pageNumber="513">Figs 1</figureCitation>
, 
<figureCitation id="1327C5FD4F7EFFC4DDD6EF1CFD7C8205" box="[655,669,1632,1657]" captionStart="FIGURES 5–8" captionStartId="2.[151,264,1069,1092]" captionTargetBox="[151,1435,193,1049]" captionTargetId="figure@2.[151,1435,193,1049]" captionTargetPageId="2" captionText="FIGURES 5–8. Podolestes species females, head. P. atomarius (KAL12_MEG38): (5) dorsal; (7) frontal. P. parvus sp. nov. paratype (SAR15_ARG4): (6) dorsal; (8) frontal." httpUri="https://zenodo.org/record/2647312/files/figure.png" pageId="8" pageNumber="513">5</figureCitation>
with 
<figureCitation id="1327C5FD4F7EFFC4DD87EF1CFCC48205" box="[734,805,1632,1657]" captionStart="FIGURES 1–4" captionStartId="1.[151,264,1826,1849]" captionTargetBox="[151,1435,878,1805]" captionTargetId="figure@1.[151,1435,878,1805]" captionTargetPageId="1" captionText="FIGURES 1–4. Podolestes species males, head. P. atomarius (KAL12_MEG28): (1) dorsal; (3) frontal. P. parvus sp. nov. holotype: (2) dorsal; (4) dorsal-frontal." httpUri="https://zenodo.org/record/2647310/files/figure.png" pageId="8" pageNumber="513">Figs 2</figureCitation>
, 
<figureCitation id="1327C5FD4F7EFFC4DC68EF1CFCA08205" box="[817,833,1632,1657]" captionStart="FIGURES 5–8" captionStartId="2.[151,264,1069,1092]" captionTargetBox="[151,1435,193,1049]" captionTargetId="figure@2.[151,1435,193,1049]" captionTargetPageId="2" captionText="FIGURES 5–8. Podolestes species females, head. P. atomarius (KAL12_MEG38): (5) dorsal; (7) frontal. P. parvus sp. nov. paratype (SAR15_ARG4): (6) dorsal; (8) frontal." httpUri="https://zenodo.org/record/2647312/files/figure.png" pageId="8" pageNumber="513">6</figureCitation>
), the lack of a blue mark larger than the others on the mesepisternum near the mesinfraepisternum (present in 
<taxonomicName id="4C1CA2FB4F7EFFC4DC4AEFF9FC4382E0" authorityName=", Lieftinck" authorityYear="1950" box="[787,930,1669,1692]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="513" phylum="Arthropoda" rank="species" species="atomarius">
<emphasis id="B968056A4F7EFFC4DC4AEFF9FC4382E0" box="[787,930,1669,1692]" italics="true" pageId="8" pageNumber="513">P. atomarius</emphasis>
</taxonomicName>
) (compare 
<figureCitation id="1327C5FD4F7EFFC4DB7CEFF8FB4482E1" box="[1061,1189,1668,1693]" captionStart="FIGURES 17–20" captionStartId="5.[151,264,1357,1380]" captionTargetBox="[151,1435,481,1337]" captionTargetId="figure@5.[151,1435,481,1337]" captionTargetPageId="5" captionText="FIGURES 17–20. Podolestes species, synthorax dorsal view. P. atomarius: (17) male (KAL12_MEG28); (18) female (KAL12_MEG38). P. parvus sp. nov. (19) holotype; (20) female paratype (SAR15_ARG4)." httpUri="https://zenodo.org/record/2647318/files/figure.png" pageId="8" pageNumber="513">Figs 17, 18</figureCitation>
with 
<figureCitation id="1327C5FD4F7EFFC4DBBEEFF8FA8B82E1" box="[1255,1386,1668,1693]" captionStart="FIGURES 17–20" captionStartId="5.[151,264,1357,1380]" captionTargetBox="[151,1435,481,1337]" captionTargetId="figure@5.[151,1435,481,1337]" captionTargetPageId="5" captionText="FIGURES 17–20. Podolestes species, synthorax dorsal view. P. atomarius: (17) male (KAL12_MEG28); (18) female (KAL12_MEG38). P. parvus sp. nov. (19) holotype; (20) female paratype (SAR15_ARG4)." httpUri="https://zenodo.org/record/2647318/files/figure.png" pageId="8" pageNumber="513">Figs 19, 20</figureCitation>
), by the blue dorsal mark on S9 running the length of the segment whereas it is smaller and apically positioned in 
<taxonomicName id="4C1CA2FB4F7EFFC4DADEEFD5FEE88298" authorityName=", Lieftinck" authorityYear="1950" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="513" phylum="Arthropoda" rank="species" species="atomarius">
