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<document id="84148A449F83E7BB4177F5B04325E133" ID-CLB-Dataset="6634" ID-DOI="10.5852/ejt.2021.749.1355" ID-GBIF-Dataset="83109e54-596a-4be0-ac2c-c4e1702cfb8a" ID-ISSN="2118-9773" ID-Zenodo-Dep="4770293" ID-ZooBank="25FD5744-861D-45E4-B6AD-350716AA29BF" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1621349695117" checkinUser="felipe" docAuthor="Gao, Chuan-bu &amp; Coca-Abia, María Milagro" docDate="2021" docId="03B56743A6070619FDF5F905FE05E014" docLanguage="en" docName="ejt-749_1-94.pdf" docOrigin="European Journal of Taxonomy 749" docSource="http://dx.doi.org/10.5852/ejt.2021.749.1355" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Miridiba leucophthalma" docType="treatment" docVersion="9" lastPageNumber="52" masterDocId="FF8C1F3BA635062AFFFBFFA9FFE2E431" masterDocTitle="Revision of the genus Miridiba Reitter, 1902 (Coleoptera, Scarabaeidae, Melolonthinae): genital morphotypes and new taxonomic data" masterLastPageNumber="94" masterPageNumber="1" pageNumber="51" updateTime="1698921317801" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="2DA089B5374D058324BCF278231DDE56">Revision of the genus Miridiba Reitter, 1902 (Coleoptera, Scarabaeidae, Melolonthinae): genital morphotypes and new taxonomic data</mods:title>
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<mods:roleTerm id="CA4A8484EA91C56A3EF21E1ED17D6F1B">Author</mods:roleTerm>
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<mods:namePart id="765F2426F03E78D9CFBE331256231E04">Gao, Chuan-bu</mods:namePart>
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<mods:affiliation id="E72EA1B14B1B44DCBFFA3D6402F7A0EF">Guangdong Key Laboratory of Animal Conservation and Resource Utilization, Guangdong Public Laboratory of Wild Animal Conservation and Utilization, Institute of Zoology, Guangdong Academy of Sciences, 105 Xingang Road W., Guangzhou 510260, China. Unidad de Protección Vegetal. Centro de Investigación y Tecnología Agroalimentaria (CITA) e Instituto Agroalimentario de Aragón (IA 2) (CITA-Universidad de Zaragoza), Avenida Montañana, 930. 50059 Zaragoza, Spain. &amp; Buzh.</mods:affiliation>
<mods:nameIdentifier id="51EC3002F73512ACD4AAED8372C455C6" type="email">Buzh.Gao@msn.com</mods:nameIdentifier>
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<mods:namePart id="6239CFECB63153587E5C0A480725CCC7">Coca-Abia, María Milagro</mods:namePart>
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<mods:nameIdentifier id="62DD627AE3099934ECD3A938404AAE2F" type="email">mcoca@aragon.es</mods:nameIdentifier>
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<mods:title id="A0ABB5CC833EE803E9DAC00859E38830">European Journal of Taxonomy</mods:title>
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<mods:date id="1B90DA97AD457AA4FCEF9ACB13DDAC4C">2021</mods:date>
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<treatment id="03B56743A6070619FDF5F905FE05E014" ID-DOI="http://doi.org/10.5281/zenodo.4773276" ID-GBIF-Taxon="182237255" ID-Zenodo-Dep="4773276" LSID="urn:lsid:plazi:treatment:03B56743A6070619FDF5F905FE05E014" httpUri="http://treatment.plazi.org/id/03B56743A6070619FDF5F905FE05E014" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51">
<subSubSection id="C30685DEA6070618FDF5F905FBC7E2F6" box="[526,1061,1708,1735]" pageId="50" pageNumber="51" type="nomenclature">
<paragraph id="8BA3D655A6070618FDF5F905FBC7E2F6" blockId="50.[526,1061,1708,1770]" box="[526,1061,1708,1735]" pageId="50" pageNumber="51">
<heading id="D0EB6139A6070618FDF5F905FBC7E2F6" box="[526,1061,1708,1735]" centered="true" fontSize="11" level="2" pageId="50" pageNumber="51" reason="2">
<taxonomicName id="4C1CADD6A6070618FDF5F905FBC7E2F6" authority="(Wiedemann, 1819)" baseAuthorityName="Wiedemann" baseAuthorityYear="1819" box="[526,1061,1708,1735]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="leucophthalma">
<emphasis id="B9680A47A6070618FDF5F905FCD7E2F6" bold="true" box="[526,821,1708,1735]" italics="true" pageId="50" pageNumber="51">Miridiba leucophthalma</emphasis>
(
<bibRefCitation id="EF8DABA4A6070618FCBFF904FBFFE2F6" author="Wiedemann C. R. W." box="[836,1053,1709,1735]" pageId="50" pageNumber="51" pagination="157 - 183" refId="ref46073" refString="Wiedemann C. R. W. 1819. VI. Neue Kafer aus Bengalen und Java; befchrieben vom Herausgeber. Zoologisches Magazin, Kiel 1 (3): 157 - 183." type="journal article" year="1819">Wiedemann, 1819</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA6070618FD0CF979FCDEE2DB" box="[759,828,1744,1770]" pageId="50" pageNumber="51" type="description">
<paragraph id="8BA3D655A6070618FD0CF979FCDEE2DB" blockId="50.[526,1061,1708,1770]" box="[759,828,1744,1770]" pageId="50" pageNumber="51">
<figureCitation id="1327CAD0A6070618FD0CF979FCDEE2DB" box="[759,828,1744,1770]" captionStart="Fig" captionStartId="10.[189,232,720,746]" captionTargetBox="[259,1335,326,633]" captionTargetId="figure-17@10.[238,1349,305,646]" captionTargetPageId="10" captionText="Fig. 8. Morphotype III “Leucophthalma”. Male genitalia of M. leucophthalma (Wiedemann, 1819) (non-type, MFNB). Parameres in dorsal (A) and lateral (B) views. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770305" httpUri="https://zenodo.org/record/4770305/files/figure.png" pageId="50" pageNumber="51">Fig. 8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA6070618FF46F8B8FAA2E385" pageId="50" pageNumber="51" type="reference_group">
<paragraph id="8BA3D655A6070618FF46F8B8FBBCE31D" blockId="50.[189,1118,1809,1836]" box="[189,1118,1809,1836]" pageId="50" pageNumber="51">
<treatmentCitationGroup id="AB0CF17BA6070618FF46F8B8FBBCE31D" box="[189,1118,1809,1836]" pageId="50" pageNumber="51">
<treatmentCitation id="0ABDF044A6070618FF46F8B8FCF0E31D" author="Wiedemann C. R. W." box="[189,786,1809,1836]" page="170" pageId="50" pageNumber="51" year="1819">
<taxonomicName id="4C1CADD6A6070618FF46F8B8FCF0E31D" authority="Wiedemann, 1819: 170" authorityName="Wiedemann" authorityPageNumber="170" authorityYear="1819" box="[189,786,1809,1836]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="leucophthalma">
<emphasis id="B9680A47A6070618FF46F8B8FE18E31A" box="[189,506,1809,1835]" italics="true" pageId="50" pageNumber="51">Melolontha leucophthalma</emphasis>
<bibRefCitation id="EF8DABA4A6070618FDFAF8B8FCF0E31D" author="Wiedemann C. R. W." box="[513,786,1809,1836]" pageId="50" pageNumber="51" pagination="157 - 183" refId="ref46073" refString="Wiedemann C. R. W. 1819. VI. Neue Kafer aus Bengalen und Java; befchrieben vom Herausgeber. Zoologisches Magazin, Kiel 1 (3): 157 - 183." type="journal article" year="1819">Wiedemann, 1819: 170</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<typeStatus id="54A768F7A6070618FCDAF8BBFCB7E31D" box="[801,853,1810,1836]" pageId="50" pageNumber="51">type</typeStatus>
loc.:
<collectingRegion id="49D818B7A6070618FC63F8BBFC2DE31D" box="[920,975,1810,1836]" country="Indonesia" name="Jawa Timur" pageId="50" pageNumber="51">Java</collectingRegion>
,
<collectingCountry id="F30B96C5A6070618FC21F8B8FBB0E31D" box="[986,1106,1809,1836]" name="Indonesia" pageId="50" pageNumber="51">Indonesia</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA3D655A6070618FF46F8FAFC3FE35F" blockId="50.[189,1344,1875,1972]" box="[189,989,1875,1902]" pageId="50" pageNumber="51">
<treatmentCitationGroup id="AB0CF17BA6070618FF46F8FAFC3FE35F" box="[189,989,1875,1902]" pageId="50" pageNumber="51">
<taxonomicName id="4C1CADD6A6070618FF46F8FAFDE8E35C" box="[189,522,1875,1901]" class="Insecta" family="Scarabaeidae" genus="Ancylonycha" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="leucophthalma">
<emphasis id="B9680A47A6070618FF46F8FAFDE8E35C" box="[189,522,1875,1901]" italics="true" pageId="50" pageNumber="51">Ancylonycha leucophthalma</emphasis>
</taxonomicName>
–
<treatmentCitation id="0ABDF044A6070618FDDCF8FAFCC5E35C" author="Burmeister H. C. C." box="[551,807,1875,1902]" page="316" pageId="50" pageNumber="51" year="1855">
<bibRefCitation id="EF8DABA4A6070618FDDCF8FAFCC5E35C" author="Burmeister H. C. C." box="[551,807,1875,1902]" pageId="50" pageNumber="51" refId="ref43411" refString="Burmeister H. C. C. 1855. Handbuch der Entomologie. Vierter Band. Besondere Entomologie. Fortsetzung. Zweite Abtheilung. Coleoptera Lamellicornia Phyllophaga chaenochela. Theodor Christian Friedrich Enslin, Berlin. https: // doi. org / 10.5962 / bhl. title. 8135" type="book" year="1855">Burmeister 1855: 316</bibRefCitation>
</treatmentCitation>
(combination).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA3D655A6070618FF46F8DFFC4DE3A0" blockId="50.[189,1344,1875,1972]" box="[189,943,1910,1937]" pageId="50" pageNumber="51">
<treatmentCitationGroup id="AB0CF17BA6070618FF46F8DFFC4DE3A0" box="[189,943,1910,1937]" pageId="50" pageNumber="51">
<taxonomicName id="4C1CADD6A6070618FF46F8DFFE19E3A1" box="[189,507,1910,1936]" class="Insecta" family="Melolonthidae" genus="Holotrichia" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="leucophthalma">
<emphasis id="B9680A47A6070618FF46F8DFFE19E3A1" box="[189,507,1910,1936]" italics="true" pageId="50" pageNumber="51">Holotrichia leucophthalma</emphasis>
</taxonomicName>
–
<treatmentCitation id="0ABDF044A6070618FDE2F8DFFCFEE3A1" author="Dalla Torre K. W. von" box="[537,796,1910,1937]" page="204" pageId="50" pageNumber="51" year="1912">
<bibRefCitation id="EF8DABA4A6070618FDE2F8DFFCFEE3A1" author="Dalla Torre K. W. von" box="[537,796,1910,1937]" pageId="50" pageNumber="51" pagination="135 - 290" refId="ref43620" refString="Dalla Torre K. W. von. 1912. Pars 49. Scarabaeidae: Melolonthidae III. In: Junk W. &amp; Schenkling S. (eds) Coleopterorum Catalogus. Vol. XX: 135 - 290. W. Junk, Berlin, Germany." type="book chapter" year="1912">Dalla Torre 1912: 204</bibRefCitation>
</treatmentCitation>
(catalogue).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA3D655A6070618FF46F830FAA2E385" blockId="50.[189,1344,1875,1972]" box="[189,1344,1945,1972]" pageId="50" pageNumber="51">
<treatmentCitationGroup id="AB0CF17BA6070618FF46F830FAA2E385" box="[189,1344,1945,1972]" pageId="50" pageNumber="51">
<taxonomicName id="4C1CADD6A6070618FF46F830FE39E382" baseAuthorityName="Wiedemann" baseAuthorityYear="1819" box="[189,475,1945,1971]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="leucophthalma">
<emphasis id="B9680A47A6070618FF46F830FE39E382" box="[189,475,1945,1971]" italics="true" pageId="50" pageNumber="51">Miridiba leucophthalma</emphasis>
</taxonomicName>
–
<treatmentCitation id="0ABDF044A6070618FE03F830FD1AE382" author="Coca-Abia M. M." box="[504,760,1945,1971]" page="681" pageId="50" pageNumber="51" year="2008">
<bibRefCitation id="EF8DABA4A6070618FE03F830FD1AE382" author="Coca-Abia M. M." box="[504,760,1945,1971]" pageId="50" pageNumber="51" pagination="673 - 685" refId="ref43499" refString="Coca-Abia M. M. 2008. Revision of the genus Miridiba Reitter, 1901 (Coleoptera, Scarabaeidae, Melolonthinae). Zoological Science 25 (6): 673 - 685. https: // doi. org / 10.2108 / zsj. 25.673" type="journal article" year="2008">Coca-Abia 2008: 681</bibRefCitation>
</treatmentCitation>
, 683, fig. 17 (combination;
<typeStatus id="54A768F7A6070618FBC7F833FB8DE385" box="[1084,1135,1946,1972]" pageId="50" pageNumber="51">type</typeStatus>
material; in key).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA6060619FF46FEA1FB9EE6BF" pageId="51" pageNumber="52" type="diagnosis">
<paragraph id="8BA3D655A6060619FF46FEA1FED4E513" blockId="51.[189,310,264,290]" box="[189,310,264,290]" pageId="51" pageNumber="52">
<heading id="D0EB6139A6060619FF46FEA1FED4E513" bold="true" box="[189,310,264,290]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
<emphasis id="B9680A47A6060619FF46FEA1FED4E513" bold="true" box="[189,310,264,290]" pageId="51" pageNumber="52">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A6060619FF46FE9EFB9EE6BF" blockId="51.[189,1400,311,654]" pageId="51" pageNumber="52">
External morphology of adult (
<bibRefCitation id="EF8DABA4A6060619FDD0FE9EFD16E560" author="Coca-Abia M. M." box="[555,756,311,337]" pageId="51" pageNumber="52" pagination="673 - 685" refId="ref43499" refString="Coca-Abia M. M. 2008. Revision of the genus Miridiba Reitter, 1901 (Coleoptera, Scarabaeidae, Melolonthinae). Zoological Science 25 (6): 673 - 685. https: // doi. org / 10.2108 / zsj. 25.673" type="journal article" year="2008">Coca-Abia 2008</bibRefCitation>
). Dorsal surface glabrous. Clypeus with oblique sides, anterior edge rather emarginate at middle. Frons with strong carina. Pronotal surface densely punctate, glabrous; anterior margin strongly flanged; posterior margin finely flanged except at middle; lateral margins smooth; anterior angles sharp and elevated, not projected forward; posterior angles obtuse and rounded. Epipleuron sparsely pubescent. Meso- and metatibia with transverse carina interrupted at middle, inner margin dorsal surface with very few spines moderately developed. Pygidium with pubescence hardly visible, conspicuous pubescence on apex. Tegmen (
<bibRefCitation id="EF8DABA4A6060619FBC9FDA3FAE2E615" author="Coca-Abia M. M." box="[1074,1280,522,548]" pageId="51" pageNumber="52" pagination="673 - 685" refId="ref43499" refString="Coca-Abia M. M. 2008. Revision of the genus Miridiba Reitter, 1901 (Coleoptera, Scarabaeidae, Melolonthinae). Zoological Science 25 (6): 673 - 685. https: // doi. org / 10.2108 / zsj. 25.673" type="journal article" year="2008">Coca-Abia 2008</bibRefCitation>
: fig. 17): Parameres (
<figureCitation id="1327CAD0A6060619FEB3FD84FE6DE679" box="[328,399,557,584]" captionStart="Fig" captionStartId="10.[189,232,720,746]" captionTargetBox="[259,1335,326,633]" captionTargetId="figure-17@10.[238,1349,305,646]" captionTargetPageId="10" captionText="Fig. 8. Morphotype III “Leucophthalma”. Male genitalia of M. leucophthalma (Wiedemann, 1819) (non-type, MFNB). Parameres in dorsal (A) and lateral (B) views. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770305" httpUri="https://zenodo.org/record/4770305/files/figure.png" pageId="51" pageNumber="52">Fig. 8</figureCitation>
) with dorsal branches shorter than ventral ones, wrinkled at distal end. Endophallus with strong spines, larger towards distal end, placed helicoidally surrounding internal walls of sac. Temones reduced, apophysis separated at distal end. Female genitalia not studied.
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA6060619FF46FD1CFCCCE6CF" pageId="51" pageNumber="52" type="materials_examined">
<paragraph id="8BA3D655A6060619FF46FD1CFE48E6FE" blockId="51.[189,426,693,719]" box="[189,426,693,719]" pageId="51" pageNumber="52">
<heading id="D0EB6139A6060619FF46FD1CFE48E6FE" bold="true" box="[189,426,693,719]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
<emphasis id="B9680A47A6060619FF46FD1CFE48E6FE" bold="true" box="[189,426,693,719]" pageId="51" pageNumber="52">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A6060619FF46FD4DFCCCE6CF" blockId="51.[189,814,738,766]" box="[189,814,738,766]" pageId="51" pageNumber="52">
<materialsCitation id="3B74DC08A6060619FF46FD4DFCCBE6CF" ID-GBIF-Occurrence="3114121318" box="[189,809,738,766]" collectionCode="MFNB" country="Indonesia" location="Indonesia" pageId="51" pageNumber="52" specimenCount="1" specimenCount-male="1" stateProvince="Java">
<collectingCountry id="F30B96C5A6060619FF46FD4DFEB9E6CF" box="[189,347,740,766]" name="Indonesia" pageId="51" pageNumber="52">INDONESIA</collectingCountry>
•
<specimenCount id="9D1A1DDCA6060619FE8FFD4DFE42E6CF" box="[372,416,738,766]" pageId="51" pageNumber="52" type="male">
1
<emphasis id="B9680A47A6060619FE71FD4BFE42E6CF" box="[394,416,738,766]" italics="true" pageId="51" pageNumber="52">♂</emphasis>
</specimenCount>
;
<location id="8EC3808EA6060619FE54FD4DFDC6E6CF" LSID="urn:lsid:plazi:treatment:03B56743A6070619FDF5F905FE05E014:8EC3808EA6060619FE54FD4DFDC6E6CF" box="[431,548,740,766]" country="Indonesia" name="Indonesia" pageId="51" pageNumber="52" stateProvince="Java">Indonesia</location>
,
<collectingRegion id="49D818B7A6060619FDCBFD4DFD8AE6CF" box="[560,616,740,766]" country="Indonesia" name="Jawa Timur" pageId="51" pageNumber="52">Java</collectingRegion>
;
<collectionCode id="ED0D4E90A6060619FD8FFD4DFD25E6CF" box="[628,711,740,766]" collectionName="Italy, Udine, Museo Friulano di Storia Naturale" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="51" pageNumber="52">MFNB</collectionCode>
(
<figureCitation id="1327CAD0A6060619FD2CFD4DFCFCE6CF" box="[727,798,740,766]" captionStart="Fig" captionStartId="10.[189,232,720,746]" captionTargetBox="[259,1335,326,633]" captionTargetId="figure-17@10.[238,1349,305,646]" captionTargetPageId="10" captionText="Fig. 8. Morphotype III “Leucophthalma”. Male genitalia of M. leucophthalma (Wiedemann, 1819) (non-type, MFNB). Parameres in dorsal (A) and lateral (B) views. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4770305" httpUri="https://zenodo.org/record/4770305/files/figure.png" pageId="51" pageNumber="52">Fig. 8</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA6060619FF46FC8CFC8AE785" pageId="51" pageNumber="52" type="discussion">
<paragraph id="8BA3D655A6060619FF46FC8CFECDE70E" blockId="51.[189,303,805,831]" box="[189,303,805,831]" pageId="51" pageNumber="52">
<heading id="D0EB6139A6060619FF46FC8CFECDE70E" bold="true" box="[189,303,805,831]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
<emphasis id="B9680A47A6060619FF46FC8CFECDE70E" bold="true" box="[189,303,805,831]" pageId="51" pageNumber="52">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A6060619FF46FCFDFC8AE785" blockId="51.[189,1398,852,949]" pageId="51" pageNumber="52">
<bibRefCitation id="EF8DABA4A6060619FF46FCFDFE76E75F" author="Coca-Abia M. M." box="[189,404,852,878]" pageId="51" pageNumber="52" pagination="673 - 685" refId="ref43499" refString="Coca-Abia M. M. 2008. Revision of the genus Miridiba Reitter, 1901 (Coleoptera, Scarabaeidae, Melolonthinae). Zoological Science 25 (6): 673 - 685. https: // doi. org / 10.2108 / zsj. 25.673" type="journal article" year="2008">Coca-Abia (2008)</bibRefCitation>
studied this species and transferred it to
<taxonomicName id="4C1CADD6A6060619FC72FCFDFC13E75F" baseAuthorityName="Nomura" baseAuthorityYear="1977" box="[905,1009,852,878]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B9680A47A6060619FC72FCFDFC13E75F" box="[905,1009,852,878]" italics="true" pageId="51" pageNumber="52">Miridiba</emphasis>
</taxonomicName>
.
<taxonomicName id="4C1CADD6A6060619FBF9FCFDFAC1E75F" baseAuthorityName="Wiedemann" baseAuthorityYear="1819" box="[1026,1315,852,878]" class="Insecta" family="Melolonthidae" genus="Miridiba" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="leucophthalma">
<emphasis id="B9680A47A6060619FBF9FCFDFAC1E75F" box="[1026,1315,852,878]" italics="true" pageId="51" pageNumber="52">Miridiba leucophthalma</emphasis>
</taxonomicName>
shows features of external morphology and male genitalia that characterize morphotype III “
<emphasis id="B9680A47A6060619FB57FCDEFA81E7A0" box="[1196,1379,887,913]" italics="true" pageId="51" pageNumber="52">Leucophthalma</emphasis>
”. Hence, this species is included in this genital morphotype.
</paragraph>
</subSubSection>
<subSubSection id="C30685DEA6060619FF46FC72FE05E014" box="[189,487,987,1061]" pageId="51" pageNumber="52" type="distribution">
<paragraph id="8BA3D655A6060619FF46FC72FEB4E7C4" blockId="51.[189,342,987,1013]" box="[189,342,987,1013]" pageId="51" pageNumber="52">
<heading id="D0EB6139A6060619FF46FC72FEB4E7C4" bold="true" box="[189,342,987,1013]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
<emphasis id="B9680A47A6060619FF46FC72FEB4E7C4" bold="true" box="[189,342,987,1013]" pageId="51" pageNumber="52">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA3D655A6060619FF46FBA3FE05E014" blockId="51.[189,487,1034,1061]" box="[189,487,1034,1061]" pageId="51" pageNumber="52">
<collectingCountry id="F30B96C5A6060619FF46FBA3FECDE015" box="[189,303,1034,1060]" name="Indonesia" pageId="51" pageNumber="52">Indonesia</collectingCountry>
(
<collectingRegion id="49D818B7A6060619FEC5FBA3FE74E015" box="[318,406,1034,1060]" country="Indonesia" name="Jakarta Raya" pageId="51" pageNumber="52">Jakarta</collectingRegion>
,
<collectingRegion id="49D818B7A6060619FE59FBA2FE39E014" box="[418,475,1035,1061]" country="Indonesia" name="Jawa Timur" pageId="51" pageNumber="52">Java</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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