301 lines
32 KiB
XML
301 lines
32 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1142.90097" ID-Pensoft-Pub="1313-2970-1142-1" ID-Pensoft-UUID="1D4BDF04F72B59B9984E00D0B4FB338D" ID-ZooBank="A1129EE50F9941CFB73AE771B66E2486" ModsDocID="1313-2970-1142-1" checkinTime="1674655519517" checkinUser="pensoft" docAuthor="Tongkerd, Piyoros, Tumpeesuwan, Sakboworn, Inkhavilay, Khamla, Prasankok, Pongpun, Jeratthitikul, Ekgachai, Panha, Somsak & Sutcharit, Chirasak" docDate="2023" docId="7279C3DCA2715E4297DDD938D43BAF25" docLanguage="en" docName="ZooKeys 1142: 1-144" docOrigin="ZooKeys 1142" docPubDate="2023-01-24" docSource="http://dx.doi.org/10.3897/zookeys.1142.90097" docTitle="Pterocyclos jousseaumei" docType="treatment" docVersion="1" id="1D4BDF04F72B59B9984E00D0B4FB338D" lastPageNumber="1" masterDocId="1D4BDF04F72B59B9984E00D0B4FB338D" masterDocTitle="Systematic revision of the snorkel snail genus Rhiostoma Benson, 1860 (Gastropoda, Caenogastropoda, Cyclophoridae) with descriptions of new species" masterLastPageNumber="144" masterPageNumber="1" pageNumber="1" updateTime="1674655519517" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Systematic revision of the snorkel snail genus Rhiostoma Benson, 1860 (Gastropoda, Caenogastropoda, Cyclophoridae) with descriptions of new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tongkerd, Piyoros</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tumpeesuwan, Sakboworn</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Present address: Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai, Maha Sarakham 44150 Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Inkhavilay, Khamla</mods:namePart>
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<mods:affiliation>Department of Biology, Faculty of Natural Science, National University of Laos, P. O. Box 7322, Dongdok, Vientiane, Laos</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Prasankok, Pongpun</mods:namePart>
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<mods:affiliation>School of Biology, Institute of Science, Suranaree University of Technology, Nakhon Ratchasima 30000, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jeratthitikul, Ekgachai</mods:namePart>
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<mods:affiliation>Animal Systematics and Molecular Ecology Laboratory, Department of Biology, Faculty of Science, Mahidol University, Bangkok 10400, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panha, Somsak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-01-24</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1142</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>144</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1142.90097</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1142.90097</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1142-1</mods:identifier>
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<mods:identifier type="ZooBank">A1129EE50F9941CFB73AE771B66E2486</mods:identifier>
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<mods:identifier type="Pensoft-UUID">1D4BDF04F72B59B9984E00D0B4FB338D</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:7279C3DCA2715E4297DDD938D43BAF25" httpUri="http://treatment.plazi.org/id/7279C3DCA2715E4297DDD938D43BAF25" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="7279C3DC-A271-5E42-97DD-D938D43BAF25" authority="(Morgan, 1885)" baseAuthorityName="Morgan" baseAuthorityYear="1885" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pterocyclos jousseaumei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="jousseaumei">Pterocyclos jousseaumei (Morgan, 1885)</taxonomicName>
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</paragraph>
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||
</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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||
<paragraph pageId="0" pageNumber="1">
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||
<figureCitation captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Shell of Opisthoporus jousseaumei A original figure (after Morgan 1885: pl. 8, fig. 2) B specimen CUMZ 4752 from Ipoh, Perak, Malaysia C specimen CUMZ 7457 from Kinta, Perak, Malaysia D lectotype of " Rhiostoma macalpinewoodsi Laidlaw, 1939 " NHMUK 1939.4.13.23 from Sungei Siput, Perak E, F specimens CUMZ 10213 from Gunung Dato, Ipoh, Perak, Malaysia." figureDoi="10.3897/zookeys.1142.90097.figure56" httpUri="https://binary.pensoft.net/fig/799914" pageId="0" pageNumber="1">Fig. 56</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="0" pageNumber="1" type="reference_group">
|
||
<paragraph pageId="0" pageNumber="1">
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||
<taxonomicName authorityName="de Morgan" authorityYear="1885" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma jousseaumei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="jousseaumei">Rhiostoma jousseaumei</taxonomicName>
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Morgan, 1885: 400, pl. 8, fig. 2. Type locality: la haute
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<normalizedToken originalValue="vallée">vallee</normalizedToken>
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de Kinta (
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<normalizedToken originalValue="Pérak">Perak</normalizedToken>
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) [Kinta valley, Perak, Malaysia].