<emphasis id="B968056A4F7EFFC4DADEEFD5FEE88298" italics="true" pageId="8" pageNumber="513">P. atomarius</emphasis>
</taxonomicName>
(compare 
<figureCitation id="1327C5FD4F7EFFC4DEDDEFB0FDE58299" box="[388,516,1740,1765]" captionStart="FIGURES 33–36" captionStartId="7.[151,264,1978,2001]" captionTargetBox="[151,1435,1316,1958]" captionTargetId="figure@7.[151,1435,1316,1958]" captionTargetPageId="7" captionText="FIGURES 33–36. Podolestes species, markings of terminal abdominal segments, dorsal view. P. atomarius: (33) male (KAL12_MEG28); (34) female (KAL12_MEG38). P. parvus sp. nov. (35) holotype; (36) female paratype (SAR15_ARG4)." httpUri="https://zenodo.org/record/2647330/files/figure.png" pageId="8" pageNumber="513">Figs 33, 34</figureCitation>
with 
<figureCitation id="1327C5FD4F7EFFC4DD11EFB0FD2D8299" box="[584,716,1740,1765]" captionStart="FIGURES 33–36" captionStartId="7.[151,264,1978,2001]" captionTargetBox="[151,1435,1316,1958]" captionTargetId="figure@7.[151,1435,1316,1958]" captionTargetPageId="7" captionText="FIGURES 33–36. Podolestes species, markings of terminal abdominal segments, dorsal view. P. atomarius: (33) male (KAL12_MEG28); (34) female (KAL12_MEG38). P. parvus sp. nov. (35) holotype; (36) female paratype (SAR15_ARG4)." httpUri="https://zenodo.org/record/2647330/files/figure.png" pageId="8" pageNumber="513">Figs 35, 36</figureCitation>
), and by the ground-colour of all body parts brown rather than black. Both sexes are smaller in 
<taxonomicName id="4C1CA2FB4F7EFFC4DD4DEF8DFD9F8374" authorityName="Dow &amp; Ngiam" authorityYear="2019" box="[532,638,1777,1800]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="513" phylum="Arthropoda" rank="species" species="parvus">
<emphasis id="B968056A4F7EFFC4DD4DEF8DFD9F8374" box="[532,638,1777,1800]" italics="true" pageId="8" pageNumber="513">P. parvus</emphasis>
</taxonomicName>
than 
<taxonomicName id="4C1CA2FB4F7EFFC4DD9AEF8DFBAC8375" authority=", Lieftinck (1950: 44)" authorityName=", Lieftinck" authorityPageNumber="44" authorityYear="1950" box="[707,1101,1776,1801]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="513" phylum="Arthropoda" rank="species" species="atomarius">
<emphasis id="B968056A4F7EFFC4DD9AEF8DFCB38374" box="[707,850,1777,1800]" italics="true" pageId="8" pageNumber="513">P. atomarius</emphasis>
, 
<bibRefCitation id="EF8DA4894F7EFFC4DC3DEF8CFBFB8375" author="Lieftinck, M. A." box="[868,1050,1776,1801]" pageId="8" pageNumber="513" pagination="39 - 47" refId="ref4184" refString="Lieftinck, M. A. (1950) Two new species of Podolestes Selys from Borneo, with a key for the identification of the known species (Odonata, Megapodagriidae). Zoologische Mededelingen, Leiden, 31 (4), 39 - 47." type="journal article" year="1950">Lieftinck (1950</bibRefCitation>
: 44)
</taxonomicName>
gives the length of the male abdomen with appendages in 
<taxonomicName id="4C1CA2FB4F7EFFC4DD5DEE69FD788350" authorityName=", Lieftinck" authorityYear="1950" box="[516,665,1813,1836]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="513" phylum="Arthropoda" rank="species" species="atomarius">
<emphasis id="B968056A4F7EFFC4DD5DEE69FD788350" box="[516,665,1813,1836]" italics="true" pageId="8" pageNumber="513">P. atomarius</emphasis>
</taxonomicName>
as 26.5– 
<quantity id="4CE4749D4F7EFFC4DC57EE68FCB98351" box="[782,856,1812,1837]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="513" unit="mm" value="30.0">30mm</quantity>
, but in 
<taxonomicName id="4C1CA2FB4F7EFFC4DC9AEE69FBD38350" authorityName="Dow &amp; Ngiam" authorityYear="2019" box="[963,1074,1813,1836]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="513" phylum="Arthropoda" rank="species" species="parvus">
<emphasis id="B968056A4F7EFFC4DC9AEE69FBD38350" box="[963,1074,1813,1836]" italics="true" pageId="8" pageNumber="513">P. parvus</emphasis>
</taxonomicName>
it is ca 
<quantity id="4CE4749D4F7EFFC4DBF3EE68FAFC8351" box="[1194,1309,1812,1837]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="8" pageNumber="513" unit="mm" value="24.0" valueMax="25.0" valueMin="23.0">23–25mm</quantity>
, the same measurements for the female are 25.5–27.5mm in 
<taxonomicName id="4C1CA2FB4F7EFFC4DD83EE45FC88832C" authorityName=", Lieftinck" authorityYear="1950" box="[730,873,1849,1872]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="513" phylum="Arthropoda" rank="species" species="atomarius">
<emphasis id="B968056A4F7EFFC4DD83EE45FC88832C" box="[730,873,1849,1872]" italics="true" pageId="8" pageNumber="513">P. atomarius</emphasis>
</taxonomicName>
compared with ca 21.5– 
<quantity id="4CE4749D4F7EFFC4DBDFEE44FB2E832D" box="[1158,1231,1848,1873]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="8" pageNumber="513" unit="mm" value="22.0">22mm</quantity>
in 
<taxonomicName id="4C1CA2FB4F7EFFC4DBAFEE45FA81832C" authorityName="Dow &amp; Ngiam" authorityYear="2019" box="[1270,1376,1849,1872]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="513" phylum="Arthropoda" rank="species" species="parvus">
<emphasis id="B968056A4F7EFFC4DBAFEE45FA81832C" box="[1270,1376,1849,1872]" italics="true" pageId="8" pageNumber="513">P. parvus</emphasis>
</taxonomicName>
. The males of the two species are further separated by the differently shaped ends of the arms of the terminal segment of the genital ligula, more abruptly and further expanded in 
<taxonomicName id="4C1CA2FB4F7EFFC4DC42EEFDFC6283E4" authorityName="Dow &amp; Ngiam" authorityYear="2019" box="[795,899,1921,1944]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="513" phylum="Arthropoda" rank="species" species="parvus">
<emphasis id="B968056A4F7EFFC4DC42EEFDFC6283E4" box="[795,899,1921,1944]" italics="true" pageId="8" pageNumber="513">P. parvus</emphasis>
</taxonomicName>
than in 
<taxonomicName id="4C1CA2FB4F7EFFC4DC86EEFDFB8C83E4" authorityName=", Lieftinck" authorityYear="1950" box="[991,1133,1921,1944]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="8" pageNumber="513" phylum="Arthropoda" rank="species" species="atomarius">
<emphasis id="B968056A4F7EFFC4DC86EEFDFB8C83E4" box="[991,1133,1921,1944]" italics="true" pageId="8" pageNumber="513">P. atomarius</emphasis>
</taxonomicName>
(compare 
<figureCitation id="1327C5FD4F7EFFC4DBBCEEFCFA6E83E5" box="[1253,1423,1920,1945]" captionStart="FIGURES 29–32" captionStartId="7.[151,264,1212,1235]" captionTargetBox="[151,1435,193,1192]" captionTargetId="figure@7.[151,1435,193,1192]" captionTargetPageId="7" captionText="FIGURES 29–32. Genital ligula of Podolestes species. P. atomarius (KAL12_MEG1): (29) ventral; (31) lateral. P. parvus sp. nov. holotype: (30) ventral; (32) lateral. All scale bars 0.2mm." httpUri="https://zenodo.org/record/2647328/files/figure.png" pageId="8" pageNumber="513">Figs 31 and 32</figureCitation>
). The differences in the position of the arms of the terminal segment of the genital ligula apparent in 
<figureCitation id="1327C5FD4F7EFFC4DBACEED8FA9283C1" box="[1269,1395,1956,1981]" captionStart="FIGURES 29–32" captionStartId="7.[151,264,1212,1235]" captionTargetBox="[151,1435,193,1192]" captionTargetId="figure@7.[151,1435,193,1192]" captionTargetPageId="7" captionText="FIGURES 29–32. Genital ligula of Podolestes species. P. atomarius (KAL12_MEG1): (29) ventral; (31) lateral. P. parvus sp. nov. holotype: (30) ventral; (32) lateral. All scale bars 0.2mm." httpUri="https://zenodo.org/record/2647328/files/figure.png" pageId="8" pageNumber="513">Figs 29–32</figureCitation>
