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<document id="9AF54696D5E4EF99DCBA7CD35776BE43" ID-CLB-Dataset="26049" ID-DOI="10.11646/zootaxa.4654.1.1" ID-GBIF-Dataset="9ce1d103-9234-4fbe-9544-f1c294bb8783" ID-ISSN="1175-5326" ID-Zenodo-Dep="3365803" ID-ZooBank="CFD82CC0-3110-472E-972B-7ADC0C523A04" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1565592061838" checkinUser="plazi" docAuthor="Hansen, Thomas" docDate="2019" docId="BF18F633A9A5FF402B9CC005FBADFA79" docLanguage="en" docName="zootaxa.4654.1.1.pdf" docOrigin="Zootaxa 4654 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Cylichna stevniensis Hansen 2019, n. sp." docType="treatment" docUuid="75C19A0D-E889-407D-B920-71856F9E151A" docUuidSource="ZooBank" docVersion="24" lastPageNumber="166" masterDocId="43218E4BA900FFE52B0BC14DFFA9FFE3" masterDocTitle="Gastropods from the Cretaceous-Palaeogene boundary in Denmark" masterLastPageNumber="196" masterPageNumber="1" pageNumber="166" updateTime="1698750564626" updateUser="ExternalLinkService">
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<mods:title id="6DF03E99004E172B5271B665123620C6">Gastropods from the Cretaceous-Palaeogene boundary in Denmark</mods:title>
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<mods:namePart id="96A70641E5FBA85020AEE8588189F939">Hansen, Thomas</mods:namePart>
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<treatment id="BF18F633A9A5FF402B9CC005FBADFA79" ID-DOI="http://doi.org/10.5281/zenodo.5583001" ID-GBIF-Taxon="159074647" ID-Zenodo-Dep="5583001" LSID="urn:lsid:zoobank.org:act:75C19A0D-E889-407D-B920-71856F9E151A" httpUri="http://treatment.plazi.org/id/BF18F633A9A5FF402B9CC005FBADFA79" lastPageNumber="166" pageId="165" pageNumber="166">
<subSubSection id="7FAB14AEA9A5FF402B9CC005FE71FE81" box="[151,472,328,355]" pageId="165" pageNumber="166" type="nomenclature">
<paragraph id="370E4725A9A5FF402B9CC005FE71FE81" blockId="165.[151,472,328,390]" box="[151,472,328,355]" pageId="165" pageNumber="166">
<heading id="6C46F049A9A5FF402B9CC005FE71FE81" bold="true" box="[151,472,328,355]" fontSize="11" level="1" pageId="165" pageNumber="166" reason="1">
<emphasis id="05C59B37A9A5FF402B9CC005FE71FE81" bold="true" box="[151,472,328,355]" pageId="165" pageNumber="166">
<taxonomicName id="F0B13CA6A9A5FF402B9CC005FE27FE81" authority="Hansen, 2019" authorityName="Hansen" authorityYear="2019" box="[151,398,328,355]" class="Gastropoda" family="Cylichnidae" genus="Cylichna" kingdom="Animalia" order="Cephalaspidea" pageId="165" pageNumber="166" phylum="Mollusca" rank="species" species="stevniensis" status="sp. nov.">
<emphasis id="05C59B37A9A5FF402B9CC005FE27FE81" bold="true" box="[151,398,328,355]" italics="true" pageId="165" pageNumber="166">Cylichna stevniensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1EF6264CA9A5FF402A9EC005FE71FE81" box="[405,472,328,354]" pageId="165" pageNumber="166" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7FAB14AEA9A5FF402B9CC021FEB7FE65" box="[151,286,363,390]" pageId="165" pageNumber="166" type="description">
<paragraph id="370E4725A9A5FF402B9CC021FEB7FE65" blockId="165.[151,472,328,390]" box="[151,286,363,390]" pageId="165" pageNumber="166">
