572 lines
78 KiB
XML
572 lines
78 KiB
XML
<document id="0FA376E3353A4DCF87EBBE236C1F8452" ID-CLB-Dataset="133821" ID-DOI="10.1093/zoolinnean/zlac009" ID-GBIF-Dataset="cab69ef3-0fe3-4309-8df7-f8584c027c48" ID-ISSN="0024-4082" ID-Zenodo-Dep="7037746" ID-ZooBank="C288BAB2-A92C-4F13-B04D-D6D4510461F5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1661767826270" checkinUser="plazi" docAuthor="Smirnoff, Dimitri S., Donohoo, Samantha A. & Gosliner, Terrence M." docDate="2022" docId="03E487E4FF95FB3FEA11FB32FEA7FB45" docLanguage="en" docName="zlac009.pdf" docOrigin="Zoological Journal of the Linnean Society 196" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Trapania tatsulok Smirnoff & Donohoo & Gosliner 2022, SP. NOV." docType="treatment" docUuid="9DF25EE2-655D-485B-92A7-45B7D23B188D" docUuidSource="ZooBank" docVersion="9" lastPageNumber="286" masterDocId="FFDDFF9CFF98FB2FEB05FFF4FFD8FD72" masterDocTitle="Extra-branchial processes manifest extra diversity: systematics of the genus Trapania (Nudibranchia: Goniodorididae) and nine new species descriptions" masterLastPageNumber="313" masterPageNumber="270" pageNumber="283" updateTime="1699374017120" updateUser="ExternalLinkService">
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<mods:titleInfo id="BDFB1BB15EA3E3F6F23F3084114FD431">
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<mods:title id="F0878E74752E14D921F2D09152FF439A">Extra-branchial processes manifest extra diversity: systematics of the genus Trapania (Nudibranchia: Goniodorididae) and nine new species descriptions</mods:title>
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<mods:roleTerm id="4DA640F740E40746A791A5A6976FBFF2">Author</mods:roleTerm>
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<mods:namePart id="81BAD7D314E6C969F4FDE94081486B54">Smirnoff, Dimitri S.</mods:namePart>
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<mods:roleTerm id="F91AD77C8E10009A189CECEDB96FCB2B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="DF922CC72056542FF7E1675644AD5A4A">Donohoo, Samantha A.</mods:namePart>
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</mods:name>
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<mods:name id="B00072DC110400B229255579FC7035BC" type="personal">
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<mods:role id="33BD6CF480937EBC42463ABFDA7D6640">
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<mods:roleTerm id="1E72A5D64CC6676D3C1E92C099CD5414">Author</mods:roleTerm>
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<mods:namePart id="BFFB052118613EE95673CB4DC7EB3ED4">Gosliner, Terrence M.</mods:namePart>
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<mods:typeOfResource id="712E4479D36B774D3860C279789BB2E3">text</mods:typeOfResource>
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<mods:title id="7636BDE6DBD999478F4E61FB76050FFF">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="771CACE516C1AB699A719BDFC4121D6A">
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<mods:date id="C9EACE39C077A54D29975C4CF9440E0B">2022</mods:date>
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<mods:detail id="B813AA2BF024A10E7487D0CB71EE0CB5" type="volume">
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<mods:number id="DD5A3EA9EAD04ECE4479A585E0BD1819">196</mods:number>
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<mods:classification id="ABDA53E3BAE9B9CD0504A362C8038C3D">journal article</mods:classification>
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<mods:identifier id="5C26530C8732C094A9B05273A48CABE7" type="DOI">10.1093/zoolinnean/zlac009</mods:identifier>
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<mods:identifier id="B0013E519CACF33B8FB0E74BC265B0D7" type="ISSN">0024-4082</mods:identifier>
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<treatment id="03E487E4FF95FB3FEA11FB32FEA7FB45" ID-DOI="http://doi.org/10.5281/zenodo.7043800" ID-GBIF-Taxon="199901981" ID-Zenodo-Dep="7043800" LSID="urn:lsid:zoobank.org:act:9DF25EE2-655D-485B-92A7-45B7D23B188D" httpUri="http://treatment.plazi.org/id/03E487E4FF95FB3FEA11FB32FEA7FB45" lastPageId="16" lastPageNumber="286" pageId="13" pageNumber="283">
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<subSubSection id="C3576579FF95FB22EA11FB32FDADF9AC" box="[276,629,1222,1246]" pageId="13" pageNumber="283" type="nomenclature">
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<paragraph id="8BF236F2FF95FB22EA11FB32FDADF9AC" blockId="13.[276,629,1222,1246]" box="[276,629,1222,1246]" pageId="13" pageNumber="283">
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<heading id="D0BA819EFF95FB22EA11FB32FDADF9AC" box="[276,629,1222,1246]" centered="true" fontSize="9" level="2" pageId="13" pageNumber="283" reason="2">
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<taxonomicName id="4C4D4D71FF95FB22EA11FB32FDD3F9AF" ID-CoL="992SF" authority="Smirnoff & Donohoo & Gosliner, 2022" authorityName="Smirnoff & Donohoo & Gosliner" authorityYear="2022" box="[276,523,1222,1245]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="13" pageNumber="285" phylum="Mollusca" rank="species" species="tatsulok" status="sp. nov.">TRAPANIA TATSULOK</taxonomicName>
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<emphasis id="B939EAE0FF95FB22E917FB3DFDADF9AC" bold="true" box="[530,629,1222,1246]" pageId="13" pageNumber="283">
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<taxonomicNameLabel id="A20A579BFF95FB22E917FB3DFDADF9AC" box="[530,629,1222,1246]" pageId="13" pageNumber="283" rank="species">SP. NOV.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3576579FF95FB22EA6FFB19FE8BF81A" pageId="13" pageNumber="283" type="description">
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<paragraph id="8BF236F2FF95FB22EA6FFB19FDC7F874" blockId="13.[362,543,1261,1286]" box="[362,543,1261,1286]" pageId="13" pageNumber="283">
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(
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<figureCitation id="13762A77FF95FB22EA76FB19FE17F874" box="[371,463,1261,1286]" captionStart="Figure 2" captionStartId="12.[163,241,1701,1723]" captionTargetBox="[259,1346,197,1660]" captionTargetId="figure-27@12.[257,1348,195,1661]" captionTargetPageId="12" captionText="Figure 2. Living animals. A, Trapania japonica (Baba, 1935), CASIZ 221116, Mabini, Batangas, Philippines, photo by Peri Paleracio. B, Trapania tatsulok sp. nov., holotype, NMP 041333 (formerly CASIZ 200530), Calatagan, Batangas, Philippines. C, Trapania kahel sp. nov., holotype, NMP 041330 (formerly CASIZ 186131), Mabini, Batangas, Philippines. D, Trapania lemanioides sp. nov., holotype, NMP 041335 (formerly CASIZ 208433), Puerto Galera, Mindoro Oriental, Philippines. E, Trapania lemanioides sp. nov., paratype, CASIZ 221979, Romblon, Philippines. F, Trapania aurata Rudman, 1987, CASIZ 186205, Mabini, Batangas, Philippines. G, Trapania reticulata Rudman, 1987, CASIZ 191431, Madang, Papua New Guinea, photo by Vanessa Knutson. H, Trapania tigger sp. nov., holotype, NMP 041328 (formerly CASIZ 180412), Mabini, Batangas, Philippines, photo by Alicia McKowen. Photo by T. Gosliner unless indicated." figureDoi="http://doi.org/10.5281/zenodo.7037751" httpUri="https://zenodo.org/record/7037751/files/figure.png" pageId="13" pageNumber="283">FIGS 2B</figureCitation>
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,
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<figureCitation id="13762A77FF95FB22EAD9FB19FE23F874" box="[476,507,1261,1286]" captionStart="Figure 4" captionStartId="14.[163,241,1790,1812]" captionTargetBox="[258,1346,197,1747]" captionTargetId="figure-27@14.[255,1349,194,1750]" captionTargetPageId="14" captionText="Figure 4. Reproductive systems. A, Trapania japonica (Baba, 1935), CASIZ 221116, scale = 0.50 mm. B, Trapania tatsulok sp. nov., holotype, NMP 041333 (formerly CASIZ 200530), scale = 0.33 mm. C, Trapania kahel sp. nov., holotype, NMP 041330 (formerly CASIZ 186131), scale = 0.28 mm. D, Trapania lemanioides sp. nov., holotype, NMP 041335 (formerly" figureDoi="http://doi.org/10.5281/zenodo.7037757" httpUri="https://zenodo.org/record/7037757/files/figure.png" pageId="13" pageNumber="283">4B</figureCitation>
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,
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<figureCitation id="13762A77FF95FB22E90CFB19FDCFF874" box="[521,535,1261,1286]" captionStart="Figure 5" captionStartId="16.[163,241,1111,1133]" captionTargetBox="[163,1443,195,1071]" captionTargetId="figure-314@16.[163,1443,195,1071]" captionTargetPageId="16" captionText="Figure 5. Trapania tatsulok sp. nov. Scanning electron micrographs of buccal armature of holotype NMP 041333 (formerly CASIZ 200530). A, entire jaw plate. B, detail of armature of jaws. C, majority of radula. D, individual teeth." figureDoi="http://doi.org/10.5281/zenodo.7037759" httpUri="https://zenodo.org/record/7037759/files/figure.png" pageId="13" pageNumber="283">5</figureCitation>
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)
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</paragraph>
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<paragraph id="8BF236F2FF95FB22EB94FAE2FE8BF81A" blockId="13.[145,762,1301,1384]" pageId="13" pageNumber="283">
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<emphasis id="B939EAE0FF95FB22EB94FAE2FE06F859" box="[145,478,1301,1323]" italics="true" pageId="13" pageNumber="283">Z o o b a n k r e g i s t r a t i o n:</emphasis>
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u r n: l s i d: z o o b a n k. o r g: a c t:
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<uuid id="FFEB0C27FF95FB22EA12FAC0FE97F81A" pageId="13" pageNumber="283">
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9 D F
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E 2 - 6 5 5 D - 4 8 5 B - 9 2 A 7 - 45B7D23B188D
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3576579FF95FB22EB94FA76FDFDFBD5" pageId="13" pageNumber="283" type="reference_group">
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<paragraph id="8BF236F2FF95FB22EB94FA76FEECF8C4" blockId="13.[145,761,1409,1523]" pageId="13" pageNumber="283">
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<taxonomicName id="4C4D4D71FF95FB22EB94FA76FEF9F8E5" box="[145,289,1410,1431]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="13" pageNumber="283" phylum="Mollusca" rank="species" species="undetermined">
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<emphasis id="B939EAE0FF95FB22EB94FA76FF24F8E5" box="[145,252,1410,1431]" italics="true" pageId="13" pageNumber="283">Trapania</emphasis>
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||
sp.