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<bibRefCitation DOI="https://doi.org/10.1111/j.1096-3642.1891.tb01757.x" author="Moellendorff, OF von" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" pagination="330 - 348" refId="B96" refString="Moellendorff, OF von, 1891. On the land and freshwater shells of Perak. Proceedings of the Zoological Society of London 59 (3): 330 - 348, DOI: https://doi.org/10.1111/j.1096-3642.1891.tb01757.x" title="On the land and freshwater shells of Perak." url="https://doi.org/10.1111/j.1096-3642.1891.tb01757.x" volume="59" year="1891">
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<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
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1891
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</bibRefCitation>
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: 340.
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<bibRefCitation author="Kobelt, W" journalOrPublisher="Das Tierreich" pageId="0" pageNumber="1" pagination="1 - 663" refId="B71" refString="Kobelt, W, 1902. Cyclophoridae. Das Tierreich 16: 1 - 663" title="Cyclophoridae." volume="16" year="1902">Kobelt 1902</bibRefCitation>
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: 178. Kobelt 1911: 758, pl. 110, figs 11-13.
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<bibRefCitation author="Laidlaw, FF" journalOrPublisher="Journal of the Malaysian Branch of the Royal Asiatic Society" pageId="0" pageNumber="1" pagination="25 - 37" refId="B80" refString="Laidlaw, FF, 1928. A list of land and fresh-water Mollusca of the Malay Peninsula with note. Journal of the Malaysian Branch of the Royal Asiatic Society 3: 25 - 37" title="A list of land and fresh-water Mollusca of the Malay Peninsula with note." volume="3" year="1928">Laidlaw 1928</bibRefCitation>
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: 31.
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<bibRefCitation author="Chang, S-Y" journalOrPublisher="The Papustyla" pageId="0" pageNumber="1" refId="B27" refString="Chang, S-Y, 1995. A brief collection of land shells of Desaru anjung Penawar. The Papustyla 9: 4, 5." title="A brief collection of land shells of Desaru anjung Penawar. The Papustyla 9: 4, 5." year="1995">Chang 1995</bibRefCitation>
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: 5 with text figure.
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<bibRefCitation DOI="https://doi.org/10.3897/zookeys.682.12999" author="Foon, JK" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 94" refId="B45" refString="Foon, JK, Clements, GR, Liew, T-S, 2017. Diversity and biogeography of land snails (Mollusca, Gastropoda) in the limestone hills of Perak, Peninsular Malaysia. ZooKeys 682: 1 - 94, DOI: https://doi.org/10.3897/zookeys.682.12999" title="Diversity and biogeography of land snails (Mollusca, Gastropoda) in the limestone hills of Perak, Peninsular Malaysia." url="https://doi.org/10.3897/zookeys.682.12999" volume="682" year="2017">Foon et al. 2017</bibRefCitation>
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||
: 24, fig. 8f.
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||
</paragraph>
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||
<paragraph pageId="0" pageNumber="1">
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||
<taxonomicName authorityName="Tongkerd & Tumpeesuwan & Inkhavilay & Prasankok & Jeratthitikul & Panha & Sutcharit" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Rhiostoma</taxonomicName>
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<taxonomicName authorityName="Tongkerd & Tumpeesuwan & Inkhavilay & Prasankok & Jeratthitikul & Panha & Sutcharit" authorityYear="2023" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rhiostoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Rhiostoma</taxonomicName>
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macalpine-woodsi Laidlaw, 1939: 166, with text figure. Type locality: Sungei Siput, Perak.