<figureCitation id="AF8A5BA0A9A5FF402B9CC021FF40FE65" box="[151,233,363,390]" captionStart="FIGURE 35" captionStartId="167.[151,250,188,213]" captionTargetId="figure@166.[151,1435,179,1918]" captionTargetPageId="166" captionText="FIGURE 35. AB. Pseudomalaxis sp., Silicone cast of MGUH 33306. Width 4.3 mm. CD. Climacopoma? sp., Silicone cast of MGUH 33307. Width 13.1 and 9.9 mm respectively. EI. Mathilda dania n. sp. E. Silicone cast of holotype MGUH 33308. Width 1.9 mm. F. Silicone cast of paratype MGUH 33309. Width 2.0 mm. G. Silicone cast of paratype MGUH 33310. Width 0.8 mm. H. Silicone cast of paratype MGUH 33311. Width 0.7 mm. I. Silicone cast of paratype MGUH 33312. Width 1.7 mm. JK. Clathrobaculus sp., Silicone cast of MGUH 33313. Width 0.5 mm. LP. Ringicula sp. L. Silicone cast of MGUH 33314. Width 7.0 mm. MN. Silicone cast of MGUH 33315 and a partly hidden second specimen. Width 7.3 and 6.6 mm respectively. OP. Silicone cast of MGUH 33316. Width 5.6, 5.3 mm respectively. QT. Cylichna stevniensis n. sp. Q. Silicone cast of holotype MGUH 33317. Width 1.4 mm. RS. Internal mould of paratype MGUH 33318. Width 1.5 mm. T. Internal mould of paratype MGUH 33319. Width 2.8 mm. UV. Cylichna? sp. A. Cast of MGUH 33320. Width 2.3, 2.4 mm respectively. WY. Cylichna? sp. B. W. Wax cast of MGUH 33321. Width 2.4 mm. XY. Inner mould of MGUH 33322. Width 1.3 mm. ZAE. Roxania sp. Cast of MGUH 33323. Width 3.2, 3.3 mm respectively." figureDoi="http://doi.org/10.5281/zenodo.3442126" httpUri="https://zenodo.org/record/3442126/files/figure.png" pageId="165" pageNumber="166">Figs 35</figureCitation>
QT
</paragraph>
</subSubSection>
<subSubSection id="7FAB14AEA9A5FF402B9CC0FEFBADFA79" pageId="165" pageNumber="166" type="materials_examined">
<paragraph id="370E4725A9A5FF402B9CC0FEFE52FDF4" blockId="165.[151,1437,435,1435]" pageId="165" pageNumber="166">
<emphasis id="05C59B37A9A5FF402B9CC0FEFEBBFE2D" bold="true" box="[151,274,435,462]" pageId="165" pageNumber="166">Diagnosis.</emphasis>
Shell rather elongate with pointed posterior end, aperture distinctly higher than aperturally strongly convex penultimate whorl; apex hidden on adult specimens, not sharply truncated. Columellar fold very weak. Surface sculpture of fine spiral grooves.
</paragraph>
<paragraph id="370E4725A9A5FF402BCCC352FCDEFDD9" blockId="165.[151,1437,435,1435]" box="[199,887,543,571]" pageId="165" pageNumber="166">
<emphasis id="05C59B37A9A5FF402BCCC352FE1DFDD9" bold="true" box="[199,436,543,570]" pageId="165" pageNumber="166">Derivation of name.</emphasis>
Refers to the
<typeStatus id="E80AF987A9A5FF40295AC36CFD28FDD8" box="[593,641,545,571]" pageId="165" pageNumber="166">type</typeStatus>
locality Stevns Klint.
</paragraph>
<paragraph id="370E4725A9A5FF402BCCC309FD7EFD45" blockId="165.[151,1437,435,1435]" pageId="165" pageNumber="166">
<emphasis id="05C59B37A9A5FF402BCCC309FEDEFDBD" bold="true" box="[199,375,579,606]" pageId="165" pageNumber="166">Type material.</emphasis>
<materialsCitation id="87D94D78A9A5FF402A8BC309FB96FDBD" ID-GBIF-Occurrence="2331574371" box="[384,1087,580,607]" collectionCode="MGUH" pageId="165" pageNumber="166" specimenCode="MGUH 33317" specimenCount="1" typeStatus="holotype">
<typeStatus id="E80AF987A9A5FF402A8BC309FE41FDBD" box="[384,488,580,606]" pageId="165" pageNumber="166" type="holotype">Holotype</typeStatus>
<specimenCode id="6717EF5EA9A5FF402AF9C309FD3EFDBD" box="[498,663,580,606]" collectionCode="MGUH" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="165" pageNumber="166">MGUH 33317</specimenCode>
is a nearly complete external mould
</materialsCitation>
.
<materialsCitation id="87D94D78A9A5FF402F47C308FB86FD61" ID-GBIF-Occurrence="2331574411" collectionCode="MGUH" pageId="165" pageNumber="166" specimenCode="MGUH 33318" specimenCount="1" typeStatus="paratype">
<typeStatus id="E80AF987A9A5FF402F47C308FB07FDBC" box="[1100,1198,581,607]" pageId="165" pageNumber="166" type="paratype">Paratype</typeStatus>
<specimenCode id="6717EF5EA9A5FF402FB3C309FAF5FDBD" box="[1208,1372,580,606]" collectionCode="MGUH" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="165" pageNumber="166">MGUH 33318</specimenCode>
is an internal mould and comes from the
<taxonomicName id="F0B13CA6A9A5FF402938C325FC25FD61" authority="Limestone Member" authorityName="Gilleaudeau et al." authorityYear="2018" box="[563,908,616,642]" class="Gastropoda" family="Cerithiidae" genus="Cerithium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="165" pageNumber="166" phylum="Mollusca" rank="genus">Cerithium Limestone Member</taxonomicName>
at Skeldervig
</materialsCitation>
.
<materialsCitation id="87D94D78A9A5FF402F37C324FD7AFD45" ID-GBIF-Occurrence="2331574375" collectionCode="MGUH" pageId="165" pageNumber="166" specimenCode="MGUH 33319" specimenCount="2" typeStatus="paratype">
<typeStatus id="E80AF987A9A5FF402F37C324FB37FD60" box="[1084,1182,617,643]" pageId="165" pageNumber="166" type="paratype">Paratype</typeStatus>
<specimenCode id="6717EF5EA9A5FF402FACC325FAE5FD61" box="[1191,1356,616,642]" collectionCode="MGUH" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="165" pageNumber="166">MGUH 33319</specimenCode>
comes from the same horizon and locality as the
<typeStatus id="E80AF987A9A5FF402966C3C1FD7AFD45" box="[621,723,652,678]" pageId="165" pageNumber="166" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
<paragraph id="370E4725A9A5FF402BCCC3E2FD67FCD5" blockId="165.[151,1437,435,1435]" pageId="165" pageNumber="166">
<emphasis id="05C59B37A9A5FF402BCCC3E2FE12FD29" bold="true" box="[199,443,687,714]" pageId="165" pageNumber="166">Additional material.</emphasis>
Six specimens with the informal sample numbers SH.179, SH.192.B, SR.608 (
<specimenCount id="21B78CACA9A5FF402E49C3FDFF7FFD0C" pageId="165" pageNumber="166" type="generic">2 specimens</specimenCount>
), SR.680 and SO.191, together with three uncatalogued external and internal moulds from Stevns and a single mould from Northern Jutland, which is found in the private collection of K. I. Schnetler. The Stevns material belongs to the Natural History Museum of
<collectingCountry id="4FA607B5A9A5FF402955C251FD63FCD5" box="[606,714,796,822]" name="Denmark" pageId="165" pageNumber="166">Denmark</collectingCountry>
.
</paragraph>
<paragraph id="370E4725A9A5FF402BCCC20DFF71FC9D" blockId="165.[151,1437,435,1435]" pageId="165" pageNumber="166">
<emphasis id="05C59B37A9A5FF402BCCC20DFDEBFCB9" bold="true" box="[199,578,831,858]" pageId="165" pageNumber="166">
<typeStatus id="E80AF987A9A5FF402BCCC20DFEABFCB9" box="[199,258,832,858]" pageId="165" pageNumber="166">Type</typeStatus>
stratum and
<typeStatus id="E80AF987A9A5FF402AA2C20DFE75FCB9" box="[425,476,832,858]" pageId="165" pageNumber="166">type</typeStatus>
locality.
</emphasis>
<taxonomicName id="F0B13CA6A9A5FF402940C20DFC0DFCB9" authority="Limestone Member" authorityName="Gilleaudeau et al." authorityYear="2018" box="[587,932,832,858]" class="Gastropoda" family="Cerithiidae" genus="Cerithium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="165" pageNumber="166" phylum="Mollusca" rank="genus">Cerithium Limestone Member</taxonomicName>
of the Rødvig Formation at Rødvig, Stevns Klint.
</paragraph>
<paragraph id="370E4725A9A5FF402BCCC2CAFDB7FC25" blockId="165.[151,1437,435,1435]" pageId="165" pageNumber="166">
<emphasis id="05C59B37A9A5FF402BCCC2CAFEF3FC41" bold="true" box="[199,346,903,930]" pageId="165" pageNumber="166">Occurrence.</emphasis>
The
<taxonomicName id="F0B13CA6A9A5FF402A9AC2C5FD4CFC41" authority="Limestone Member" authorityName="Gilleaudeau et al." authorityYear="2018" box="[401,741,904,930]" class="Gastropoda" family="Cerithiidae" genus="Cerithium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="165" pageNumber="166" phylum="Mollusca" rank="genus">Cerithium Limestone Member</taxonomicName>
at Stevns Klint and the stratigrapically equivalent dead layer in Dania Quarry, Northern Jutland.
</paragraph>
<paragraph id="370E4725A9A5FF402BCCC282FB11FBB5" blockId="165.[151,1437,435,1435]" pageId="165" pageNumber="166">
<emphasis id="05C59B37A9A5FF402BCCC282FEF0FC09" bold="true" box="[199,345,975,1002]" pageId="165" pageNumber="166">Description.</emphasis>
Protoconch unknown. Teleoconch moderately thick-shelled, involute, elongated in shape with weakly curved outer lip and aperturally strongly convex penultimate whorl; apex covered by last whorl on adult specimens. Aperture apically pointed, widened and rounded abapically. Width of last whorl, excluding aperture, corresponding to half the spire height. Columella with very weak plait partly hidden by callus.
</paragraph>
<paragraph id="370E4725A9A5FF402BCCC52DFE03FB7D" blockId="165.[151,1437,435,1435]" pageId="165" pageNumber="166">
Sculpture, if present, consisting of weak, fine spiral grooves without pitting, grooves numbering round 30 on
<typeStatus id="E80AF987A9A5FF402B9CC5C9FF50FB7D" box="[151,249,1156,1182]" pageId="165" pageNumber="166" type="holotype">holotype</typeStatus>
<collectionCode id="51A0DFE0A9A5FF402A0BC5C9FEFFFB7D" box="[256,342,1156,1182]" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="165" pageNumber="166">MGUH</collectionCode>
33317.
</paragraph>
<paragraph id="370E4725A9A5FF402BCCC5E5FB61FB21" blockId="165.[151,1437,435,1435]" box="[199,1224,1191,1219]" pageId="165" pageNumber="166">
<emphasis id="05C59B37A9A5FF402BCCC5E5FED7FB21" bold="true" box="[199,382,1192,1218]" pageId="165" pageNumber="166">Measurements.</emphasis>
Largest specimen
<collectionCode id="51A0DFE0A9A5FF40295BC5E5FD0FFB21" box="[592,678,1192,1218]" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="165" pageNumber="166">MGUH</collectionCode>
33319 is
<quantity id="F049EAC0A9A5FF402818C5EAFCC0FB20" box="[787,873,1191,1219]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="165" pageNumber="166" unit="mm" value="5.2">5.2 mm</quantity>
high and at least
<quantity id="F049EAC0A9A5FF402F26C5E5FB2AFB20" box="[1069,1155,1192,1219]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="165" pageNumber="166" unit="mm" value="2.8">2.8 mm</quantity>
wide.
</paragraph>
<paragraph id="370E4725A9A5FF402BCCC581FBADFA79" blockId="165.[151,1437,435,1435]" pageId="165" pageNumber="166">
<emphasis id="05C59B37A9A5FF402BCCC581FE92FB05" bold="true" box="[199,315,1228,1254]" pageId="165" pageNumber="166">Remarks.</emphasis>
As noted by
<bibRefCitation id="53203AD4A9A5FF402ADCC581FD78FB05" author="Kaim, A. &amp; Beisel, A. L." box="[471,721,1227,1254]" pageId="165" pageNumber="166" pagination="41 - 64" refId="ref133059" refString="Kaim, A. &amp; Beisel, A. L. (2005) Mesozoic gastropods from Siberia and Timan (Russia). Part 2: Neogastropoda and Heterobranchia. Polish Polar Research, 26, 41 - 64." type="journal article" year="2005">Kaim &amp; Beisel (2005</bibRefCitation>
: p. 61) the species of
<taxonomicName id="F0B13CA6A9A5FF4028DBC581FB9CFB05" authorityName="Loven" authorityYear="1846" box="[976,1077,1228,1254]" class="Gastropoda" family="Cylichnidae" genus="Cylichna" kingdom="Animalia" order="Cephalaspidea" pageId="165" pageNumber="166" phylum="Mollusca" rank="genus">
<emphasis id="05C59B37A9A5FF4028DBC581FB9CFB05" box="[976,1077,1228,1254]" italics="true" pageId="165" pageNumber="166">Cylichna</emphasis>
</taxonomicName>
are extremely similar and only few features may be used for differentiation on fossil specimens. Among the stratigraphically and geographically comparable taxa
<taxonomicName id="F0B13CA6A9A5FF402A5DC459FD92FACD" authority="Hansen, 2019" authorityName="Hansen" authorityYear="2019" box="[342,571,1300,1326]" class="Gastropoda" family="Cylichnidae" genus="Cylichna" kingdom="Animalia" order="Cephalaspidea" pageId="165" pageNumber="166" phylum="Mollusca" rank="species" species="stevniensis" status="sp. nov.">
<emphasis id="05C59B37A9A5FF402A5DC459FD92FACD" box="[342,571,1300,1326]" italics="true" pageId="165" pageNumber="166">Cylichna stevniensis</emphasis>
</taxonomicName>
<emphasis id="05C59B37A9A5FF402949C459FD28FACD" bold="true" box="[578,641,1300,1326]" pageId="165" pageNumber="166">
<taxonomicNameLabel id="1EF6264CA9A5FF402949C459FD28FACD" box="[578,641,1300,1326]" pageId="165" pageNumber="166" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is distinguished from the Danian
<taxonomicName id="F0B13CA6A9A5FF4028F0C458FF70FAB1" authority="(Briart &amp; Cornet, 1887)" baseAuthorityName="Briart &amp; Cornet" baseAuthorityYear="1887" class="Gastropoda" family="Scaphandridae" genus="Roxania" kingdom="Animalia" order="Cephalaspidea" pageId="165" pageNumber="166" phylum="Mollusca" rank="species" species="raristriata">
<emphasis id="05C59B37A9A5FF4028F0C458FB7DFACD" box="[1019,1236,1301,1326]" italics="true" pageId="165" pageNumber="166">Roxania raristriata</emphasis>
(
<bibRefCitation id="53203AD4A9A5FF402FE9C459FF79FAB1" author="Briart, A. &amp; Cornet, F. L." pageId="165" pageNumber="166" pagination="1 - 118" refId="ref126106" refString="Briart, A. &amp; Cornet, F. L. (1887) Description des fossiles du calcaire grossier de Mons. Gasteropodes. Partie IV. Memoires couronnes et Memoires des Savants Etrangers, Academie Royale des Sciences, des Lettres et des Beaux-Arts de Belgique, 47, 1 - 118, pls. 1 - 26." type="journal article" year="1887">Briart &amp; Cornet, 1887</bibRefCitation>
)
</taxonomicName>
from
<collectingCountry id="4FA607B5A9A5FF402A16C475FED4FAB1" box="[285,381,1336,1362]" name="Belgium" pageId="165" pageNumber="166">Belgium</collectingCountry>
as figured by
<bibRefCitation id="53203AD4A9A5FF402915C475FD13FAB1" author="Glibert, M." box="[542,698,1336,1362]" pageId="165" pageNumber="166" pagination="1 - 116" refId="ref130648" refString="Glibert, M. (1973) Revision des Gastropoda du Danien et du Montien de la Belgique. 1. Les Gastropoda du Calcaire de Mons. Institut Royal des Sciences Naturelles de Belgique Memoire, 173, 1 - 116, pls. 1 - 11." type="journal article" year="1973">Glibert (1973</bibRefCitation>
: pl.
<collectionCode id="51A0DFE0A9A5FF4029E2C474FCA2FAB0" box="[745,779,1337,1363]" pageId="165" pageNumber="166">XI</collectionCode>
: 1) by the distinctly higher aperture opening relative to the apex. It resembles somewhat
<taxonomicName id="F0B13CA6A9A5FF402AE0C411FDF9FA95" authorityName="Loven" authorityYear="1846" box="[491,592,1372,1398]" class="Gastropoda" family="Cylichnidae" genus="Cylichna" kingdom="Animalia" order="Cephalaspidea" pageId="165" pageNumber="166" phylum="Mollusca" rank="genus">
<emphasis id="05C59B37A9A5FF402AE0C411FDF9FA95" box="[491,592,1372,1398]" italics="true" pageId="165" pageNumber="166">Cylichna</emphasis>
</taxonomicName>
sp. 1 of
<bibRefCitation id="53203AD4A9A5FF4029B0C411FC71FA95" author="Kollmann, H. A. &amp; Peel, J. S." box="[699,984,1372,1398]" pageId="165" pageNumber="166" pagination="1 - 115" refId="ref134023" refString="Kollmann, H. A. &amp; Peel, J. S. (1983) Paleocene gastropods from Nugssuaq, West Greenland. GrOnlands Geologiske UndersOgelse, Bulletin, 146, 1 - 115." type="journal article" year="1983">Kollmann &amp; Peel (1983)</bibRefCitation>
from the Danian of
<collectingCountry id="4FA607B5A9A5FF402FC3C411FAE9FA95" box="[1224,1344,1372,1398]" name="Greenland" pageId="165" pageNumber="166">Greenland</collectingCountry>
, but the columella is generally longer with a slightly better developed columellar plait.
</paragraph>
</subSubSection>
</treatment>
</document>