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</taxonomicName>
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||
15
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||
<treatmentCitation id="0AEC10E3FF95FB22EA42FA75FDB8F8E5" author="Gosliner TM & Behrens DW & Valdes A" box="[327,608,1409,1431]" page="145" pageId="13" pageNumber="283" year="2015">
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<bibRefCitation id="EFDC4B03FF95FB22EA42FA75FDB8F8E5" author="Gosliner TM & Behrens DW & Valdes A" box="[327,608,1409,1431]" pageId="13" pageNumber="283" refId="ref28995" refString="Gosliner TM, Behrens DW, Valdes A. 2015. Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna. Jacksonville: New World Publications, Inc." type="book" year="2015">
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||
Gosliner
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<emphasis id="B939EAE0FF95FB22EAAAFA76FE3DF8E4" box="[431,485,1409,1431]" italics="true" pageId="13" pageNumber="283">et al.</emphasis>
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, 2015: 145
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</bibRefCitation>
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||
</treatmentCitation>
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||
, middle-right photograph.
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</treatmentCitationGroup>
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</paragraph>
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||
<paragraph id="8BF236F2FF95FB22EB94FA4BFEECF881" blockId="13.[145,761,1409,1523]" pageId="13" pageNumber="283">
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<treatmentCitationGroup id="AB5D11DCFF95FB22EB94FA4BFEECF881" pageId="13" pageNumber="283">
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<taxonomicName id="4C4D4D71FF95FB22EB94FA4BFEFFF8A6" box="[145,295,1471,1492]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="13" pageNumber="283" phylum="Mollusca" rank="species" species="undetermined">
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||
<emphasis id="B939EAE0FF95FB22EB94FA4BFF26F8A6" box="[145,254,1471,1492]" italics="true" pageId="13" pageNumber="283">Trapania</emphasis>
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||
sp.
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||
</taxonomicName>
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||
17
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||
<treatmentCitation id="0AEC10E3FF95FB22EA56FA4BFDB3F8A6" author="Gosliner TM & Valdes A & Behrens DW" box="[339,619,1470,1493]" page="63" pageId="13" pageNumber="283" year="2018">
|
||
<bibRefCitation id="EFDC4B03FF95FB22EA56FA4BFDB3F8A6" author="Gosliner TM & Valdes A & Behrens DW" box="[339,619,1470,1493]" pageId="13" pageNumber="283" refId="ref29030" refString="Gosliner TM, Valdes A, Behrens DW. 2018. Indo-Pacific nudibranchs and sea slugs: a field guide to the world's most diverse fauna. Jacksonville: New World Publications, Inc." type="book" year="2018">
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||
Gosliner
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<emphasis id="B939EAE0FF95FB22EABBFA4BFE21F8A6" box="[446,505,1471,1492]" italics="true" pageId="13" pageNumber="283">et al.</emphasis>
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, 2018: 63
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</bibRefCitation>
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||
</treatmentCitation>
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||
, bottom-left photograph.
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||
</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BF236F2FF95FB22EB94F9E3FDFDFBD5" blockId="13.[145,761,1559,1703]" pageId="13" pageNumber="283">
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<emphasis id="B939EAE0FF95FB22EB94F9E3FEF7FB5E" box="[145,303,1559,1580]" italics="true" pageId="13" pageNumber="283">Typematerial:</emphasis>
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||
<materialsCitation id="3B253CAFFF95FB22EA39F9E3FDFDFBD5" ID-GBIF-Occurrence="3897192330" collectingDate="2014-05-19" collectionCode="NMP" collectorName="Shayle Matstuda." country="Philippines" county="Luzon Island" latitude="13.92816" location="Verde Island Passage" longLatPrecision="5" longitude="120.6109" municipality="Calatagan" pageId="13" pageNumber="283" specimenCode="NMP041333" specimenCount="1" stateProvince="Batangas" typeStatus="holotype">
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<emphasis id="B939EAE0FF95FB22EA39F9E3FE7EFB5E" box="[316,422,1559,1580]" italics="true" pageId="13" pageNumber="283">
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<typeStatus id="54F68850FF95FB22EA39F9E3FE78FB5E" box="[316,416,1559,1580]" pageId="13" pageNumber="283" type="holotype">Holotype</typeStatus>
|
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:
|
||
</emphasis>
|
||
<specimenCode id="DBEB9E89FF95FB22EAADF9E3FDE1FB5E" box="[424,569,1559,1580]" collectionCode="NMP" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" pageId="13" pageNumber="283">NMP041333</specimenCode>
|
||
,originallyCASIZ 200530,
|
||
<specimenCount id="9D4BFD7BFF95FB22EBF5F9C2FE52FB39" box="[240,394,1589,1611]" pageId="13" pageNumber="283" type="generic">one specimen</specimenCount>
|
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, dissected and sequenced,
|
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<location id="8E926029FF95FB22E9B2F9C1FEE7FB1B" LSID="urn:lsid:plazi:treatment:03E487E4FF95FB3FEA11FB32FEA7FB45:8E926029FF95FB22E9B2F9C1FEE7FB1B" country="Philippines" county="Luzon Island" latitude="13.92816" longLatPrecision="5" longitude="120.6109" municipality="Calatagan" name="Verde Island Passage" pageId="13" pageNumber="283" stateProvince="Batangas">
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<location id="8E926029FF95FB22E9B2F9C1FF02FB18" LSID="urn:lsid:plazi:treatment:03E487E4FF95FB3FEA11FB32FEA7FB45:8E926029FF95FB22E9B2F9C1FF02FB18" country="Philippines" county="Luzon Island" latitude="13.92816" longLatPrecision="5" longitude="120.6109" municipality="Calatagan" name="Verde Island" pageId="13" pageNumber="283" stateProvince="Batangas">Verde Island</location>
|
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Passage
|
||
</location>
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(
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<geoCoordinate id="EE795035FF95FB22EADAF9A0FDBEFB18" box="[479,614,1620,1642]" degrees="120.6109" direction="east" orientation="longitude" pageId="13" pageNumber="283" precision="5" value="120.6109">120.6109°E</geoCoordinate>
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),
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<collectingMunicipality id="6B96AC88FF95FB22E97FF9A0FD2CFB18" box="[634,756,1620,1642]" pageId="13" pageNumber="283">Calatagan</collectingMunicipality>
|
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,
|
||
<collectingRegion id="4989F810FF95FB22EB94F987FEB0FBFA" box="[145,360,1650,1672]" country="Philippines" name="Batangas" pageId="13" pageNumber="283">Batangas Province</collectingRegion>
|
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,
|
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<collectingCounty id="62934E7EFF95FB22EA71F987FDD3FBFA" box="[372,523,1650,1672]" pageId="13" pageNumber="283">Luzon Island</collectingCounty>
|
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,
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<collectingCountry id="F35A7662FF95FB22E912F986FD42FBFA" box="[535,666,1650,1672]" name="Philippines" pageId="13" pageNumber="283">Philippines</collectingCountry>
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,
|
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<date id="FFF31032FF95FB22E9A1F987FF11FBD4" pageId="13" pageNumber="283" value="2014-05-19">
|
||
<collectingDate id="EFB7E9DAFF95FB22E9A1F987FF11FBD4" pageId="13" pageNumber="283" value="2014-05-19">19 May 2014</collectingDate>
|
||
</date>
|
||
, collected by
|
||
<collectorName id="26B85324FF95FB22EA58F965FDFDFBD5" box="[349,549,1681,1703]" pageId="13" pageNumber="283">Shayle Matstuda.</collectorName>
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</materialsCitation>
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</paragraph>
|
||
</subSubSection>
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<subSubSection id="C3576579FF95FB3FEB94F925FEA7FB45" lastPageId="16" lastPageNumber="286" pageId="13" pageNumber="283" type="materials_examined">
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<paragraph id="8BF236F2FF95FB22EB94F925FEB6FA51" blockId="13.[145,760,1744,1827]" pageId="13" pageNumber="283">
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<materialsCitation id="3B253CAFFF95FB22EB94F925FEB3FA51" ID-GBIF-Occurrence="3897192307" country="Philippines" county="Luzon Island" latitude="13.92816" location="Verde Island Passage" longLatPrecision="5" longitude="120.6109" municipality="Calatagan" pageId="13" pageNumber="283" specimenCount="1" stateProvince="Batangas" typeStatus="holotype">
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<typeStatus id="54F68850FF95FB22EB94F925FF12FB94" box="[145,202,1745,1766]" pageId="13" pageNumber="283">Type</typeStatus>
|
||
locality:
|
||
</emphasis>
|
||
<location id="8E926029FF95FB22EA42F924FD8EFB97" LSID="urn:lsid:plazi:treatment:03E487E4FF95FB3FEA11FB32FEA7FB45:8E926029FF95FB22EA42F924FD8EFB97" box="[327,598,1744,1766]" country="Philippines" county="Luzon Island" latitude="13.92816" longLatPrecision="5" longitude="120.6109" municipality="Calatagan" name="Verde Island Passage" pageId="13" pageNumber="283" stateProvince="Batangas">
|
||
<location id="8E926029FF95FB22EA42F924FE3FFB94" LSID="urn:lsid:plazi:treatment:03E487E4FF95FB3FEA11FB32FEA7FB45:8E926029FF95FB22EA42F924FE3FFB94" box="[327,487,1744,1766]" country="Philippines" county="Luzon Island" latitude="13.92816" longLatPrecision="5" longitude="120.6109" municipality="Calatagan" name="Verde Island" pageId="13" pageNumber="283" stateProvince="Batangas">Verde Island</location>
|
||
Passage
|
||
</location>
|
||
(
|
||
<geoCoordinate id="EE795035FF95FB22E96CF924FD2FFB94" box="[617,759,1744,1766]" degrees="13.92816" direction="north" orientation="latitude" pageId="13" pageNumber="283" precision="1" value="13.92816">13.92816ºN</geoCoordinate>
|
||
<geoCoordinate id="EE795035FF95FB22EB94F91AFEC4FA76" box="[145,284,1774,1796]" degrees="120.6109" direction="east" orientation="longitude" pageId="13" pageNumber="283" precision="5" value="120.6109">120.6109°E</geoCoordinate>
|
||
),
|
||
<collectingMunicipality id="6B96AC88FF95FB22EA37F91BFE68FA77" box="[306,432,1775,1797]" pageId="13" pageNumber="283">Calatagan</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="4989F810FF95FB22EABBF91BFD7AFA77" box="[446,674,1775,1797]" country="Philippines" name="Batangas" pageId="13" pageNumber="283">Batangas Province</collectingRegion>
|
||
,
|
||
<collectingCounty id="62934E7EFF95FB22E9AAF91BFF03FA51" pageId="13" pageNumber="283">Luzon Island</collectingCounty>
|
||
,
|
||
<collectingCountry id="F35A7662FF95FB22EBE3F8F9FEB3FA51" box="[230,363,1805,1827]" name="Philippines" pageId="13" pageNumber="283">Philippines</collectingCountry>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF236F2FF95FB22EB94F8B3FE10FA09" blockId="13.[145,761,1862,1915]" pageId="13" pageNumber="283">
|
||
<emphasis id="B939EAE0FF95FB22EB94F8B3FE23FA2E" box="[145,507,1862,1884]" italics="true" pageId="13" pageNumber="283">G e o g r a p h i c a l d i s t r i b u t i o n:</emphasis>
|
||
K n o w n o n l y f r o m Calatagan, the
|
||
<collectingCountry id="F35A7662FF95FB22EA45F891FE1DFA09" box="[320,453,1893,1915]" name="Philippines" pageId="13" pageNumber="283">Philippines</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF236F2FF95FB22E82CFCE7FA59FE14" blockId="13.[809,1425,787,870]" pageId="13" pageNumber="283">
|
||
<emphasis id="B939EAE0FF95FB22E82CFCE7FBC7FE5A" box="[809,1055,787,808]" italics="true" pageId="13" pageNumber="283">Etymology: Tatsulok</emphasis>
|
||
is the Tagalog word for triangle and it refers to the black equilateral triangle found on the dorsum of this species. It is a noun in apposition.