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||
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.842.29243" author="Sutcharit, C" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 65" refId="B160" refString="Sutcharit, C, Ablett, JD, Panha, S, 2019. An annotated type catalogue of seven genera of operculate land snails (Caenogastropoda, Cyclophoridae) in the Natural History Museum, London. ZooKeys 842: 1 - 65, DOI: https://doi.org/10.3897/zookeys.842.29243" title="An annotated type catalogue of seven genera of operculate land snails (Caenogastropoda, Cyclophoridae) in the Natural History Museum, London." url="https://doi.org/10.3897/zookeys.842.29243" volume="842" year="2019">Sutcharit et al. 2019</bibRefCitation>
|
||
: 37, fig. 8g.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="type material">
|
||
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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||
<paragraph pageId="0" pageNumber="1">
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||
The type specimens
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||
<taxonomicName authorityName="de Morgan" authorityYear="1885" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma jousseaumei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="jousseaumei">
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||
<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma jousseaumei</emphasis>
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||
</taxonomicName>
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Morgan, 1885 could not be located in the MNHN collection.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Lectotype</emphasis>
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(designation in
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<bibRefCitation DOI="https://doi.org/10.3897/zookeys.842.29243" author="Sutcharit, C" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 65" refId="B160" refString="Sutcharit, C, Ablett, JD, Panha, S, 2019. An annotated type catalogue of seven genera of operculate land snails (Caenogastropoda, Cyclophoridae) in the Natural History Museum, London. ZooKeys 842: 1 - 65, DOI: https://doi.org/10.3897/zookeys.842.29243" title="An annotated type catalogue of seven genera of operculate land snails (Caenogastropoda, Cyclophoridae) in the Natural History Museum, London." url="https://doi.org/10.3897/zookeys.842.29243" volume="842" year="2019">Sutcharit et al. (2019)</bibRefCitation>
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||
) NHMUK 1939.4.13.23 (Fig.
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||
<figureCitation captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Shell of Opisthoporus jousseaumei A original figure (after Morgan 1885: pl. 8, fig. 2) B specimen CUMZ 4752 from Ipoh, Perak, Malaysia C specimen CUMZ 7457 from Kinta, Perak, Malaysia D lectotype of " Rhiostoma macalpinewoodsi Laidlaw, 1939 " NHMUK 1939.4.13.23 from Sungei Siput, Perak E, F specimens CUMZ 10213 from Gunung Dato, Ipoh, Perak, Malaysia." figureDoi="10.3897/zookeys.1142.90097.figure56" httpUri="https://binary.pensoft.net/fig/799914" pageId="0" pageNumber="1">56D</figureCitation>
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||
) of
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||
<taxonomicName authorityName="Laidlaw" authorityYear="1939" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma macalpinewoodsi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="macalpinewoodsi">
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<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma macalpinewoodsi</emphasis>
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</taxonomicName>
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Laidlaw, 1939, from Sungei Siput, Perak.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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||
<paragraph pageId="0" pageNumber="1">Other material examined.</paragraph>
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||
<paragraph pageId="0" pageNumber="1">
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||
<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Malaysia" location="Perak State" specimenCount="1" stateProvince="Perak State">
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||
<emphasis bold="true" pageId="0" pageNumber="1">
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||
<collectingCountry name="Malaysia">Malaysia</collectingCountry>
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</emphasis>
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: Specimen CUMZ 4752 (2 shells; Fig.
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<figureCitation captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Shell of Opisthoporus jousseaumei A original figure (after Morgan 1885: pl. 8, fig. 2) B specimen CUMZ 4752 from Ipoh, Perak, Malaysia C specimen CUMZ 7457 from Kinta, Perak, Malaysia D lectotype of " Rhiostoma macalpinewoodsi Laidlaw, 1939 " NHMUK 1939.4.13.23 from Sungei Siput, Perak E, F specimens CUMZ 10213 from Gunung Dato, Ipoh, Perak, Malaysia." figureDoi="10.3897/zookeys.1142.90097.figure56" httpUri="https://binary.pensoft.net/fig/799914" pageId="0" pageNumber="1">56B</figureCitation>
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) from Ipoh,
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<collectingRegion country="Malaysia" name="Perak">Perak State</collectingRegion>
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||
</materialsCitation>
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||
. CUMZ 7457 (15 shells; Fig.
