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<document ID-DOI="http://doi.org/10.5281/zenodo.4763728" ID-GBIF-Dataset="0420e2c2-2025-486d-96a2-c661dc998d66" ID-GBIF-Taxon="182074750" ID-ISSN="1854-0392" ID-Zenodo-Dep="4763728" approvalRequired="3" approvalRequired_for_taxonomicNames="1" approvalRequired_for_textStreams="2" checkinTime="1620963536357" checkinUser="felipe" docAuthor="Stark, Bill P." docDate="2012" docId="03E487F6023F937D0A64FC924953848C" docLanguage="en" docName="Illiesia.8.10.114-118.pdf" docOrigin="Illiesia 8 (10)" docStyle="DocumentStyle:3ED4ECBC8873166DC296BE619562C54F.3:Illiesia.2009-.journal_article" docStyleId="3ED4ECBC8873166DC296BE619562C54F" docStyleName="Illiesia.2009-.journal_article" docStyleVersion="3" docTitle="Anacroneuria" docType="key" docVersion="4" lastPageId="4" lastPageNumber="118" masterDocId="FFDDFF8E023C93790923FFAE4A4E876F" masterDocTitle="Anacroneuria Pakaraima And A. Wokomung, Two New Stonefly Species From Guyana (Plecoptera: Perlidae)" masterLastPageNumber="118" masterPageNumber="114" pageId="3" pageNumber="117" updateTime="1643545951448" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title>Anacroneuria Pakaraima And A. Wokomung, Two New Stonefly Species From Guyana (Plecoptera: Perlidae)</mods:title>
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<mods:namePart>Stark, Bill P.</mods:namePart>
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<mods:title>Illiesia</mods:title>
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<mods:date>2012</mods:date>
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<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4760603</mods:identifier>
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Provisional key for male
<taxonomicName box="[1144,1305,829,853]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="117" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[1144,1305,829,853]" italics="true" pageId="3" pageNumber="117">Anacroneuria</emphasis>
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known for
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(Male
<taxonomicName authority="Stark 1999" authorityName="Stark" authorityYear="1999" box="[974,1249,900,924]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="117" phylum="Arthropoda" rank="species" species="wapishana">
<emphasis box="[974,1121,900,924]" italics="true" pageId="3" pageNumber="117">A. wapishana</emphasis>
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unknown)
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1 Forewing patterned with a longitudinal amber band, narrow at the base and expanded around an unpigmented circular area beyond cord (
<figureCitation box="[1361,1428,1042,1067]" captionStart="Figs" captionStartId="2.[189,241,1823,1847]" captionTargetBox="[335,1249,510,1716]" captionTargetId="figure-49@2.[325,1308,464,1765]" captionTargetPageId="2" captionText="Figs. 6-13. Anacroneuria wokomung structures. 6. Head and pronotum. 7. Forewing. 8. Male sternum 9. 9. Male aedeagus, ventral. 10. Male aedeagus, lateral. 11. Male aedeagus, dorsal. 12. Egg outline. 13. Female abdominal sterna 8-9." figureDoi="http://doi.org/10.5281/zenodo.4760609" httpUri="https://zenodo.org/record/4760609/files/figure.png" pageId="3" pageNumber="117">Fig. 7</figureCitation>
), pattern obscure in some specimens.….…….…. 2