|
||
</paragraph>
|
||
<paragraph id="8BF236F2FF95FB22E82CFC7DFC47FB0D" blockId="13.[809,1426,905,1663]" pageId="13" pageNumber="283">
|
||
<emphasis id="B939EAE0FF95FB22E82CFC7DFBFEFEEC" box="[809,1062,905,926]" italics="true" pageId="13" pageNumber="283">External morphology:</emphasis>
|
||
The living animal (
|
||
<figureCitation id="13762A77FF95FB22EE12FC7DFAB4FEEC" box="[1303,1388,905,927]" captionStart="Figure 2" captionStartId="12.[163,241,1701,1723]" captionTargetBox="[259,1346,197,1660]" captionTargetId="figure-27@12.[257,1348,195,1661]" captionTargetPageId="12" captionText="Figure 2. Living animals. A, Trapania japonica (Baba, 1935), CASIZ 221116, Mabini, Batangas, Philippines, photo by Peri Paleracio. B, Trapania tatsulok sp. nov., holotype, NMP 041333 (formerly CASIZ 200530), Calatagan, Batangas, Philippines. C, Trapania kahel sp. nov., holotype, NMP 041330 (formerly CASIZ 186131), Mabini, Batangas, Philippines. D, Trapania lemanioides sp. nov., holotype, NMP 041335 (formerly CASIZ 208433), Puerto Galera, Mindoro Oriental, Philippines. E, Trapania lemanioides sp. nov., paratype, CASIZ 221979, Romblon, Philippines. F, Trapania aurata Rudman, 1987, CASIZ 186205, Mabini, Batangas, Philippines. G, Trapania reticulata Rudman, 1987, CASIZ 191431, Madang, Papua New Guinea, photo by Vanessa Knutson. H, Trapania tigger sp. nov., holotype, NMP 041328 (formerly CASIZ 180412), Mabini, Batangas, Philippines, photo by Alicia McKowen. Photo by T. Gosliner unless indicated." figureDoi="http://doi.org/10.5281/zenodo.7037751" httpUri="https://zenodo.org/record/7037751/files/figure.png" pageId="13" pageNumber="283">Fig. 2B</figureCitation>
|
||
) is
|
||
<quantity id="4CB59B17FF95FB22E82FFC5CFCB6FECF" box="[810,878,936,958]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="283" unit="mm" value="5.0">5 mm</quantity>
|
||
. The body coloration is opaque white with dark orange pigmentation on the outer three quarters of the extra-rhinophoral and extra-branchial appendages. There is a gradation of colour such that the basal portion is more yellow orange. The extra-rhinophoral and extra-branchial appendages are somewhat curved and cylindrical with a uniform diameter throughout. The conical, perfoliate rhinophores have five to six lamellae and are dark maroon/brown except for an opaque-white base. The oral tentacles are identical to the rhinophore coloration, with black spanning between the oral tentacles colouring the anterior margin of the head with a semicircle protruding posteriorly at the medial line. The anterior margins of the foot extend laterally as elongate appendages and are more translucent than the rest of the body. This specimen has a distinct black, equilateral triangle on its dorsum centred on the body midline, midway between the gill plume and rhinophores, with its tip pointing anteriorly. There are three other black markings adorning each side of the lateral flanks, starting at the middle of the body and extending to the posterior extension of the foot. There is another black spot situated medially, posterior to the gill plume.
|
||
</paragraph>
|
||
<paragraph id="8BF236F2FF95FB20E82CF959FD08FFD4" blockId="13.[809,1425,1709,1915]" lastBlockId="15.[145,761,197,679]" lastPageId="15" lastPageNumber="285" pageId="13" pageNumber="283">
|
||
<emphasis id="B939EAE0FF95FB22E82CF959FC11FBB1" box="[809,969,1709,1731]" italics="true" pageId="13" pageNumber="283">Buccal mass:</emphasis>
|
||
The buccal mass is muscular with a moderately developed buccal pump on the dorsal surface. Inside the anterior portion of the mass is a pair of well-developed jaws (
|
||
<figureCitation id="13762A77FF95FB22EF77F8FDFB1CFA6D" box="[1138,1220,1801,1823]" captionStart="Figure 5" captionStartId="16.[163,241,1111,1133]" captionTargetBox="[163,1443,195,1071]" captionTargetId="figure-314@16.[163,1443,195,1071]" captionTargetPageId="16" captionText="Figure 5. Trapania tatsulok sp. nov. Scanning electron micrographs of buccal armature of holotype NMP 041333 (formerly CASIZ 200530). A, entire jaw plate. B, detail of armature of jaws. C, majority of radula. D, individual teeth." figureDoi="http://doi.org/10.5281/zenodo.7037759" httpUri="https://zenodo.org/record/7037759/files/figure.png" pageId="13" pageNumber="283">Fig. 5A</figureCitation>
|
||
). The jaw surface is covered with small semi-spheres, each covered with numerous short and blunt spikes resembling a medieval flail or mace (
|
||
<figureCitation id="13762A77FF95FB22EF3FF891FB55FA09" box="[1082,1165,1893,1915]" captionStart="Figure 5" captionStartId="16.[163,241,1111,1133]" captionTargetBox="[163,1443,195,1071]" captionTargetId="figure-314@16.[163,1443,195,1071]" captionTargetPageId="16" captionText="Figure 5. Trapania tatsulok sp. nov. Scanning electron micrographs of buccal armature of holotype NMP 041333 (formerly CASIZ 200530). A, entire jaw plate. B, detail of armature of jaws. C, majority of radula. D, individual teeth." figureDoi="http://doi.org/10.5281/zenodo.7037759" httpUri="https://zenodo.org/record/7037759/files/figure.png" pageId="13" pageNumber="283">Fig. 5B</figureCitation>
|
||
). The radular formula of the
|
||
<typeStatus id="54F68850FF97FB20EBDFFF31FEE4FDA9" box="[218,316,197,219]" pageId="15" pageNumber="285" type="holotype">holotype</typeStatus>
|
||
is 25 × 1.0.1 (
|
||
<figureCitation id="13762A77FF97FB20EADAFF31FDE9FDA9" box="[479,561,197,219]" captionStart="Figure 5" captionStartId="16.[163,241,1111,1133]" captionTargetBox="[163,1443,195,1071]" captionTargetId="figure-314@16.[163,1443,195,1071]" captionTargetPageId="16" captionText="Figure 5. Trapania tatsulok sp. nov. Scanning electron micrographs of buccal armature of holotype NMP 041333 (formerly CASIZ 200530). A, entire jaw plate. B, detail of armature of jaws. C, majority of radula. D, individual teeth." figureDoi="http://doi.org/10.5281/zenodo.7037759" httpUri="https://zenodo.org/record/7037759/files/figure.png" pageId="15" pageNumber="285">Fig. 5C</figureCitation>
|
||
) in the specimen examined and the radula was fragmented into three pieces. The older teeth are much smaller than the newer ones and lack denticles on the outer side of the radular tooth (
|
||
<figureCitation id="13762A77FF97FB20EBDCFEB4FEF0FC24" box="[217,296,320,342]" captionStart="Figure 5" captionStartId="16.[163,241,1111,1133]" captionTargetBox="[163,1443,195,1071]" captionTargetId="figure-314@16.[163,1443,195,1071]" captionTargetPageId="16" captionText="Figure 5. Trapania tatsulok sp. nov. Scanning electron micrographs of buccal armature of holotype NMP 041333 (formerly CASIZ 200530). A, entire jaw plate. B, detail of armature of jaws. C, majority of radula. D, individual teeth." figureDoi="http://doi.org/10.5281/zenodo.7037759" httpUri="https://zenodo.org/record/7037759/files/figure.png" pageId="15" pageNumber="285">Fig. 5C</figureCitation>
|
||
). The teeth bear numerous denticles with the smallest ones being found on the inner edge of the tooth. In the newer, larger teeth, which were separated from the main part of the radula during preparation, there are approximately five to seven denticles on the inner side of the larger primary cusp with the ones nearest the primary cusp being the largest (almost the same size as the primary cusp) and a variable number of denticles on the outer side of the cusp with some teeth entirely lacking an outer denticle. There may be as few as three larger denticles or as many as eight smaller denticles, depending on the tooth (
|
||
<figureCitation id="13762A77FF97FB20E96AFD65FD18FFD5" box="[623,704,657,679]" captionStart="Figure 5" captionStartId="16.[163,241,1111,1133]" captionTargetBox="[163,1443,195,1071]" captionTargetId="figure-314@16.[163,1443,195,1071]" captionTargetPageId="16" captionText="Figure 5. Trapania tatsulok sp. nov. Scanning electron micrographs of buccal armature of holotype NMP 041333 (formerly CASIZ 200530). A, entire jaw plate. B, detail of armature of jaws. C, majority of radula. D, individual teeth." figureDoi="http://doi.org/10.5281/zenodo.7037759" httpUri="https://zenodo.org/record/7037759/files/figure.png" pageId="15" pageNumber="285">Fig. 5D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF32667AFF96FB21EBA6F90AFA7AFA3D" ID-DOI="http://doi.org/10.5281/zenodo.7037757" ID-Zenodo-Dep="7037757" httpUri="https://zenodo.org/record/7037757/files/figure.png" pageId="14" pageNumber="284" startId="14.[163,241,1790,1812]" targetBox="[258,1346,197,1747]" targetPageId="14">
|
||
<paragraph id="8BF236F2FF96FB21EBA6F90AFA7AFA3D" blockId="14.[163,1442,1790,1871]" pageId="14" pageNumber="284">
|
||
<emphasis id="B939EAE0FF96FB21EBA6F90AFED3FA61" bold="true" box="[163,267,1790,1812]" pageId="14" pageNumber="284">Figure 4.</emphasis>
|
||
Reproductive systems. A,
|
||
<taxonomicName id="4C4D4D71FF96FB21E925F90AFCB3FA66" authority="(Baba, 1935)" authorityName="Rudman" authorityYear="1987" baseAuthorityName="Baba" baseAuthorityYear="1935" box="[544,875,1790,1812]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="14" pageNumber="284" phylum="Mollusca" rank="species" species="japonica">
|
||
<emphasis id="B939EAE0FF96FB21E925F90AFD39FA66" box="[544,737,1790,1812]" italics="true" pageId="14" pageNumber="284">Trapania japonica</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDC4B03FF96FB21E9EAF90AFCBAFA66" author="Baba K" box="[751,866,1790,1812]" pageId="14" pageNumber="284" pagination="331 - 360" refId="ref28201" refString="Baba K. 1935. Nudibranchia of Mutsu Bay. Science Reports of Tohoku University 10: 331 - 360." type="journal article" year="1935">Baba, 1935</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, CASIZ 221116, scale = 0.50 mm. B,
