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<mods:title id="53DB584F91E1DB957A050CFD512ECE50">Revision of the genus Miridiba Reitter, 1902 (Coleoptera, Scarabaeidae, Melolonthinae): genital morphotypes and new taxonomic data</mods:title>
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<mods:namePart id="92153A555305B1B7C7E56A3F3706E9F7">Gao, Chuan-bu</mods:namePart>
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<mods:affiliation id="903BC77804569FF23FF73C5FB180A595">Guangdong Key Laboratory of Animal Conservation and Resource Utilization, Guangdong Public Laboratory of Wild Animal Conservation and Utilization, Institute of Zoology, Guangdong Academy of Sciences, 105 Xingang Road W., Guangzhou 510260, China. Unidad de Protección Vegetal. Centro de Investigación y Tecnología Agroalimentaria (CITA) e Instituto Agroalimentario de Aragón (IA 2) (CITA-Universidad de Zaragoza), Avenida Montañana, 930. 50059 Zaragoza, Spain. &amp; Buzh.</mods:affiliation>
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<mods:namePart id="6F058DFABE8E20CB9531EC5D920CAE55">Coca-Abia, María Milagro</mods:namePart>
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<mods:nameIdentifier id="A371291F45BE5AA618CA718F43444BBC" type="email">mcoca@aragon.es</mods:nameIdentifier>
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<mods:title id="2205301CD63F44CA3FD132A6FEA713A9">European Journal of Taxonomy</mods:title>
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<mods:date id="EDBE4AB00E9CABBCC29D5E076FA30DD1">2021</mods:date>
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<treatment id="03B56743A611060CFDA3FB6BFBCFE1A7" ID-DOI="http://doi.org/10.5281/zenodo.4773251" ID-GBIF-Taxon="182237246" ID-Zenodo-Dep="4773251" LSID="urn:lsid:plazi:treatment:03B56743A611060CFDA3FB6BFBCFE1A7" httpUri="http://treatment.plazi.org/id/03B56743A611060CFDA3FB6BFBCFE1A7" lastPageId="38" lastPageNumber="39" pageId="36" pageNumber="37">
<subSubSection id="C30685DEA611060EFDA3FB6BFC39E0ED" box="[600,987,1218,1245]" pageId="36" pageNumber="37" type="nomenclature">
<paragraph id="8BA3D655A611060EFDA3FB6BFC39E0ED" blockId="36.[600,987,1218,1281]" box="[600,987,1218,1245]" pageId="36" pageNumber="37">
<heading id="D0EB6139A611060EFDA3FB6BFC39E0ED" box="[600,987,1218,1245]" centered="true" fontSize="11" level="2" pageId="36" pageNumber="37" reason="2">
<taxonomicName id="4C1CADD6A611060EFDA3FB6BFC39E0ED" authority="(Moser, 1908)" baseAuthorityName="Moser" baseAuthorityYear="1908" box="[600,987,1218,1245]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="pilosella">
<emphasis id="B9680A47A611060EFDA3FB6BFCCCE0EC" bold="true" box="[600,814,1218,1245]" italics="true" pageId="36" pageNumber="37">Miridiba pilosella</emphasis>
(
<bibRefCitation id="EF8DABA4A611060EFCC6FB6BFC31E0ED" author="Moser J." box="[829,979,1218,1244]" pageId="36" pageNumber="37" pagination="325 - 343" refId="ref45266" refString="Moser J. 1908. Verzeichnis der von H. Fruhstorfer in Tonkin gesammelten Melolonthiden. Annales de la Societe entomologique de Belgique 52: 325 - 343." type="journal article" year="1908">Moser, 1908</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA611060EFD23FB4FFCB9E130" box="[728,859,1254,1281]" pageId="36" pageNumber="37" type="description">
<paragraph id="8BA3D655A611060EFD23FB4FFCB9E130" blockId="36.[600,987,1218,1281]" box="[728,859,1254,1281]" pageId="36" pageNumber="37">
<figureCitation id="1327CAD0A611060EFD23FB4FFCB9E130" box="[728,859,1254,1281]" captionStart="Figs 51–54" captionStartId="37.[189,241,1883,1909]" captionTargetBox="[284,1310,917,1831]" captionTargetId="figure-247@37.[264,1323,898,1844]" captionTargetPageId="37" captionText="Figs 51–54. Miridiba pilosella (Moser, 1908). 51. Lectotype (MFNB). 52–54. Paralectotypes (MFNB). 51. Habitus in dorsal view. 52–53. Male genitalia. 52. Parameres in dorsal (A) and lateral (B) views. 53. Endophallus and temones in dorsal view. 54. Female genitalia in dorsal view. Scale bars: 51 = 5.0 mm; 52–54 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770362" httpUri="https://zenodo.org/record/4770362/files/figure.png" pageId="36" pageNumber="37">Figs 51–54</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA611060EFF46FA81FD0DE33F" pageId="36" pageNumber="37" type="reference_group">
<paragraph id="8BA3D655A611060EFF46FA81FBB0E172" blockId="36.[189,1106,1320,1383]" box="[189,1106,1320,1349]" pageId="36" pageNumber="37">
<treatmentCitationGroup id="AB0CF17BA611060EFF46FA81FBB0E172" box="[189,1106,1320,1349]" pageId="36" pageNumber="37">
<treatmentCitation id="0ABDF044A611060EFF46FA81FD6AE173" author="Moser J." box="[189,648,1320,1347]" page="336" pageId="36" pageNumber="37" year="1908">
<taxonomicName id="4C1CADD6A611060EFF46FA81FD6AE173" authority="Moser, 1908: 336" authorityName="Moser" authorityPageNumber="336" authorityYear="1908" box="[189,648,1320,1347]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="pilosella">
<emphasis id="B9680A47A611060EFF46FA81FE50E173" box="[189,434,1320,1346]" italics="true" pageId="36" pageNumber="37">Holotrichia pilosella</emphasis>
<bibRefCitation id="EF8DABA4A611060EFE41FA80FD6AE173" author="Moser J." box="[442,648,1320,1347]" pageId="36" pageNumber="37" pagination="325 - 343" refId="ref45266" refString="Moser J. 1908. Verzeichnis der von H. Fruhstorfer in Tonkin gesammelten Melolonthiden. Annales de la Societe entomologique de Belgique 52: 325 - 343." type="journal article" year="1908">Moser, 1908: 336</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<typeStatus id="54A768F7A611060EFD63FA81FD2EE173" box="[664,716,1320,1346]" pageId="36" pageNumber="37">type</typeStatus>
loc.: Mount Mẫu SƠn,
<collectingCountry id="F30B96C5A611060EFC26FA81FBA4E172" box="[989,1094,1320,1347]" name="Vietnam" pageId="36" pageNumber="37">Vietnam</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA3D655A611060EFF46FAE5FBB0E157" blockId="36.[189,1106,1320,1383]" box="[189,1106,1356,1383]" pageId="36" pageNumber="37">
<treatmentCitationGroup id="AB0CF17BA611060EFF46FAE5FBB0E157" box="[189,1106,1356,1383]" pageId="36" pageNumber="37">
<treatmentCitation id="0ABDF044A611060EFF46FAE5FD43E157" author="Moser J." box="[189,673,1356,1382]" page="470" pageId="36" pageNumber="37" year="1909">
<taxonomicName id="4C1CADD6A611060EFF46FAE5FD43E157" authority="Moser, 1909: 470" authorityName="Moser" authorityPageNumber="470" authorityYear="1909" box="[189,673,1356,1382]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B9680A47A611060EFF46FAE5FE28E157" box="[189,458,1356,1382]" italics="true" pageId="36" pageNumber="37">Holotrichia formosana</emphasis>
<bibRefCitation id="EF8DABA4A611060EFE29FAE5FD43E157" author="Moser J." box="[466,673,1356,1382]" pageId="36" pageNumber="37" pagination="468 - 488" refId="ref45295" refString="Moser J. 1909. Neue arten der Melolonthiden-Gattungen Holotrichia und Brahmina. Annales de la Societe entomologique de Belgique 53: 468 - 488." type="journal article" year="1909">Moser, 1909: 470</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<typeStatus id="54A768F7A611060EFD4BFAE5FD06E157" box="[688,740,1356,1382]" pageId="36" pageNumber="37">type</typeStatus>
loc.:
<collectingCountry id="F30B96C5A611060EFCDDFAE5FC9DE157" box="[806,895,1356,1382]" name="Taiwan" pageId="36" pageNumber="37">Taiwan</collectingCountry>
,
<collectingCountry id="F30B96C5A611060EFC71FAE5FC37E156" box="[906,981,1356,1383]" name="China" pageId="36" pageNumber="37">China</collectingCountry>
). syn. nov.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA3D655A611060EFF46FA27FCC5E1FD" blockId="36.[189,1400,1422,1806]" pageId="36" pageNumber="37">
<treatmentCitationGroup id="AB0CF17BA611060EFF46FA27FCC5E1FD" pageId="36" pageNumber="37">
<taxonomicName id="4C1CADD6A611060EFF46FA27FE56E199" authorityName="Moser" authorityYear="1908" box="[189,436,1422,1448]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="pilosella">
<emphasis id="B9680A47A611060EFF46FA27FE56E199" box="[189,436,1422,1448]" italics="true" pageId="36" pageNumber="37">Holotrichia pilosella</emphasis>
</taxonomicName>
–
<treatmentCitation id="0ABDF044A611060EFE23FA27FD03E198" author="Dalla Torre K. W. von" box="[472,737,1422,1449]" page="205" pageId="36" pageNumber="37" year="1912">
<bibRefCitation id="EF8DABA4A611060EFE23FA27FD03E198" author="Dalla Torre K. W. von" box="[472,737,1422,1449]" pageId="36" pageNumber="37" pagination="135 - 290" refId="ref43620" refString="Dalla Torre K. W. von. 1912. Pars 49. Scarabaeidae: Melolonthidae III. In: Junk W. &amp; Schenkling S. (eds) Coleopterorum Catalogus. Vol. XX: 135 - 290. W. Junk, Berlin, Germany." type="book chapter" year="1912">Dalla Torre 1912: 205</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="0ABDF044A611060EFC5EFA27FB94E199" author="Chang Y. - W." box="[933,1142,1422,1448]" page="146" pageId="36" pageNumber="37" year="1964">
<bibRefCitation id="EF8DABA4A611060EFC5EFA27FB94E199" author="Chang Y. - W." box="[933,1142,1422,1448]" pageId="36" pageNumber="37" pagination="139 - 152" refId="ref43461" refString="Chang Y. - W. 1964. Revision of Chinese may beetles of the genus Holotrichia Hope I. (Coleoptera, Scarabaeidae). Acta Zootaxonomica Sinica 1 (1): 139 - 152." type="journal article" year="1964">Chang 1964: 146</bibRefCitation>
</treatmentCitation>
, 149–150, figs 48–49 (species list; in key). —
<treatmentCitation id="0ABDF044A611060EFDF2FA1BFD5EE1FD" author="Frey G." box="[521,700,1458,1484]" page="248" pageId="36" pageNumber="37" year="1970">
<bibRefCitation id="EF8DABA4A611060EFDF2FA1BFD5EE1FD" author="Frey G." box="[521,700,1458,1484]" pageId="36" pageNumber="37" pagination="244 - 265" refId="ref43949" refString="Frey G. 1970. Bestimmungstabelle der Holotrichia - Arten Hinterindiens (Vietnam, Laos, Thailand, Kambodscha, Burma, Malaya) (Col., Scarab, Melolonthinae). Entomologische Arbeiten aus dem Museum G. Frey Tutzing bei Munchen 21: 244 - 265." type="journal article" year="1970">Frey 1970: 248</bibRefCitation>
</treatmentCitation>
(in key).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA3D655A611060EFF46FA7CFD0DE225" blockId="36.[189,1400,1422,1806]" pageId="36" pageNumber="37">
<treatmentCitationGroup id="AB0CF17BA611060EFF46FA7CFD0DE225" pageId="36" pageNumber="37">
<taxonomicName id="4C1CADD6A611060EFF46FA7CFE70E1DE" baseAuthorityName="Moser" baseAuthorityYear="1908" box="[189,402,1493,1519]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="pilosella">
<emphasis id="B9680A47A611060EFF46FA7CFE70E1DE" box="[189,402,1493,1519]" italics="true" pageId="36" pageNumber="37">Miridiba pilosella</emphasis>
</taxonomicName>
–
<treatmentCitation id="0ABDF044A611060EFE4AFA7FFD66E1DE" author="Nomura S." box="[433,644,1493,1520]" page="88" pageId="36" pageNumber="37" year="1977">
<bibRefCitation id="EF8DABA4A611060EFE4AFA7FFD66E1DE" author="Nomura S." box="[433,644,1493,1520]" pageId="36" pageNumber="37" pagination="81 - 95" refId="ref45479" refString="Nomura S. 1977. On the Melolonthini of Taiwan (Coleoptera, Scarabaeidae). Toho-Gakuho 27: 85 - 109. Nonfried A. F. 1892. Verzeichniss der um Nienghali in Sudchina gesammelten Lucanoiden, Scarabaeiden, Buprestiden, und Cerambyciden, nebst Beschreibung neuer arten. Entomologische Nachrichten 18 (6): 81 - 95." type="journal article" year="1977">Nomura 1977: 88</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0ABDF044A611060EFD4FFA7CFC50E1DE" author="Coca-Abia M. M." box="[692,946,1493,1519]" page="681" pageId="36" pageNumber="37" year="2008">
<bibRefCitation id="EF8DABA4A611060EFD4FFA7CFC50E1DE" author="Coca-Abia M. M." box="[692,946,1493,1519]" pageId="36" pageNumber="37" pagination="673 - 685" refId="ref43499" refString="Coca-Abia M. M. 2008. Revision of the genus Miridiba Reitter, 1901 (Coleoptera, Scarabaeidae, Melolonthinae). Zoological Science 25 (6): 673 - 685. https: // doi. org / 10.2108 / zsj. 25.673" type="journal article" year="2008">Coca-Abia 2008: 681</bibRefCitation>
</treatmentCitation>
(combination). —
<treatmentCitation id="0ABDF044A611060EFB6FFA7CFA94E1C1" ID-DOI="http://doi.org/10.5281/zenodo.5971124" ID-Zenodo-Dep="5971124" author="Gao C. - B. &amp; Bai M. &amp; Fang H. &amp; Yu Z. - G." box="[1172,1398,1493,1520]" httpUri="http://treatment.plazi.org/id/03B587C5ED3BF34D28EDF88E5D3CDECF" page="15" pageId="36" pageNumber="37" year="2018">
<bibRefCitation id="EF8DABA4A611060EFB6FFA7CFA94E1C1" author="Gao C. - B. &amp; Bai M. &amp; Fang H. &amp; Yu Z. - G." box="[1172,1398,1493,1520]" pageId="36" pageNumber="37" pagination="1 - 20" refId="ref44032" refString="Gao C. - B., Bai M., Fang H. &amp; Yu Z. - G. 2018. Four new species of the genus Miridiba Reitter (Coleoptera: Scarabaeidae: Melolonthinae) from China. Zootaxa 4527 (1): 1 - 20. https: // doi. org / 10.11646 / zootaxa. 4527.1.1" type="journal article" year="2018">
Gao
<emphasis id="B9680A47A611060EFB37FA7FFB1DE1DE" box="[1228,1279,1493,1519]" italics="true" pageId="36" pageNumber="37">et al</emphasis>
. 2018: 15
</bibRefCitation>
</treatmentCitation>
(catalogue); 2019: 461 (species list; in key).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA3D655A611060EFF46F9B4FE3FE26A" blockId="36.[189,1400,1422,1806]" pageId="36" pageNumber="37">
<treatmentCitationGroup id="AB0CF17BA611060EFF46F9B4FE3FE26A" pageId="36" pageNumber="37">
<taxonomicName id="4C1CADD6A611060EFF46F9B4FE28E206" authorityName="Moser" authorityYear="1909" box="[189,458,1565,1591]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B9680A47A611060EFF46F9B4FE28E206" box="[189,458,1565,1591]" italics="true" pageId="36" pageNumber="37">Holotrichia formosana</emphasis>
</taxonomicName>
–
<treatmentCitation id="0ABDF044A611060EFE12F9B4FD0FE206" author="Dalla Torre K. W. von" box="[489,749,1565,1591]" page="202" pageId="36" pageNumber="37" year="1912">
<bibRefCitation id="EF8DABA4A611060EFE12F9B4FD0FE206" author="Dalla Torre K. W. von" box="[489,749,1565,1591]" pageId="36" pageNumber="37" pagination="135 - 290" refId="ref43620" refString="Dalla Torre K. W. von. 1912. Pars 49. Scarabaeidae: Melolonthidae III. In: Junk W. &amp; Schenkling S. (eds) Coleopterorum Catalogus. Vol. XX: 135 - 290. W. Junk, Berlin, Germany." type="book chapter" year="1912">Dalla Torre 1912: 202</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="0ABDF044A611060EFC57F9B4FB98E206" author="Chang Y. - W." box="[940,1146,1565,1591]" page="146" pageId="36" pageNumber="37" year="1964">
<bibRefCitation id="EF8DABA4A611060EFC57F9B4FB98E206" author="Chang Y. - W." box="[940,1146,1565,1591]" pageId="36" pageNumber="37" pagination="139 - 152" refId="ref43461" refString="Chang Y. - W. 1964. Revision of Chinese may beetles of the genus Holotrichia Hope I. (Coleoptera, Scarabaeidae). Acta Zootaxonomica Sinica 1 (1): 139 - 152." type="journal article" year="1964">Chang 1964: 146</bibRefCitation>
</treatmentCitation>
, 149–150, figs 46–47 (species list; in key).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA3D655A611060EFF46F9CDFC6FE24E" blockId="36.[189,1400,1422,1806]" box="[189,909,1636,1663]" pageId="36" pageNumber="37">
<treatmentCitationGroup id="AB0CF17BA611060EFF46F9CDFC6FE24E" box="[189,909,1636,1663]" pageId="36" pageNumber="37">
<taxonomicName id="4C1CADD6A611060EFF46F9CDFDE2E24F" authorityName="Moser" authorityYear="1913" box="[189,512,1636,1662]" class="Insecta" family="Melolonthidae" genus="Neodontocnema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B9680A47A611060EFF46F9CDFDE2E24F" box="[189,512,1636,1662]" italics="true" pageId="36" pageNumber="37">Neodontocnema formosana</emphasis>
</taxonomicName>
–
<treatmentCitation id="0ABDF044A611060EFDE7F9CDFD35E24E" author="Arrow G. J." box="[540,727,1636,1663]" page="51" pageId="36" pageNumber="37" year="1948">
<bibRefCitation id="EF8DABA4A611060EFDE7F9CDFD35E24E" author="Arrow G. J." box="[540,727,1636,1663]" pageId="36" pageNumber="37" pagination="49 - 54" refId="ref43061" refString="Arrow G. J. 1948. Further notes on the beetle genus Lachnosterna (Col.: Melolonthinae), with descriptions of three new genera. The Proceedings of the Royal Entomological Society of London. Series B, Taxonomy 17: 49 - 54. https: // doi. org / 10.1111 / j. 1365 - 3113.1948. tb 00885. x" type="journal article" year="1948">Arrow 1948: 51</bibRefCitation>
</treatmentCitation>
(combination).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA3D655A611060EFF46F921FD0DE33F" blockId="36.[189,1400,1422,1806]" pageId="36" pageNumber="37">
<treatmentCitationGroup id="AB0CF17BA611060EFF46F921FD0DE33F" pageId="36" pageNumber="37">
<taxonomicName id="4C1CADD6A611060EFF46F921FE48E293" authorityName="Coca-Abia" authorityYear="2008" baseAuthorityName="Moser" baseAuthorityYear="1909" box="[189,426,1672,1698]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B9680A47A611060EFF46F921FE48E293" box="[189,426,1672,1698]" italics="true" pageId="36" pageNumber="37">Miridiba formosana</emphasis>
</taxonomicName>
–
<treatmentCitation id="0ABDF044A611060EFE31F921FD78E293" author="Nomura S." box="[458,666,1672,1698]" page="88" pageId="36" pageNumber="37" year="1977">
<bibRefCitation id="EF8DABA4A611060EFE31F921FD78E293" author="Nomura S." box="[458,666,1672,1698]" pageId="36" pageNumber="37" pagination="81 - 95" refId="ref45479" refString="Nomura S. 1977. On the Melolonthini of Taiwan (Coleoptera, Scarabaeidae). Toho-Gakuho 27: 85 - 109. Nonfried A. F. 1892. Verzeichniss der um Nienghali in Sudchina gesammelten Lucanoiden, Scarabaeiden, Buprestiden, und Cerambyciden, nebst Beschreibung neuer arten. Entomologische Nachrichten 18 (6): 81 - 95." type="journal article" year="1977">Nomura 1977: 88</bibRefCitation>
</treatmentCitation>
(combination). —
<treatmentCitation id="0ABDF044A611060EFC85F921FB5CE293" author="Smetana A. &amp; Kral D." box="[894,1214,1672,1698]" page="222" pageId="36" pageNumber="37" year="2006">
<bibRefCitation id="EF8DABA4A611060EFC85F921FB5CE293" author="Smetana A. &amp; Kral D." box="[894,1214,1672,1698]" pageId="36" pageNumber="37" pagination="207 - 228" refId="ref45912" refString="Smetana A. &amp; Kral D. 2006. Tribe Rhizotrogini Burmeister, 1855. In: Lobl I. &amp; Smetana A. (eds) Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea: 207 - 228. Apollo Books, Stenstrup, Denmark." type="book chapter" year="2006">Smetana &amp; Král 2006: 222</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="0ABDF044A611060EFF10F905FE0CE2F4" author="Coca-Abia M. M." box="[235,494,1707,1734]" page="679" pageId="36" pageNumber="37" year="2008">
<bibRefCitation id="EF8DABA4A611060EFF10F905FE0CE2F4" author="Coca-Abia M. M." box="[235,494,1707,1734]" pageId="36" pageNumber="37" pagination="673 - 685" refId="ref43499" refString="Coca-Abia M. M. 2008. Revision of the genus Miridiba Reitter, 1901 (Coleoptera, Scarabaeidae, Melolonthinae). Zoological Science 25 (6): 673 - 685. https: // doi. org / 10.2108 / zsj. 25.673" type="journal article" year="2008">Coca-Abia 2008: 679</bibRefCitation>
</treatmentCitation>
, 683, figs 14–16 (
<typeStatus id="54A768F7A611060EFD3DF905FD18E2F7" box="[710,762,1708,1734]" pageId="36" pageNumber="37">type</typeStatus>
material; in key). —
<treatmentCitation id="0ABDF044A611060EFBFBF905FB01E2F7" ID-DOI="http://doi.org/10.5281/zenodo.6115519" ID-Zenodo-Dep="6115519" author="Li C. - L. &amp; Yang P. - S. &amp; Wang C. - C." box="[1024,1251,1707,1734]" httpUri="http://treatment.plazi.org/id/03CA87D3FFBEFF8098808E92FAEECEEA" page="523" pageId="36" pageNumber="37" year="2015">
<bibRefCitation id="EF8DABA4A611060EFBFBF905FB01E2F7" author="Li C. - L. &amp; Yang P. - S. &amp; Wang C. - C." box="[1024,1251,1707,1734]" pageId="36" pageNumber="37" pagination="521 - 536" refId="ref44907" refString="Li C. - L., Yang P. - S. &amp; Wang C. - C. 2015. A review of the genus Miridiba Reitter (Coleoptera: Scarabaeidae: Melolonthinae) of Taiwan. Zootaxa 3955 (4): 521 - 536. https: // doi. org / 10.11646 / zootaxa. 3955.4.4" type="journal article" year="2015">
Li
<emphasis id="B9680A47A611060EFBD9F905FBBAE2F7" box="[1058,1112,1708,1734]" italics="true" pageId="36" pageNumber="37">et al</emphasis>
. 2015: 523
</bibRefCitation>
</treatmentCitation>
, 528, figs 4, 10, 16, 22, 31–32 (redescription; in key). —
<treatmentCitation id="0ABDF044A611060EFD07F966FC30E2D8" author="Bezdek A." box="[764,978,1743,1769]" page="271" pageId="36" pageNumber="37" year="2016">
<bibRefCitation id="EF8DABA4A611060EFD07F966FC30E2D8" author="Bezdek A." box="[764,978,1743,1769]" pageId="36" pageNumber="37" pagination="249 - 280" refId="ref43171" refString="Bezdek A. 2016. Tribe Rhizotrogini Burmeister, 1855. In: Lobl I. &amp; Lobl D. (eds) Catalogue of Palaearctic Coleoptera. Vol. 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition: 249 - 280. Koninklijke Brill NV, Leiden, The Netherlands." type="book chapter" year="2016">Bezděk 2016: 271</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="0ABDF044A611060EFB69F966FA94E2D8" ID-DOI="http://doi.org/10.5281/zenodo.5971114" ID-Zenodo-Dep="5971114" author="Gao C. - B. &amp; Bai M. &amp; Fang H. &amp; Yu Z. - G." box="[1170,1398,1743,1770]" httpUri="http://treatment.plazi.org/id/03B587C5ED39F34E28EDFC6A5A8FDAD9" page="13" pageId="36" pageNumber="37" year="2018">
<bibRefCitation id="EF8DABA4A611060EFB69F966FA94E2D8" author="Gao C. - B. &amp; Bai M. &amp; Fang H. &amp; Yu Z. - G." box="[1170,1398,1743,1770]" pageId="36" pageNumber="37" pagination="1 - 20" refId="ref44032" refString="Gao C. - B., Bai M., Fang H. &amp; Yu Z. - G. 2018. Four new species of the genus Miridiba Reitter (Coleoptera: Scarabaeidae: Melolonthinae) from China. Zootaxa 4527 (1): 1 - 20. https: // doi. org / 10.11646 / zootaxa. 4527.1.1" type="journal article" year="2018">
Gao
<emphasis id="B9680A47A611060EFB30F979FB1DE2D8" box="[1227,1279,1743,1769]" italics="true" pageId="36" pageNumber="37">et al</emphasis>
. 2018: 13
</bibRefCitation>
</treatmentCitation>
(catalogue); 2019: 461 (species list; in key).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA611060FFF46F89CFCB8E70D" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37" type="diagnosis">
<paragraph id="8BA3D655A611060EFF46F89CFED4E37E" blockId="36.[189,310,1845,1871]" box="[189,310,1845,1871]" pageId="36" pageNumber="37">
<heading id="D0EB6139A611060EFF46F89CFED4E37E" bold="true" box="[189,310,1845,1871]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B9680A47A611060EFF46F89CFED4E37E" bold="true" box="[189,310,1845,1871]" pageId="36" pageNumber="37">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A611060FFF46F8CDFCB8E70D" blockId="36.[189,1399,1892,1990]" lastBlockId="37.[187,1401,265,828]" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
External morphology of adult (
<figureCitation id="1327CAD0A611060EFDCCF8CDFD70E34E" box="[567,658,1892,1919]" captionStart="Figs 51–54" captionStartId="37.[189,241,1883,1909]" captionTargetBox="[284,1310,917,1831]" captionTargetId="figure-247@37.[264,1323,898,1844]" captionTargetPageId="37" captionText="Figs 51–54. Miridiba pilosella (Moser, 1908). 51. Lectotype (MFNB). 52–54. Paralectotypes (MFNB). 51. Habitus in dorsal view. 52–53. Male genitalia. 52. Parameres in dorsal (A) and lateral (B) views. 53. Endophallus and temones in dorsal view. 54. Female genitalia in dorsal view. Scale bars: 51 = 5.0 mm; 52–54 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770362" httpUri="https://zenodo.org/record/4770362/files/figure.png" pageId="36" pageNumber="37">Fig. 51</figureCitation>
). Body size 17.4–19.0 mm. Dorsal surface with conspicuous short pubescence. Clypeus flat with anterior edge moderately emarginate at middle, oblique laterally. Frons with sharp carina, not interrupted at middle. Pronotal anterior margin with short setae, without concavity at lateral ends; posterior margin glabrous; lateral margins strongly serrated with short setae; anterior angles acute almost right, rounded, not projected forward; posterior angles obtuse and rounded. Prosternal process with a ventral bulge and posterior margin rounded. Scutellum densely punctate. Elytra with punctures regularly distributed, more scattered and larger than those of pronotum; epipleuron with long marginal setae except on apex. Foretibia with sharp dorsal carina, insertion of inner spur closer third outer tooth than second one. Meso- and metatibia with interrupted and complete transverse carina, respectively; inner margin of dorsal surface with strong spines. Pygidium with rough punctures, dense pubescence and irregular in length. Abdominal ventrites covered with dense, short and decumbent setae and punctures regularly distributed. Male genitalia: parameres (
<figureCitation id="1327CAD0A610060FFC65FD8EFC10E673" box="[926,1010,551,578]" captionStart="Figs 51–54" captionStartId="37.[189,241,1883,1909]" captionTargetBox="[284,1310,917,1831]" captionTargetId="figure-247@37.[264,1323,898,1844]" captionTargetPageId="37" captionText="Figs 51–54. Miridiba pilosella (Moser, 1908). 51. Lectotype (MFNB). 52–54. Paralectotypes (MFNB). 51. Habitus in dorsal view. 52–53. Male genitalia. 52. Parameres in dorsal (A) and lateral (B) views. 53. Endophallus and temones in dorsal view. 54. Female genitalia in dorsal view. Scale bars: 51 = 5.0 mm; 52–54 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770362" httpUri="https://zenodo.org/record/4770362/files/figure.png" pageId="37" pageNumber="38">Fig. 52</figureCitation>
) with sparse setae on ventral side of dorsal branches and on dorsal side of ventral ones; in lateral view dorsal branches moderately bent at two stretches towards apices, which are curved downward with a small apical tip; ventral branches with apices curved outward. Endophallus (
<figureCitation id="1327CAD0A610060FFD3AFD3BFCFBE69C" box="[705,793,658,685]" captionStart="Figs 51–54" captionStartId="37.[189,241,1883,1909]" captionTargetBox="[284,1310,917,1831]" captionTargetId="figure-247@37.[264,1323,898,1844]" captionTargetPageId="37" captionText="Figs 51–54. Miridiba pilosella (Moser, 1908). 51. Lectotype (MFNB). 52–54. Paralectotypes (MFNB). 51. Habitus in dorsal view. 52–53. Male genitalia. 52. Parameres in dorsal (A) and lateral (B) views. 53. Endophallus and temones in dorsal view. 54. Female genitalia in dorsal view. Scale bars: 51 = 5.0 mm; 52–54 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770362" httpUri="https://zenodo.org/record/4770362/files/figure.png" pageId="37" pageNumber="38">Fig. 53</figureCitation>
) with spines arranged helicoidally around internal walls towards distal end, with a raspula of dark sensillae at proximal half of sac. Apophysis of temones (
<figureCitation id="1327CAD0A610060FFF3DFD70FEFAE6C5" box="[198,280,729,756]" captionStart="Figs 51–54" captionStartId="37.[189,241,1883,1909]" captionTargetBox="[284,1310,917,1831]" captionTargetId="figure-247@37.[264,1323,898,1844]" captionTargetPageId="37" captionText="Figs 51–54. Miridiba pilosella (Moser, 1908). 51. Lectotype (MFNB). 52–54. Paralectotypes (MFNB). 51. Habitus in dorsal view. 52–53. Male genitalia. 52. Parameres in dorsal (A) and lateral (B) views. 53. Endophallus and temones in dorsal view. 54. Female genitalia in dorsal view. Scale bars: 51 = 5.0 mm; 52–54 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770362" httpUri="https://zenodo.org/record/4770362/files/figure.png" pageId="37" pageNumber="38">Fig. 53</figureCitation>
) with distal ends separated from each other. Female genitalia as describe at morphotype I. Genital chamber with a pair of vestigial sternites reduced to few sensillae, median oviduct very developed with hardened epithelium and dark sclerotized areas (
<figureCitation id="1327CAD0A610060FFD0FFC88FCAEE70D" box="[756,844,801,828]" captionStart="Figs 51–54" captionStartId="37.[189,241,1883,1909]" captionTargetBox="[284,1310,917,1831]" captionTargetId="figure-247@37.[264,1323,898,1844]" captionTargetPageId="37" captionText="Figs 51–54. Miridiba pilosella (Moser, 1908). 51. Lectotype (MFNB). 52–54. Paralectotypes (MFNB). 51. Habitus in dorsal view. 52–53. Male genitalia. 52. Parameres in dorsal (A) and lateral (B) views. 53. Endophallus and temones in dorsal view. 54. Female genitalia in dorsal view. Scale bars: 51 = 5.0 mm; 52–54 = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770362" httpUri="https://zenodo.org/record/4770362/files/figure.png" pageId="37" pageNumber="38">Fig. 54</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF6386DDA610060FFF46F8F2FE12E3D1" ID-DOI="http://doi.org/10.5281/zenodo.4770362" ID-Zenodo-Dep="4770362" httpUri="https://zenodo.org/record/4770362/files/figure.png" pageId="37" pageNumber="38" startId="37.[189,241,1883,1909]" targetBox="[284,1310,917,1831]" targetPageId="37">
<paragraph id="8BA3D655A610060FFF46F8F2FE12E3D1" blockId="37.[189,1399,1883,2016]" pageId="37" pageNumber="38">
<emphasis id="B9680A47A610060FFF46F8F2FEABE344" bold="true" box="[189,329,1883,1909]" pageId="37" pageNumber="38">Figs 51–54.</emphasis>
<taxonomicName id="4C1CADD6A610060FFEABF8F2FD34E347" authority="(Moser, 1908)" baseAuthorityName="Moser" baseAuthorityYear="1908" box="[336,726,1883,1910]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="pilosella">
<emphasis id="B9680A47A610060FFEABF8F2FDC7E344" box="[336,549,1883,1909]" italics="true" pageId="37" pageNumber="38">Miridiba pilosella</emphasis>
(
<bibRefCitation id="EF8DABA4A610060FFDCFF8F5FD2FE347" author="Moser J." box="[564,717,1884,1910]" pageId="37" pageNumber="38" pagination="325 - 343" refId="ref45266" refString="Moser J. 1908. Verzeichnis der von H. Fruhstorfer in Tonkin gesammelten Melolonthiden. Annales de la Societe entomologique de Belgique 52: 325 - 343." type="journal article" year="1908">Moser, 1908</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="B9680A47A610060FFD1AF8F2FD1CE344" bold="true" box="[737,766,1883,1909]" pageId="37" pageNumber="38">51</emphasis>
. Lectotype (MFNB).
<emphasis id="B9680A47A610060FFC05F8F2FBAAE344" bold="true" box="[1022,1096,1883,1909]" pageId="37" pageNumber="38">52–54</emphasis>
. Paralectotypes (MFNB).
<emphasis id="B9680A47A610060FFF46F8D6FF38E3A8" bold="true" box="[189,218,1919,1945]" pageId="37" pageNumber="38">51</emphasis>
. Habitus in dorsal view.
<emphasis id="B9680A47A610060FFDF0F8D6FDB0E3A8" bold="true" box="[523,594,1919,1945]" pageId="37" pageNumber="38">52–53</emphasis>
. Male genitalia.
<emphasis id="B9680A47A610060FFCDAF8D6FCDEE3A8" bold="true" box="[801,828,1919,1945]" pageId="37" pageNumber="38">52</emphasis>
. Parameres in dorsal (A) and lateral (B) views.
<emphasis id="B9680A47A610060FFF44F80BFF38E38D" bold="true" box="[191,218,1954,1980]" pageId="37" pageNumber="38">53</emphasis>
. Endophallus and temones in dorsal view.
<emphasis id="B9680A47A610060FFD16F80BFCE8E38D" bold="true" box="[749,778,1954,1980]" pageId="37" pageNumber="38">54</emphasis>
. Female genitalia in dorsal view. Scale bars: 51 = 5.0 mm; 52–54 = 1.0 mm.
</paragraph>
</caption>
<subSubSection id="C30685DEA613060CFF46FEA1FCD8E6DC" pageId="38" pageNumber="39" type="materials_examined">
<paragraph id="8BA3D655A613060CFF46FEA1FE48E513" blockId="38.[189,426,264,290]" box="[189,426,264,290]" pageId="38" pageNumber="39">
<heading id="D0EB6139A613060CFF46FEA1FE48E513" bold="true" box="[189,426,264,290]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="B9680A47A613060CFF46FEA1FE48E513" bold="true" box="[189,426,264,290]" pageId="38" pageNumber="39">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A613060CFF11FE91FDDAE563" blockId="38.[188,1398,311,445]" box="[234,568,311,338]" pageId="38" pageNumber="39">
<emphasis id="B9680A47A613060CFF11FE91FE84E563" bold="true" box="[234,358,312,338]" pageId="38" pageNumber="39">
<typeStatus id="54A768F7A613060CFF11FE91FE84E563" box="[234,358,312,338]" pageId="38" pageNumber="39" type="lectotype">Lectotype</typeStatus>
</emphasis>
(here designated)
</paragraph>
<paragraph id="8BA3D655A613060CFF46FEF2FB61E58C" blockId="38.[188,1398,311,445]" pageId="38" pageNumber="39">
<materialsCitation id="3B74DC08A613060CFF46FEF2FB61E58C" ID-GBIF-Occurrence="3114121332" collectionCode="MFNB" collectorName="H. Fruhstorfer" country="Vietnam" determinerName="Chuan-bu Gao &amp; Coca-Abia" location="VIETNAM" pageId="38" pageNumber="39" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<collectingCountry id="F30B96C5A613060CFF46FEF2FEA6E544" box="[189,324,347,373]" name="Vietnam" pageId="38" pageNumber="39">VIETNAM</collectingCountry>
•
<specimenCount id="9D1A1DDCA613060CFE9DFEF2FE7BE547" box="[358,409,347,374]" pageId="38" pageNumber="39" type="male">1 ♂</specimenCount>
; “Tonkin, Montes Mauson” [Tonkin, centre of Vietnam; Montes Mauson, Mount Mẫu SƠn]; “April Mai” [Apr.–May]; “2-3000’
<collectorName id="26E9B383A613060CFD25FE29FC6BE5A8" box="[734,905,383,410]" pageId="38" pageNumber="39">H. Fruhstorfer</collectorName>
; 100753”; “
<taxonomicName id="4C1CADD6A613060CFBEEFE29FB3CE5A8" authority="Mos" authorityName="Mos" box="[1045,1246,383,409]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="pilosella">
<emphasis id="B9680A47A613060CFBEEFE29FB7FE5A8" box="[1045,1181,383,409]" italics="true" pageId="38" pageNumber="39">H. pilosella</emphasis>
Mos
</taxonomicName>
”; “Lectotype
<taxonomicName id="4C1CADD6A613060CFF47FE0AFDDDE58C" authority="(Moser, 1908)" baseAuthorityName="Moser" baseAuthorityYear="1908" box="[188,575,419,445]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="pilosella">
<emphasis id="B9680A47A613060CFF47FE0AFE73E58C" box="[188,401,419,445]" italics="true" pageId="38" pageNumber="39">Miridiba pilosella</emphasis>
(
<bibRefCitation id="EF8DABA4A613060CFE5AFE0AFDD5E58C" author="Moser J." box="[417,567,419,445]" pageId="38" pageNumber="39" pagination="325 - 343" refId="ref45266" refString="Moser J. 1908. Verzeichnis der von H. Fruhstorfer in Tonkin gesammelten Melolonthiden. Annales de la Societe entomologique de Belgique 52: 325 - 343." type="journal article" year="1908">Moser, 1908</bibRefCitation>
)
</taxonomicName>
<determinerName id="7E7F3A91A613060CFDBCFE0AFD11E58C" box="[583,755,419,445]" pageId="38" pageNumber="39">Chuan-bu Gao</determinerName>
&amp;
<determinerName id="7E7F3A91A613060CFCE3FE0AFC75E58C" box="[792,919,419,445]" pageId="38" pageNumber="39">Coca-Abia</determinerName>
det. 2018”;
<collectionCode id="ED0D4E90A613060CFBD3FE0AFB9DE58C" box="[1064,1151,419,445]" collectionName="Italy, Udine, Museo Friulano di Storia Naturale" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="38" pageNumber="39">MFNB</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BA3D655A613060CFF11FE4CFD81E631" blockId="38.[189,1398,485,583]" box="[234,611,485,512]" pageId="38" pageNumber="39">
<emphasis id="B9680A47A613060CFF11FE4CFE40E5CE" bold="true" box="[234,418,485,511]" pageId="38" pageNumber="39">
<typeStatus id="54A768F7A613060CFF11FE4CFE40E5CE" box="[234,418,485,511]" pageId="38" pageNumber="39" type="paralectotype">Paralectotypes</typeStatus>
</emphasis>
(here indicated)
</paragraph>
<paragraph id="8BA3D655A613060CFF46FDA0FC45E676" blockId="38.[189,1398,485,583]" pageId="38" pageNumber="39">
<materialsCitation id="3B74DC08A613060CFF46FDA0FC41E676" ID-GBIF-Occurrence="3114121347" collectionCode="MFNB" collectorName="H. Fruhstorfer" country="Vietnam" determinerName="Chuan-bu Gao &amp; Coca-Abia" location="VIETNAM" pageId="38" pageNumber="39" specimenCount="5" specimenCount-female="3" specimenCount-male="2" typeStatus="paralectotype">
<collectingCountry id="F30B96C5A613060CFF46FDA0FEA6E612" box="[189,324,521,547]" name="Vietnam" pageId="38" pageNumber="39">VIETNAM</collectingCountry>
•
<specimenCount id="9D1A1DDCA613060CFE9FFDA0FE4EE612" box="[356,428,521,547]" pageId="38" pageNumber="39" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D1A1DDCA613060CFE46FDA0FDE1E612" box="[445,515,521,547]" pageId="38" pageNumber="39" type="female">3 ♀♀</specimenCount>
; same collection data as for lectotype; “Paralectotype
<taxonomicName id="4C1CADD6A613060CFB65FDA1FE81E677" authority="(Moser, 1908)" baseAuthorityName="Moser" baseAuthorityYear="1908" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="pilosella">
<emphasis id="B9680A47A613060CFB65FDA1FA94E613" box="[1182,1398,520,546]" italics="true" pageId="38" pageNumber="39">Miridiba pilosella</emphasis>
(
<bibRefCitation id="EF8DABA4A613060CFF3EFD85FEB9E677" author="Moser J." box="[197,347,556,582]" pageId="38" pageNumber="39" pagination="325 - 343" refId="ref45266" refString="Moser J. 1908. Verzeichnis der von H. Fruhstorfer in Tonkin gesammelten Melolonthiden. Annales de la Societe entomologique de Belgique 52: 325 - 343." type="journal article" year="1908">Moser, 1908</bibRefCitation>
)
</taxonomicName>
<determinerName id="7E7F3A91A613060CFE90FD85FDF5E677" box="[363,535,556,582]" pageId="38" pageNumber="39">Chuan-bu Gao</determinerName>
&amp;
<determinerName id="7E7F3A91A613060CFDC7FD85FD59E677" box="[572,699,556,582]" pageId="38" pageNumber="39">Coca-Abia</determinerName>
det. 2018”;
<collectionCode id="ED0D4E90A613060CFCB7FD84FC41E676" box="[844,931,557,583]" collectionName="Italy, Udine, Museo Friulano di Storia Naturale" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="38" pageNumber="39">MFNB</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BA3D655A613060CFF11FDC4FE44E6B6" blockId="38.[189,814,621,683]" box="[234,422,621,647]" pageId="38" pageNumber="39">
<emphasis id="B9680A47A613060CFF11FDC4FE44E6B6" bold="true" box="[234,422,621,647]" pageId="38" pageNumber="39">Other material</emphasis>
</paragraph>
<paragraph id="8BA3D655A613060CFF46FD38FCCCE69A" blockId="38.[189,814,621,683]" box="[189,814,657,683]" pageId="38" pageNumber="39">
<materialsCitation id="3B74DC08A613060CFF46FD38FCC8E69A" ID-GBIF-Occurrence="3114121403" box="[189,810,657,683]" collectionCode="MFNB" collectorName="Guangzhou &amp; Lehmann S." country="China" location="CHINA" pageId="38" pageNumber="39" specimenCount="1" specimenCount-female="1">
<collectingCountry id="F30B96C5A613060CFF46FD38FEFAE69A" box="[189,280,657,683]" name="China" pageId="38" pageNumber="39">CHINA</collectingCountry>
•
<specimenCount id="9D1A1DDCA613060CFECAFD38FEBCE69A" box="[305,350,657,683]" pageId="38" pageNumber="39" type="female">1 ♀</specimenCount>
;
<collectorName id="26E9B383A613060CFE96FD38FE1AE69A" box="[365,504,657,683]" pageId="38" pageNumber="39">Guangzhou</collectorName>
;
<collectorName id="26E9B383A613060CFDF8FD38FD73E69A" box="[515,657,657,683]" pageId="38" pageNumber="39">Lehmann S.</collectorName>
leg.;
<collectionCode id="ED0D4E90A613060CFD28FD38FCC8E69A" box="[723,810,657,683]" collectionName="Italy, Udine, Museo Friulano di Storia Naturale" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="38" pageNumber="39">MFNB</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BA3D655A613060CFF46FD7BFCD8E6DC" blockId="38.[189,826,722,749]" box="[189,826,722,749]" pageId="38" pageNumber="39">
VIETNAN •
<materialsCitation id="3B74DC08A613060CFEA3FD7BFCD4E6DC" ID-GBIF-Occurrence="3114121401" box="[344,822,722,749]" collectionCode="MFNB" collectorName="H. Fruhstorfer" country="Vietnam" location="Tonkin" pageId="38" pageNumber="39" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D1A1DDCA613060CFEA3FD7BFE65E6DC" box="[344,391,722,749]" pageId="38" pageNumber="39" type="male">1 ♂</specimenCount>
;
<collectingCountry id="F30B96C5A613060CFE68FD7BFE08E6DD" box="[403,490,722,748]" name="Vietnam" pageId="38" pageNumber="39">Tonkin</collectingCountry>
;
<collectorName id="26E9B383A613060CFE0EFD7AFD7FE6DD" box="[501,669,722,749]" pageId="38" pageNumber="39">H. Fruhstorfer</collectorName>
leg.;
<collectionCode id="ED0D4E90A613060CFD24FD7AFCD4E6DC" box="[735,822,723,749]" collectionName="Italy, Udine, Museo Friulano di Storia Naturale" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="38" pageNumber="39">MFNB</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA613060CFF46FCBDFDD0E132" pageId="38" pageNumber="39" type="discussion">
<paragraph id="8BA3D655A613060CFF46FCBDFECDE71F" blockId="38.[189,303,788,814]" box="[189,303,788,814]" pageId="38" pageNumber="39">
<heading id="D0EB6139A613060CFF46FCBDFECDE71F" bold="true" box="[189,303,788,814]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="B9680A47A613060CFF46FCBDFECDE71F" bold="true" box="[189,303,788,814]" pageId="38" pageNumber="39">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A613060CFF46FCEAFDD0E132" blockId="38.[187,1400,834,1283]" pageId="38" pageNumber="39">
<bibRefCitation id="EF8DABA4A613060CFF46FCEAFE82E76D" author="Moser J." box="[189,352,834,861]" pageId="38" pageNumber="39" pagination="325 - 343" refId="ref45266" refString="Moser J. 1908. Verzeichnis der von H. Fruhstorfer in Tonkin gesammelten Melolonthiden. Annales de la Societe entomologique de Belgique 52: 325 - 343." type="journal article" year="1908">Moser (1908)</bibRefCitation>
did not indicate the number of specimens used in describing
<taxonomicName id="4C1CADD6A613060CFBB1FCEBFAA2E76D" authorityName="Moser" authorityYear="1908" box="[1098,1344,834,860]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="pilosella">
<emphasis id="B9680A47A613060CFBB1FCEBFAA2E76D" box="[1098,1344,834,860]" italics="true" pageId="38" pageNumber="39">Holotrichia pilosella</emphasis>
</taxonomicName>
. We have studied a series of specimens (MFNB), two of them labelled as types by the author and another
<specimenCount id="9D1A1DDCA613060CFAB7FCCCFED5E792" pageId="38" pageNumber="39" type="generic">five specimens</specimenCount>
with the same collection data, which belong to the type series. We designate the
<typeStatus id="54A768F7A613060CFB20FC20FAA5E792" box="[1243,1351,905,931]" pageId="38" pageNumber="39" type="lectotype">lectotype</typeStatus>
and
<specimenCount id="9D1A1DDCA613060CFF47FC05FE7FE7F7" box="[188,413,940,966]" pageId="38" pageNumber="39" type="generic" typeStatus="paralectotypes">five paralectotypes</specimenCount>
. On the other hand,
<taxonomicName id="4C1CADD6A613060CFD75FC05FBC8E7F7" authority="(Moser, 1909)" authorityName="Coca-Abia" authorityYear="2008" baseAuthorityName="Moser" baseAuthorityYear="1909" box="[654,1066,940,966]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B9680A47A613060CFD75FC05FC99E7F7" box="[654,891,940,966]" italics="true" pageId="38" pageNumber="39">Miridiba formosana</emphasis>
(
<bibRefCitation id="EF8DABA4A613060CFC77FC05FBC0E7F7" author="Moser J." box="[908,1058,940,966]" pageId="38" pageNumber="39" pagination="468 - 488" refId="ref45295" refString="Moser J. 1909. Neue arten der Melolonthiden-Gattungen Holotrichia und Brahmina. Annales de la Societe entomologique de Belgique 53: 468 - 488." type="journal article" year="1909">Moser, 1909</bibRefCitation>
)
</taxonomicName>
was transferred to
<taxonomicName id="4C1CADD6A613060CFAF4FC05FA95E7F7" baseAuthorityName="Nomura" baseAuthorityYear="1977" box="[1295,1399,940,966]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A613060CFAF4FC05FA95E7F7" box="[1295,1399,940,966]" italics="true" pageId="38" pageNumber="39">Miridiba</emphasis>
</taxonomicName>
by
<bibRefCitation id="EF8DABA4A613060CFF19FC79FE79E7D8" author="Nomura S." box="[226,411,975,1002]" pageId="38" pageNumber="39" pagination="81 - 95" refId="ref45479" refString="Nomura S. 1977. On the Melolonthini of Taiwan (Coleoptera, Scarabaeidae). Toho-Gakuho 27: 85 - 109. Nonfried A. F. 1892. Verzeichniss der um Nienghali in Sudchina gesammelten Lucanoiden, Scarabaeiden, Buprestiden, und Cerambyciden, nebst Beschreibung neuer arten. Entomologische Nachrichten 18 (6): 81 - 95." type="journal article" year="1977">Nomura (1977)</bibRefCitation>
, and a
<typeStatus id="54A768F7A613060CFE16FC66FDBBE7D8" box="[493,601,975,1001]" pageId="38" pageNumber="39" type="lectotype">lectotype</typeStatus>
and
<specimenCount id="9D1A1DDCA613060CFD69FC79FC8DE7D8" box="[658,879,975,1002]" pageId="38" pageNumber="39" type="generic" typeStatus="paralectotypes">two paralectotypes</specimenCount>
were studied and were designated by Coca- Abia (2008). After studying the types of
<taxonomicName id="4C1CADD6A613060CFD6DFC5AFCC0E03D" authorityName="Moser" authorityYear="1908" box="[662,802,1010,1036]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="pilosella">
<emphasis id="B9680A47A613060CFD6DFC5AFCC0E03D" box="[662,802,1010,1036]" italics="true" pageId="38" pageNumber="39">M. pilosella</emphasis>
</taxonomicName>
, we noted that features of the external morphology and male genitalia of
<taxonomicName id="4C1CADD6A613060CFE47FBBFFC0EE001" authority="(Moser, 1909) (Coca-Abia 2008)" authorityName="Coca-Abia" authorityYear="2008" baseAuthorityName="Moser" baseAuthorityYear="1909" box="[444,1004,1045,1072]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B9680A47A613060CFE47FBBFFD83E001" box="[444,609,1046,1072]" italics="true" pageId="38" pageNumber="39">M. formosana</emphasis>
(
<bibRefCitation id="EF8DABA4A613060CFD8AFBBFFCE5E001" author="Moser J." box="[625,775,1046,1072]" pageId="38" pageNumber="39" pagination="468 - 488" refId="ref45295" refString="Moser J. 1909. Neue arten der Melolonthiden-Gattungen Holotrichia und Brahmina. Annales de la Societe entomologique de Belgique 53: 468 - 488." type="journal article" year="1909">Moser, 1909</bibRefCitation>
) (
<bibRefCitation id="EF8DABA4A613060CFCE4FBBCFC06E001" author="Coca-Abia M. M." box="[799,996,1045,1072]" pageId="38" pageNumber="39" pagination="673 - 685" refId="ref43499" refString="Coca-Abia M. M. 2008. Revision of the genus Miridiba Reitter, 1901 (Coleoptera, Scarabaeidae, Melolonthinae). Zoological Science 25 (6): 673 - 685. https: // doi. org / 10.2108 / zsj. 25.673" type="journal article" year="2008">Coca-Abia 2008</bibRefCitation>
)
</taxonomicName>
match well those of
<taxonomicName id="4C1CADD6A613060CFB18FBBFFA8DE01E" authorityName="Moser" authorityYear="1908" box="[1251,1391,1045,1071]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="pilosella">
<emphasis id="B9680A47A613060CFB18FBBFFA8DE01E" box="[1251,1391,1045,1071]" italics="true" pageId="38" pageNumber="39">M. pilosella</emphasis>
</taxonomicName>
. These characters are body pubescence (short and decumbent), head (clypeus shape, frontal suture and carina), pronotum (pubescent anterior margin, crenulate and pubescent lateral margins), foretibiae (strongly carinate), pygidium (conspicuous punctures and pubescence), male genitalia (parameres shape, internal structures of endophallus and temones shape). Therefore, we here synonymize
<taxonomicName id="4C1CADD6A613060CFB29FB0AFA9AE08D" authorityName="Coca-Abia" authorityYear="2008" baseAuthorityName="Moser" baseAuthorityYear="1909" box="[1234,1400,1186,1212]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="formosana">
<emphasis id="B9680A47A613060CFB29FB0AFA9AE08D" box="[1234,1400,1186,1212]" italics="true" pageId="38" pageNumber="39">M. formosana</emphasis>
</taxonomicName>
with
<taxonomicName id="4C1CADD6A613060CFF03FB6FFE61E0EE" authorityName="Moser" authorityYear="1908" box="[248,387,1221,1247]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="pilosella">
<emphasis id="B9680A47A613060CFF03FB6FFE61E0EE" box="[248,387,1221,1247]" italics="true" pageId="38" pageNumber="39">M. pilosella</emphasis>
</taxonomicName>
and include it in morphotype I “
<taxonomicName id="4C1CADD6A613060CFD05FB6CFC73E0EE" box="[766,913,1221,1247]" class="Insecta" family="Tachinidae" genus="Trichophora" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A613060CFD05FB6CFC73E0EE" box="[766,913,1221,1247]" italics="true" pageId="38" pageNumber="39">Trichophora</emphasis>
</taxonomicName>
”, from its antenna 9-segmented and male and female genital morphology.
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA613060CFF46FA83FBCFE1A7" pageId="38" pageNumber="39" type="distribution">
<paragraph id="8BA3D655A613060CFF46FA83FEB4E175" blockId="38.[189,342,1322,1348]" box="[189,342,1322,1348]" pageId="38" pageNumber="39">
<heading id="D0EB6139A613060CFF46FA83FEB4E175" bold="true" box="[189,342,1322,1348]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="B9680A47A613060CFF46FA83FEB4E175" bold="true" box="[189,342,1322,1348]" pageId="38" pageNumber="39">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A613060CFF46FAF0FBCFE1A7" blockId="38.[189,1398,1369,1431]" pageId="38" pageNumber="39">
<collectingCountry id="F30B96C5A613060CFF46FAF0FEE1E142" box="[189,259,1369,1395]" name="China" pageId="38" pageNumber="39">China</collectingCountry>
(
<collectingRegion id="49D818B7A613060CFEE1FAF0FE8AE142" box="[282,360,1369,1395]" country="China" name="Anhui" pageId="38" pageNumber="39">Anhui</collectingRegion>
, Guangzhou,
<collectingRegion id="49D818B7A613060CFDE2FAF0FD85E142" box="[537,615,1369,1395]" country="China" name="Fujian" pageId="38" pageNumber="39">Fujian</collectingRegion>
,
<collectingRegion id="49D818B7A613060CFD81FAF0FD00E142" box="[634,738,1369,1395]" country="China" name="Guangxi" pageId="38" pageNumber="39">Guangxi</collectingRegion>
,
<collectingRegion id="49D818B7A613060CFD0DFAF0FCBCE142" box="[758,862,1369,1395]" country="China" name="Guizhou" pageId="38" pageNumber="39">Guizhou</collectingRegion>
,
<collectingRegion id="49D818B7A613060CFC8AFAF0FC58E142" box="[881,954,1369,1395]" country="China" name="Hebei" pageId="38" pageNumber="39">Hebei</collectingRegion>
,
<collectingRegion id="49D818B7A613060CFC35FAF0FBFBE142" box="[974,1049,1369,1395]" country="China" name="Hubei" pageId="38" pageNumber="39">Hubei</collectingRegion>
,
<collectingRegion id="49D818B7A613060CFBD7FAF0FB65E142" box="[1068,1159,1369,1395]" country="China" name="Jiangsu" pageId="38" pageNumber="39">Jiangsu</collectingRegion>
,
<collectingRegion id="49D818B7A613060CFB61FAF0FB10E142" box="[1178,1266,1369,1395]" country="China" name="Jiangxi" pageId="38" pageNumber="39">Jiangxi</collectingRegion>
,
<collectingRegion id="49D818B7A613060CFAFEFAF0FA90E142" box="[1285,1394,1369,1395]" country="China" name="Liaoning" pageId="38" pageNumber="39">Liaoning</collectingRegion>
,
<collectingRegion id="49D818B7A613060CFF46FAD5FED7E1A7" box="[189,309,1404,1430]" country="China" name="Shandong" pageId="38" pageNumber="39">Shandong</collectingRegion>
,
<collectingRegion id="49D818B7A613060CFEBAFAD5FE77E1A7" box="[321,405,1404,1430]" country="China" name="Shanxi" pageId="38" pageNumber="39">Shanxi</collectingRegion>
,
<collectingRegion id="49D818B7A613060CFE5AFAD5FDE0E1A7" box="[417,514,1404,1430]" country="China" name="Sichuan" pageId="38" pageNumber="39">Sichuan</collectingRegion>
,
<collectingCountry id="F30B96C5A613060CFDF5FAD5FD85E1A7" box="[526,615,1404,1430]" name="Taiwan" pageId="38" pageNumber="39">Taiwan</collectingCountry>
, Zhejiang) (
<bibRefCitation id="EF8DABA4A613060CFD0FFAD5FC4DE1A7" author="Gao C. - B. &amp; Bai M. &amp; Fang H. &amp; Yu Z. - G." box="[756,943,1404,1430]" pageId="38" pageNumber="39" pagination="1 - 20" refId="ref44032" refString="Gao C. - B., Bai M., Fang H. &amp; Yu Z. - G. 2018. Four new species of the genus Miridiba Reitter (Coleoptera: Scarabaeidae: Melolonthinae) from China. Zootaxa 4527 (1): 1 - 20. https: // doi. org / 10.11646 / zootaxa. 4527.1.1" type="journal article" year="2018">
Gao
<emphasis id="B9680A47A613060CFCD5FAD4FC83E1A7" box="[814,865,1404,1430]" italics="true" pageId="38" pageNumber="39">et al</emphasis>
. 2018
</bibRefCitation>
),
<collectingCountry id="F30B96C5A613060CFC39FAD5FBCBE1A7" box="[962,1065,1404,1430]" name="Vietnam" pageId="38" pageNumber="39">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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