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<paragraph blockId="3.[835,1443,971,1919]" box="[835,1443,1113,1138]" pageId="3" pageNumber="117">
<keyLead box="[835,1443,1113,1138]" pageId="3" pageNumber="117">1 Forewing pattern not as above …………………. 3</keyLead>
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2 Hammer width greater than height (
<figureCitation box="[1352,1428,1149,1173]" captionStart="Figs" captionStartId="2.[189,241,1823,1847]" captionTargetBox="[335,1249,510,1716]" captionTargetId="figure-49@2.[325,1308,464,1765]" captionTargetPageId="2" captionText="Figs. 6-13. Anacroneuria wokomung structures. 6. Head and pronotum. 7. Forewing. 8. Male sternum 9. 9. Male aedeagus, ventral. 10. Male aedeagus, lateral. 11. Male aedeagus, dorsal. 12. Egg outline. 13. Female abdominal sterna 8-9." figureDoi="http://doi.org/10.5281/zenodo.4760609" httpUri="https://zenodo.org/record/4760609/files/figure.png" pageId="3" pageNumber="117">Fig. 8</figureCitation>
); aedeagal apex somewhat circular in lateral aspect (
<figureCitation box="[881,961,1220,1244]" captionStart="Figs" captionStartId="2.[189,241,1823,1847]" captionTargetBox="[335,1249,510,1716]" captionTargetId="figure-49@2.[325,1308,464,1765]" captionTargetPageId="2" captionText="Figs. 6-13. Anacroneuria wokomung structures. 6. Head and pronotum. 7. Forewing. 8. Male sternum 9. 9. Male aedeagus, ventral. 10. Male aedeagus, lateral. 11. Male aedeagus, dorsal. 12. Egg outline. 13. Female abdominal sterna 8-9." figureDoi="http://doi.org/10.5281/zenodo.4760609" httpUri="https://zenodo.org/record/4760609/files/figure.png" pageId="3" pageNumber="117">Fig. 10</figureCitation>
); dorsal keel line short and reaching apex (
<figureCitation box="[881,958,1256,1280]" captionStart="Figs" captionStartId="2.[189,241,1823,1847]" captionTargetBox="[335,1249,510,1716]" captionTargetId="figure-49@2.[325,1308,464,1765]" captionTargetPageId="2" captionText="Figs. 6-13. Anacroneuria wokomung structures. 6. Head and pronotum. 7. Forewing. 8. Male sternum 9. 9. Male aedeagus, ventral. 10. Male aedeagus, lateral. 11. Male aedeagus, dorsal. 12. Egg outline. 13. Female abdominal sterna 8-9." figureDoi="http://doi.org/10.5281/zenodo.4760609" httpUri="https://zenodo.org/record/4760609/files/figure.png" pageId="3" pageNumber="117">Fig. 11</figureCitation>
) …………………...………….
<taxonomicName authorityName="Stark" authorityYear="2012" box="[1296,1443,1256,1280]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="117" phylum="Arthropoda" rank="species" species="wokomung">
<emphasis box="[1296,1443,1256,1280]" italics="true" pageId="3" pageNumber="117">A. wokomung</emphasis>
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2 Hammer laterally compressed, height greater than width (
<figureCitation box="[1014,1076,1327,1351]" captionStart="Figs" captionStartId="1.[189,241,1326,1350]" captionTargetBox="[382,1253,304,1237]" captionTargetId="figure-49@1.[325,1308,251,1292]" captionTargetPageId="1" captionText="Figs. 1-5. Anacroneuria pakaraima structures. 1. Male sternum 9. 2. Female abdominal sterna 8-9. 3. Male aedeagus, ventral. 4. Male aedeagus, lateral. 5. Male aedeagus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4760607" httpUri="https://zenodo.org/record/4760607/files/figure.png" pageId="3" pageNumber="117">Fig. 1</figureCitation>
); aedeagal apex curved ventrad but flattened across ventroapical margin (