|
||
<taxonomicName id="4C4D4D71FF96FB21EFE3F90AFA7AFA66" authority="Smirnoff & Donohoo & Gosliner, 2022" authorityName="Smirnoff & Donohoo & Gosliner" authorityYear="2022" box="[1254,1442,1790,1812]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="14" pageNumber="284" phylum="Mollusca" rank="species" species="tatsulok" status="sp. nov.">
|
||
<emphasis id="B939EAE0FF96FB21EFE3F90AFA7AFA66" box="[1254,1442,1790,1812]" italics="true" pageId="14" pageNumber="284">Trapania tatsulok</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20A579BFF96FB21EBA6F8E8FF2DFA40" box="[163,245,1820,1842]" pageId="14" pageNumber="284" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, NMP 041333 (formerly CASIZ 200530), scale = 0.33 mm. C,
|
||
<taxonomicName id="4C4D4D71FF96FB21E8F2F8E8FB46FA40" authority="Smirnoff & Donohoo & Gosliner, 2022" authorityName="Smirnoff & Donohoo & Gosliner" authorityYear="2022" box="[1015,1182,1820,1842]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="14" pageNumber="284" phylum="Mollusca" rank="species" species="kahel" status="sp. nov.">
|
||
<emphasis id="B939EAE0FF96FB21E8F2F8E8FB46FA40" box="[1015,1182,1820,1842]" italics="true" pageId="14" pageNumber="284">Trapania kahel</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20A579BFF96FB21EFA0F8E8FB2FFA40" box="[1189,1271,1820,1842]" pageId="14" pageNumber="284" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, NMP 041330 (formerly CASIZ 186131), scale = 0.28 mm. D,
|
||
<taxonomicName id="4C4D4D71FF96FB21E9F5F8CDFC06FA3D" authority="Smirnoff & Donohoo & Gosliner, 2022" authorityName="Smirnoff & Donohoo & Gosliner" authorityYear="2022" box="[752,990,1849,1871]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="14" pageNumber="284" phylum="Mollusca" rank="species" species="lemanioides" status="sp. nov.">
|
||
<emphasis id="B939EAE0FF96FB21E9F5F8CDFC06FA3D" box="[752,990,1849,1871]" italics="true" pageId="14" pageNumber="284">Trapania lemanioides</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20A579BFF96FB21E8E1F8CDFBEEFA3D" box="[996,1078,1849,1871]" pageId="14" pageNumber="284" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, NMP 041335 (formerly
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF236F2FF97FB20EB94FD21FDADF861" blockId="15.[145,762,725,1299]" pageId="15" pageNumber="285">
|
||
<emphasis id="B939EAE0FF97FB20EB94FD21FEA4FF99" box="[145,380,725,747]" italics="true" pageId="15" pageNumber="285">Reproductive system:</emphasis>
|
||
The mature reproductive system is triaulic (
|
||
<figureCitation id="13762A77FF97FB20EA0FFD00FE81FE7B" box="[266,345,756,778]" captionStart="Figure 4" captionStartId="14.[163,241,1790,1812]" captionTargetBox="[258,1346,197,1747]" captionTargetId="figure-27@14.[255,1349,194,1750]" captionTargetPageId="14" captionText="Figure 4. Reproductive systems. A, Trapania japonica (Baba, 1935), CASIZ 221116, scale = 0.50 mm. B, Trapania tatsulok sp. nov., holotype, NMP 041333 (formerly CASIZ 200530), scale = 0.33 mm. C, Trapania kahel sp. nov., holotype, NMP 041330 (formerly CASIZ 186131), scale = 0.28 mm. D, Trapania lemanioides sp. nov., holotype, NMP 041335 (formerly" figureDoi="http://doi.org/10.5281/zenodo.7037757" httpUri="https://zenodo.org/record/7037757/files/figure.png" pageId="15" pageNumber="285">Fig. 4B</figureCitation>
|
||
). The pre-ampullary is narrow at the distal end where it connects to the ovotestis. It then enters the saccate, elongate ampulla near its distal end. The ampulla narrows to the point where it divides into the short oviduct and the elongate vas deferens. The oviduct enters the female gland mass. The vas deferens widens into a curved prostatic portion that narrows into a short ejaculatory segment that is contiguous with the relatively narrow penial sac.The penial sac exits adjacent to the narrow vagina. The vagina is narrow throughout its length and joins the almost spherical bursa copulatrix adjacent to the long, narrow receptaculum duct. The receptaculum duct joins the spherical receptaculum seminist adjacent to the moderately long uterine duct, which enters the female gland mass. The bursa and receptaculum are roughly the same size. The female gland mass is composed of the large mucous gland and the smaller albumen and membrane glands.
|
||
</paragraph>
|
||
<paragraph id="8BF236F2FF97FB20EB94FAC0FB03FF39" blockId="15.[145,762,1332,1691]" lastBlockId="15.[807,1426,197,1691]" pageId="15" pageNumber="285">
|
||
<emphasis id="B939EAE0FF97FB20EB94FAC0FF27F83B" box="[145,255,1332,1353]" italics="true" pageId="15" pageNumber="285">Remarks:</emphasis>
|
||
Our molecular phylogeny, the 16S
|
||
<collectionCode id="ED5CAE37FF97FB20E9ABFAC0FD21F83B" box="[686,761,1332,1353]" pageId="15" pageNumber="285">ABGD</collectionCode>
|
||
analysis and the bPTP analysis reveal that
|
||
<taxonomicName id="4C4D4D71FF97FB20E989FAA7FF28F8F4" authorityName="Smirnoff & Donohoo & Gosliner" authorityYear="2022" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="tatsulok">
|
||
<emphasis id="B939EAE0FF97FB20E989FAA7FF28F8F4" italics="true" pageId="15" pageNumber="285">Trapania tatsulok</emphasis>
|
||
</taxonomicName>
|
||
is a distinct species of
|
||
<taxonomicName id="4C4D4D71FF97FB20E901FA86FDAAF8F5" authorityName="Pruvot-Fol" authorityYear="1931" box="[516,626,1394,1415]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B939EAE0FF97FB20E901FA86FDAAF8F5" box="[516,626,1394,1415]" italics="true" pageId="15" pageNumber="285">Trapania</emphasis>
|
||
</taxonomicName>
|
||
and can be distinguished by its unique colour pattern of white pigment on the body with black pigment on the oral tentacles, anterior margin of the head, on the rhinophores, medially in front and behind the gill and laterally on the sides of the body. This species is a member of a clade that includes
|
||
<taxonomicName id="4C4D4D71FF97FB20E901F9DEFDB1FB4D" authorityName="Gosliner & Fahey" authorityYear="2008" box="[516,617,1578,1599]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="scurra">
|
||
<emphasis id="B939EAE0FF97FB20E901F9DEFDB1FB4D" box="[516,617,1578,1599]" italics="true" pageId="15" pageNumber="285">T. scurra</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4D4D71FF97FB20E973F9DEFD2AFB4C" authorityName="Gosliner & Fahey" authorityYear="2008" box="[630,754,1577,1599]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="palmula">
|
||
<emphasis id="B939EAE0FF97FB20E973F9DEFD2AFB4C" box="[630,754,1577,1599]" italics="true" pageId="15" pageNumber="285">T. palmula</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4D4D71FF97FB20EB94F9BDFEC8FB2F" authorityName="Rudman" authorityYear="1987" baseAuthorityName="Baba" baseAuthorityYear="1935" box="[145,272,1608,1630]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="japonica">
|
||
<emphasis id="B939EAE0FF97FB20EB94F9BDFEC8FB2F" box="[145,272,1608,1630]" italics="true" pageId="15" pageNumber="285">T. japonica</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4D4D71FF97FB20EA4CF9BDFE7CFB2F" authorityName="Smirnoff & Donohoo & Gosliner" authorityYear="2022" box="[329,420,1608,1630]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="kahel">
|
||
<emphasis id="B939EAE0FF97FB20EA4CF9BDFE7CFB2F" box="[329,420,1608,1630]" italics="true" pageId="15" pageNumber="285">T. kahel</emphasis>
|
||
</taxonomicName>
|
||
(see Remarks section below). Both
|
||
<taxonomicName id="4C4D4D71FF97FB20EBD3F993FE9DFB0E" authorityName="Gosliner & Fahey" authorityYear="2008" box="[214,325,1639,1660]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="scurra">
|
||
<emphasis id="B939EAE0FF97FB20EBD3F993FE9DFB0E" box="[214,325,1639,1660]" italics="true" pageId="15" pageNumber="285">T. scurra</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4D4D71FF97FB20EA8DF993FDD7FB09" authorityName="Gosliner & Fahey" authorityYear="2008" box="[392,527,1638,1660]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="palmula">
|
||
<emphasis id="B939EAE0FF97FB20EA8DF993FDD7FB09" box="[392,527,1638,1660]" italics="true" pageId="15" pageNumber="285">T. palmula</emphasis>
|
||
</taxonomicName>
|
||
have a stout body with proportionately thicker extra-rhinophoral and extra-branchial processes, whereas
|
||
<taxonomicName id="4C4D4D71FF97FB20EFDEFF32FA86FDA9" authorityName="Rudman" authorityYear="1987" baseAuthorityName="Baba" baseAuthorityYear="1935" box="[1243,1374,198,219]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="japonica">
|
||
<emphasis id="B939EAE0FF97FB20EFDEFF32FA86FDA9" box="[1243,1374,198,219]" italics="true" pageId="15" pageNumber="285">T. japonica</emphasis>
|
||
</taxonomicName>
|
||
has a more delicate, elongate body with thin, elongate extra-rhinophoral and extra-branchial as in
|
||
<taxonomicName id="4C4D4D71FF97FB20E82CFED6FC7AFC44" authorityName="Smirnoff & Donohoo & Gosliner" authorityYear="2022" box="[809,930,289,311]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="tatsulok">
|
||
<emphasis id="B939EAE0FF97FB20E82CFED6FC7AFC44" box="[809,930,289,311]" italics="true" pageId="15" pageNumber="285">T. tatsulok</emphasis>
|
||
</taxonomicName>
|
||
. These last two species also have a similar colour pattern of a white body with dark brown to black markings. In
|
||
<taxonomicName id="4C4D4D71FF97FB20EF0CFEABFB50FC06" authorityName="Rudman" authorityYear="1987" baseAuthorityName="Baba" baseAuthorityYear="1935" box="[1033,1160,351,372]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="japonica">