are not likely to be diagnostic but are probably due to the process of extraction. The male anal appendages are very similar in the two species, the most notable difference is that in 
<taxonomicName id="4C1CA2FB4F7FFFC5DC04E9E5FC2584CC" authorityName="Dow &amp; Ngiam" authorityYear="2019" box="[861,964,153,176]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="514" phylum="Arthropoda" rank="species" species="parvus">
<emphasis id="B968056A4F7FFFC5DC04E9E5FC2584CC" box="[861,964,153,176]" italics="true" pageId="9" pageNumber="514">P. parvus</emphasis>
</taxonomicName>
the cerci are slightly longer relative to the paraprocts than in 
<taxonomicName id="4C1CA2FB4F7FFFC5DE3FE9C1FE1584A8" authorityName=", Lieftinck" authorityYear="1950" box="[358,500,189,212]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="514" phylum="Arthropoda" rank="species" species="atomarius">
<emphasis id="B968056A4F7FFFC5DE3FE9C1FE1584A8" box="[358,500,189,212]" italics="true" pageId="9" pageNumber="514">P. atomarius</emphasis>
</taxonomicName>
(compare 
<figureCitation id="1327C5FD4F7FFFC5DD32E9C0FCF484A8" box="[619,789,188,213]" captionStart="FIGURES 37–40" captionStartId="8.[151,264,1426,1449]" captionTargetBox="[151,1435,193,1405]" captionTargetId="figure@8.[151,1435,193,1405]" captionTargetPageId="8" captionText="FIGURES 37–40. Anal appendages of Podolestes species males. P. atomarius (KAL12_MEG2): (37) dorsal; (39) lateral. P. parvus sp. nov. holotype: (38) dorsal; (40) lateral. All scale bars 0.5mm." httpUri="https://zenodo.org/record/2647332/files/figure.png" pageId="9" pageNumber="514">Figs 39 and 40</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3068AF34F7FFFC5DF9EE9A3FA7A85AC" pageId="9" pageNumber="514" type="discussion">
<paragraph id="8BA3D9784F7FFFC5DF9EE9A3FB018518" blockId="9.[151,1436,151,464]" pageId="9" pageNumber="514">
<emphasis id="B968056A4F7FFFC5DF9EE9A3FED58484" bold="true" box="[199,308,223,248]" pageId="9" pageNumber="514">Remarks</emphasis>
. The pterostigma is brown or greyish brown in both sexes of 
<taxonomicName id="4C1CA2FB4F7FFFC5DB5BE99DFB8D8484" authorityName="Dow &amp; Ngiam" authorityYear="2019" box="[1026,1132,225,248]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="514" phylum="Arthropoda" rank="species" species="parvus">
<emphasis id="B968056A4F7FFFC5DB5BE99DFB8D8484" box="[1026,1132,225,248]" italics="true" pageId="9" pageNumber="514">P. parvus</emphasis>
</taxonomicName>
but yellow cream in most individuals of 
<taxonomicName id="4C1CA2FB4F7FFFC5DE63E879FE298560" authorityName=", Lieftinck" authorityYear="1950" box="[314,456,261,284]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="514" phylum="Arthropoda" rank="species" species="atomarius">
<emphasis id="B968056A4F7FFFC5DE63E879FE298560" box="[314,456,261,284]" italics="true" pageId="9" pageNumber="514">P. atomarius</emphasis>
</taxonomicName>
, becoming darker with age as noted by 
<bibRefCitation id="EF8DA4894F7FFFC5DCD7E878FBA08561" author="Lieftinck, M. A." box="[910,1089,260,285]" pageId="9" pageNumber="514" pagination="39 - 47" refId="ref4184" refString="Lieftinck, M. A. (1950) Two new species of Podolestes Selys from Borneo, with a key for the identification of the known species (Odonata, Megapodagriidae). Zoologische Mededelingen, Leiden, 31 (4), 39 - 47." type="journal article" year="1950">Lieftinck (1950</bibRefCitation>
: 42). However in more mature individuals of 
<taxonomicName id="4C1CA2FB4F7FFFC5DE63E855FE29853C" authorityName=", Lieftinck" authorityYear="1950" box="[314,456,297,320]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="514" phylum="Arthropoda" rank="species" species="atomarius">
<emphasis id="B968056A4F7FFFC5DE63E855FE29853C" box="[314,456,297,320]" italics="true" pageId="9" pageNumber="514">P. atomarius</emphasis>
</taxonomicName>
examined by the first author the underside of the pterostigma (so the side most visible in unpinned specimens) is brown but the upper side is whitish (compare 
<figureCitation id="1327C5FD4F7FFFC5DC9AE830FBA18519" box="[963,1088,332,357]" captionStart="FIGURES 25–28" captionStartId="6.[151,264,1978,2001]" captionTargetBox="[151,1435,1316,1958]" captionTargetId="figure@6.[151,1435,1316,1958]" captionTargetPageId="6" captionText="FIGURES 25–28. Pterostigma of Podolestes species. P. atomarius: (25) male Fw (KAL12_MEG28); (26) male Hw (KAL12_MEG28); (27) female Fw (KAL12_MEG38). (28) P. parvus female (SAR15_ARG4) Hw." httpUri="https://zenodo.org/record/2647322/files/figure.png" pageId="9" pageNumber="514">Figs 25–27</figureCitation>
with 
<figureCitation id="1327C5FD4F7FFFC5DBD9E830FB328518" box="[1152,1235,332,357]" captionStart="FIGURES 25–28" captionStartId="6.[151,264,1978,2001]" captionTargetBox="[151,1435,1316,1958]" captionTargetId="figure@6.[151,1435,1316,1958]" captionTargetPageId="6" captionText="FIGURES 25–28. Pterostigma of Podolestes species. P. atomarius: (25) male Fw (KAL12_MEG28); (26) male Hw (KAL12_MEG28); (27) female Fw (KAL12_MEG38). (28) P. parvus female (SAR15_ARG4) Hw." httpUri="https://zenodo.org/record/2647322/files/figure.png" pageId="9" pageNumber="514">Fig. 28</figureCitation>
).
</paragraph>
<paragraph id="8BA3D9784F7FFFC5DF9EE813FB2A85D1" blockId="9.[151,1436,151,464]" pageId="9" pageNumber="514">
The female is associated with the male by reasonable supposition; it has been found in the same locations and habitats as the males and is not that of any other 
<taxonomicName id="4C1CA2FB4F7FFFC5DDE1E8E8FCD085D1" authorityName="Selys" authorityYear="1862" box="[696,817,404,429]" class="Insecta" family="Megapodagrionidae" genus="Podolestes" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="514" phylum="Arthropoda" rank="genus">
<emphasis id="B968056A4F7FFFC5DDE1E8E8FCD085D1" box="[696,817,404,429]" italics="true" pageId="9" pageNumber="514">Podolestes</emphasis>
</taxonomicName>
species occurring at these locations.
</paragraph>
<paragraph id="8BA3D9784F7FFFC5DF9EE8CBFA7A85AC" blockId="9.[151,1436,151,464]" box="[199,1435,439,464]" pageId="9" pageNumber="514">
The 
<typeStatus id="54A767DA4F7FFFC5DFA0E8CBFEBA85AC" box="[249,347,439,464]" pageId="9" pageNumber="514" type="holotype">holotype</typeStatus>
male and one of the 
<typeStatus id="54A767DA4F7FFFC5DD1FE8C4FD4785AC" box="[582,678,440,464]" pageId="9" pageNumber="514" type="paratype">paratype</typeStatus>
females are shown in life in 
<figureCitation id="1327C5FD4F7FFFC5DCB2E8CBFB8A85AC" box="[1003,1131,439,464]" captionStart="FIGURES 42–45" captionStartId="10.[151,264,962,985]" captionTargetBox="[151,1435,193,942]" captionTargetId="figure@10.[151,1435,193,942]" captionTargetPageId="10" captionText="FIGURES 42–45. Podolestes parvus sp. nov. in life. Holotype male: (42) whole animal in lateral view; (44) head and thorax in dorsal view. Female paratype (SAR11_12_MEG95): (43) whole animal in lateral view; (45) head and thorax in dorsal view. Photographs by R.W.J. Ngiam." httpUri="https://zenodo.org/record/2647336/files/figure.png" pageId="9" pageNumber="514">Figs 42–45</figureCitation>
, the distribution in 
<figureCitation id="1327C5FD4F7FFFC5DA1CE8CBFA7685AC" box="[1349,1431,439,464]" captionStart="FIGURE 41" captionStartId="9.[151,250,1912,1934]" captionTargetBox="[151,1435,519,1890]" captionTargetId="figure@9.[151,1435,519,1890]" captionTargetPageId="9" captionText="FIGURE 41. Distribution of Podolestes species. Yellow circles: P. atomarius; red squares: P. parvus." httpUri="https://zenodo.org/record/2647334/files/figure.png" pageId="9" pageNumber="514">Fig. 41</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>