<figureCitation box="[1362,1430,1362,1386]" captionStart="Figs" captionStartId="1.[189,241,1326,1350]" captionTargetBox="[382,1253,304,1237]" captionTargetId="figure-49@1.[325,1308,251,1292]" captionTargetPageId="1" captionText="Figs. 1-5. Anacroneuria pakaraima structures. 1. Male sternum 9. 2. Female abdominal sterna 8-9. 3. Male aedeagus, ventral. 4. Male aedeagus, lateral. 5. Male aedeagus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4760607" httpUri="https://zenodo.org/record/4760607/files/figure.png" pageId="3" pageNumber="117">Fig. 4</figureCitation>
); dorsal keel line obscure, but weak arms diverge near tip (
<figureCitation box="[977,1042,1433,1457]" captionStart="Figs" captionStartId="1.[189,241,1326,1350]" captionTargetBox="[382,1253,304,1237]" captionTargetId="figure-49@1.[325,1308,251,1292]" captionTargetPageId="1" captionText="Figs. 1-5. Anacroneuria pakaraima structures. 1. Male sternum 9. 2. Female abdominal sterna 8-9. 3. Male aedeagus, ventral. 4. Male aedeagus, lateral. 5. Male aedeagus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4760607" httpUri="https://zenodo.org/record/4760607/files/figure.png" pageId="3" pageNumber="117">Fig. 5</figureCitation>
) ………………………
<taxonomicName authorityName="Stark" authorityYear="2012" box="[1301,1443,1433,1457]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="117" phylum="Arthropoda" rank="species" species="pakaraima">
<emphasis box="[1301,1443,1433,1457]" italics="true" pageId="3" pageNumber="117">A. pakaraima</emphasis>
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3 Apical aedeagal section with a prominent pair of membranous lobes (
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Fig.
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in
<bibRefCitation author="Stark, B. P." box="[1212,1341,1504,1528]" pageId="3" pageNumber="117" pagination="70 - 93" refId="ref2353" refString="Stark, B. P. 1999. Anacroneuria from northeastern South America (Insecta: Plecoptera: Perlidae). Proceedings of the Biological Society of Washington, 112: 70 - 93." type="journal article" year="1999">Stark, 1999</bibRefCitation>
) …….. 4
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3 Apical aedeagal section without membranous lobes (
<figureCitation box="[949,1013,1575,1599]" captionStart="Figs" captionStartId="1.[189,241,1326,1350]" captionTargetBox="[382,1253,304,1237]" captionTargetId="figure-49@1.[325,1308,251,1292]" captionTargetPageId="1" captionText="Figs. 1-5. Anacroneuria pakaraima structures. 1. Male sternum 9. 2. Female abdominal sterna 8-9. 3. Male aedeagus, ventral. 4. Male aedeagus, lateral. 5. Male aedeagus, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4760607" httpUri="https://zenodo.org/record/4760607/files/figure.png" pageId="3" pageNumber="117">
Fig.
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</figureCitation>
in
<bibRefCitation author="Stark, B. P." box="[1049,1177,1575,1599]" pageId="3" pageNumber="117" pagination="305 - 310" refId="ref2388" refString="Stark, B. P. 2000. Notes on the Anacroneuria (Plecoptera: Perlidae) of Guyana with the description of a new species. Aquatic Insects, 22: 305 - 310." type="journal article" year="2000">Stark, 2000</bibRefCitation>
) …………………….. 6
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4 Aedeagal hooks stout (Fig.
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); body of aedeagus with a dorsomesal, transverse, arcuate process (Fig.
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<bibRefCitation author="Stark, B. P." box="[1285,1435,1682,1706]" pageId="3" pageNumber="117" pagination="70 - 93" refId="ref2353" refString="Stark, B. P. 1999. Anacroneuria from northeastern South America (Insecta: Plecoptera: Perlidae). Proceedings of the Biological Society of Washington, 112: 70 - 93." type="journal article" year="1999">Stark, 1999</bibRefCitation>
) …..…………….……………….……..…
<taxonomicName authorityName="Stark" authorityYear="1999" box="[1318,1441,1717,1741]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="117" phylum="Arthropoda" rank="species" species="makushi">
<emphasis box="[1318,1441,1717,1741]" italics="true" pageId="3" pageNumber="117">A. makushi</emphasis>
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4 Aedeagal hooks relatively slender (
<figureCitation box="[1293,1372,1753,1777]" captionStart="Figs" captionStartId="2.[189,241,1823,1847]" captionTargetBox="[335,1249,510,1716]" captionTargetId="figure-49@2.[325,1308,464,1765]" captionTargetPageId="2" captionText="Figs. 6-13. Anacroneuria wokomung structures. 6. Head and pronotum. 7. Forewing. 8. Male sternum 9. 9. Male aedeagus, ventral. 10. Male aedeagus, lateral. 11. Male aedeagus, dorsal. 12. Egg outline. 13. Female abdominal sterna 8-9." figureDoi="http://doi.org/10.5281/zenodo.4760609" httpUri="https://zenodo.org/record/4760609/files/figure.png" pageId="3" pageNumber="117">Figs. 9</figureCitation>
, 45 in
<bibRefCitation author="Stark, B. P." box="[873,1027,1788,1813]" pageId="3" pageNumber="117" pagination="70 - 93" refId="ref2353" refString="Stark, B. P. 1999. Anacroneuria from northeastern South America (Insecta: Plecoptera: Perlidae). Proceedings of the Biological Society of Washington, 112: 70 - 93." type="journal article" year="1999">Stark, 1999</bibRefCitation>
); body of aedeagus without dorsomesal, transverse, arcuate process …..…… 5
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5 Apex of aedeagus gradually narrowed from hooks to tip (
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Fig.
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</figureCitation>
in
<bibRefCitation author="Stark, B. P." box="[1128,1259,1895,1919]" pageId="3" pageNumber="117" pagination="70 - 93" refId="ref2353" refString="Stark, B. P. 1999. Anacroneuria from northeastern South America (Insecta: Plecoptera: Perlidae). Proceedings of the Biological Society of Washington, 112: 70 - 93." type="journal article" year="1999">Stark, 1999</bibRefCitation>
); inner margins of hooks smooth …………………...……
<taxonomicName authorityName="Stark" authorityYear="1999" box="[689,797,225,249]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="118" phylum="Arthropoda" rank="species" species="arawak">
<emphasis box="[689,797,225,249]" italics="true" pageId="4" pageNumber="118">A. arawak</emphasis>
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5 Apex of aedeagus flared subapically (Fig.
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); inner margins of hooks irregularly scalloped …….……………………….
<taxonomicName baseAuthorityName="Banks" baseAuthorityYear="1914" box="[658,797,331,355]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="118" phylum="Arthropoda" rank="species" species="phantoma">
<emphasis box="[658,797,331,355]" italics="true" pageId="4" pageNumber="118">A. phantoma</emphasis>
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6 Dorsal aedeagal keel a basally directed triangular process (Fig.
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); R-vein of forewing conspicuously darker than others (Fig.
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) …………………….………………
<taxonomicName authorityName="Klapalek" authorityYear="1923" box="[682,796,474,497]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="4" pageNumber="118" phylum="Arthropoda" rank="species" species="pictipes">
<emphasis box="[682,796,474,497]" italics="true" pageId="4" pageNumber="118">A. pictipes</emphasis>
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6 Dorsal aedeagal keel consists of a pair of narrowly separated ridges (
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Fig.
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); R-vein similarly pigmented to other forewing veins ……………………………………………….. 7
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7 Hooks almost straight and crossing near aedeagal midlength (Fig.
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).….…..
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<emphasis box="[706,797,687,710]" italics="true" pageId="4" pageNumber="118">A. cruza</emphasis>
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7 Hooks evenly curved and crossing at base of aedeagal apex (
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Fig.
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) ….……….. 8
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8 Aedeagal apex truncate, width of aedeagal base ca. 2.6 times width of aedeagal apex (
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) ………………………………
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<emphasis box="[695,797,864,888]" italics="true" pageId="4" pageNumber="118">A. takutu</emphasis>
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8 Aedeagal apex rounded, width of aedeagal base ca. 6.5 times width of aedeagal apex (Fig.
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) ……………………..………….
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<emphasis box="[713,797,971,995]" italics="true" pageId="4" pageNumber="118">A. llana</emphasis>
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