|
||
<emphasis id="B939EAE0FF97FB20EF0CFEABFB50FC06" box="[1033,1160,351,372]" italics="true" pageId="15" pageNumber="285">T. japonica</emphasis>
|
||
</taxonomicName>
|
||
, the extra-rhinophoral and extra-branchial process are yellow, while in
|
||
<taxonomicName id="4C4D4D71FF97FB20E82CFE69FC68FCC3" authorityName="Smirnoff & Donohoo & Gosliner" authorityYear="2022" box="[809,944,412,434]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="tatsulok">
|
||
<emphasis id="B939EAE0FF97FB20E82CFE69FC68FCC3" box="[809,944,412,434]" italics="true" pageId="15" pageNumber="285">T. tatsulok</emphasis>
|
||
</taxonomicName>
|
||
they are orange. The dark markings of
|
||
<taxonomicName id="4C4D4D71FF97FB20E842FE4FFC16FCA2" authorityName="Rudman" authorityYear="1987" baseAuthorityName="Baba" baseAuthorityYear="1935" box="[839,974,443,464]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="japonica">
|
||
<emphasis id="B939EAE0FF97FB20E842FE4FFC16FCA2" box="[839,974,443,464]" italics="true" pageId="15" pageNumber="285">T. japonica</emphasis>
|
||
</taxonomicName>
|
||
are rounded spots, whereas they are irregular in shape in
|
||
<taxonomicName id="4C4D4D71FF97FB20EF19FE2EFB4DFC9C" authorityName="Smirnoff & Donohoo & Gosliner" authorityYear="2022" box="[1052,1173,473,495]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="tatsulok">
|
||
<emphasis id="B939EAE0FF97FB20EF19FE2EFB4DFC9C" box="[1052,1173,473,495]" italics="true" pageId="15" pageNumber="285">T. tatsulok</emphasis>
|
||
</taxonomicName>
|
||
, with the exception of the triangular marking anterior to the gill. The gill is white in
|
||
<taxonomicName id="4C4D4D71FF97FB20E888FDE3FBDEFF59" authorityName="Smirnoff & Donohoo & Gosliner" authorityYear="2022" box="[909,1030,534,556]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="tatsulok">
|
||
<emphasis id="B939EAE0FF97FB20E888FDE3FBDEFF59" box="[909,1030,534,556]" italics="true" pageId="15" pageNumber="285">T. tatsulok</emphasis>
|
||
</taxonomicName>
|
||
, whereas the outer portions of the gill branches are black in
|
||
<taxonomicName id="4C4D4D71FF97FB20EF53FDC2FB0DFF39" authorityName="Rudman" authorityYear="1987" baseAuthorityName="Baba" baseAuthorityYear="1935" box="[1110,1237,566,587]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="japonica">
|
||
<emphasis id="B939EAE0FF97FB20EF53FDC2FB0DFF39" box="[1110,1237,566,587]" italics="true" pageId="15" pageNumber="285">T. japonica</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF236F2FF97FB20E844FDA0FAFBF9DD" blockId="15.[807,1426,197,1691]" pageId="15" pageNumber="285">
|
||
Internally,
|
||
<taxonomicName id="4C4D4D71FF97FB20E8C9FDA1FB96FF1B" authorityName="Smirnoff & Donohoo & Gosliner" authorityYear="2022" box="[972,1102,596,618]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="tatsulok">
|
||
<emphasis id="B939EAE0FF97FB20E8C9FDA1FB96FF1B" box="[972,1102,596,618]" italics="true" pageId="15" pageNumber="285">T. tatsulok</emphasis>
|
||
</taxonomicName>
|
||
has several features that clearly distinguish it from other members of the clade of which it is a member. The jaw elements of
|
||
<taxonomicName id="4C4D4D71FF97FB20EE19FD66FA49FFD4" authorityName="Smirnoff & Donohoo & Gosliner" authorityYear="2022" box="[1308,1425,657,679]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="tatsulok">
|
||
<emphasis id="B939EAE0FF97FB20EE19FD66FA49FFD4" box="[1308,1425,657,679]" italics="true" pageId="15" pageNumber="285">T. tatsulok</emphasis>
|
||
</taxonomicName>
|
||
contain numerous minute spikes resembling a flail, while
|
||
<taxonomicName id="4C4D4D71FF97FB20E869FD3BFC15FF96" authorityName="Gosliner & Fahey" authorityYear="2008" box="[876,973,719,740]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="scurra">
|
||
<emphasis id="B939EAE0FF97FB20E869FD3BFC15FF96" box="[876,973,719,740]" italics="true" pageId="15" pageNumber="285">T. scurra</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4D4D71FF97FB20E8DCFD3BFB89FF91" authorityName="Gosliner & Fahey" authorityYear="2008" box="[985,1105,718,740]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="palmula">
|
||
<emphasis id="B939EAE0FF97FB20E8DCFD3BFB89FF91" box="[985,1105,718,740]" italics="true" pageId="15" pageNumber="285">T. palmula</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4D4D71FF97FB20EF8DFD3BFAD8FF96" authorityName="Rudman" authorityYear="1987" baseAuthorityName="Baba" baseAuthorityYear="1935" box="[1160,1280,719,740]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="japonica">
|
||
<emphasis id="B939EAE0FF97FB20EF8DFD3BFAD8FF96" box="[1160,1280,719,740]" italics="true" pageId="15" pageNumber="285">T. japonica</emphasis>
|
||
</taxonomicName>
|
||
have simple, acutely pointed jaw elements (
|
||
<bibRefCitation id="EFDC4B03FF97FB20EF86FD19FA52FE70" author="Gosliner TM & Fahey SJ" box="[1155,1418,749,771]" pageId="15" pageNumber="285" pagination="53 - 98" refId="ref28925" refString="Gosliner TM, Fahey SJ. 2008. Systematics of Trapania (Mollusca: Nudibranchia: Goniodorididae) with descriptions of 16 new species. Systematics and Biodiversity 6: 53 - 98." type="journal article" year="2008">Gosliner & Fahey, 2008</bibRefCitation>
|
||
, fig. 8a, fig. 10a;
|
||
<figureCitation id="13762A77FF97FB20E8D1FCF8FBFBFE50" box="[980,1059,780,802]" captionStart="Figure 5" captionStartId="16.[163,241,1111,1133]" captionTargetBox="[163,1443,195,1071]" captionTargetId="figure-314@16.[163,1443,195,1071]" captionTargetPageId="16" captionText="Figure 5. Trapania tatsulok sp. nov. Scanning electron micrographs of buccal armature of holotype NMP 041333 (formerly CASIZ 200530). A, entire jaw plate. B, detail of armature of jaws. C, majority of radula. D, individual teeth." figureDoi="http://doi.org/10.5281/zenodo.7037759" httpUri="https://zenodo.org/record/7037759/files/figure.png" pageId="15" pageNumber="285">Fig. 5B</figureCitation>
|
||
). The radular teeth of
|
||
<taxonomicName id="4C4D4D71FF97FB20EE18FCF9FA48FE53" authorityName="Smirnoff & Donohoo & Gosliner" authorityYear="2022" box="[1309,1424,780,802]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="tatsulok">
|
||
<emphasis id="B939EAE0FF97FB20EE18FCF9FA48FE53" box="[1309,1424,780,802]" italics="true" pageId="15" pageNumber="285">T. tatsulok</emphasis>
|
||
</taxonomicName>
|
||
have one or two large denticles on the inner side of the primary cusp and numerous smaller denticles, whereas the teeth of
|
||
<taxonomicName id="4C4D4D71FF97FB20E8ADFC9DFBD1FE0C" authorityName="Gosliner & Fahey" authorityYear="2008" box="[936,1033,873,894]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="scurra">
|
||
<emphasis id="B939EAE0FF97FB20E8ADFC9DFBD1FE0C" box="[936,1033,873,894]" italics="true" pageId="15" pageNumber="285">T. scurra</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4D4D71FF97FB20EF11FC9DFB53FE0F" authorityName="Gosliner & Fahey" authorityYear="2008" box="[1044,1163,872,894]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="palmula">
|
||
<emphasis id="B939EAE0FF97FB20EF11FC9DFB53FE0F" box="[1044,1163,872,894]" italics="true" pageId="15" pageNumber="285">T. palmula</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4D4D71FF97FB20EFB8FC9DFAEDFE0F" authorityName="Rudman" authorityYear="1987" baseAuthorityName="Baba" baseAuthorityYear="1935" box="[1213,1333,872,894]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="japonica">
|
||
<emphasis id="B939EAE0FF97FB20EFB8FC9DFAEDFE0F" box="[1213,1333,872,894]" italics="true" pageId="15" pageNumber="285">T. japonica</emphasis>
|
||
</taxonomicName>
|
||
all have a single prominent cusp and numerous smaller denticles (
|
||
<bibRefCitation id="EFDC4B03FF97FB20E834FC51FBE2FEC8" author="Gosliner TM & Fahey SJ" box="[817,1082,933,955]" pageId="15" pageNumber="285" pagination="53 - 98" refId="ref28925" refString="Gosliner TM, Fahey SJ. 2008. Systematics of Trapania (Mollusca: Nudibranchia: Goniodorididae) with descriptions of 16 new species. Systematics and Biodiversity 6: 53 - 98." type="journal article" year="2008">Gosliner & Fahey, 2008</bibRefCitation>
|
||
: fig. 8b–d, fig. 10b–d;
|
||
<figureCitation id="13762A77FF97FB20EE34FC51FA5BFEC9" box="[1329,1411,933,955]" captionStart="Figure 5" captionStartId="16.[163,241,1111,1133]" captionTargetBox="[163,1443,195,1071]" captionTargetId="figure-314@16.[163,1443,195,1071]" captionTargetPageId="16" captionText="Figure 5. Trapania tatsulok sp. nov. Scanning electron micrographs of buccal armature of holotype NMP 041333 (formerly CASIZ 200530). A, entire jaw plate. B, detail of armature of jaws. C, majority of radula. D, individual teeth." figureDoi="http://doi.org/10.5281/zenodo.7037759" httpUri="https://zenodo.org/record/7037759/files/figure.png" pageId="15" pageNumber="285">Fig. 5D</figureCitation>
|
||
). The reproductive system of
|
||
<taxonomicName id="4C4D4D71FF97FB20EF59FC31FB08FEAB" authorityName="Smirnoff & Donohoo & Gosliner" authorityYear="2022" box="[1116,1232,964,986]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="tatsulok">
|
||
<emphasis id="B939EAE0FF97FB20EF59FC31FB08FEAB" box="[1116,1232,964,986]" italics="true" pageId="15" pageNumber="285">T. tatsulok</emphasis>
|
||
</taxonomicName>
|
||
is similar to that of
|
||
<taxonomicName id="4C4D4D71FF97FB20E843FC17FC1EFE85" authorityName="Gosliner & Fahey" authorityYear="2008" box="[838,966,994,1016]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="palmula">
