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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.529.6139" ID-GBIF-Dataset="db8513fd-e2ce-4f63-a857-3530566adade" ID-PMC="PMC4668899" ID-Pensoft-Pub="1313-2970-529-1" ID-PubMed="26692792" ID-ZBK="AD4323B4913C447A88A7CE05EC8862A3" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-529-1" ModsDocOrigin="ZooKeys 529" ModsDocTitle="Review of the genus Endothyrella Zilch, 1960 with description of five new species (Gastropoda, Pulmonata, Plectopylidae)" checkinTime="1451243868489" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Budha, Prem B., Naggs, Fred, Backeljau, Thierry &amp; Asami, Takahiro" docDate="2015" docId="FE7941F5FE83936454160B6E228A3847" docLanguage="en" docName="ZooKeys 529: 1-70" docOrigin="ZooKeys 529" docSource="http://dx.doi.org/10.3897/zookeys.529.6139" docTitle="Endothyrella dolakhaensis Budha &amp; Pall-Gergely, sp. n." docType="treatment" docUuid="B1043A93-8B29-4E3E-A291-5AEED2E3B66F" docUuidSource="ZooBank" docVersion="4" lastPageNumber="27" masterDocId="E206FFD8FFF5FFFCFFF16833B258FB6B" masterDocTitle="Review of the genus Endothyrella Zilch, 1960 with description of five new species (Gastropoda, Pulmonata, Plectopylidae)" masterLastPageNumber="70" masterPageNumber="1" pageNumber="27" updateTime="1668160861707" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Review of the genus Endothyrella Zilch, 1960 with description of five new species (Gastropoda, Pulmonata, Plectopylidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pall-Gergely, Barna</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Budha, Prem B.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Naggs, Fred</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Backeljau, Thierry</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Asami, Takahiro</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="volume">
<mods:number>529</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>70</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.529.6139</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.529.6139</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-529-1</mods:identifier>
<mods:identifier type="ZBK">AD4323B4913C447A88A7CE05EC8862A3</mods:identifier>
<mods:identifier type="ZooBank">AD4323B4913C447A88A7CE05EC8862A3</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152062802" LSID="urn:lsid:zoobank.org:act:B1043A93-8B29-4E3E-A291-5AEED2E3B66F" httpUri="http://treatment.plazi.org/id/FE7941F5FE83936454160B6E228A3847" lastPageNumber="27" pageId="26" pageNumber="27">
<subSubSection pageId="26" pageNumber="27" type="multiple">
<paragraph pageId="26" pageNumber="27">Taxon classification Animalia Pulmonata Plectopylidae</paragraph>
</subSubSection>
<subSubSection pageId="26" pageNumber="27" type="nomenclature">
<paragraph pageId="26" pageNumber="27">
<taxonomicName LSID="http://zoobank.org/B1043A93-8B29-4E3E-A291-5AEED2E3B66F" authority="Budha &amp; Pall-Gergely" class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella dolakhaensis" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="dolakhaensis">
Endothyrella dolakhaensis Budha &amp;
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
</taxonomicName>
<taxonomicNameLabel pageId="26" pageNumber="27">sp. n.</taxonomicNameLabel>
Figures 9
<normalizedToken originalValue="GH">G-H</normalizedToken>
, 14A
</paragraph>
</subSubSection>
<subSubSection pageId="26" pageNumber="27" type="type material">
<paragraph pageId="26" pageNumber="27">Type material.</paragraph>
<paragraph pageId="26" pageNumber="27">
Nepal, Suridobhan, Dolakha, 1023 m,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="27.758852">27.758852°N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="86.19789">86.197894°E</geoCoordinate>
, leg. Budha, P., 03.02.2009., holotype (CDZMTU001, Figure 14A), CDZMTU002 (2 paratypes = shells from the same locality); Nepal, Bhorle, Dolakha, 800 m,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="27.696651">27.696652°N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="86.129585">86.129583°E</geoCoordinate>
, leg. Budha, P., 03.02.2009., 11 paratypes = shells (CDZMTU003).
</paragraph>
</subSubSection>
<subSubSection pageId="26" pageNumber="27" type="diagnosis">
<paragraph pageId="26" pageNumber="27">Diagnosis.</paragraph>
<paragraph pageId="26" pageNumber="27">Shell small with rather conical dorsal surface; body whorl slightly angulated with five rows of hairs; parietal lamella simple with one or two denticles posteriorly and a plica below; middle palatal plicae divided or almost divided.</paragraph>
</subSubSection>
<subSubSection pageId="26" pageNumber="27" type="description">
<paragraph pageId="26" pageNumber="27">Description.</paragraph>
<paragraph pageId="26" pageNumber="27">Shell very small, sinistral, with somewhat elevated spire and rather conical apex; protoconch elevated from the dorsal surface; colour brownish or greyish; protoconch conspicuously large, consists of 2.25-2.5 whorls (n = 2), very finely, regularly ribbed; teleoconch with clearly visible reticulated sculpture dominated by radial growth lines; sculpture somewhat weaker on the ventral surface; very slender, long periostracal folds (hairs) standing in five spiral lines along the body whorl; whorls 5.25-5.5 (n = 3) moderately bulging, separated by relatively deep suture; umbilicus wide and deep; apertural lip whitish, thin, slightly reflexed; callus also very weak, slightly S-shaped; no fold in the aperture.</paragraph>
<paragraph pageId="26" pageNumber="27">
One specimen from the type locality was opened. Parietal wall with one rather straight lamella with slight lower arms pointing in both directions; small denticle near the upper end posteriorly, connected to the lamella; two short horizontal plicae under the lamella; palatal wall with six plicae; first slim and short, the second-fifth plicae are divided in the middle and are of the same length; last plica also short, rather straight (Figures 9
<normalizedToken originalValue="GH">G-H</normalizedToken>
).
</paragraph>
</subSubSection>
<subSubSection pageId="26" pageNumber="27" type="measurements">
<paragraph pageId="26" pageNumber="27">Measurements</paragraph>
<paragraph pageId="26" pageNumber="27">(in mm): D: 6.5-9.0, H: 4.0-5.0., Wh: 5.5-6.0 (n = 5).</paragraph>
</subSubSection>
<subSubSection pageId="26" pageNumber="27" type="differential diagnosis">
<paragraph pageId="26" pageNumber="27">Differential diagnosis.</paragraph>
<paragraph pageId="26" pageNumber="27">
The most similar species are
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella affinis" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="affinis">Endothyrella affinis</taxonomicName>
and
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella plectostoma" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="plectostoma">Endothyrella plectostoma</taxonomicName>
, which are larger, have a higher spire, and a deeper, narrower umbilicus.
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella dolakhaensis" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="dolakhaensis">Endothyrella dolakhaensis</taxonomicName>
sp. n. has a more elevated spire and more rounded body whorl than
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella angulata" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="angulata">Endothyrella angulata</taxonomicName>
sp. n. Moreover,
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella dolakhaensis" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="dolakhaensis">Endothyrella dolakhaensis</taxonomicName>
sp. n. has five rows of periostracal folds, whereas
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella angulata" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="angulata">Endothyrella angulata</taxonomicName>
sp. n. has four. See also under
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella macromphalus" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="macromphalus">Endothyrella macromphalus</taxonomicName>
,
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella minor" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="minor">Endothyrella minor</taxonomicName>
and
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella nepalica" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="nepalica">Endothyrella nepalica</taxonomicName>
sp. n. and Table 5.
</paragraph>
</subSubSection>
<subSubSection pageId="26" pageNumber="27" type="etymology">
<paragraph pageId="26" pageNumber="27">Etymology.</paragraph>
<paragraph pageId="26" pageNumber="27">The new species is named after the district name (Dolakha).</paragraph>
</subSubSection>
<subSubSection pageId="26" pageNumber="27" type="type locality">
<paragraph pageId="26" pageNumber="27">Type locality.</paragraph>
<paragraph pageId="26" pageNumber="27">
Nepal, Suridobhan, Dolakha, 1023 m,
<geoCoordinate direction="north" orientation="latitude" precision="1" value="27.758852">27.758852°N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="1" value="86.19789">86.197894°E</geoCoordinate>
.
</paragraph>
</subSubSection>
<subSubSection pageId="26" pageNumber="27" type="distribution">
<paragraph pageId="26" pageNumber="27">Distribution.</paragraph>
<paragraph pageId="26" pageNumber="27">
<taxonomicName class="Gastropoda" family="Plectopylidae" genus="Endothyrella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Endothyrella dolakhaensis" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="dolakhaensis">Endothyrella dolakhaensis</taxonomicName>
sp. n. is known from two localities in the valley of the Tamakoshi River, Dolakha district, Central Nepal (Figure 15).
</paragraph>
</subSubSection>
</treatment>
</document>