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<figureCitation captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Shell of Opisthoporus jousseaumei A original figure (after Morgan 1885: pl. 8, fig. 2) B specimen CUMZ 4752 from Ipoh, Perak, Malaysia C specimen CUMZ 7457 from Kinta, Perak, Malaysia D lectotype of " Rhiostoma macalpinewoodsi Laidlaw, 1939 " NHMUK 1939.4.13.23 from Sungei Siput, Perak E, F specimens CUMZ 10213 from Gunung Dato, Ipoh, Perak, Malaysia." figureDoi="10.3897/zookeys.1142.90097.figure56" httpUri="https://binary.pensoft.net/fig/799914" pageId="0" pageNumber="1">56C</figureCitation>
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||
) from Tanjung Rambutan, Kinta,
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<collectingRegion country="Malaysia" name="Perak">Perak State</collectingRegion>
|
||
. CUMZ 10213 (5 shells; Fig.
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<figureCitation captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Shell of Opisthoporus jousseaumei A original figure (after Morgan 1885: pl. 8, fig. 2) B specimen CUMZ 4752 from Ipoh, Perak, Malaysia C specimen CUMZ 7457 from Kinta, Perak, Malaysia D lectotype of " Rhiostoma macalpinewoodsi Laidlaw, 1939 " NHMUK 1939.4.13.23 from Sungei Siput, Perak E, F specimens CUMZ 10213 from Gunung Dato, Ipoh, Perak, Malaysia." figureDoi="10.3897/zookeys.1142.90097.figure56" httpUri="https://binary.pensoft.net/fig/799914" pageId="0" pageNumber="1">56E, F</figureCitation>
|
||
) from Gunung Dato, Ipoh,
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<collectingRegion country="Malaysia" name="Perak">Perak State</collectingRegion>
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.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1142.90097.figure56" httpUri="https://binary.pensoft.net/fig/799914" pageId="0" pageNumber="1" start="Figure 56" startId="F56">
|
||
<paragraph pageId="0" pageNumber="1">
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||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 56.</emphasis>
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Shell of
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<taxonomicName class="Gastropoda" family="Cyclophoridae" genus="Opisthoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Opisthoporus jousseaumei" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="jousseaumei">
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||
<emphasis italics="true" pageId="0" pageNumber="1">Opisthoporus jousseaumei</emphasis>
|
||
</taxonomicName>
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||
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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||
original figure (after
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.14301" author="Morgan, J de" journalOrPublisher="Bulletin de la Societe Zoologique de France" pageId="0" pageNumber="1" pagination="353 - 428" refId="B104" refString="Morgan, J de, 1885. Mollusques terrestres & fluviatiles du royaume de Perak et des pays voisins (presqueIle Malaise). Bulletin de la Societe Zoologique de France 10: 353 - 428, DOI: https://doi.org/10.5962/bhl.part.14301" title="Mollusques terrestres & fluviatiles du royaume de Perak et des pays voisins (presqueIle Malaise)." url="https://doi.org/10.5962/bhl.part.14301" volume="10" year="1885">Morgan 1885</bibRefCitation>
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||
: pl. 8, fig. 2)
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||
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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||
specimen CUMZ 4752 from Ipoh,
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<collectingRegion country="Malaysia" name="Perak">Perak</collectingRegion>
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,
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<collectingCountry name="Malaysia">Malaysia</collectingCountry>
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||
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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||
specimen CUMZ 7457 from Kinta,
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<collectingRegion country="Malaysia" name="Perak">Perak</collectingRegion>
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||
,
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<collectingCountry name="Malaysia">Malaysia</collectingCountry>
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<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
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||
<typeStatus>lectotype</typeStatus>
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||
of "
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||
<taxonomicName authorityName="Laidlaw" authorityYear="1939" class="Gastropoda" family="Cyclophoridae" genus="Rhiostoma" kingdom="Animalia" lsidName="Rhiostoma macalpinewoodsi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="macalpinewoodsi">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Rhiostoma macalpinewoodsi</emphasis>
|
||
</taxonomicName>
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||
Laidlaw, 1939" NHMUK 1939.4.13.23 from Sungei Siput,
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||
<collectingRegion country="Malaysia" name="Perak">Perak</collectingRegion>
|
||
<emphasis bold="true" pageId="0" pageNumber="1">E, F</emphasis>
|
||
specimens CUMZ 10213 from Gunung Dato, Ipoh,
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||
<collectingRegion country="Malaysia" name="Perak">Perak</collectingRegion>
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||
,
|
||
<collectingCountry name="Malaysia">Malaysia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="remarks">
|
||
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">Periostracum thin, corneous, shell varying from uniform colour to dark brown zigzag pattern; periphery with a thin spiral band. Detached-whorl length similar to apertural width. Peristome circular and double or boundary hardly visible; lip thick or thin, and not expanded to slightly expanded. Breathing device with incomplete tube or tubular shape, curved posteriorly and its tip usually attached to preceding whorl. Operculum calcareous, multispiral, and slightly concave on both surfaces.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
An operculum accompanied the examined specimen and was also figured in
|
||
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.14301" author="Morgan, J de" journalOrPublisher="Bulletin de la Societe Zoologique de France" pageId="0" pageNumber="1" pagination="353 - 428" refId="B104" refString="Morgan, J de, 1885. Mollusques terrestres & fluviatiles du royaume de Perak et des pays voisins (presqueIle Malaise). Bulletin de la Societe Zoologique de France 10: 353 - 428, DOI: https://doi.org/10.5962/bhl.part.14301" title="Mollusques terrestres & fluviatiles du royaume de Perak et des pays voisins (presqueIle Malaise)." url="https://doi.org/10.5962/bhl.part.14301" volume="10" year="1885">Morgan (1885)</bibRefCitation>
|
||
: it was multispiral and plate-shaped (Fig.
|
||
<figureCitation captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Shell of Opisthoporus jousseaumei A original figure (after Morgan 1885: pl. 8, fig. 2) B specimen CUMZ 4752 from Ipoh, Perak, Malaysia C specimen CUMZ 7457 from Kinta, Perak, Malaysia D lectotype of " Rhiostoma macalpinewoodsi Laidlaw, 1939 " NHMUK 1939.4.13.23 from Sungei Siput, Perak E, F specimens CUMZ 10213 from Gunung Dato, Ipoh, Perak, Malaysia." figureDoi="10.3897/zookeys.1142.90097.figure56" httpUri="https://binary.pensoft.net/fig/799914" pageId="0" pageNumber="1">56A, C</figureCitation>
|
||
). With these observations, we suggest the re-assignment of this species to the
|
||
<taxonomicName authorityName="Benson" authorityYear="1832" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" lsidName="Pterocyclos" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Pterocyclos</emphasis>
|
||
</taxonomicName>
|
||
. This species can be separated from
|
||
<taxonomicName authorityName="Sykes" authorityYear="1903" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" lsidName="Pterocyclos subalatus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="subalatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Pterocyclos subalatus</emphasis>
|
||
</taxonomicName>
|
||
Sykes, 1903,
|
||
<taxonomicName baseAuthorityName="Tomlin" baseAuthorityYear="1931" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" lsidName="Pterocyclos spelaeotes" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="spelaeotes">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Pterocyclos spelaeotes</emphasis>
|
||
</taxonomicName>
|
||
(Tomlin, 1931), and
|
||
<taxonomicName baseAuthorityName="van Benthem Jutting" baseAuthorityYear="1949" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" lsidName="Pterocyclos umbraticus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="umbraticus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Pterocyclos umbraticus</emphasis>
|
||
</taxonomicName>
|
||
(Benthem Jutting, 1949) from peninsular Malaysia in having the apertural lip thickened and usually not expanded (rarely slightly expanded), while the three species (Fig.
|
||
<figureCitation captionStart="Figure 57" captionStartId="F57" captionText="Figure 57. A Pterocyclos subalatus Sykes, 1903, holotype UMZC 1032 from Gunong Inas, Perak, Malay Peninsula B Pterocyclos spelaeotes (Tomlin, 1931), lectotype NMW 1955.158.01107 from Batu Cave, Kula Lumpur, Selangor C Pterocyclos umbraticus (Benthem Jutting, 1949), holotype ZMA 135622 from Maxwell's Hill, Perak, Malaysia D Cyclotus boxalli syntype NHMUK 1894.5.23.1 from Kina Balu E Opisthoporus battambangensis holotype MNHN-IM- 2000 - 35506 from Battambang, Cambodia F Opisthoporus grohi holotype MNHN-IM- 2000 - 35507 from Dak Nong, Vietnam. Photograph: H Wood, NMW (B), M Caballer, A Lardeur and V Heros, MNHN (E, F)." figureDoi="10.3897/zookeys.1142.90097.figure57" httpUri="https://binary.pensoft.net/fig/799915" pageId="0" pageNumber="1">57A-C</figureCitation>
|
||
) tend to have a widely expanded lip (see also
|
||
<bibRefCitation author="Sutcharit, C" journalOrPublisher="The Raffles Bulletin of Zoology" pageId="0" pageNumber="1" pagination="330 - 338" refId="B159" refString="Sutcharit, C, Tongkerd, P, Panha, S, 2014. The land snail genus Pterocyclos Benson, 1832 (Caenogastropoda: Cyclophoridae) from Thailand and Peninsular Malaysia, with description of two new species. The Raffles Bulletin of Zoology 62: 330 - 338" title="The land snail genus Pterocyclos Benson, 1832 (Caenogastropoda: Cyclophoridae) from Thailand and Peninsular Malaysia, with description of two new species." volume="62" year="2014">Sutcharit et al. 2014</bibRefCitation>
|
||
). Furthermore,
|
||
<taxonomicName lsidName="P. subalatus" pageId="0" pageNumber="1" rank="species" species="subalatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. subalatus</emphasis>
|
||
</taxonomicName>
|
||
has no detached whorl but has a wide canal-shaped breathing device attached to the preceding whorl;
|
||
<taxonomicName lsidName="P. umbraticus" pageId="0" pageNumber="1" rank="species" species="umbraticus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">P. umbraticus</emphasis>
|
||
</taxonomicName>
|
||
has a larger shell (width 27 mm), without a detached whorl, and an undulated shell surface.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1142.90097.figure57" httpUri="https://binary.pensoft.net/fig/799915" pageId="0" pageNumber="1" start="Figure 57" startId="F57">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 57.</emphasis>
|
||
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
<taxonomicName authorityName="Sykes" authorityYear="1903" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" lsidName="Pterocyclos subalatus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="subalatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Pterocyclos subalatus</emphasis>
|
||
</taxonomicName>
|
||
Sykes, 1903, holotype UMZC 1032 from Gunong Inas, Perak, Malay Peninsula
|
||
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
<taxonomicName baseAuthorityName="Tomlin" baseAuthorityYear="1931" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" lsidName="Pterocyclos spelaeotes" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="spelaeotes">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Pterocyclos spelaeotes</emphasis>
|
||
</taxonomicName>
|
||
(Tomlin, 1931), lectotype NMW 1955.158.01107 from Batu Cave, Kula Lumpur, Selangor
|
||
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
<taxonomicName baseAuthorityName="van Benthem Jutting" baseAuthorityYear="1949" class="Gastropoda" family="Cyclophoridae" genus="Pterocyclos" kingdom="Animalia" lsidName="Pterocyclos umbraticus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="umbraticus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Pterocyclos umbraticus</emphasis>
|
||
</taxonomicName>
|
||
(Benthem Jutting, 1949), holotype ZMA 135622 from
|
||
<normalizedToken originalValue="Maxwell’s">Maxwell's</normalizedToken>
|
||
Hill, Perak, Malaysia
|
||
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
<taxonomicName baseAuthorityName="Godwin-Austen" baseAuthorityYear="1893" class="Gastropoda" family="Cyclophoridae" genus="Cyclotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyclotus boxalli" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="boxalli">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Cyclotus boxalli</emphasis>