|
||
<emphasis id="B939EAE0FF97FB20E843FC17FC1EFE85" box="[838,966,994,1016]" italics="true" pageId="15" pageNumber="285">T. palmula</emphasis>
|
||
</taxonomicName>
|
||
, with a relatively elongate vagina and ejaculatory portion of the vas deferens, but in
|
||
<taxonomicName id="4C4D4D71FF97FB20EE1CFBF6FA49F964" authorityName="Gosliner & Fahey" authorityYear="2008" box="[1305,1425,1025,1047]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="palmula">
|
||
<emphasis id="B939EAE0FF97FB20EE1CFBF6FA49F964" box="[1305,1425,1025,1047]" italics="true" pageId="15" pageNumber="285">T. palmula</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFDC4B03FF97FB20E834FBD4FB9EF947" author="Gosliner TM & Fahey SJ" box="[817,1094,1056,1078]" pageId="15" pageNumber="285" pagination="53 - 98" refId="ref28925" refString="Gosliner TM, Fahey SJ. 2008. Systematics of Trapania (Mollusca: Nudibranchia: Goniodorididae) with descriptions of 16 new species. Systematics and Biodiversity 6: 53 - 98." type="journal article" year="2008">Gosliner & Fahey, 2008</bibRefCitation>
|
||
: fig. 9) both the vagina and ejaculatory duct are more elongate than in
|
||
<taxonomicName id="4C4D4D71FF97FB20EFFFFBCBFAB5F921" authorityName="Smirnoff & Donohoo & Gosliner" authorityYear="2022" box="[1274,1389,1086,1108]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="tatsulok">
|
||
<emphasis id="B939EAE0FF97FB20EFFFFBCBFAB5F921" box="[1274,1389,1086,1108]" italics="true" pageId="15" pageNumber="285">T. tatsulok</emphasis>
|
||
</taxonomicName>
|
||
. In
|
||
<taxonomicName id="4C4D4D71FF97FB20E822FBAAFC51F901" authorityName="Gosliner & Fahey" authorityYear="2008" box="[807,905,1118,1139]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="scurra">
|
||
<emphasis id="B939EAE0FF97FB20E822FBAAFC51F901" box="[807,905,1118,1139]" italics="true" pageId="15" pageNumber="285">T. scurra</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFDC4B03FF97FB20E890FBA9FB4AF900" author="Gosliner TM & Fahey SJ" box="[917,1170,1117,1139]" pageId="15" pageNumber="285" pagination="53 - 98" refId="ref28925" refString="Gosliner TM, Fahey SJ. 2008. Systematics of Trapania (Mollusca: Nudibranchia: Goniodorididae) with descriptions of 16 new species. Systematics and Biodiversity 6: 53 - 98." type="journal article" year="2008">Gosliner & Fahey, 2008</bibRefCitation>
|
||
: fig. 11) and
|
||
<taxonomicName id="4C4D4D71FF97FB20EE1DFBAAFA49F901" authorityName="Rudman" authorityYear="1987" baseAuthorityName="Baba" baseAuthorityYear="1935" box="[1304,1425,1118,1139]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="japonica">
|
||
<emphasis id="B939EAE0FF97FB20EE1DFBAAFA49F901" box="[1304,1425,1118,1139]" italics="true" pageId="15" pageNumber="285">T. japonica</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13762A77FF97FB20E834FB88FC5AF9E3" box="[817,898,1148,1170]" captionStart="Figure 4" captionStartId="14.[163,241,1790,1812]" captionTargetBox="[258,1346,197,1747]" captionTargetId="figure-27@14.[255,1349,194,1750]" captionTargetPageId="14" captionText="Figure 4. Reproductive systems. A, Trapania japonica (Baba, 1935), CASIZ 221116, scale = 0.50 mm. B, Trapania tatsulok sp. nov., holotype, NMP 041333 (formerly CASIZ 200530), scale = 0.33 mm. C, Trapania kahel sp. nov., holotype, NMP 041330 (formerly CASIZ 186131), scale = 0.28 mm. D, Trapania lemanioides sp. nov., holotype, NMP 041335 (formerly" figureDoi="http://doi.org/10.5281/zenodo.7037757" httpUri="https://zenodo.org/record/7037757/files/figure.png" pageId="15" pageNumber="285">Fig. 4A</figureCitation>
|
||
) the vagina and ejaculatory portion of the vas deferens are much shorter than in
|
||
<emphasis id="B939EAE0FF97FB20EFACFB6FFAFBF9DD" box="[1193,1315,1178,1200]" italics="true" pageId="15" pageNumber="285">
|
||
<taxonomicName id="4C4D4D71FF97FB20EFACFB6FFAC7F9DD" authorityName="Smirnoff & Donohoo & Gosliner" authorityYear="2022" box="[1193,1311,1178,1200]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="tatsulok">T. tatsulok</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF236F2FF97FB20E844FB4DFA49FBE9" blockId="15.[807,1426,197,1691]" pageId="15" pageNumber="285">
|
||
Three other described species of
|
||
<taxonomicName id="4C4D4D71FF97FB20EFBEFB4EFAFEF9BD" authorityName="Pruvot-Fol" authorityYear="1931" box="[1211,1318,1210,1231]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B939EAE0FF97FB20EFBEFB4EFAFEF9BD" box="[1211,1318,1210,1231]" italics="true" pageId="15" pageNumber="285">Trapania</emphasis>
|
||
</taxonomicName>
|
||
from the eastern Pacific have similar colour patterns to
|
||
<taxonomicName id="4C4D4D71FF97FB20EE1FFB2DFA53F99F" authorityName="Smirnoff & Donohoo & Gosliner" authorityYear="2022" box="[1306,1419,1240,1262]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="tatsulok">
|
||
<emphasis id="B939EAE0FF97FB20EE1FFB2DFA53F99F" box="[1306,1419,1240,1262]" italics="true" pageId="15" pageNumber="285">T. tatsulok</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C4D4D71FF97FB20E82CFB03FC17F879" baseAuthorityName="Cockerell" baseAuthorityYear="1901" box="[809,975,1270,1292]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="velox">
|
||
<emphasis id="B939EAE0FF97FB20E82CFB03FC17F879" box="[809,975,1270,1292]" italics="true" pageId="15" pageNumber="285">Trapania velox</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4D4D71FF97FB20E8D9FB03FAB8F87E" authority="Millen & Bertsch, 2000" authorityName="Millen & Bertsch" authorityYear="2000" box="[988,1376,1270,1292]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="goslineri">
|
||
<emphasis id="B939EAE0FF97FB20E8D9FB03FB8FF879" box="[988,1111,1270,1292]" italics="true" pageId="15" pageNumber="285">T. goslineri</emphasis>
|
||
<bibRefCitation id="EFDC4B03FF97FB20EF58FB02FAB8F87E" author="Millen SV & Bertsch H" box="[1117,1376,1270,1292]" pageId="15" pageNumber="285" pagination="354 - 366" refId="ref29792" refString="Millen SV, Bertsch H. 2000. Three new species of dorid nudibranchs from southern California, USA, and the Baja California Peninsula, Mexico. The Veliger 43: 354 - 366." type="journal article" year="2000">Millen & Bertsch, 2000</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4D4D71FF97FB20E82CFAE2FB71F858" authority="Gosliner & Fahey, 2008" authorityName="Gosliner & Fahey" authorityYear="2008" box="[809,1193,1301,1323]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="darwini">
|
||
<emphasis id="B939EAE0FF97FB20E82CFAE2FC45F858" box="[809,925,1301,1323]" italics="true" pageId="15" pageNumber="285">T. darwini</emphasis>
|
||
<bibRefCitation id="EFDC4B03FF97FB20E8A1FAE1FB71F858" author="Gosliner TM & Fahey SJ" box="[932,1193,1301,1323]" pageId="15" pageNumber="285" pagination="53 - 98" refId="ref28925" refString="Gosliner TM, Fahey SJ. 2008. Systematics of Trapania (Mollusca: Nudibranchia: Goniodorididae) with descriptions of 16 new species. Systematics and Biodiversity 6: 53 - 98." type="journal article" year="2008">Gosliner & Fahey, 2008</bibRefCitation>
|
||
</taxonomicName>
|
||
, all have white body pigment with brown or black markings and orange pigment, but differ in the pattern and distribution of these pigments. In these three species, there is both brown and orange pigment on the extra-rhinophoral and extrabranchial processes, while in
|
||
<taxonomicName id="4C4D4D71FF97FB20EF7AFA5BFB2FF8B1" authorityName="Smirnoff & Donohoo & Gosliner" authorityYear="2022" box="[1151,1271,1454,1476]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="tatsulok">
|
||
<emphasis id="B939EAE0FF97FB20EF7AFA5BFB2FF8B1" box="[1151,1271,1454,1476]" italics="true" pageId="15" pageNumber="285">T. tatsulok</emphasis>
|
||
</taxonomicName>
|
||
there is only orange. These three species also have orange rhinophores, while in
|
||
<taxonomicName id="4C4D4D71FF97FB20E889FA19FBDAFB73" authorityName="Smirnoff & Donohoo & Gosliner" authorityYear="2022" box="[908,1026,1516,1538]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="tatsulok">
|
||
<emphasis id="B939EAE0FF97FB20E889FA19FBDAFB73" box="[908,1026,1516,1538]" italics="true" pageId="15" pageNumber="285">T. tatsulok</emphasis>
|
||
</taxonomicName>
|
||
the rhinophores are dark brown to black. In
|
||
<taxonomicName id="4C4D4D71FF97FB20E895F9FFFC39FB6D" baseAuthorityName="Cockerell" baseAuthorityYear="1901" box="[912,993,1546,1568]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="velox">
|
||
<emphasis id="B939EAE0FF97FB20E895F9FFFC39FB6D" box="[912,993,1546,1568]" italics="true" pageId="15" pageNumber="285">T. velox</emphasis>
|
||
</taxonomicName>
|
||
, the translucent, white body has darkbrown longitudinal lines, while
|
||
<taxonomicName id="4C4D4D71FF97FB20EF90F9DEFACAFB4C" authorityName="Millen & Bertsch" authorityYear="2000" box="[1173,1298,1577,1599]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="goslineri">
|
||
<emphasis id="B939EAE0FF97FB20EF90F9DEFACAFB4C" box="[1173,1298,1577,1599]" italics="true" pageId="15" pageNumber="285">T. goslineri</emphasis>
|
||
</taxonomicName>
|
||
has brown spots. In
|
||
<taxonomicName id="4C4D4D71FF97FB20E882F9BDFC20FB2F" authorityName="Gosliner & Fahey" authorityYear="2008" box="[903,1016,1608,1630]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="darwini">
|
||
<emphasis id="B939EAE0FF97FB20E882F9BDFC20FB2F" box="[903,1016,1608,1630]" italics="true" pageId="15" pageNumber="285">T. darwini</emphasis>
|
||
</taxonomicName>
|
||
there is a series of longitudinal brown markings interrupted by white spots.
|
||
<bibRefCitation id="EFDC4B03FF97FB20EFBDF992FA9EFB0E" author="Bhave VJ" box="[1208,1350,1638,1660]" pageId="15" pageNumber="285" refId="ref28356" refString="Bhave VJ. 2010. Trapania goslineri? from Ratnagiri, India. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / 23291. Accessed 29 November 2021." type="url" year="2010">Bhave (2010)</bibRefCitation>
|
||
posted a photo of another species from
|
||
<collectingCountry id="F35A7662FF97FB20EF7DF971FB6AFBE9" box="[1144,1202,1669,1691]" name="India" pageId="15" pageNumber="285">India</collectingCountry>
|
||
that has yet another
|
||
</paragraph>
|
||
<paragraph id="8BF236F2FF97FB20EB8AF93FFBCAFA06" blockId="15.[143,1425,1739,1908]" pageId="15" pageNumber="285">
|
||
<materialsCitation id="3B253CAFFF97FB20EB8AF93FFC88FB93" ID-GBIF-Occurrence="3897192336" box="[143,848,1739,1761]" collectionCode="CASIZ" pageId="15" pageNumber="285" specimenCode="CASIZ 208433" specimenCount="1">
|
||
<specimenCode id="DBEB9E89FF97FB20EB8AF93FFEF0FB93" box="[143,296,1739,1761]" collectionCode="CASIZ" pageId="15" pageNumber="285">CASIZ 208433</specimenCode>
|
||
), scale =
|
||
<quantity id="4CB59B17FF97FB20EA80F93FFE3BFB93" box="[389,483,1739,1761]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="15" pageNumber="285" unit="mm" value="0.33">0.33 mm</quantity>
|
||
. E,
|
||
<taxonomicName id="4C4D4D71FF97FB20E900F93FFC88FB93" authority="Rudman, 1987" authorityName="Rudman" authorityYear="1987" box="[517,848,1739,1761]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="aurata">
|
||
<emphasis id="B939EAE0FF97FB20E900F93FFD6BFB93" box="[517,691,1739,1761]" italics="true" pageId="15" pageNumber="285">Trapania aurata</emphasis>
|
||
<bibRefCitation id="EFDC4B03FF97FB20E9BCF93FFC88FB93" author="Rudman WB" box="[697,848,1739,1761]" pageId="15" pageNumber="285" pagination="189 - 212" refId="ref30529" refString="Rudman WB. 1987. The genus Trapania (Nudibranchia: Goniodorididae) in the Indo-West Pacific. Journal of Molluscan Studies 53: 189 - 212." type="journal article" year="1987">Rudman, 1987</bibRefCitation>
|
||
</taxonomicName>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B253CAFFF97FB20E85EF93FFEF4FB8C" ID-GBIF-Occurrence="3897192326" collectionCode="CASIZ" pageId="15" pageNumber="285" specimenCode="CASIZ 186205" specimenCount="1">
|
||
<specimenCode id="DBEB9E89FF97FB20E85EF93FFC2BFB93" box="[859,1011,1739,1761]" collectionCode="CASIZ" pageId="15" pageNumber="285">CASIZ 186205</specimenCode>
|
||
, scale =
|
||
<quantity id="4CB59B17FF97FB20EF4DF93FFB7DFB93" box="[1096,1189,1739,1761]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" pageId="15" pageNumber="285" unit="mm" value="0.46">0.46 mm</quantity>
|
||
. F,
|
||
<taxonomicName id="4C4D4D71FF97FB20EFC6F93FFEF4FB8C" authority="Rudman, 1987" authorityName="Rudman" authorityYear="1987" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="reticulata">
|
||
<emphasis id="B939EAE0FF97FB20EFC6F93FFA48FB93" box="[1219,1424,1739,1761]" italics="true" pageId="15" pageNumber="285">Trapania reticulata</emphasis>
|
||
<bibRefCitation id="EFDC4B03FF97FB20EB94F91CFEF4FB8C" author="Rudman WB" box="[145,300,1768,1790]" pageId="15" pageNumber="285" pagination="189 - 212" refId="ref30529" refString="Rudman WB. 1987. The genus Trapania (Nudibranchia: Goniodorididae) in the Indo-West Pacific. Journal of Molluscan Studies 53: 189 - 212." type="journal article" year="1987">Rudman, 1987</bibRefCitation>
|
||
</taxonomicName>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B253CAFFF97FB20EA3CF91CFBCFFB8C" ID-GBIF-Occurrence="3897192328" box="[313,1047,1768,1790]" collectionCode="CASIZ" pageId="15" pageNumber="285" specimenCode="CASIZ 191431" specimenCount="1">
|
||
<specimenCode id="DBEB9E89FF97FB20EA3CF91CFE0EFB8C" box="[313,470,1768,1790]" collectionCode="CASIZ" pageId="15" pageNumber="285">CASIZ 191431</specimenCode>
|
||
, scale =
|
||
<quantity id="4CB59B17FF97FB20E935F91CFD48FB8C" box="[560,656,1768,1790]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="285" unit="mm" value="0.5">0.50 mm</quantity>
|
||
. G,
|
||
<taxonomicName id="4C4D4D71FF97FB20E9BDF91CFBCFFB8C" authority="Rudman, 1987" authorityName="Rudman" authorityYear="1987" box="[696,1047,1768,1790]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="darvelli">
|
||
<emphasis id="B939EAE0FF97FB20E9BDF91CFCAEFB8C" box="[696,886,1768,1790]" italics="true" pageId="15" pageNumber="285">Trapania darvelli</emphasis>
|
||
<bibRefCitation id="EFDC4B03FF97FB20E879F91CFBCFFB8C" author="Rudman WB" box="[892,1047,1768,1790]" pageId="15" pageNumber="285" pagination="189 - 212" refId="ref30529" refString="Rudman WB. 1987. The genus Trapania (Nudibranchia: Goniodorididae) in the Indo-West Pacific. Journal of Molluscan Studies 53: 189 - 212." type="journal article" year="1987">Rudman, 1987</bibRefCitation>
|
||
</taxonomicName>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B253CAFFF97FB20EF20F91CFE2EFA6E" ID-GBIF-Occurrence="3897192309" collectionCode="CASIZ" pageId="15" pageNumber="285" specimenCode="CASIZ 222004" specimenCount="1" typeStatus="holotype">
|
||
<specimenCode id="DBEB9E89FF97FB20EF20F91CFB1AFB8C" box="[1061,1218,1768,1790]" collectionCode="CASIZ" pageId="15" pageNumber="285">CASIZ 222004</specimenCode>
|
||
, scale =
|
||
<quantity id="4CB59B17FF97FB20EE19F91CFAB7FB8C" box="[1308,1391,1768,1790]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="15" pageNumber="285" unit="mm" value="0.4">0.4 mm</quantity>
|
||
. H,
|
||
<taxonomicName id="4C4D4D71FF97FB20EB94F8F2FEE1FA6E" authority="Smirnoff & Donohoo & Gosliner, 2022" authorityName="Smirnoff & Donohoo & Gosliner" authorityYear="2022" box="[145,313,1798,1820]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="tigger" status="sp. nov.">
|
||
<emphasis id="B939EAE0FF97FB20EB94F8F2FEE1FA6E" box="[145,313,1798,1820]" italics="true" pageId="15" pageNumber="285">Trapania tigger</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20A579BFF97FB20EA3AF8F2FE57FA6E" box="[319,399,1798,1820]" pageId="15" pageNumber="285" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<typeStatus id="54F68850FF97FB20EA9FF8F2FE2EFA6E" box="[410,502,1798,1820]" pageId="15" pageNumber="285" type="holotype">holotype</typeStatus>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B253CAFFF97FB20E905F8F2FA52FA6E" ID-GBIF-Occurrence="3897192304" box="[512,1418,1798,1820]" collectionCode="NMP, CASIZ" pageId="15" pageNumber="285" specimenCode="NMP 041328, CASIZ 180412" specimenCount="1" typeStatus="holotype">
|
||
<specimenCode id="DBEB9E89FF97FB20E905F8F2FD54FA6E" box="[512,652,1798,1820]" collectionCode="NMP" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" pageId="15" pageNumber="285">NMP 041328</specimenCode>
|
||
(formerly
|
||
<specimenCode id="DBEB9E89FF97FB20E9F8F8F2FC41FA6E" box="[765,921,1798,1820]" collectionCode="CASIZ" pageId="15" pageNumber="285">CASIZ 180412</specimenCode>
|
||
), scale = 1.0 mm. I,
|
||
<taxonomicName id="4C4D4D71FF97FB20EF68F8F2FAEFFA69" authority="Smirnoff & Donohoo & Gosliner, 2022" authorityName="Smirnoff & Donohoo & Gosliner" authorityYear="2022" box="[1133,1335,1798,1820]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="tamaraw" status="sp. nov.">
|
||
<emphasis id="B939EAE0FF97FB20EF68F8F2FAEFFA69" box="[1133,1335,1798,1820]" italics="true" pageId="15" pageNumber="285">Trapania tamaraw</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20A579BFF97FB20EE3BF8F2FA52FA6E" box="[1342,1418,1798,1820]" pageId="15" pageNumber="285" rank="species">sp. nov</taxonomicNameLabel>
|
||
</materialsCitation>
|
||
.,
|
||
<materialsCitation id="3B253CAFFF97FB20EB94F8D7FB4BFA4B" ID-GBIF-Occurrence="3897192305" box="[145,1171,1827,1849]" collectionCode="NMP, CASIZ" pageId="15" pageNumber="285" specimenCode="NMP 041334, CASIZ 208391" specimenCount="1" typeStatus="holotype">
|
||
<specimenCode id="DBEB9E89FF97FB20EB94F8D7FEC5FA4B" box="[145,285,1827,1849]" collectionCode="NMP" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" pageId="15" pageNumber="285">NMP 041334</specimenCode>
|
||
(formerly
|
||
<specimenCode id="DBEB9E89FF97FB20EA8AF8D7FDF5FA4B" box="[399,557,1827,1849]" collectionCode="CASIZ" pageId="15" pageNumber="285">CASIZ 208391</specimenCode>
|
||
), scale =
|
||
<quantity id="4CB59B17FF97FB20E98BF8D7FD34FA4B" box="[654,748,1827,1849]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="285" unit="mm" value="0.5">0.50 mm</quantity>
|
||
. J,
|
||
<taxonomicName id="4C4D4D71FF97FB20E80AF8D7FC0FFA4B" authority="Smirnoff & Donohoo & Gosliner, 2022" authorityName="Smirnoff & Donohoo & Gosliner" authorityYear="2022" box="[783,983,1827,1849]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="15" pageNumber="285" phylum="Mollusca" rank="species" species="stegodon" status="sp. nov.">
|
||
<emphasis id="B939EAE0FF97FB20E80AF8D7FC0FFA4B" box="[783,983,1827,1849]" italics="true" pageId="15" pageNumber="285">Trapania stegodon</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20A579BFF97FB20E8D8F8D7FBF6FA4B" box="[989,1070,1827,1849]" pageId="15" pageNumber="285" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<typeStatus id="54F68850FF97FB20EF3DF8D7FB4BFA4B" box="[1080,1171,1827,1849]" pageId="15" pageNumber="285" type="holotype">holotype</typeStatus>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B253CAFFF97FB20EF9FF8D7FBD7FA06" ID-GBIF-Occurrence="3897192323" collectionCode="NMP, CASIZ" pageId="15" pageNumber="285" specimenCode="NMP 041332, CASIZ 186206" specimenCount="1" typeStatus="holotype">
|
||
<specimenCode id="DBEB9E89FF97FB20EF9FF8D7FAFEFA4B" box="[1178,1318,1827,1849]" collectionCode="NMP" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" pageId="15" pageNumber="285">NMP 041332</specimenCode>
|
||
(formerly
|
||
<specimenCode id="DBEB9E89FF97FB20EB94F8B4FEF3FA24" box="[145,299,1856,1878]" collectionCode="CASIZ" pageId="15" pageNumber="285">CASIZ 186206</specimenCode>
|
||
), scale =
|
||
<quantity id="4CB59B17FF97FB20EA83F8B4FE3BFA24" box="[390,483,1856,1878]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="285" unit="mm" value="0.5">0.50 mm</quantity>
|
||
. Abbreviations: am, ampulla; bc, bursa copulatrix; fgm, female gland mass; p, penial sac; pr, prostatic portion of vas deferens; rs, receptaculum seminis; ud, uterine duct; v, vagina
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF236F2FF88FB3FEBA6FF74FDC8FDE8" blockId="16.[163,528,128,155]" box="[163,528,128,155]" pageId="16" pageNumber="286">
|
||
286 D. S. SMIRNOFF
|
||
<emphasis id="B939EAE0FF88FB3FEABAFF71FDD2FDE8" box="[447,522,133,154]" italics="true" pageId="16" pageNumber="286">
|
||
<collectionCode id="ED5CAE37FF88FB3FEABAFF71FE3AFDE8" box="[447,482,133,154]" country="0" name="East Texas State University" pageId="16" pageNumber="286">ET</collectionCode>
|
||
AL
|
||
</emphasis>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF32667AFF88FB3FEBA6FBA3FB25F9F9" ID-DOI="http://doi.org/10.5281/zenodo.7037759" ID-Zenodo-Dep="7037759" httpUri="https://zenodo.org/record/7037759/files/figure.png" pageId="16" pageNumber="286" startId="16.[163,241,1111,1133]" targetBox="[163,1443,195,1071]" targetPageId="16">
|
||
<paragraph id="8BF236F2FF88FB3FEBA6FBA3FB25F9F9" blockId="16.[163,1442,1111,1163]" pageId="16" pageNumber="286">
|
||
<emphasis id="B939EAE0FF88FB3FEBA6FBA3FED2F91E" bold="true" box="[163,266,1111,1133]" pageId="16" pageNumber="286">Figure 5.</emphasis>
|
||
<taxonomicName id="4C4D4D71FF88FB3FEA10FBA3FE09F91F" authority="Smirnoff & Donohoo & Gosliner, 2022" authorityName="Smirnoff & Donohoo & Gosliner" authorityYear="2022" box="[277,465,1111,1133]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="16" pageNumber="286" phylum="Mollusca" rank="species" species="tatsulok" status="sp. nov.">
|
||
<emphasis id="B939EAE0FF88FB3FEA10FBA3FE09F91F" box="[277,465,1111,1133]" italics="true" pageId="16" pageNumber="286">Trapania tatsulok</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20A579BFF88FB3FEAD2FBA3FDFAF91F" box="[471,546,1111,1133]" pageId="16" pageNumber="286" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Scanning electron micrographs of buccal armature of holotype NMP 041333 (formerly CASIZ 200530). A, entire jaw plate. B, detail of armature of jaws. C, majority of radula. D, individual teeth.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF236F2FF88FB3FEBA6FB45FEA7FB45" blockId="16.[161,780,1201,1591]" pageId="16" pageNumber="286">
|
||
combination of white, orange and brown markings and differs from the species discussed above.
|
||
<bibRefCitation id="EFDC4B03FF88FB3FE965FB24FCD3F997" author="Rudman WB" box="[608,779,1232,1254]" pageId="16" pageNumber="286" refId="ref30601" refString="Rudman WB. 2010. Comment on Trapania goslineri? from Ratnagiri, India by Vishal Bhave. In: Sea slug forum. Sydney: Australian Museum. Available at: http: // www. seaslugforum. net / find / 23291. Accessed 29 November 2021." type="url" year="2010">Rudman (2010)</bibRefCitation>
|
||
suggested that this specimen may be
|
||
<taxonomicName id="4C4D4D71FF88FB3FE937FB1BFD71F871" authorityName="Millen & Bertsch" authorityYear="2000" box="[562,681,1262,1284]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="16" pageNumber="286" phylum="Mollusca" rank="species" species="goslineri">
|
||
<emphasis id="B939EAE0FF88FB3FE937FB1BFD71F871" box="[562,681,1262,1284]" italics="true" pageId="16" pageNumber="286">T. goslineri</emphasis>
|
||
</taxonomicName>
|
||
and that
|
||
<taxonomicName id="4C4D4D71FF88FB3FEBA4FAFAFECFF850" authorityName="Gosliner & Fahey" authorityYear="2008" box="[161,279,1293,1315]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="16" pageNumber="286" phylum="Mollusca" rank="species" species="darwini">
|
||
<emphasis id="B939EAE0FF88FB3FEBA4FAFAFECFF850" box="[161,279,1293,1315]" italics="true" pageId="16" pageNumber="286">T. darwini</emphasis>
|
||
</taxonomicName>
|
||
may also be the same species as
|
||
<taxonomicName id="4C4D4D71FF88FB3FE98FFAFAFCDDF850" authorityName="Millen & Bertsch" authorityYear="2000" box="[650,773,1293,1315]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="16" pageNumber="286" phylum="Mollusca" rank="species" species="goslineri">
|
||
<emphasis id="B939EAE0FF88FB3FE98FFAFAFCDDF850" box="[650,773,1293,1315]" italics="true" pageId="16" pageNumber="286">T. goslineri</emphasis>
|
||
</taxonomicName>
|
||
, despite the morphological distinctions clearly noted by
|
||
<bibRefCitation id="EFDC4B03FF88FB3FEBC2FABEFE31F82D" author="Gosliner TM & Fahey SJ" box="[199,489,1354,1376]" pageId="16" pageNumber="286" pagination="53 - 98" refId="ref28925" refString="Gosliner TM, Fahey SJ. 2008. Systematics of Trapania (Mollusca: Nudibranchia: Goniodorididae) with descriptions of 16 new species. Systematics and Biodiversity 6: 53 - 98." type="journal article" year="2008">Gosliner & Fahey (2008)</bibRefCitation>
|
||
. Another feature clearly distinguishes the specimen photographed by Bhave. In this specimen from
|
||
<collectingCountry id="F35A7662FF88FB3FEA72FA73FE6AF8EF" box="[375,434,1415,1437]" name="India" pageId="16" pageNumber="286">India</collectingCountry>
|
||
, the base of the rhinophore club is broad and bulbous with an abruptly narrow apex. All of the other species of
|
||
<taxonomicName id="4C4D4D71FF88FB3FEA76FA31FE01F8A8" authorityName="Pruvot-Fol" authorityYear="1931" box="[371,473,1477,1498]" class="Gastropoda" family="Goniodorididae" genus="Trapania" kingdom="Animalia" order="Nudibranchia" pageId="16" pageNumber="286" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B939EAE0FF88FB3FEA76FA31FE01F8A8" box="[371,473,1477,1498]" italics="true" pageId="16" pageNumber="286">Trapania</emphasis>
|
||
</taxonomicName>
|
||
with similar colour patterns have narrow, evenly graded rhinophores and it is likely that the specimen from
|
||
<collectingCountry id="F35A7662FF88FB3FEAA7F9F6FE04FB6A" box="[418,476,1538,1560]" name="India" pageId="16" pageNumber="286">India</collectingCountry>
|
||
represents another distinct, undescribed species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |