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<document id="96990093159106423E01CCDA1A7BB8CC" ID-CLB-Dataset="6634" ID-DOI="10.5852/ejt.2021.749.1355" ID-GBIF-Dataset="83109e54-596a-4be0-ac2c-c4e1702cfb8a" ID-ISSN="2118-9773" ID-Zenodo-Dep="4770293" ID-ZooBank="25FD5744-861D-45E4-B6AD-350716AA29BF" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1621349695117" checkinUser="felipe" docAuthor="Gao, Chuan-bu &amp; Coca-Abia, María Milagro" docDate="2021" docId="03B56743A6290637FD8DFBF6FECEE19B" docLanguage="en" docName="ejt-749_1-94.pdf" docOrigin="European Journal of Taxonomy 749" docSource="http://dx.doi.org/10.5852/ejt.2021.749.1355" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Miridiba hirsuta Itoh 2001" docType="treatment" docVersion="8" lastPageNumber="30" masterDocId="FF8C1F3BA635062AFFFBFFA9FFE2E431" masterDocTitle="Revision of the genus Miridiba Reitter, 1902 (Coleoptera, Scarabaeidae, Melolonthinae): genital morphotypes and new taxonomic data" masterLastPageNumber="94" masterPageNumber="1" pageNumber="29" updateTime="1698921317801" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="EF3510528E3EC43BDD516D8EEAE3F7B2">Revision of the genus Miridiba Reitter, 1902 (Coleoptera, Scarabaeidae, Melolonthinae): genital morphotypes and new taxonomic data</mods:title>
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<mods:name id="606DEDDB2B27FAFF76C3B171AF8720B9" type="personal">
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<mods:namePart id="9969DB8090F2D7C234362FCB8592FCF8">Gao, Chuan-bu</mods:namePart>
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<mods:affiliation id="BBA16C65FAF832681BD1CAA6B616B05F">Guangdong Key Laboratory of Animal Conservation and Resource Utilization, Guangdong Public Laboratory of Wild Animal Conservation and Utilization, Institute of Zoology, Guangdong Academy of Sciences, 105 Xingang Road W., Guangzhou 510260, China. Unidad de Protección Vegetal. Centro de Investigación y Tecnología Agroalimentaria (CITA) e Instituto Agroalimentario de Aragón (IA 2) (CITA-Universidad de Zaragoza), Avenida Montañana, 930. 50059 Zaragoza, Spain. &amp; Buzh.</mods:affiliation>
<mods:nameIdentifier id="E933E05A347893EE04F3CEE6979067E8" type="email">Buzh.Gao@msn.com</mods:nameIdentifier>
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<mods:namePart id="64E8A28019363F871D3A4EFC58159800">Coca-Abia, María Milagro</mods:namePart>
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<mods:nameIdentifier id="A42CCD271966091C7A5FF936522C3EAF" type="email">mcoca@aragon.es</mods:nameIdentifier>
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<mods:title id="F8D3CEA746CA8F89E4C2FFAC6FFA70D8">European Journal of Taxonomy</mods:title>
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<mods:part id="98F36D762C6F8BAA48055D9DFA09CCB0">
<mods:date id="75CE28B6CFB9C32EAED89F029AB9A180">2021</mods:date>
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<mods:number id="CF76532770611AF8A5E26A1DAB14F8AD">2021-05-12</mods:number>
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<treatment id="03B56743A6290637FD8DFBF6FECEE19B" ID-DOI="http://doi.org/10.5281/zenodo.4773974" ID-GBIF-Taxon="182237220" ID-Zenodo-Dep="4773974" LSID="urn:lsid:plazi:treatment:03B56743A6290637FD8DFBF6FECEE19B" httpUri="http://treatment.plazi.org/id/03B56743A6290637FD8DFBF6FECEE19B" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
<subSubSection id="C30685DEA6290636FD8DFBF6FC5CE048" box="[630,958,1119,1146]" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="8BA3D655A6290636FD8DFBF6FC5CE048" blockId="28.[630,958,1119,1181]" box="[630,958,1119,1146]" pageId="28" pageNumber="29">
<heading id="D0EB6139A6290636FD8DFBF6FC5CE048" box="[630,958,1119,1146]" centered="true" fontSize="11" level="2" pageId="28" pageNumber="29" reason="2">
<taxonomicName id="4C1CADD6A6290636FD8DFBF6FC5CE048" authority="Itoh, 2001" authorityName="Itoh" authorityYear="2001" box="[630,958,1119,1146]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="B9680A47A6290636FD8DFBF6FCDFE04B" bold="true" box="[630,829,1119,1146]" italics="true" pageId="28" pageNumber="29">Miridiba hirsuta</emphasis>
<bibRefCitation id="EF8DABA4A6290636FCBEFBF6FC5CE048" author="Itoh T." box="[837,958,1119,1145]" pageId="28" pageNumber="29" pagination="435 - 439" refId="ref44461" refString="Itoh T. 2001. A new species of the genus Miridiba (Scarabaeidae: Melolonthinae: Melolonthini) from the Yoyama Islands, Southwest Japan. Elytra, Tokyo 29 (2): 435 - 439." type="journal article" year="2001">Itoh, 2001</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA6290636FD23FB2BFCB9E0AC" box="[728,859,1154,1181]" pageId="28" pageNumber="29" type="description">
<paragraph id="8BA3D655A6290636FD23FB2BFCB9E0AC" blockId="28.[630,958,1119,1181]" box="[728,859,1154,1181]" pageId="28" pageNumber="29">
<figureCitation id="1327CAD0A6290636FD23FB2BFCB9E0AC" box="[728,859,1154,1181]" captionStart="Figs 37–40" captionStartId="29.[187,239,1221,1247]" captionTargetBox="[279,1316,284,1175]" captionTargetId="figure-124@29.[264,1323,273,1182]" captionTargetPageId="29" captionText="Figs 37–40. Miridiba hirsuta Itoh, 2001 (non-types, TMC). 37. Habitus in dorsal view. 38. Parameres in dorsal (A) and lateral (B) views. 39. Endophallus and temones in dorsal view. 40. Female genitalia in dorsal view. Scale bars: 38–40 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770348" httpUri="https://zenodo.org/record/4770348/files/figure.png" pageId="28" pageNumber="29">Figs 37–40</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA6290636FF46FB6DFDB6E175" pageId="28" pageNumber="29" type="reference_group">
<paragraph id="8BA3D655A6290636FF46FB6DFBA5E0EE" blockId="28.[189,1095,1220,1247]" box="[189,1095,1220,1247]" pageId="28" pageNumber="29">
<treatmentCitationGroup id="AB0CF17BA6290636FF46FB6DFBA5E0EE" box="[189,1095,1220,1247]" pageId="28" pageNumber="29">
<treatmentCitation id="0ABDF044A6290636FF46FB6DFDDCE0EE" author="Itoh T." box="[189,574,1220,1247]" page="435" pageId="28" pageNumber="29" year="2001">
<taxonomicName id="4C1CADD6A6290636FF46FB6DFDDCE0EE" authority="Itoh, 2001: 435" authorityName="Itoh" authorityPageNumber="435" authorityYear="2001" box="[189,574,1220,1247]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="B9680A47A6290636FF46FB6DFE62E0EF" box="[189,384,1220,1246]" italics="true" pageId="28" pageNumber="29">Miridiba hirsuta</emphasis>
<bibRefCitation id="EF8DABA4A6290636FE7CFB6DFDDCE0EE" author="Itoh T." box="[391,574,1220,1247]" pageId="28" pageNumber="29" pagination="435 - 439" refId="ref44461" refString="Itoh T. 2001. A new species of the genus Miridiba (Scarabaeidae: Melolonthinae: Melolonthini) from the Yoyama Islands, Southwest Japan. Elytra, Tokyo 29 (2): 435 - 439." type="journal article" year="2001">Itoh, 2001: 435</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 1–9 (
<typeStatus id="54A768F7A6290636FD4CFB6DFD09E0EF" box="[695,747,1220,1246]" pageId="28" pageNumber="29">type</typeStatus>
loc.: Ishigakijima, Ryukyus).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA3D655A6290636FF46FAACFDB6E175" blockId="28.[189,1400,1285,1348]" pageId="28" pageNumber="29">
<treatmentCitationGroup id="AB0CF17BA6290636FF46FAACFDB6E175" pageId="28" pageNumber="29">
<taxonomicName id="4C1CADD6A6290636FF46FAACFE63E12E" authorityName="Itoh" authorityYear="2001" box="[189,385,1285,1311]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="B9680A47A6290636FF46FAACFE63E12E" box="[189,385,1285,1311]" italics="true" pageId="28" pageNumber="29">Miridiba hirsuta</emphasis>
</taxonomicName>
–
<treatmentCitation id="0ABDF044A6290636FE58FAAFFD07E111" author="Smetana A. &amp; Kral D." box="[419,741,1285,1312]" page="222" pageId="28" pageNumber="29" year="2006">
<bibRefCitation id="EF8DABA4A6290636FE58FAAFFD07E111" author="Smetana A. &amp; Kral D." box="[419,741,1285,1312]" pageId="28" pageNumber="29" pagination="207 - 228" refId="ref45912" refString="Smetana A. &amp; Kral D. 2006. Tribe Rhizotrogini Burmeister, 1855. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea: 207 - 228. Apollo Books, Stenstrup, Denmark." type="book chapter" year="2006">Smetana &amp; Král 2006: 222</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="0ABDF044A6290636FC53FAACFB49E12E" author="Coca-Abia M. M." box="[936,1195,1285,1312]" page="680" pageId="28" pageNumber="29" year="2008">
<bibRefCitation id="EF8DABA4A6290636FC53FAACFB49E12E" author="Coca-Abia M. M." box="[936,1195,1285,1312]" pageId="28" pageNumber="29" pagination="673 - 685" refId="ref43499" refString="Coca-Abia M. M. 2008. Revision of the genus Miridiba Reitter, 1901 (Coleoptera, Scarabaeidae, Melolonthinae). Zoological Science 25 (6): 673 - 685. https: // doi. org / 10.2108 / zsj. 25.673" type="journal article" year="2008">Coca-Abia 2008: 680</bibRefCitation>
</treatmentCitation>
, 683 (in key). —
<treatmentCitation id="0ABDF044A6290636FF17FA80FE23E172" author="Bezdek A." box="[236,449,1320,1347]" page="271" pageId="28" pageNumber="29" year="2016">
<bibRefCitation id="EF8DABA4A6290636FF17FA80FE23E172" author="Bezdek A." box="[236,449,1320,1347]" pageId="28" pageNumber="29" pagination="249 - 280" refId="ref43171" refString="Bezdek A. 2016. Tribe Rhizotrogini Burmeister, 1855. In: Lobl I. &amp; Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Vol. 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition: 249 - 280. Koninklijke Brill NV, Leiden, The Netherlands." type="book chapter" year="2016">Bezděk 2016: 271</bibRefCitation>
</treatmentCitation>
(catalogue).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA6290636FF46FAC3FB80E182" box="[189,1122,1386,1459]" pageId="28" pageNumber="29" type="diagnosis">
<paragraph id="8BA3D655A6290636FF46FAC3FED4E1B5" blockId="28.[189,310,1386,1412]" box="[189,310,1386,1412]" pageId="28" pageNumber="29">
<heading id="D0EB6139A6290636FF46FAC3FED4E1B5" bold="true" box="[189,310,1386,1412]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="B9680A47A6290636FF46FAC3FED4E1B5" bold="true" box="[189,310,1386,1412]" pageId="28" pageNumber="29">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A6290636FF46FA30FB80E182" blockId="28.[189,1122,1433,1459]" box="[189,1122,1433,1459]" pageId="28" pageNumber="29">
See
<bibRefCitation id="EF8DABA4A6290636FF14FA30FE95E182" author="Itoh T." box="[239,375,1433,1459]" pageId="28" pageNumber="29" pagination="435 - 439" refId="ref44461" refString="Itoh T. 2001. A new species of the genus Miridiba (Scarabaeidae: Melolonthinae: Melolonthini) from the Yoyama Islands, Southwest Japan. Elytra, Tokyo 29 (2): 435 - 439." type="journal article" year="2001">Itoh (2001)</bibRefCitation>
. External morphology of adult and male genitalia (
<figureCitation id="1327CAD0A6290636FC37FA30FBB4E182" box="[972,1110,1433,1459]" captionStart="Figs 37–40" captionStartId="29.[187,239,1221,1247]" captionTargetBox="[279,1316,284,1175]" captionTargetId="figure-124@29.[264,1323,273,1182]" captionTargetPageId="29" captionText="Figs 37–40. Miridiba hirsuta Itoh, 2001 (non-types, TMC). 37. Habitus in dorsal view. 38. Parameres in dorsal (A) and lateral (B) views. 39. Endophallus and temones in dorsal view. 40. Female genitalia in dorsal view. Scale bars: 38–40 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770348" httpUri="https://zenodo.org/record/4770348/files/figure.png" pageId="28" pageNumber="29">Figs 37–40</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA6290636FF46FA73FC1EE277" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="8BA3D655A6290636FF46FA73FE48E1C5" blockId="28.[189,426,1498,1524]" box="[189,426,1498,1524]" pageId="28" pageNumber="29">
<heading id="D0EB6139A6290636FF46FA73FE48E1C5" bold="true" box="[189,426,1498,1524]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="B9680A47A6290636FF46FA73FE48E1C5" bold="true" box="[189,426,1498,1524]" pageId="28" pageNumber="29">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A6290636FF46F9A0FC1EE277" blockId="28.[189,1399,1544,1606]" pageId="28" pageNumber="29">
<materialsCitation id="3B74DC08A6290636FF46F9A0FED5E277" ID-GBIF-Occurrence="3114121313" collectingDate="2004-02-29" collectionCode="TMC" collectorName="Is. Ishigakijima &amp; T. Nakata" country="Japan" determinerName="Matsumoto" location="Tonoshiro-Takeda" pageId="28" pageNumber="29" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa">
<collectingCountry id="F30B96C5A6290636FF46F9A0FEF4E212" box="[189,278,1545,1571]" name="Japan" pageId="28" pageNumber="29">JAPAN</collectingCountry>
•
<specimenCount id="9D1A1DDCA6290636FECBF9A0FEBDE212" box="[304,351,1545,1571]" pageId="28" pageNumber="29" type="male">1 ♂</specimenCount>
;
<collectingRegion id="49D818B7A6290636FE96F9A1FE38E213" box="[365,474,1544,1570]" country="Japan" name="Okinawa" pageId="28" pageNumber="29">Okinawa</collectingRegion>
,
<collectorName id="26E9B383A6290636FE1DF9A0FD42E213" box="[486,672,1544,1571]" pageId="28" pageNumber="29">Is. Ishigakijima</collectorName>
,
<location id="8EC3808EA6290636FD57F9A1FC67E213" LSID="urn:lsid:plazi:treatment:03B56743A6290637FD8DFBF6FECEE19B:8EC3808EA6290636FD57F9A1FC67E213" box="[684,901,1544,1570]" country="Japan" name="Tonoshiro-Takeda" pageId="28" pageNumber="29" stateProvince="Okinawa">Tonoshiro-Takeda</location>
;
<date id="FFA2F095A6290636FC6BF9A0FBCDE212" box="[912,1071,1544,1571]" pageId="28" pageNumber="29" value="2004-02-29">
<collectingDate id="EFE6097DA6290636FC6BF9A0FBCDE212" box="[912,1071,1544,1571]" pageId="28" pageNumber="29" value="2004-02-29">29 Feb. 2004</collectingDate>
</date>
;
<collectorName id="26E9B383A6290636FBC0F9A0FB4FE213" box="[1083,1197,1544,1571]" pageId="28" pageNumber="29">T. Nakata</collectorName>
leg.;
<determinerName id="7E7F3A91A6290636FB0AF9A0FA95E212" box="[1265,1399,1545,1571]" pageId="28" pageNumber="29">Matsumoto</determinerName>
det.;
<collectionCode id="ED0D4E90A6290636FF0CF985FED5E277" box="[247,311,1580,1606]" country="USA" httpUri="http://grbio.org/cool/vnt5-s7fx" name="Trudeau Mycobacterial Culture Collection, Trudeau Institute" pageId="28" pageNumber="29">TMC</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B74DC08A6290636FEABF985FC15E277" ID-GBIF-Occurrence="3114121344" box="[336,1015,1580,1606]" collectingDate="2004-02-29" collectionCode="TMC" collectorName="Is. Ishigakijima &amp; T. Nakata" country="Japan" determinerName="Matsumoto" location="Tonoshiro-Takeda" pageId="28" pageNumber="29" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Okinawa">
<specimenCount id="9D1A1DDCA6290636FEABF985FE76E277" box="[336,404,1580,1606]" pageId="28" pageNumber="29" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D1A1DDCA6290636FE5BF985FE01E277" box="[416,483,1580,1606]" pageId="28" pageNumber="29" type="female">2 ♀♀</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED0D4E90A6290636FC4FF985FC15E277" box="[948,1015,1580,1606]" country="USA" httpUri="http://grbio.org/cool/vnt5-s7fx" name="Trudeau Mycobacterial Culture Collection, Trudeau Institute" pageId="28" pageNumber="29">TMC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA6290636FF46F9C4FAE1E3FE" pageId="28" pageNumber="29" type="discussion">
<paragraph id="8BA3D655A6290636FF46F9C4FECDE2B6" blockId="28.[189,303,1645,1671]" box="[189,303,1645,1671]" pageId="28" pageNumber="29">
<heading id="D0EB6139A6290636FF46F9C4FECDE2B6" bold="true" box="[189,303,1645,1671]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="B9680A47A6290636FF46F9C4FECDE2B6" bold="true" box="[189,303,1645,1671]" pageId="28" pageNumber="29">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A6290636FF46F935FAE1E3FE" blockId="28.[188,1399,1692,2000]" pageId="28" pageNumber="29">
<bibRefCitation id="EF8DABA4A6290636FF46F935FEA4E287" author="Itoh T." box="[189,326,1692,1718]" pageId="28" pageNumber="29" pagination="435 - 439" refId="ref44461" refString="Itoh T. 2001. A new species of the genus Miridiba (Scarabaeidae: Melolonthinae: Melolonthini) from the Yoyama Islands, Southwest Japan. Elytra, Tokyo 29 (2): 435 - 439." type="journal article" year="2001">Itoh (2001)</bibRefCitation>
described
<taxonomicName id="4C1CADD6A6290636FE34F935FD75E287" authorityName="Itoh" authorityYear="2001" box="[463,663,1692,1718]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="B9680A47A6290636FE34F935FD75E287" box="[463,663,1692,1718]" italics="true" pageId="28" pageNumber="29">Miridiba hirsuta</emphasis>
</taxonomicName>
based on a male
<typeStatus id="54A768F7A6290636FC82F935FC02E287" box="[889,992,1692,1718]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
and
<specimenCount id="9D1A1DDCA6290636FBD9F935FB29E287" box="[1058,1227,1692,1718]" pageId="28" pageNumber="29" type="generic" typeStatus="paratypes">five paratypes</specimenCount>
(
<specimenCount id="9D1A1DDCA6290636FB1BF935FAA2E287" box="[1248,1344,1692,1718]" pageId="28" pageNumber="29" type="male">2 males</specimenCount>
and
<specimenCount id="9D1A1DDCA6290636FF46F916FED5E2EB" box="[189,311,1727,1754]" pageId="28" pageNumber="29" type="female">3 females</specimenCount>
). According to the author (
<bibRefCitation id="EF8DABA4A6290636FD86F916FD15E2E8" author="Itoh T." box="[637,759,1727,1753]" pageId="28" pageNumber="29" pagination="435 - 439" refId="ref44461" refString="Itoh T. 2001. A new species of the genus Miridiba (Scarabaeidae: Melolonthinae: Melolonthini) from the Yoyama Islands, Southwest Japan. Elytra, Tokyo 29 (2): 435 - 439." type="journal article" year="2001">Itoh 2001</bibRefCitation>
), this species is closely allied to
<taxonomicName id="4C1CADD6A6290636FB72F969FADDE2E8" authorityName="CH. Bu Gao &amp; M. Coca-Abia" baseAuthorityName="Fairmaire" baseAuthorityYear="1891" box="[1161,1343,1727,1753]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="trichophora">
<emphasis id="B9680A47A6290636FB72F969FADDE2E8" box="[1161,1343,1727,1753]" italics="true" pageId="28" pageNumber="29">M. trichophora</emphasis>
</taxonomicName>
, but
<taxonomicName id="4C1CADD6A6290636FF46F94AFED9E2CD" authorityName="Itoh" authorityYear="2001" box="[189,315,1762,1788]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="B9680A47A6290636FF46F94AFED9E2CD" box="[189,315,1762,1788]" italics="true" pageId="28" pageNumber="29">M. hirsuta</emphasis>
</taxonomicName>
is distinguishable by the larger size, sharper apices of upper branches of parameres, and antennal club weakly more elongate in male. We have studied
<specimenCount id="9D1A1DDCA6290636FC6FF8ACFBA3E32E" box="[916,1089,1797,1823]" pageId="28" pageNumber="29" type="generic">five specimens</specimenCount>
(TMC) collected from the type locality and identified by Mr Matsumoto as
<taxonomicName id="4C1CADD6A6290636FD08F880FC8EE373" authorityName="Itoh" authorityYear="2001" box="[755,876,1832,1858]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="B9680A47A6290636FD08F880FC8EE373" box="[755,876,1832,1858]" italics="true" pageId="28" pageNumber="29">M. hirsuta</emphasis>
</taxonomicName>
, observing some differences with the type of
<taxonomicName id="4C1CADD6A6290636FF47F8E4FE8CE357" authorityName="CH. Bu Gao &amp; M. Coca-Abia" baseAuthorityName="Fairmaire" baseAuthorityYear="1891" box="[188,366,1868,1894]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="trichophora">
<emphasis id="B9680A47A6290636FF47F8E4FE8CE357" box="[188,366,1868,1894]" italics="true" pageId="28" pageNumber="29">M. trichophora</emphasis>
</taxonomicName>
in endophallus. However, given that we were unable to examine the types of
<taxonomicName id="4C1CADD6A6290636FB0EF8E4FA8CE357" authorityName="Itoh" authorityYear="2001" box="[1269,1390,1868,1894]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="B9680A47A6290636FB0EF8E4FA8CE357" box="[1269,1390,1868,1894]" italics="true" pageId="28" pageNumber="29">M. hirsuta</emphasis>
</taxonomicName>
, it is not possible to confirm that these differences characterize
<taxonomicName id="4C1CADD6A6290636FC66F8D9FBF5E3B8" authorityName="Itoh" authorityYear="2001" box="[925,1047,1903,1929]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="B9680A47A6290636FC66F8D9FBF5E3B8" box="[925,1047,1903,1929]" italics="true" pageId="28" pageNumber="29">M. hirsuta</emphasis>
</taxonomicName>
. Until the types of
<taxonomicName id="4C1CADD6A6290636FB07F8D9FA95E3B8" authorityName="Itoh" authorityYear="2001" box="[1276,1399,1903,1929]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="B9680A47A6290636FB07F8D9FA95E3B8" box="[1276,1399,1903,1929]" italics="true" pageId="28" pageNumber="29">M. hirsuta</emphasis>
</taxonomicName>
are studied, we consider it a valid species. According to the antenna 9-segmented and the parameres described by
<bibRefCitation id="EF8DABA4A6290636FEA2F81CFE03E3FE" author="Itoh T." box="[345,481,1973,1999]" pageId="28" pageNumber="29" pagination="435 - 439" refId="ref44461" refString="Itoh T. 2001. A new species of the genus Miridiba (Scarabaeidae: Melolonthinae: Melolonthini) from the Yoyama Islands, Southwest Japan. Elytra, Tokyo 29 (2): 435 - 439." type="journal article" year="2001">Itoh (2001)</bibRefCitation>
, this species is included in this genital morphotype I “
<taxonomicName id="4C1CADD6A6290636FBA7F81CFB0DE3FE" box="[1116,1263,1973,1999]" class="Insecta" family="Tachinidae" genus="Trichophora" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A6290636FBA7F81CFB0DE3FE" box="[1116,1263,1973,1999]" italics="true" pageId="28" pageNumber="29">Trichophora</emphasis>
</taxonomicName>
”.
</paragraph>
</subSubSection>
<caption id="DF6386DDA6280637FF40FB6CFD40E117" ID-DOI="http://doi.org/10.5281/zenodo.4770348" ID-Zenodo-Dep="4770348" httpUri="https://zenodo.org/record/4770348/files/figure.png" pageId="29" pageNumber="30" startId="29.[187,239,1221,1247]" targetBox="[279,1316,284,1175]" targetPageId="29">
<paragraph id="8BA3D655A6280637FF40FB6CFD40E117" blockId="29.[187,1399,1221,1318]" pageId="29" pageNumber="30">
<emphasis id="B9680A47A6280637FF40FB6CFEAAE0EE" bold="true" box="[187,328,1221,1247]" pageId="29" pageNumber="30">Figs 37–40.</emphasis>
<taxonomicName id="4C1CADD6A6280637FEA8FB6CFD79E0EE" authority="Itoh, 2001" authorityName="Itoh" authorityYear="2001" box="[339,667,1221,1247]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="B9680A47A6280637FEA8FB6CFDF5E0EE" box="[339,535,1221,1247]" italics="true" pageId="29" pageNumber="30">Miridiba hirsuta</emphasis>
<bibRefCitation id="EF8DABA4A6280637FDDAFB6CFD79E0EE" author="Itoh T." box="[545,667,1221,1247]" pageId="29" pageNumber="30" pagination="435 - 439" refId="ref44461" refString="Itoh T. 2001. A new species of the genus Miridiba (Scarabaeidae: Melolonthinae: Melolonthini) from the Yoyama Islands, Southwest Japan. Elytra, Tokyo 29 (2): 435 - 439." type="journal article" year="2001">Itoh, 2001</bibRefCitation>
</taxonomicName>
(non-types, TMC).
<emphasis id="B9680A47A6280637FC72FB6CFC47E0EE" bold="true" box="[905,933,1221,1247]" pageId="29" pageNumber="30">37</emphasis>
. Habitus in dorsal view.
<emphasis id="B9680A47A6280637FB2BFB6CFB0EE0EE" bold="true" box="[1232,1260,1221,1247]" pageId="29" pageNumber="30">38</emphasis>
. Parameres in dorsal (A) and lateral (B) views.
<emphasis id="B9680A47A6280637FDA1FB41FD95E133" bold="true" box="[602,631,1256,1282]" pageId="29" pageNumber="30">39</emphasis>
. Endophallus and temones in dorsal view.
<emphasis id="B9680A47A6280637FB90FB41FB6AE133" bold="true" box="[1131,1160,1256,1282]" pageId="29" pageNumber="30">40</emphasis>
. Female genitalia in dorsal view. Scale bars: 38–40 = 1.0 mm.
</paragraph>
</caption>
<subSubSection id="C30685DEA6280637FF46FAC8FECEE19B" box="[189,342,1377,1450]" pageId="29" pageNumber="30" type="distribution">
<paragraph id="8BA3D655A6280637FF46FAC8FEB4E14A" blockId="29.[189,342,1377,1403]" box="[189,342,1377,1403]" pageId="29" pageNumber="30">
<heading id="D0EB6139A6280637FF46FAC8FEB4E14A" bold="true" box="[189,342,1377,1403]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="B9680A47A6280637FF46FAC8FEB4E14A" bold="true" box="[189,342,1377,1403]" pageId="29" pageNumber="30">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A6280637FF46FA39FECEE19B" blockId="29.[189,300,1424,1450]" box="[189,300,1424,1450]" pageId="29" pageNumber="30">Ryukyus.</paragraph>
</subSubSection>
</treatment>
</document>
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