|
||
</taxonomicName>
|
||
syntype NHMUK 1894.5.23.1 from Kina Balu
|
||
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||
<taxonomicName baseAuthorityName="Thach & Huber" baseAuthorityYear="2020" class="Gastropoda" family="Cyclophoridae" genus="Opisthoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Opisthoporus battambangensis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="battambangensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Opisthoporus battambangensis</emphasis>
|
||
</taxonomicName>
|
||
holotype MNHN-IM-2000-35506 from Battambang, Cambodia
|
||
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||
<taxonomicName baseAuthorityName="Thach" baseAuthorityYear="2020" class="Gastropoda" family="Cyclophoridae" genus="Opisthoporus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Opisthoporus grohi" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="grohi">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Opisthoporus grohi</emphasis>
|
||
</taxonomicName>
|
||
holotype MNHN-IM-2000-35507 from Dak Nong, Vietnam. Photograph: H Wood, NMW (B), M Caballer, A Lardeur and V Heros, MNHN (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">E, F</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<bibRefCitation author="Laidlaw, FF" journalOrPublisher="Journal of the Malaysian Branch of the Royal Asiatic Society" pageId="0" pageNumber="1" refId="B81" refString="Laidlaw, FF, 1939. A new Rhiostoma from Malay. Journal of Conchology 21: 116." title="A new Rhiostoma from Malay. Journal of Conchology 21: 116." year="1939">Laidlaw (1939)</bibRefCitation>
|
||
stated that
|
||
<taxonomicName lsidName="R. macalpinewoodsi" pageId="0" pageNumber="1" rank="species" species="macalpinewoodsi">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">R. macalpinewoodsi</emphasis>
|
||
</taxonomicName>
|
||
differed from
|
||
<taxonomicName lsidName="R. jousseaumei" pageId="0" pageNumber="1" rank="species" species="jousseaumei">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">R. jousseaumei</emphasis>
|
||
</taxonomicName>
|
||
in having a relatively small shell (height 6 mm, width 12 mm), with slightly thickened but not expanded apertural lip, a breathing device channel-like to short tubular [= incomplete tube] shape. Later,
|
||
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.682.12999" author="Foon, JK" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 94" refId="B45" refString="Foon, JK, Clements, GR, Liew, T-S, 2017. Diversity and biogeography of land snails (Mollusca, Gastropoda) in the limestone hills of Perak, Peninsular Malaysia. ZooKeys 682: 1 - 94, DOI: https://doi.org/10.3897/zookeys.682.12999" title="Diversity and biogeography of land snails (Mollusca, Gastropoda) in the limestone hills of Perak, Peninsular Malaysia." url="https://doi.org/10.3897/zookeys.682.12999" volume="682" year="2017">Foon et al. (2017)</bibRefCitation>
|
||
examined several specimens and found an intermediate shell form; the type locality of both nominal species is situated in the Kinta Valley, Perak State, peninsular Malaysia. In addition, the specimens examined herein show a mix between smaller shell (with incomplete tube) and larger shell (with tubular) forms in a single collection locality, with the intermediates as reported in
|
||
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.682.12999" author="Foon, JK" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 94" refId="B45" refString="Foon, JK, Clements, GR, Liew, T-S, 2017. Diversity and biogeography of land snails (Mollusca, Gastropoda) in the limestone hills of Perak, Peninsular Malaysia. ZooKeys 682: 1 - 94, DOI: https://doi.org/10.3897/zookeys.682.12999" title="Diversity and biogeography of land snails (Mollusca, Gastropoda) in the limestone hills of Perak, Peninsular Malaysia." url="https://doi.org/10.3897/zookeys.682.12999" volume="682" year="2017">Foon et al. (2017)</bibRefCitation>
|
||
. This distinctive shell size and breathing device could be attributed to intraspecific variation, and examination of more specimens and systematic study will help resolve this issue.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |