treatments-xml/data/03/E4/87/03E48797FFE6FF9FFF214944C62CFACC.xml

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<mods:title id="6DE9063CC45713C3132454F2D8FCE159">On two species of Aniptumnus Ng, 2002 (Crustacea: Brachyura: Pilumnidae) from India, with a description of a new species from the coast of Kerala</mods:title>
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<mods:namePart id="C12E354CAC110AEFB5D1C925679B7AD2">Hari, P. Praved</mods:namePart>
<mods:affiliation id="F5CA8A018BAB5B3232EEF46073BA54EA">Department of Marine Biology, Microbiology and Biochemistry, School of Marine Sciences, Cochin University of Science and Technology, Cochin 682016 Kerala, India</mods:affiliation>
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<mods:namePart id="C624B9D60308EE60B6D62D06E43D6DC7">Hershey, N. Regina</mods:namePart>
<mods:affiliation id="36E2788392FBDF564306D9DC5F011D52">Department of Marine Biology, Microbiology and Biochemistry, School of Marine Sciences, Cochin University of Science and Technology, Cochin 682016 Kerala, India &amp; Department of Zoology, N. S. S. Hindu College, Changanassery 686 102 Kerala, India</mods:affiliation>
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<mods:namePart id="B1D0ECE428E4CC5B9C9F2B83DC511E48">Mendoza, Jose Christopher E.</mods:namePart>
<mods:affiliation id="8880A4AE6DB4547D332B2F917B0FFCA5">Lee Kong Chian Natural History Museum, Faculty of Science, National University of Singapore, 2 Conservatory Drive, 117377, Singapore</mods:affiliation>
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<treatment id="03E48797FFE6FF9FFF214944C62CFACC" ID-DOI="http://doi.org/10.5281/zenodo.7386400" ID-GBIF-Taxon="205344592" ID-Zenodo-Dep="7386400" LSID="urn:lsid:plazi:treatment:03E48797FFE6FF9FFF214944C62CFACC" httpUri="http://treatment.plazi.org/id/03E48797FFE6FF9FFF214944C62CFACC" lastPageId="9" lastPageNumber="270" pageId="2" pageNumber="263">
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<paragraph id="8BF23681FFE6FF94FF214944C39BF9F3" blockId="2.[151,476,1579,1642]" box="[151,476,1579,1606]" pageId="2" pageNumber="263">
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<taxonomicName id="4C4D4D02FFE6FF94FF214944C33FF9F3" ID-CoL="a2549b20-f23f-403a-afc0-e9b90cc72e71" authority="Hari &amp; Hershey &amp; Mendoza, 2022" authorityName="Hari &amp; Hershey &amp; Mendoza" authorityYear="2022" box="[151,376,1579,1606]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="263" phylum="Arthropoda" rank="species" species="bijoyi" status="sp. nov.">
<emphasis id="B939EA93FFE6FF94FF214944C33FF9F3" bold="true" box="[151,376,1579,1606]" italics="true" pageId="2" pageNumber="263">Aniptumnus bijoyi</emphasis>
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<taxonomicNameLabel id="A20A57E8FFE6FF94FEC94944C39BF9F3" box="[383,476,1580,1606]" pageId="2" pageNumber="263" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C357650AFFE6FF94FF214938C347F9DF" box="[151,256,1616,1642]" pageId="2" pageNumber="263" type="description">
<paragraph id="8BF23681FFE6FF94FF214938C347F9DF" blockId="2.[151,476,1579,1642]" box="[151,256,1616,1642]" pageId="2" pageNumber="263">
<figureCitation id="13762A04FFE6FF94FF214938C347F9DF" box="[151,256,1616,1642]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="3.[151,250,1950,1974]" captionStartId-1="5.[151,250,1891,1915]" captionStartId-2="6.[151,250,1891,1915]" captionStartId-3="7.[151,250,1490,1514]" captionTargetBox-0="[223,1366,993,1927]" captionTargetBox-1="[151,1435,181,1867]" captionTargetBox-2="[151,1435,181,1867]" captionTargetBox-3="[151,1435,705,1467]" captionTargetId-0="figure-382@3.[222,1366,993,1927]" captionTargetId-1="figure-20@5.[151,1435,181,1867]" captionTargetId-2="figure-21@6.[151,1435,181,1867]" captionTargetId-3="figure-241@7.[151,1435,705,1467]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="FIGURE 1. Aniptumnus bijoyi sp. nov., paratype, male, 10.1 by 6.7 mm (ZRC 2022.0137), Kochi, Kerala, SW India. Dorsal habitus and live coloration." captionText-1="FIGURE 2. Aniptumnus bijoyi sp. nov., holotype, male, 10.7 by 7.1 mm (CUSAT/SBN/HRJ/1), Kochi, Kerala, SW India. A, carapace, dorsal view; B, left third maxilliped, external view; cephalothorax and chelipeds, frontal view; D, anterior thoracic sternum and telson, ventral view; E, thoracic sternum and sternopleonal cavity, ventral view; F, pleon, ventral view; G, pleonal somites 1–3, posteroventral view. Scale bars: 2.0 mm (A, C); 1.0 mm (B, D–G)." captionText-2="FIGURE 3. Aniptumnus bijoyi sp. nov., holotype, male, 10.7 by 7.1 mm (CUSAT/SBN/HRJ/1), Kochi, Kerala, SW India. A, minor cheliped, external view; B, major cheliped, external view; C, right P5, dorsal view; D, E, left G1 (D, pleonal/ventral view; E, mesiosternal/mesiodorsal view); F, left G2, sternal/dorsal view; G, H, distal tip of left G1 (G, pleonal/ventral view; H, mesiosternal/mesiodorsal view). Scale bars: 1.0 mm (A–C); 0.5 mm (D–F); 0.25 mm (G, H)." captionText-3="FIGURE 4. Aniptumnus bijoyi sp. nov., paratype, female, 8.2 by 5.1 mm (ZRC 2022.0137), Kochi, Kerala, SW India. A, detached pleon, external view; B, thoracic sternum and vulvae, ventral view. Scale bars: 1.0 mm (A, B)." figureDoi-0="http://doi.org/10.5281/zenodo.7385090" figureDoi-1="http://doi.org/10.5281/zenodo.7385093" figureDoi-2="http://doi.org/10.5281/zenodo.7385098" figureDoi-3="http://doi.org/10.5281/zenodo.7385100" httpUri-0="https://zenodo.org/record/7385090/files/figure.png" httpUri-1="https://zenodo.org/record/7385093/files/figure.png" httpUri-2="https://zenodo.org/record/7385098/files/figure.png" httpUri-3="https://zenodo.org/record/7385100/files/figure.png" pageId="2" pageNumber="263">Figs. 1–4</figureCitation>
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<paragraph id="8BF23681FFE6FF94FF2149F0C6CEF94F" blockId="2.[151,1437,1688,2039]" pageId="2" pageNumber="263">
<emphasis id="B939EA93FFE6FF94FF2149F0C30EF907" bold="true" box="[151,329,1688,1714]" pageId="2" pageNumber="263">Type material.</emphasis>
<materialsCitation id="3B253CDCFFE6FF94FEE249F0C6CEF94F" ID-GBIF-Occurrence="3979393311" collectingDate="2021-09-29" collectionCode="CUSAT, SBN, R" collectorName="P. H. Praved &amp; Hershey" country="India" county="Marine Science Boat Jetty" latitude="9.964114" location="University of Science" longLatPrecision="1" longitude="76.28244" municipality="School of Marine Sciences" pageId="2" pageNumber="263" specimenCount="1" stateProvince="Kerala" typeStatus="holotype">
<emphasis id="B939EA93FFE6FF94FEE249F0C3FCF907" box="[340,443,1688,1714]" italics="true" pageId="2" pageNumber="263">
<typeStatus id="54F68823FFE6FF94FEE249F0C3FCF907" box="[340,443,1688,1714]" pageId="2" pageNumber="263" type="holotype">Holotype</typeStatus>
</emphasis>
: male, 10.7 by 
<quantity id="4CB59B64FFE6FF94FDCF49F0C094F906" box="[633,723,1688,1715]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="2" pageNumber="263" unit="mm" value="7.1">7.1 mm</quantity>
(
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/ 
<collectionCode id="ED5CAE44FFE6FF94FCF849F0C1C2F907" box="[846,901,1688,1714]" pageId="2" pageNumber="263">SBN</collectionCode>
/HRJ/1), 
<collectingCounty id="62934E0DFFE6FF94FC5F49F0C75AF906" box="[1001,1309,1688,1715]" pageId="2" pageNumber="263">Marine Science Boat Jetty</collectingCounty>
, 
<collectingMunicipality id="6B96ACFBFFE6FF94FA9A49F0C31DF963" pageId="2" pageNumber="263">School of Marine Sciences</collectingMunicipality>
, Cochin 
<location id="8E92605AFFE6FF94FE7C49D4C08EF963" LSID="urn:lsid:plazi:treatment:03E48797FFE6FF9FFF214944C62CFACC:8E92605AFFE6FF94FE7C49D4C08EF963" box="[458,713,1724,1750]" country="India" county="Marine Science Boat Jetty" latitude="9.964114" longLatPrecision="1" longitude="76.28244" municipality="School of Marine Sciences" name="University of Science" pageId="2" pageNumber="263" stateProvince="Kerala">University of Science</location>
and 
<location id="8E92605AFFE6FF94FCBA49D4C1D6F963" LSID="urn:lsid:plazi:treatment:03E48797FFE6FF9FFF214944C62CFACC:8E92605AFFE6FF94FCBA49D4C1D6F963" box="[780,913,1724,1750]" country="India" county="Marine Science Boat Jetty" latitude="9.964114" longLatPrecision="1" longitude="76.28244" municipality="School of Marine Sciences" name="Technology" pageId="2" pageNumber="263" stateProvince="Kerala">Technology</location>
, 
<collectingCountry id="F35A7611FFE6FF94FC1449D4C1AEF963" box="[930,1001,1724,1750]" name="India" pageId="2" pageNumber="263">Kochi</collectingCountry>
, 
<collectingRegion id="4989F863FFE6FF94FC4D49D4C60FF963" box="[1019,1096,1724,1750]" country="India" name="Kerala" pageId="2" pageNumber="263">Kerala</collectingRegion>
, SW 
<collectingCountry id="F35A7611FFE6FF94FB2749D4C68AF963" box="[1169,1229,1724,1750]" name="India" pageId="2" pageNumber="263">India</collectingCountry>
, 
<geoCoordinate id="EE795046FFE6FF94FB6949D5C7DFF96D" box="[1247,1432,1725,1752]" degrees="9" direction="north" minutes="57" orientation="latitude" pageId="2" pageNumber="263" precision="1" seconds="50.8104" value="9.964114">9°57′50.8104″N</geoCoordinate>
, 
<geoCoordinate id="EE795046FFE6FF94FF214989C31CF949" box="[151,347,1761,1788]" degrees="76" direction="east" minutes="16" orientation="longitude" pageId="2" pageNumber="263" precision="1" seconds="56.7834" value="76.28244">76°16′56.7834″E</geoCoordinate>
, coll. 
<collectorName id="26B85357FFE6FF94FE284989C065F94F" box="[414,546,1760,1787]" pageId="2" pageNumber="263">P.H. Praved</collectorName>
&amp; N. 
<collectionCode id="ED5CAE44FFE6FF94FDD74989C030F94E" box="[609,631,1761,1787]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="263">R</collectionCode>
. 
<collectorName id="26B85357FFE6FF94FD374988C098F94F" box="[641,735,1760,1786]" pageId="2" pageNumber="263">Hershey</collectorName>
, 
<date id="FFF31041FFE6FF94FD5D4988C139F94F" box="[747,894,1760,1786]" pageId="2" pageNumber="263" value="2021-09-29">
<collectingDate id="EFB7E9A9FFE6FF94FD5D4988C139F94F" box="[747,894,1760,1786]" pageId="2" pageNumber="263" value="2021-09-29">29 Sep. 2021</collectingDate>
</date>
(fixed in 5% formalin).
</materialsCitation>
</paragraph>
<paragraph id="8BF23681FFE6FF94FF71486DC290F81B" blockId="2.[151,1437,1688,2039]" pageId="2" pageNumber="263">
<emphasis id="B939EA93FFE6FF94FF71486DC37EF8AB" box="[199,313,1797,1822]" italics="true" pageId="2" pageNumber="263">
<typeStatus id="54F68823FFE6FF94FF71486DC37EF8AB" box="[199,313,1797,1822]" pageId="2" pageNumber="263" type="paratype">Paratypes</typeStatus>
</emphasis>
: 
<specimenCount id="9D4BFD08FFE6FF94FEFF486CC3E4F8AB" box="[329,419,1796,1822]" pageId="2" pageNumber="263" type="male">6 males</specimenCount>
, 8.1 by 
<quantity id="4CB59B64FFE6FF94FE48486CC012F8AA" box="[510,597,1796,1823]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="2" pageNumber="263" unit="mm" value="5.9">5.9 mm</quantity>
to 10.5 by 
<quantity id="4CB59B64FFE6FF94FD6E486CC176F8AA" box="[728,817,1796,1823]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="2" pageNumber="263" unit="mm" value="7.1">7.1 mm</quantity>
, 
<materialsCitation id="3B253CDCFFE6FF94FC8B486CC010F8F7" ID-GBIF-Occurrence="3979393304" collectingDate="2021-09-29" collectionCode="CUSAT, SBN" collectorName="P. H. Praved &amp; Hershey" country="India" county="Marine Science Boat Jetty" latitude="9.964114" location="University of Science" longLatPrecision="1" longitude="76.28244" municipality="School of Marine Sciences" pageId="2" pageNumber="263" specimenCount="4" specimenCount-female="4" stateProvince="Kerala" typeStatus="paratype">
<specimenCount id="9D4BFD08FFE6FF94FC8B486CC1EAF8AB" box="[829,941,1796,1822]" pageId="2" pageNumber="263" type="female">4 females</specimenCount>
, 7.6 by 
<quantity id="4CB59B64FFE6FF94FBBE486CC618F8AA" box="[1032,1119,1796,1823]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="2" pageNumber="263" unit="mm" value="5.6">5.6 mm</quantity>
to 8.7 by 
<quantity id="4CB59B64FFE6FF94FB62486CC76CF8AA" box="[1236,1323,1796,1823]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="2" pageNumber="263" unit="mm" value="5.7">5.7 mm</quantity>
(
<collectionCode id="ED5CAE44FFE6FF94FA8C486CC7D3F8AB" box="[1338,1428,1796,1822]" pageId="2" pageNumber="263">CUSAT</collectionCode>
/ 
<collectionCode id="ED5CAE44FFE6FF94FF214840C288F8F7" box="[151,207,1832,1858]" pageId="2" pageNumber="263">SBN</collectionCode>
/HRJ/2–11), same data as holotype
</materialsCitation>
; 
<specimenCount id="9D4BFD08FFE6FF94FDD74840C0E8F8F7" box="[609,687,1832,1858]" pageId="2" pageNumber="263" type="male">1 male</specimenCount>
, 10.1 by 
<quantity id="4CB59B64FFE6FF94FCA24840C12BF8F6" box="[788,876,1832,1859]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="2" pageNumber="263" unit="mm" value="6.7">6.7 mm</quantity>
, 
<materialsCitation id="3B253CDCFFE6FF94FCC14840C315F8D3" ID-GBIF-Occurrence="3979393306" collectingDate="2021-09-29" collectionCode="ZRC" collectorName="P. H. Praved &amp; Hershey" country="India" county="Marine Science Boat Jetty" latitude="9.964114" location="University of Science" longLatPrecision="1" longitude="76.28244" municipality="School of Marine Sciences" pageId="2" pageNumber="263" specimenCode="ZRC 2022.0137" specimenCount="1" specimenCount-female="1" stateProvince="Kerala" typeStatus="paratype">
<specimenCount id="9D4BFD08FFE6FF94FCC14840C19CF8F7" box="[887,987,1832,1858]" pageId="2" pageNumber="263" type="female">1 female</specimenCount>
, 8.2 by 
<quantity id="4CB59B64FFE6FF94FB844840C6CFF8F6" box="[1074,1160,1832,1859]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="2" pageNumber="263" unit="mm" value="5.1">5.1 mm</quantity>
(
<specimenCode id="DBEB9EFAFFE6FF94FB204840C716F8F7" box="[1174,1361,1832,1858]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="263">ZRC 2022.0137</specimenCode>
), same data as holotype
</materialsCitation>
; 
<materialsCitation id="3B253CDCFFE6FF94FEE84824C294F81B" ID-GBIF-Occurrence="3979393307" collectingDate="2019-02" collectionCode="R" country="India" county="School of Marine Sciences" location="Technology" municipality="University of Science" pageId="2" pageNumber="263" specimenCode="ZRC 2019.1091" specimenCount="1" specimenCount-male="1" stateProvince="Kerala" typeStatus="paratype">
<specimenCount id="9D4BFD08FFE6FF94FEE84824C3EBF8D3" box="[350,428,1868,1894]" pageId="2" pageNumber="263" type="male">1 male</specimenCount>
, 5.0 by 
<quantity id="4CB59B64FFE6FF94FDB04824C01AF8D2" box="[518,605,1868,1895]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="2" pageNumber="263" unit="mm" value="3.2">3.2 mm</quantity>
(
<specimenCode id="DBEB9EFAFFE6FF94FDDD4824C161F8D3" box="[619,806,1868,1894]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="263">ZRC 2019.1091</specimenCode>
), intertidal rocky area with algae, just outside 
<collectingCounty id="62934E0DFFE6FF94FA864824C314F83F" pageId="2" pageNumber="263">School of Marine Sciences</collectingCounty>
, Cochin 
<collectingMunicipality id="6B96ACFBFFE6FF94FE024818C0E2F83F" box="[436,677,1904,1930]" pageId="2" pageNumber="263">University of Science</collectingMunicipality>
and 
<location id="8E92605AFFE6FF94FD6C4818C118F83F" LSID="urn:lsid:plazi:treatment:03E48797FFE6FF9FFF214944C62CFACC:8E92605AFFE6FF94FD6C4818C118F83F" box="[730,863,1904,1930]" country="India" county="School of Marine Sciences" municipality="University of Science" name="Technology" pageId="2" pageNumber="263" stateProvince="Kerala">Technology</location>
, 
<collectingCountry id="F35A7611FFE6FF94FCDF4818C1F7F83F" box="[873,944,1904,1930]" name="India" pageId="2" pageNumber="263">Kochi</collectingCountry>
, 
<collectingRegion id="4989F863FFE6FF94FC0C4818C640F83F" box="[954,1031,1904,1930]" country="India" name="Kerala" pageId="2" pageNumber="263">Kerala</collectingRegion>
, SW 
<collectingCountry id="F35A7611FFE6FF94FBF44818C639F83F" box="[1090,1150,1904,1930]" name="India" pageId="2" pageNumber="263">India</collectingCountry>
, coll. N. 
<collectionCode id="ED5CAE44FFE6FF94FB6D4819C6B6F83E" box="[1243,1265,1905,1931]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="2" pageNumber="263">R</collectionCode>
. 
<location id="8E92605AFFE6FF94FB4D4818C71FF83F" LSID="urn:lsid:plazi:treatment:03E48797FFE6FF9FFF214944C62CFACC:8E92605AFFE6FF94FB4D4818C71FF83F" box="[1275,1368,1904,1930]" country="India" county="School of Marine Sciences" municipality="University of Science" name="Hershey" pageId="2" pageNumber="263" stateProvince="Kerala">Hershey</location>
, 
<date id="FFF31041FFE6FF94FADD4818C294F81B" pageId="2" pageNumber="263" value="2019-02">
<collectingDate id="EFB7E9A9FFE6FF94FADD4818C294F81B" pageId="2" pageNumber="263" value="2019-02">Feb. 2019</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8BF23681FFE6FF95FF7148D0C1FDFE3C" blockId="2.[151,1437,1688,2039]" lastBlockId="3.[151,1437,151,969]" lastPageId="3" lastPageNumber="264" pageId="2" pageNumber="263">
<emphasis id="B939EA93FFE6FF94FF7148D0C396F867" bold="true" box="[199,465,1976,2002]" pageId="2" pageNumber="263">Comparative material</emphasis>
. 
<emphasis id="B939EA93FFE6FF94FE5648D0C1A8F867" bold="true" box="[480,1007,1975,2002]" pageId="2" pageNumber="263">
<taxonomicName id="4C4D4D02FFE6FF94FE5648D0C1ADF867" authority="(De Man, 1895)" baseAuthorityName="De Man" baseAuthorityYear="1895" box="[480,1002,1975,2002]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="263" phylum="Arthropoda" rank="species" species="quadridentatus">
<emphasis id="B939EA93FFE6FF94FE5648D0C161F867" bold="true" box="[480,806,1975,2002]" italics="true" pageId="2" pageNumber="263">Aniptumnus quadridentatus</emphasis>
(
<bibRefCitation id="EFDC4B70FFE6FF94FC8048D0C1A5F867" author="De Man, J. G." box="[822,994,1976,2002]" pageId="2" pageNumber="263" pagination="485 - 609" refId="ref7241" refString="De Man, J. G. (1895) Bericht uber die von Herrn Schiffscapitan Storm zur Atjeh, an den westlichen Kusten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8 (4), 485 - 609. https: // doi. org / 10.5962 / bhl. title. 16084" type="journal article" year="1895">De Man, 1895</bibRefCitation>
)
</taxonomicName>
.
</emphasis>
<materialsCitation id="3B253CDCFFE6FF94FC4E48D0C32BF842" ID-GBIF-Occurrence="3979393308" country="Singapore" location="Singapore" pageId="2" pageNumber="263" specimenCount="30" specimenCount-male="30">
<emphasis id="B939EA93FFE6FF94FC4E48D0C63DF867" bold="true" box="[1016,1146,1976,2002]" pageId="2" pageNumber="263">
<collectingCountry id="F35A7611FFE6FF94FC4E48D0C632F867" box="[1016,1141,1976,2002]" name="Singapore" pageId="2" pageNumber="263">Singapore</collectingCountry>
:
</emphasis>
<specimenCount id="9D4BFD08FFE6FF94FB3448D0C6ADF867" box="[1154,1258,1976,2002]" pageId="2" pageNumber="263" type="male">30 males</specimenCount>
, 5.5 by 
<quantity id="4CB59B64FFE6FF94FAF048D0C7DAF866" box="[1350,1437,1976,2003]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="2" pageNumber="263" unit="mm" value="3.9">3.9 mm</quantity>
to 13.9 by 
<quantity id="4CB59B64FFE6FF94FEA448B4C32BF842" box="[274,364,2012,2039]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="2" pageNumber="263" unit="mm" value="9.5">9.5 mm</quantity>
</materialsCitation>
, 
<materialsCitation id="3B253CDCFFE6FF95FECE48B4C64DFF60" ID-GBIF-Occurrence="3979393305" collectingDate="2017-11-07" collectionCode="MD" collectorName="Mendoza &amp; I Iesa &amp; Safaruan" country="Singapore" lastPageId="3" lastPageNumber="264" location="Pulau Ubin" pageId="2" pageNumber="263" specimenCode="ZRC 2018.0639" specimenCount="18" specimenCount-female="18">
<specimenCount id="9D4BFD08FFE6FF94FECE48B4C3B0F843" box="[376,503,2012,2038]" pageId="2" pageNumber="263" type="female">18 females</specimenCount>
, 5.6 by 4.0 mm to 12.3 by 8.0 mm, 5 ovig. females, 8.2 by 
<quantity id="4CB59B64FFE6FF94FB2948B4C6B1F842" box="[1183,1270,2012,2039]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="2" pageNumber="263" unit="mm" value="5.8">5.8 mm</quantity>
to 10.4 by 7.0 mm (
<specimenCode id="DBEB9EFAFFE7FF95FF674FFFC3CBFF04" box="[209,396,151,177]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="264">ZRC 2018.0639</specimenCode>
), off 
<location id="8E92605AFFE7FF95FE734FFFC001FF04" LSID="urn:lsid:plazi:treatment:03E48797FFE6FF9FFF214944C62CFACC:8E92605AFFE7FF95FE734FFFC001FF04" box="[453,582,151,177]" country="Singapore" name="Pulau Ubin" pageId="3" pageNumber="264">Pulau Ubin</location>
, floating fish farm (kelong FC12E), among clumps of 
<taxonomicName id="4C4D4D02FFE7FF95FB104FFFC77EFF04" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[1190,1337,151,177]" class="Insecta" family="Lasiocampidae" genus="Perna" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="viridis">
<emphasis id="B939EA93FFE7FF95FB104FFFC77EFF04" box="[1190,1337,151,177]" italics="true" pageId="3" pageNumber="264">Perna viridis</emphasis>
</taxonomicName>
attached to submerged ropes, coll. JCE 
<collectorName id="26B85357FFE7FF95FE434FD3C027FF60" box="[501,608,187,213]" pageId="3" pageNumber="264">Mendoza</collectorName>
, 
<collectorName id="26B85357FFE7FF95FDDA4FD3C0ECFF60" box="[620,683,187,213]" pageId="3" pageNumber="264">I Iesa</collectorName>
&amp; 
<collectionCode id="ED5CAE44FFE7FF95FD644FD3C0B8FF60" box="[722,767,187,213]" country="Germany" name="Museum Donaueschingen" pageId="3" pageNumber="264">MD</collectionCode>
<collectorName id="26B85357FFE7FF95FCB14FD3C12AFF60" box="[775,877,187,213]" pageId="3" pageNumber="264">Safaruan</collectorName>
, 
<date id="FFF31041FFE7FF95FCCC4FD3C64DFF60" box="[890,1034,187,213]" pageId="3" pageNumber="264" value="2017-11-07">
<collectingDate id="EFB7E9A9FFE7FF95FCCC4FD3C64DFF60" box="[890,1034,187,213]" pageId="3" pageNumber="264" value="2017-11-07">7 Nov. 2017</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B253CDCFFE7FF95FBA14FD3C311FEA8" ID-GBIF-Occurrence="3979393309" collectionCode="ZRC" collectorName="FS Tan &amp; SC Lee &amp; Ong" country="Singapore" county="Changi Point" location="Teo" municipality="Paku Buoy" pageId="3" pageNumber="264" specimenCode="ZRC 2011" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D4BFD08FFE7FF95FBA14FD3C63BFF60" box="[1047,1148,187,213]" pageId="3" pageNumber="264" type="female">1 female</specimenCount>
, 14.0 by 
<quantity id="4CB59B64FFE7FF95FB514FD3C707FF60" box="[1255,1344,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="3" pageNumber="264" unit="mm" value="8.9">8.9 mm</quantity>
, 1 ovig. female, 10.7 by 
<quantity id="4CB59B64FFE7FF95FEFA4FB7C3E5FF4C" box="[332,418,223,249]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="3" pageNumber="264" unit="mm" value="7.3">7.3 mm</quantity>
(
<specimenCode id="DBEB9EFAFFE7FF95FE064FB7C06DFF4C" box="[432,554,223,249]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="264">ZRC 2011</specimenCode>
. 0683), 
<collectingCounty id="62934E0DFFE7FF95FD324FB7C15DFF4C" box="[644,794,223,249]" pageId="3" pageNumber="264">Changi Point</collectingCounty>
, on 
<collectingMunicipality id="6B96ACFBFFE7FF95FCFE4FB7C183FF4C" box="[840,964,223,249]" pageId="3" pageNumber="264">Paku Buoy</collectingMunicipality>
(fouling), coll. SLM 
<location id="8E92605AFFE7FF95FB024FB7C6A7FF4C" LSID="urn:lsid:plazi:treatment:03E48797FFE6FF9FFF214944C62CFACC:8E92605AFFE7FF95FB024FB7C6A7FF4C" box="[1204,1248,223,249]" country="Singapore" county="Changi Point" municipality="Paku Buoy" name="Teo" pageId="3" pageNumber="264">Teo</location>
, 
<collectorName id="26B85357FFE7FF95FB5A4FB7C77AFF4C" box="[1260,1341,223,249]" pageId="3" pageNumber="264">FS Tan</collectorName>
, 
<collectorName id="26B85357FFE7FF95FAFF4FB7C7DBFF4C" box="[1353,1436,223,249]" pageId="3" pageNumber="264">SC Lee</collectorName>
&amp; JL 
<collectorName id="26B85357FFE7FF95FF6C4E6BC34AFEA8" box="[218,269,259,285]" pageId="3" pageNumber="264">Ong</collectorName>
, 2010
</materialsCitation>
; 
<specimenCount id="9D4BFD08FFE7FF95FED44E6BC3FAFEA8" box="[354,445,259,285]" pageId="3" pageNumber="264" type="male">6 males</specimenCount>
, 5.8 by 4.0 mm to 11.1 by 
<quantity id="4CB59B64FFE7FF95FD414E6BC116FEA8" box="[759,849,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="3" pageNumber="264" unit="mm" value="7.4">7.4 mm</quantity>
, 
<materialsCitation id="3B253CDCFFE7FF95FCE84E6BC6CDFEF4" ID-GBIF-Occurrence="3979393303" collectingDate="2020-09-03" collectionCode="ZRC" collectorName="BY Lee" country="Singapore" location="Pulau Ubin" pageId="3" pageNumber="264" specimenCode="ZRC 2021.0421" specimenCount="3" specimenCount-female="3">
<specimenCount id="9D4BFD08FFE7FF95FCE84E6BC188FEA8" box="[862,975,259,285]" pageId="3" pageNumber="264" type="female">3 females</specimenCount>
, 6.1 by 
<quantity id="4CB59B64FFE7FF95FB9B4E6BC6C2FEA8" box="[1069,1157,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="3" pageNumber="264" unit="mm" value="4.2">4.2 mm</quantity>
to 7.7 by 
<quantity id="4CB59B64FFE7FF95FB4A4E6AC713FEA8" box="[1276,1364,258,285]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="3" pageNumber="264" unit="mm" value="5.1">5.1 mm</quantity>
(
<specimenCode id="DBEB9EFAFFE7FF95FAD24E6BC355FEF4" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="264">ZRC 2021.0421</specimenCode>
), off 
<location id="8E92605AFFE7FF95FEF84E4FC396FEF4" LSID="urn:lsid:plazi:treatment:03E48797FFE6FF9FFF214944C62CFACC:8E92605AFFE7FF95FEF84E4FC396FEF4" box="[334,465,295,321]" country="Singapore" name="Pulau Ubin" pageId="3" pageNumber="264">Pulau Ubin</location>
, floating fish farm (kelong E63C), coll. 
<collectorName id="26B85357FFE7FF95FC2F4E4FC1B4FEF4" box="[921,1011,295,321]" pageId="3" pageNumber="264">BY Lee</collectorName>
, 
<date id="FFF31041FFE7FF95FC494E4FC6CDFEF4" box="[1023,1162,295,321]" pageId="3" pageNumber="264" value="2020-09-03">
<collectingDate id="EFB7E9A9FFE7FF95FC494E4FC6CDFEF4" box="[1023,1162,295,321]" pageId="3" pageNumber="264" value="2020-09-03">3 Sep. 2020</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="3B253CDCFFE7FF95FB234E4EC3AFFED0" ID-GBIF-Occurrence="3979393302" country="Malaysia" location="Malaysia" pageId="3" pageNumber="264" specimenCount="8" specimenCount-male="8">
<emphasis id="B939EA93FFE7FF95FB234E4EC74CFEF5" bold="true" box="[1173,1291,294,320]" pageId="3" pageNumber="264">
<collectingCountry id="F35A7611FFE7FF95FB234E4EC741FEF5" box="[1173,1286,294,320]" name="Malaysia" pageId="3" pageNumber="264">Malaysia</collectingCountry>
:
</emphasis>
<specimenCount id="9D4BFD08FFE7FF95FAA54E4FC72AFEF4" box="[1299,1389,295,321]" pageId="3" pageNumber="264" type="male">8 males</specimenCount>
, 9.5 by 
<quantity id="4CB59B64FFE7FF95FF0D4E22C356FED0" box="[187,273,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="3" pageNumber="264" unit="mm" value="6.5">6.5 mm</quantity>
to 11.0 by 
<quantity id="4CB59B64FFE7FF95FE264E23C3AFFED0" box="[400,488,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="3" pageNumber="264" unit="mm" value="7.3">7.3 mm</quantity>
</materialsCitation>
, 
<materialsCitation id="3B253CDCFFE7FF95FE424E23C1F1FE3C" ID-GBIF-Occurrence="3979393310" collectionCode="ZRC" collectorName="Lumut &amp; H. Singh" country="Malaysia" location="Perak" pageId="3" pageNumber="264" specimenCode="ZRC 1992.10966, 10978" specimenCount="3" specimenCount-female="3" stateProvince="Perak">
<specimenCount id="9D4BFD08FFE7FF95FE424E23C024FED0" box="[500,611,331,357]" pageId="3" pageNumber="264" type="female">3 females</specimenCount>
, 8.9 by 
<quantity id="4CB59B64FFE7FF95FD0A4E22C154FED0" box="[700,787,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="3" pageNumber="264" unit="mm" value="5.6">5.6 mm</quantity>
to 9.7 by 
<quantity id="4CB59B64FFE7FF95FC334E23C19AFED0" box="[901,989,331,357]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="3" pageNumber="264" unit="mm" value="6.2">6.2 mm</quantity>
, 2 ovig. females, 8.6 by 
<quantity id="4CB59B64FFE7FF95FB444E22C70DFED0" box="[1266,1354,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="3" pageNumber="264" unit="mm" value="5.8">5.8 mm</quantity>
, 9.3 by 6.0 mm (
<specimenCode id="DBEB9EFAFFE7FF95FF4D4E07C383FE3C" box="[251,452,367,393]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="264">ZRC 1992.10966</specimenCode>
– 
<specimenCode id="DBEB9EFAFFE7FF95FE654E07C05FFE3C" box="[467,536,367,393]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="264">10978</specimenCode>
), 
<collectingRegion id="4989F863FFE7FF95FD9D4E07C029FE3C" box="[555,622,367,393]" country="Malaysia" name="Perak" pageId="3" pageNumber="264">Perak</collectingRegion>
, 
<collectorName id="26B85357FFE7FF95FDCF4E07C080FE3C" box="[633,711,367,393]" pageId="3" pageNumber="264">Lumut</collectorName>
, coll. 
<collectorName id="26B85357FFE7FF95FCBF4E07C137FE3C" box="[777,880,367,393]" pageId="3" pageNumber="264">H. Singh</collectorName>
, 1989
</materialsCitation>
.
</paragraph>
<paragraph id="8BF23681FFE7FF95FF714EFBC346FE40" blockId="3.[151,1437,151,969]" pageId="3" pageNumber="264">
<emphasis id="B939EA93FFE7FF95FF714EFBC096FE19" bold="true" box="[199,721,402,429]" pageId="3" pageNumber="264">
<taxonomicName id="4C4D4D02FFE7FF95FF714EFBC08BFE19" authority="Ng &amp; Clark, 2008" authorityName="Ng &amp; Clark" authorityYear="2008" box="[199,716,402,429]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="vietnamicus">
<emphasis id="B939EA93FFE7FF95FF714EFBC3AEFE18" bold="true" box="[199,489,403,429]" italics="true" pageId="3" pageNumber="264">Aniptumnus vietnamicus</emphasis>
<bibRefCitation id="EFDC4B70FFE7FF95FE444EFBC08BFE19" author="Ng, P. K. L. &amp; Clark, P. F." box="[498,716,402,429]" pageId="3" pageNumber="264" pagination="885 - 912" refId="ref7776" refString="Ng, P. K. L. &amp; Clark, P. F. (2008) A revision of Latopilumnus Turkay &amp; Schuhmacher, 1985, and Aniptumnus Ng, 2002 (Crustacea: Decapoda: Brachyura: Pilumnidae) with larval comparisons. Journal of Natural History, 42 (9), 885 - 912. https: // doi. org / 10.1080 / 00222930701869422" type="journal article" year="2008">Ng &amp; Clark, 2008</bibRefCitation>
</taxonomicName>
.
</emphasis>
<materialsCitation id="3B253CDCFFE7FF95FD6C4EFAC346FE40" ID-GBIF-Occurrence="3979393312" collectingDate="1966-04-10" collectionCode="ZRC, R" collectorName="Nhatrang" country="Vietnam" county="Marine Science Boat Jetty" latitude="9.964114" location="Nhatrang Bay" longLatPrecision="1" longitude="76.28244" municipality="Serene" pageId="3" pageNumber="264" specimenCount="2" stateProvince="Kerala" typeStatus="holotype">
<emphasis id="B939EA93FFE7FF95FD6C4EFAC10EFE19" bold="true" box="[730,841,402,428]" pageId="3" pageNumber="264">
<collectingCountry id="F35A7611FFE7FF95FD6C4EFAC103FE19" box="[730,836,402,428]" name="Vietnam" pageId="3" pageNumber="264">Vietnam</collectingCountry>
:
</emphasis>
<typeStatus id="54F68823FFE7FF95FCE74EFBC1FBFE18" box="[849,956,403,429]" pageId="3" pageNumber="264" type="holotype">Holotype</typeStatus>
, male, 8.3 by 
<quantity id="4CB59B64FFE7FF95FBE94EFBC6F1FE18" box="[1119,1206,403,429]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="3" pageNumber="264" unit="mm" value="6.1">6.1 mm</quantity>
(
<collectionCode id="ED5CAE44FFE7FF95FB704EFBC6B9FE18" box="[1222,1278,403,429]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="264">ZRC</collectionCode>
1970.1.10.1), 
<location id="8E92605AFFE7FF95FF214EDFC370FE64" LSID="urn:lsid:plazi:treatment:03E48797FFE6FF9FFF214944C62CFACC:8E92605AFFE7FF95FF214EDFC370FE64" box="[151,311,439,465]" country="Vietnam" municipality="Serene" name="Nhatrang Bay" pageId="3" pageNumber="264">
<collectorName id="26B85357FFE7FF95FF214EDFC2B8FE64" box="[151,255,439,465]" pageId="3" pageNumber="264">Nhatrang</collectorName>
Bay
</location>
, coll. 
<collectionCode id="ED5CAE44FFE7FF95FEC84EDFC3D3FE64" box="[382,404,439,465]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="264">R</collectionCode>
. 
<collectingMunicipality id="6B96ACFBFFE7FF95FE174EDFC3B7FE64" box="[417,496,439,465]" pageId="3" pageNumber="264">Serène</collectingMunicipality>
, 
<date id="FFF31041FFE7FF95FE4B4EDFC0D1FE64" box="[509,662,439,465]" pageId="3" pageNumber="264" value="1966-04-10">
<collectingDate id="EFB7E9A9FFE7FF95FE4B4EDFC0D1FE64" box="[509,662,439,465]" pageId="3" pageNumber="264" value="1966-04-10">10 Apr. 1966</collectingDate>
</date>
; 
<typeStatus id="54F68823FFE7FF95FD124EDFC140FE64" box="[676,775,439,465]" pageId="3" pageNumber="264" type="paratype">paratype</typeStatus>
, female, 9.6 by 
<quantity id="4CB59B64FFE7FF95FC744EDFC65DFE64" box="[962,1050,439,465]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="3" pageNumber="264" unit="mm" value="6.6">6.6 mm</quantity>
(
<collectionCode id="ED5CAE44FFE7FF95FB9D4EDFC624FE64" box="[1067,1123,439,465]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="264">ZRC</collectionCode>
1970.1.10.2), same data as holotype.
</materialsCitation>
</paragraph>
<paragraph id="8BF23681FFE7FF95FF714E97C02BFD88" blockId="3.[151,1437,151,969]" pageId="3" pageNumber="264">
<emphasis id="B939EA93FFE7FF95FF714E97C3EBFDAD" bold="true" box="[199,428,510,537]" pageId="3" pageNumber="264">
Type locality. 
<collectingCountry id="F35A7611FFE7FF95FEDA4E96C3EBFDAD" box="[364,428,510,536]" name="India" pageId="3" pageNumber="264">India</collectingCountry>
</emphasis>
: Kochi, 
<collectingRegion id="4989F863FFE7FF95FDBB4E97C01CFDAC" box="[525,603,511,537]" country="India" name="Kerala" pageId="3" pageNumber="264">Kerala</collectingRegion>
; Kollam-Kottapuram National Waterway No. 3, near Marine Science Boat Jetty, CUSAT-School of Marine Sciences.
</paragraph>
</subSubSection>
<subSubSection id="C357650AFFE7FF95FF714D2EC61FFC7C" pageId="3" pageNumber="264" type="diagnosis">
<paragraph id="8BF23681FFE7FF95FF714D2EC61FFC7C" blockId="3.[151,1437,151,969]" pageId="3" pageNumber="264">
<emphasis id="B939EA93FFE7FF95FF714D2EC305FDD5" bold="true" box="[199,322,582,608]" pageId="3" pageNumber="264">Diagnosis.</emphasis>
Carapace subhexagonal, broader than long (CW/CL: 1.4–1.5); dorsal surface finely granular with small patches of granules on gastric, branchial regions, sparsely setose with tufts of longer setae on front and on granular patches of gastric, branchial regions. Anterolateral, posterolateral margins subequal in length. First tooth of carapace exorbital margin lobiform, nearly flat, second tooth broadly triangular, denticulate, third and fourth teeth acutely triangular, slightly curved. Orbits without fissures, notch at junction of supra- and infraorbital margins absent. Third maxilliped merus distolateral angle produced, rounded. Chelipeds unequal; major chela external surface sparsely setose, finely granular with scattered larger granules; minor chela more densely setose, with rows of sharp conical granules. P5 basis-ischium with conical tubercles on disto-posterior margin forming comb-like structure; merus posterior margin with sharp conical tubercles proximally. G1 slender, sinuous, distal tip with short, narrow, scoop-shaped apical lobe. G2 small, one-fifth length of G1, terminal, subterminal segments uniformly curving mesially. Penis emerging from anterior tip of coxosternal condyle of P5 coxa.
</paragraph>
</subSubSection>
<caption id="DF326609FFE7FF95FF2148F6C3EBF86F" ID-DOI="http://doi.org/10.5281/zenodo.7385090" ID-Zenodo-Dep="7385090" httpUri="https://zenodo.org/record/7385090/files/figure.png" pageId="3" pageNumber="264" startId="3.[151,250,1950,1974]" targetBox="[223,1366,993,1927]" targetPageId="3">
<paragraph id="8BF23681FFE7FF95FF2148F6C3EBF86F" blockId="3.[151,1437,1950,2010]" pageId="3" pageNumber="264">
<emphasis id="B939EA93FFE7FF95FF2148F6C349F803" bold="true" box="[151,270,1950,1974]" pageId="3" pageNumber="264">FIGURE 1</emphasis>
. 
<taxonomicName id="4C4D4D02FFE7FF95FEAD48F6C391F803" authority="Hari &amp; Hershey &amp; Mendoza, 2022" authorityName="Hari &amp; Hershey &amp; Mendoza" authorityYear="2022" box="[283,470,1950,1974]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="bijoyi" status="sp. nov.">
<emphasis id="B939EA93FFE7FF95FEAD48F6C391F803" box="[283,470,1950,1974]" italics="true" pageId="3" pageNumber="264">Aniptumnus bijoyi</emphasis>
</taxonomicName>
<emphasis id="B939EA93FFE7FF95FE6B48F6C069F803" bold="true" box="[477,558,1950,1974]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="A20A57E8FFE7FF95FE6B48F6C069F803" box="[477,558,1950,1974]" pageId="3" pageNumber="264" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype, male, 10.1 by 6.7 mm (ZRC 2022.0137), Kochi, Kerala, SW India. Dorsal habitus and live coloration.
</paragraph>
</caption>
<subSubSection id="C357650AFFE0FF91FF714FFEC69BF953" lastPageId="7" lastPageNumber="268" pageId="4" pageNumber="265" type="description">
<paragraph id="8BF23681FFE0FF92FF714FFEC1DAFDD4" blockId="4.[151,1437,150,1941]" pageId="4" pageNumber="265">
<emphasis id="B939EA93FFE0FF92FF714FFEC068FF04" bold="true" box="[199,559,150,177]" pageId="4" pageNumber="265">
Description of male 
<typeStatus id="54F68823FFE0FF92FE774FFFC06DFF04" box="[449,554,151,177]" pageId="4" pageNumber="265" type="holotype">holotype</typeStatus>
.
</emphasis>
Carapace (
<figureCitation id="13762A04FFE0FF92FD024FFFC143FF04" box="[692,772,151,177]" captionStart="FIGURE 1" captionStartId="3.[151,250,1950,1974]" captionTargetBox="[223,1366,993,1927]" captionTargetId="figure-382@3.[222,1366,993,1927]" captionTargetPageId="3" captionText="FIGURE 1. Aniptumnus bijoyi sp. nov., paratype, male, 10.1 by 6.7 mm (ZRC 2022.0137), Kochi, Kerala, SW India. Dorsal habitus and live coloration." figureDoi="http://doi.org/10.5281/zenodo.7385090" httpUri="https://zenodo.org/record/7385090/files/figure.png" pageId="4" pageNumber="265">Figs. 1</figureCitation>
, 
<figureCitation id="13762A04FFE0FF92FCA34FFFC17EFF04" box="[789,825,151,177]" captionStart="FIGURE 2" captionStartId="5.[151,250,1891,1915]" captionTargetBox="[151,1435,181,1867]" captionTargetId="figure-20@5.[151,1435,181,1867]" captionTargetPageId="5" captionText="FIGURE 2. Aniptumnus bijoyi sp. nov., holotype, male, 10.7 by 7.1 mm (CUSAT/SBN/HRJ/1), Kochi, Kerala, SW India. A, carapace, dorsal view; B, left third maxilliped, external view; cephalothorax and chelipeds, frontal view; D, anterior thoracic sternum and telson, ventral view; E, thoracic sternum and sternopleonal cavity, ventral view; F, pleon, ventral view; G, pleonal somites 1–3, posteroventral view. Scale bars: 2.0 mm (A, C); 1.0 mm (B, D–G)." figureDoi="http://doi.org/10.5281/zenodo.7385093" httpUri="https://zenodo.org/record/7385093/files/figure.png" pageId="4" pageNumber="265">2A</figureCitation>
) transversely subhexagonal, broader than long (CW/ CL = 1.5), dorsal regions poorly defined, dorsal surface convex longitudinally and transversely, generally smooth with patches of fine granules on gastric, branchial regions, sparsely setose, most setae aggregated around granular areas. Front moderately projecting, slightly deflexed ventrally, bilobed, lobes divided by median notch, frontal margins uniformly, moderately convex without any distinct lateral lobe or tooth, lined with small granules, with poorly defined row of long setae submarginally. Supraorbital margin concave in dorsal view, granular, without fissures; junction with infraorbital margin without notch. Anterolateral and posterolateral margins similar in length; anterolateral margin granular, sparsely setose, divided into 4 teeth; exorbital angle low, not prominently projecting, separated from first tooth by shallow concavity; first tooth wide, flat or weakly convex lobe, separated from second tooth by small notch; second tooth broadly subtriangular, posterior margin much longer than anterior, apex directed toward anterior; third, fourth teeth similar, acute, apices pointed obliquely; posterolateral margin nearly straight, finely granulate, convergent posteriorly; posterior margin slightly convex, with raised row of fine round granules. Suborbital, sub-branchial and pterygostomial regions finely granular.
</paragraph>
<paragraph id="8BF23681FFE0FF92FF714D03C30EFC7C" blockId="4.[151,1437,150,1941]" pageId="4" pageNumber="265">
Orbits (
<figureCitation id="13762A04FFE0FF92FEA94D03C33FFD30" box="[287,376,619,645]" captionStart="FIGURE 2" captionStartId="5.[151,250,1891,1915]" captionTargetBox="[151,1435,181,1867]" captionTargetId="figure-20@5.[151,1435,181,1867]" captionTargetPageId="5" captionText="FIGURE 2. Aniptumnus bijoyi sp. nov., holotype, male, 10.7 by 7.1 mm (CUSAT/SBN/HRJ/1), Kochi, Kerala, SW India. A, carapace, dorsal view; B, left third maxilliped, external view; cephalothorax and chelipeds, frontal view; D, anterior thoracic sternum and telson, ventral view; E, thoracic sternum and sternopleonal cavity, ventral view; F, pleon, ventral view; G, pleonal somites 1–3, posteroventral view. Scale bars: 2.0 mm (A, C); 1.0 mm (B, D–G)." figureDoi="http://doi.org/10.5281/zenodo.7385093" httpUri="https://zenodo.org/record/7385093/files/figure.png" pageId="4" pageNumber="265">Fig. 2C</figureCitation>
) transversely ovate; eyes large, filling entire orbital cavity, ocular peduncle stout, unarmed, corneas pigmented. Infraorbital margin granular, without fissures, mesial suborbital tooth absent. Antennular fossae subrectangular, antennules robust, folding transversely and obliquely. Basal antennal article short, subrectangular, filling orbital hiatus; flagellum long, exceeding orbit. Proepistome narrow, smooth. Central region of posterior border of epistome projecting medially, notched laterally. Endostome with prominent oblique ridge on either side. Third maxillipeds (mxp3; 
<figureCitation id="13762A04FFE0FF92FE704C77C05AFC8C" box="[454,541,799,825]" captionStart="FIGURE 2" captionStartId="5.[151,250,1891,1915]" captionTargetBox="[151,1435,181,1867]" captionTargetId="figure-20@5.[151,1435,181,1867]" captionTargetPageId="5" captionText="FIGURE 2. Aniptumnus bijoyi sp. nov., holotype, male, 10.7 by 7.1 mm (CUSAT/SBN/HRJ/1), Kochi, Kerala, SW India. A, carapace, dorsal view; B, left third maxilliped, external view; cephalothorax and chelipeds, frontal view; D, anterior thoracic sternum and telson, ventral view; E, thoracic sternum and sternopleonal cavity, ventral view; F, pleon, ventral view; G, pleonal somites 1–3, posteroventral view. Scale bars: 2.0 mm (A, C); 1.0 mm (B, D–G)." figureDoi="http://doi.org/10.5281/zenodo.7385093" httpUri="https://zenodo.org/record/7385093/files/figure.png" pageId="4" pageNumber="265">Fig. 2B</figureCitation>
) finely granular externally, sparsely setose; palps thick, subcylindrical; merus subquadrate, distolateral angle projecting, rounded; ischium subrectangular, about 1.5 times as long as merus, with submedian sulcus, submarginal row of long setae mesially; basis subtriangular, separated from ischium by clear suture; exopod elongate, not tapering, distal tip reaching to three-quarters lateral margin of merus, flagellum long, well developed.
</paragraph>
<paragraph id="8BF23681FFE0FF92FF714CBBC704FAB8" blockId="4.[151,1437,150,1941]" pageId="4" pageNumber="265">
Male thoracic sternum (
<figureCitation id="13762A04FFE0FF92FE684CBBC01BFC58" box="[478,604,979,1005]" captionStart="FIGURE 2" captionStartId="5.[151,250,1891,1915]" captionTargetBox="[151,1435,181,1867]" captionTargetId="figure-20@5.[151,1435,181,1867]" captionTargetPageId="5" captionText="FIGURE 2. Aniptumnus bijoyi sp. nov., holotype, male, 10.7 by 7.1 mm (CUSAT/SBN/HRJ/1), Kochi, Kerala, SW India. A, carapace, dorsal view; B, left third maxilliped, external view; cephalothorax and chelipeds, frontal view; D, anterior thoracic sternum and telson, ventral view; E, thoracic sternum and sternopleonal cavity, ventral view; F, pleon, ventral view; G, pleonal somites 1–3, posteroventral view. Scale bars: 2.0 mm (A, C); 1.0 mm (B, D–G)." figureDoi="http://doi.org/10.5281/zenodo.7385093" httpUri="https://zenodo.org/record/7385093/files/figure.png" pageId="4" pageNumber="265">Fig. 2D, E</figureCitation>
) moderately broad, surface smooth, punctate. Sternites 1, 2 completely fused, forming broad triangle abutting ischia of third maxillipeds. Sternites 2, 3 separated by distinct suture (suture 2/3); sternites 3, 4 almost completely fused, demarcated only laterally by notches that continue mesially as shallow groove. Sutures 4/5, 5/6 interrupted medially. Sutures 6/7, 7/8 complete. Sternopleonal cavity (
<figureCitation id="13762A04FFE0FF92FB134B57C6B0FBEC" box="[1189,1271,1087,1113]" captionStart="FIGURE 2" captionStartId="5.[151,250,1891,1915]" captionTargetBox="[151,1435,181,1867]" captionTargetId="figure-20@5.[151,1435,181,1867]" captionTargetPageId="5" captionText="FIGURE 2. Aniptumnus bijoyi sp. nov., holotype, male, 10.7 by 7.1 mm (CUSAT/SBN/HRJ/1), Kochi, Kerala, SW India. A, carapace, dorsal view; B, left third maxilliped, external view; cephalothorax and chelipeds, frontal view; D, anterior thoracic sternum and telson, ventral view; E, thoracic sternum and sternopleonal cavity, ventral view; F, pleon, ventral view; G, pleonal somites 1–3, posteroventral view. Scale bars: 2.0 mm (A, C); 1.0 mm (B, D–G)." figureDoi="http://doi.org/10.5281/zenodo.7385093" httpUri="https://zenodo.org/record/7385093/files/figure.png" pageId="4" pageNumber="265">Fig. 2E</figureCitation>
) deep, anterior tip on sternite 4, just reaching imaginary transverse line formed by coxo-sternal condyles of P1 coxae. Tubercle of sternopleonal lock (press-button) anteriorly positioned on sternite 5, closer to suture with sternite 4. Median line complete at level of sternite 7, 8 only; short portion visible on posterior part of sternite 4 within sternopleonal cavity. Small, median, triangular decalcified area at junction between sternite 6, 7. Sternite 8 (
<figureCitation id="13762A04FFE0FF92FBC04BA7C688FB5C" box="[1142,1231,1231,1257]" captionStart="FIGURE 2" captionStartId="5.[151,250,1891,1915]" captionTargetBox="[151,1435,181,1867]" captionTargetId="figure-20@5.[151,1435,181,1867]" captionTargetPageId="5" captionText="FIGURE 2. Aniptumnus bijoyi sp. nov., holotype, male, 10.7 by 7.1 mm (CUSAT/SBN/HRJ/1), Kochi, Kerala, SW India. A, carapace, dorsal view; B, left third maxilliped, external view; cephalothorax and chelipeds, frontal view; D, anterior thoracic sternum and telson, ventral view; E, thoracic sternum and sternopleonal cavity, ventral view; F, pleon, ventral view; G, pleonal somites 1–3, posteroventral view. Scale bars: 2.0 mm (A, C); 1.0 mm (B, D–G)." figureDoi="http://doi.org/10.5281/zenodo.7385093" httpUri="https://zenodo.org/record/7385093/files/figure.png" pageId="4" pageNumber="265">Fig. 2G</figureCitation>
) not visible when pleon folded against thoracic sternum, positioned beneath the posterolateral corner of the pleonal somite 2.
</paragraph>
<paragraph id="8BF23681FFE0FF92FF714A7FC33BF9E4" blockId="4.[151,1437,150,1941]" pageId="4" pageNumber="265">
Chelipeds (P1; 
<figureCitation id="13762A04FFE0FF92FECE4A7FC381FA84" box="[376,454,1303,1329]" captionStart="FIGURE 1" captionStartId="3.[151,250,1950,1974]" captionTargetBox="[223,1366,993,1927]" captionTargetId="figure-382@3.[222,1366,993,1927]" captionTargetPageId="3" captionText="FIGURE 1. Aniptumnus bijoyi sp. nov., paratype, male, 10.1 by 6.7 mm (ZRC 2022.0137), Kochi, Kerala, SW India. Dorsal habitus and live coloration." figureDoi="http://doi.org/10.5281/zenodo.7385090" httpUri="https://zenodo.org/record/7385090/files/figure.png" pageId="4" pageNumber="265">Figs. 1</figureCitation>
, 
<figureCitation id="13762A04FFE0FF92FE604A7FC05CFA84" box="[470,539,1303,1329]" captionStart="FIGURE 3" captionStartId="6.[151,250,1891,1915]" captionTargetBox="[151,1435,181,1867]" captionTargetId="figure-21@6.[151,1435,181,1867]" captionTargetPageId="6" captionText="FIGURE 3. Aniptumnus bijoyi sp. nov., holotype, male, 10.7 by 7.1 mm (CUSAT/SBN/HRJ/1), Kochi, Kerala, SW India. A, minor cheliped, external view; B, major cheliped, external view; C, right P5, dorsal view; D, E, left G1 (D, pleonal/ventral view; E, mesiosternal/mesiodorsal view); F, left G2, sternal/dorsal view; G, H, distal tip of left G1 (G, pleonal/ventral view; H, mesiosternal/mesiodorsal view). Scale bars: 1.0 mm (A–C); 0.5 mm (D–F); 0.25 mm (G, H)." figureDoi="http://doi.org/10.5281/zenodo.7385098" httpUri="https://zenodo.org/record/7385098/files/figure.png" pageId="4" pageNumber="265">3A, B</figureCitation>
) distinctly heterochelous. Anterior margins of basis-ischium, merus lined with subconical granules. External surface of the carpus granular, inner distal angle projecting as sharp tooth bordered with large tubercles. External surface of major palm (
<figureCitation id="13762A04FFE0FF92FD414A37C109FACC" box="[759,846,1375,1401]" captionStart="FIGURE 3" captionStartId="6.[151,250,1891,1915]" captionTargetBox="[151,1435,181,1867]" captionTargetId="figure-21@6.[151,1435,181,1867]" captionTargetPageId="6" captionText="FIGURE 3. Aniptumnus bijoyi sp. nov., holotype, male, 10.7 by 7.1 mm (CUSAT/SBN/HRJ/1), Kochi, Kerala, SW India. A, minor cheliped, external view; B, major cheliped, external view; C, right P5, dorsal view; D, E, left G1 (D, pleonal/ventral view; E, mesiosternal/mesiodorsal view); F, left G2, sternal/dorsal view; G, H, distal tip of left G1 (G, pleonal/ventral view; H, mesiosternal/mesiodorsal view). Scale bars: 1.0 mm (A–C); 0.5 mm (D–F); 0.25 mm (G, H)." figureDoi="http://doi.org/10.5281/zenodo.7385098" httpUri="https://zenodo.org/record/7385098/files/figure.png" pageId="4" pageNumber="265">Fig. 3B</figureCitation>
) finely granular, with rows of larger, widely spaced granules near upper margin, also proximally. Fingers thick, noticeably shorter than palm, cutting margins with small teeth; fixed finger with short longitudinal groove on the external surface; dactylus longer, slightly curved, upper margin granular proximally. Minor palm (
<figureCitation id="13762A04FFE0FF92FD044AA3C14DFA50" box="[690,778,1483,1509]" captionStart="FIGURE 3" captionStartId="6.[151,250,1891,1915]" captionTargetBox="[151,1435,181,1867]" captionTargetId="figure-21@6.[151,1435,181,1867]" captionTargetPageId="6" captionText="FIGURE 3. Aniptumnus bijoyi sp. nov., holotype, male, 10.7 by 7.1 mm (CUSAT/SBN/HRJ/1), Kochi, Kerala, SW India. A, minor cheliped, external view; B, major cheliped, external view; C, right P5, dorsal view; D, E, left G1 (D, pleonal/ventral view; E, mesiosternal/mesiodorsal view); F, left G2, sternal/dorsal view; G, H, distal tip of left G1 (G, pleonal/ventral view; H, mesiosternal/mesiodorsal view). Scale bars: 1.0 mm (A–C); 0.5 mm (D–F); 0.25 mm (G, H)." figureDoi="http://doi.org/10.5281/zenodo.7385098" httpUri="https://zenodo.org/record/7385098/files/figure.png" pageId="4" pageNumber="265">Fig. 3A</figureCitation>
) smaller, external surface setose, with large, sharp, conical granules, sometimes arranged in rows along long axis of palm; fingers slender, shorter than palm, cutting margins minutely dentate; fixed finger slightly deflexed; dactylus with longitudinal ridge on external surface, upper margin granular proximally.
</paragraph>
<paragraph id="8BF23681FFE0FF92FF714933C3A3F820" blockId="4.[151,1437,150,1941]" pageId="4" pageNumber="265">
Ambulatory legs (P2–P5; 
<figureCitation id="13762A04FFE0FF92FE404933C001F9C0" box="[502,582,1627,1653]" captionStart="FIGURE 1" captionStartId="3.[151,250,1950,1974]" captionTargetBox="[223,1366,993,1927]" captionTargetId="figure-382@3.[222,1366,993,1927]" captionTargetPageId="3" captionText="FIGURE 1. Aniptumnus bijoyi sp. nov., paratype, male, 10.1 by 6.7 mm (ZRC 2022.0137), Kochi, Kerala, SW India. Dorsal habitus and live coloration." figureDoi="http://doi.org/10.5281/zenodo.7385090" httpUri="https://zenodo.org/record/7385090/files/figure.png" pageId="4" pageNumber="265">Figs. 1</figureCitation>
, 
<figureCitation id="13762A04FFE0FF92FDEE4933C03EF9C0" box="[600,633,1627,1653]" captionStart="FIGURE 3" captionStartId="6.[151,250,1891,1915]" captionTargetBox="[151,1435,181,1867]" captionTargetId="figure-21@6.[151,1435,181,1867]" captionTargetPageId="6" captionText="FIGURE 3. Aniptumnus bijoyi sp. nov., holotype, male, 10.7 by 7.1 mm (CUSAT/SBN/HRJ/1), Kochi, Kerala, SW India. A, minor cheliped, external view; B, major cheliped, external view; C, right P5, dorsal view; D, E, left G1 (D, pleonal/ventral view; E, mesiosternal/mesiodorsal view); F, left G2, sternal/dorsal view; G, H, distal tip of left G1 (G, pleonal/ventral view; H, mesiosternal/mesiodorsal view). Scale bars: 1.0 mm (A–C); 0.5 mm (D–F); 0.25 mm (G, H)." figureDoi="http://doi.org/10.5281/zenodo.7385098" httpUri="https://zenodo.org/record/7385098/files/figure.png" pageId="4" pageNumber="265">3C</figureCitation>
) moderately long, slender, setose, broad surfaces finely granular; P3 longest, merus-to-dactylus length 1.02 times CW), P5 shortest, merus-to-dactylus length 0.78 times CW. Basisischium unarmed except in P5, where posterodistal margin armed with prominent spines, forming comb-like structure. Merus subrectangular, elongate, anterior margin slightly convex, finely granulate, subdistal tooth present but effaced, posterior margin, slightly convex, finely granulate except in P5, which has several spines and conical tubercles proximally, decreasing in size until mid-length of merus. Carpus subrectangular, widening distally, anterior margin convex, posterior margin straight, dorsal surface without granular ridge. Propodus subrectangular, anterior, posterior margins slightly convex. Dactylus mostly straight, tapering distally, ending in curved chitinous claw; dactylo-propodal lock absent.
</paragraph>
<caption id="DF326609FFE1FF93FF21480BC185F85D" ID-DOI="http://doi.org/10.5281/zenodo.7385093" ID-Zenodo-Dep="7385093" httpUri="https://zenodo.org/record/7385093/files/figure.png" pageId="5" pageNumber="266" startId="5.[151,250,1891,1915]" targetBox="[151,1435,181,1867]" targetPageId="5">
<paragraph id="8BF23681FFE1FF93FF21480BC185F85D" blockId="5.[151,1437,1891,2024]" pageId="5" pageNumber="266">
<emphasis id="B939EA93FFE1FF93FF21480BC349F8CE" bold="true" box="[151,270,1891,1915]" pageId="5" pageNumber="266">FIGURE 2</emphasis>
. 
<taxonomicName id="4C4D4D02FFE1FF93FEAD480CC391F8CE" authority="Hari &amp; Hershey &amp; Mendoza, 2022" authorityName="Hari &amp; Hershey &amp; Mendoza" authorityYear="2022" box="[283,470,1891,1915]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="266" phylum="Arthropoda" rank="species" species="bijoyi" status="sp. nov.">
<emphasis id="B939EA93FFE1FF93FEAD480CC391F8CE" box="[283,470,1891,1915]" italics="true" pageId="5" pageNumber="266">Aniptumnus bijoyi</emphasis>
</taxonomicName>
<emphasis id="B939EA93FFE1FF93FE6B480CC069F8C9" bold="true" box="[477,558,1892,1916]" pageId="5" pageNumber="266">
<taxonomicNameLabel id="A20A57E8FFE1FF93FE6B480CC069F8C9" box="[477,558,1892,1916]" pageId="5" pageNumber="266" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, male, 10.7 by 7.1 mm (CUSAT/SBN/HRJ/1), Kochi, Kerala, SW India. A, carapace, dorsal view; B, left third maxilliped, external view; cephalothorax and chelipeds, frontal view; D, anterior thoracic sternum and telson, ventral view; E, thoracic sternum and sternopleonal cavity, ventral view; F, pleon, ventral view; G, pleonal somites 1–3, posteroventral view. Scale bars: 2.0 mm (A, C); 1.0 mm (B, D–G).
</paragraph>
</caption>
<caption id="DF326609FFE2FF90FF21480BC603F85D" ID-DOI="http://doi.org/10.5281/zenodo.7385098" ID-Zenodo-Dep="7385098" httpUri="https://zenodo.org/record/7385098/files/figure.png" pageId="6" pageNumber="267" startId="6.[151,250,1891,1915]" targetBox="[151,1435,181,1867]" targetPageId="6">
<paragraph id="8BF23681FFE2FF90FF21480BC603F85D" blockId="6.[151,1436,1891,2024]" pageId="6" pageNumber="267">
<emphasis id="B939EA93FFE2FF90FF21480BC349F8CE" bold="true" box="[151,270,1891,1915]" pageId="6" pageNumber="267">FIGURE 3</emphasis>
. 
<taxonomicName id="4C4D4D02FFE2FF90FEAD480CC392F8CE" authority="Hari &amp; Hershey &amp; Mendoza, 2022" authorityName="Hari &amp; Hershey &amp; Mendoza" authorityYear="2022" box="[283,469,1891,1915]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="267" phylum="Arthropoda" rank="species" species="bijoyi" status="sp. nov.">
<emphasis id="B939EA93FFE2FF90FEAD480CC392F8CE" box="[283,469,1891,1915]" italics="true" pageId="6" pageNumber="267">Aniptumnus bijoyi</emphasis>
</taxonomicName>
<emphasis id="B939EA93FFE2FF90FE6A480CC069F8C9" bold="true" box="[476,558,1892,1916]" pageId="6" pageNumber="267">
<taxonomicNameLabel id="A20A57E8FFE2FF90FE6A480CC069F8C9" box="[476,558,1892,1916]" pageId="6" pageNumber="267" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, male, 10.7 by 7.1 mm (CUSAT/SBN/HRJ/1), Kochi, Kerala, SW India. A, minor cheliped, external view; B, major cheliped, external view; C, right P5, dorsal view; D, E, left G1 (D, pleonal/ventral view; E, mesiosternal/mesiodorsal view); F, left G2, sternal/dorsal view; G, H, distal tip of left G1 (G, pleonal/ventral view; H, mesiosternal/mesiodorsal view). Scale bars: 1.0 mm (A–C); 0.5 mm (D–F); 0.25 mm (G, H).
</paragraph>
</caption>
<paragraph id="8BF23681FFE3FF91FF714FFFC6B6FED0" blockId="7.[151,1437,151,681]" pageId="7" pageNumber="268">
Male pleon (
<figureCitation id="13762A04FFE3FF91FEEA4FFFC3BCFF04" box="[348,507,151,177]" captionStart="FIGURE 2" captionStartId="5.[151,250,1891,1915]" captionTargetBox="[151,1435,181,1867]" captionTargetId="figure-20@5.[151,1435,181,1867]" captionTargetPageId="5" captionText="FIGURE 2. Aniptumnus bijoyi sp. nov., holotype, male, 10.7 by 7.1 mm (CUSAT/SBN/HRJ/1), Kochi, Kerala, SW India. A, carapace, dorsal view; B, left third maxilliped, external view; cephalothorax and chelipeds, frontal view; D, anterior thoracic sternum and telson, ventral view; E, thoracic sternum and sternopleonal cavity, ventral view; F, pleon, ventral view; G, pleonal somites 1–3, posteroventral view. Scale bars: 2.0 mm (A, C); 1.0 mm (B, D–G)." figureDoi="http://doi.org/10.5281/zenodo.7385093" httpUri="https://zenodo.org/record/7385093/files/figure.png" pageId="7" pageNumber="268">Fig. 2D, F, G</figureCitation>
) smooth, with all somites and telson freely articulating, lateral margin concave. Somite 1 much wider than long, anterior margin deeply concave, posterior margin slightly sinuous, lateral margins straight. Somite 2 much wider than long, anterior margin slightly concave, posterior margin strongly convex, lateral margins rounded. Somite 3 widest, transversely subhexagonal, lateral margins strongly convex. Somites 4–6 trapezoidal, gradually decreasing in width, with somite 6 narrowest, lateral margins straight to slightly concave. Telson triangular with rounded apex, lateral margins convex, median length 0.83 times basal width.
</paragraph>
<paragraph id="8BF23681FFE3FF91FF714E07C157FE40" blockId="7.[151,1437,151,681]" pageId="7" pageNumber="268">
G1 (
<figureCitation id="13762A04FFE3FF91FF4E4E07C3E8FE3C" box="[248,431,367,393]" captionStart="FIGURE 3" captionStartId="6.[151,250,1891,1915]" captionTargetBox="[151,1435,181,1867]" captionTargetId="figure-21@6.[151,1435,181,1867]" captionTargetPageId="6" captionText="FIGURE 3. Aniptumnus bijoyi sp. nov., holotype, male, 10.7 by 7.1 mm (CUSAT/SBN/HRJ/1), Kochi, Kerala, SW India. A, minor cheliped, external view; B, major cheliped, external view; C, right P5, dorsal view; D, E, left G1 (D, pleonal/ventral view; E, mesiosternal/mesiodorsal view); F, left G2, sternal/dorsal view; G, H, distal tip of left G1 (G, pleonal/ventral view; H, mesiosternal/mesiodorsal view). Scale bars: 1.0 mm (A–C); 0.5 mm (D–F); 0.25 mm (G, H)." figureDoi="http://doi.org/10.5281/zenodo.7385098" httpUri="https://zenodo.org/record/7385098/files/figure.png" pageId="7" pageNumber="268">Fig. 3D, E, G, H</figureCitation>
) moderately long, slender, sinuous; margins lined with simple or plumose setae; distal part curved, ending in small scoop-shaped apical lobe, with stiff, simple spiniform subdistal setae; G2 (
<figureCitation id="13762A04FFE3FF91FB4F4EFBC70AFE18" box="[1273,1357,403,429]" captionStart="FIGURE 3" captionStartId="6.[151,250,1891,1915]" captionTargetBox="[151,1435,181,1867]" captionTargetId="figure-21@6.[151,1435,181,1867]" captionTargetPageId="6" captionText="FIGURE 3. Aniptumnus bijoyi sp. nov., holotype, male, 10.7 by 7.1 mm (CUSAT/SBN/HRJ/1), Kochi, Kerala, SW India. A, minor cheliped, external view; B, major cheliped, external view; C, right P5, dorsal view; D, E, left G1 (D, pleonal/ventral view; E, mesiosternal/mesiodorsal view); F, left G2, sternal/dorsal view; G, H, distal tip of left G1 (G, pleonal/ventral view; H, mesiosternal/mesiodorsal view). Scale bars: 1.0 mm (A–C); 0.5 mm (D–F); 0.25 mm (G, H)." figureDoi="http://doi.org/10.5281/zenodo.7385098" httpUri="https://zenodo.org/record/7385098/files/figure.png" pageId="7" pageNumber="268">Fig. 3F</figureCitation>
) short, about one-fifth length of G1,terminal, subterminal segments uniformly curving mesially. Penis (
<figureCitation id="13762A04FFE3FF91FB784EDFC764FE64" box="[1230,1315,439,465]" captionStart="FIGURE 2" captionStartId="5.[151,250,1891,1915]" captionTargetBox="[151,1435,181,1867]" captionTargetId="figure-20@5.[151,1435,181,1867]" captionTargetPageId="5" captionText="FIGURE 2. Aniptumnus bijoyi sp. nov., holotype, male, 10.7 by 7.1 mm (CUSAT/SBN/HRJ/1), Kochi, Kerala, SW India. A, carapace, dorsal view; B, left third maxilliped, external view; cephalothorax and chelipeds, frontal view; D, anterior thoracic sternum and telson, ventral view; E, thoracic sternum and sternopleonal cavity, ventral view; F, pleon, ventral view; G, pleonal somites 1–3, posteroventral view. Scale bars: 2.0 mm (A, C); 1.0 mm (B, D–G)." figureDoi="http://doi.org/10.5281/zenodo.7385093" httpUri="https://zenodo.org/record/7385093/files/figure.png" pageId="7" pageNumber="268">Fig. 2E</figureCitation>
) emerging from anterior portion of coxosternal condyle of P5 coxa.
</paragraph>
<paragraph id="8BF23681FFE3FF91FF714E97C00DFD1C" blockId="7.[151,1437,151,681]" pageId="7" pageNumber="268">
<emphasis id="B939EA93FFE3FF91FF714E97C362FDAC" bold="true" box="[199,293,511,537]" pageId="7" pageNumber="268">Female.</emphasis>
The female morphology is similar to the male in all non-sexual aspects. Thoracic sternum relatively wider than that of male. Pleon (
<figureCitation id="13762A04FFE3FF91FDBB4D4BC02EFD88" box="[525,617,547,573]" captionStart="FIGURE 4" captionStartId="7.[151,250,1490,1514]" captionTargetBox="[151,1435,705,1467]" captionTargetId="figure-241@7.[151,1435,705,1467]" captionTargetPageId="7" captionText="FIGURE 4. Aniptumnus bijoyi sp. nov., paratype, female, 8.2 by 5.1 mm (ZRC 2022.0137), Kochi, Kerala, SW India. A, detached pleon, external view; B, thoracic sternum and vulvae, ventral view. Scale bars: 1.0 mm (A, B)." figureDoi="http://doi.org/10.5281/zenodo.7385100" httpUri="https://zenodo.org/record/7385100/files/figure.png" pageId="7" pageNumber="268">Fig. 4A</figureCitation>
) much wider, lateral margin convex, setose, external surfaces smooth; somites and pleon freely articulating; somite 3 widest. Telson large, subtriangular, apex rounded, lateral margins convex, median length subequal to basal width. Vulvae (
<figureCitation id="13762A04FFE3FF91FC964D03C13FFD30" box="[800,888,619,645]" captionStart="FIGURE 4" captionStartId="7.[151,250,1490,1514]" captionTargetBox="[151,1435,705,1467]" captionTargetId="figure-241@7.[151,1435,705,1467]" captionTargetPageId="7" captionText="FIGURE 4. Aniptumnus bijoyi sp. nov., paratype, female, 8.2 by 5.1 mm (ZRC 2022.0137), Kochi, Kerala, SW India. A, detached pleon, external view; B, thoracic sternum and vulvae, ventral view. Scale bars: 1.0 mm (A, B)." figureDoi="http://doi.org/10.5281/zenodo.7385100" httpUri="https://zenodo.org/record/7385100/files/figure.png" pageId="7" pageNumber="268">Fig. 4B</figureCitation>
) mesially on thoracic sternite 6, abutting suture with sternite 5; semicircular in outline.
</paragraph>
<caption id="DF326609FFE3FF91FF214ABAC6F3F9BB" ID-DOI="http://doi.org/10.5281/zenodo.7385100" ID-Zenodo-Dep="7385100" httpUri="https://zenodo.org/record/7385100/files/figure.png" pageId="7" pageNumber="268" startId="7.[151,250,1490,1514]" targetBox="[151,1435,705,1467]" targetPageId="7">
<paragraph id="8BF23681FFE3FF91FF214ABAC6F3F9BB" blockId="7.[151,1436,1490,1550]" pageId="7" pageNumber="268">
<emphasis id="B939EA93FFE3FF91FF214ABAC357FA5F" bold="true" box="[151,272,1490,1514]" pageId="7" pageNumber="268">FIGURE 4</emphasis>
. 
<taxonomicName id="4C4D4D02FFE3FF91FE964ABAC39BFA5F" authority="Hari &amp; Hershey &amp; Mendoza, 2022" authorityName="Hari &amp; Hershey &amp; Mendoza" authorityYear="2022" box="[288,476,1490,1514]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="268" phylum="Arthropoda" rank="species" species="bijoyi" status="sp. nov.">
<emphasis id="B939EA93FFE3FF91FE964ABAC39BFA5F" box="[288,476,1490,1514]" italics="true" pageId="7" pageNumber="268">Aniptumnus bijoyi</emphasis>
</taxonomicName>
<emphasis id="B939EA93FFE3FF91FE534ABAC07EFA5F" bold="true" box="[485,569,1490,1514]" pageId="7" pageNumber="268">
<taxonomicNameLabel id="A20A57E8FFE3FF91FE534ABAC07EFA5F" box="[485,569,1490,1514]" pageId="7" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype, female, 8.2 by 5.1 mm (ZRC 2022.0137), Kochi, Kerala, SW India. A, detached pleon, external view; B, thoracic sternum and vulvae, ventral view. Scale bars: 1.0 mm (A, B).
</paragraph>
</caption>
<paragraph id="8BF23681FFE3FF91FF714954C69BF953" blockId="7.[151,1437,1596,2019]" pageId="7" pageNumber="268">
<emphasis id="B939EA93FFE3FF91FF714954C3D2F9E3" bold="true" box="[199,405,1596,1622]" pageId="7" pageNumber="268">Live colouration.</emphasis>
Fresh specimens are usually covered with a copious amount of sediment and debris when collected such that they need to be gently brushed clean to expose the natural coloration (
<figureCitation id="13762A04FFE3FF91FBDF4908C6EEF9CF" box="[1129,1193,1632,1658]" captionStart="FIGURE 1" captionStartId="3.[151,250,1950,1974]" captionTargetBox="[223,1366,993,1927]" captionTargetId="figure-382@3.[222,1366,993,1927]" captionTargetPageId="3" captionText="FIGURE 1. Aniptumnus bijoyi sp. nov., paratype, male, 10.1 by 6.7 mm (ZRC 2022.0137), Kochi, Kerala, SW India. Dorsal habitus and live coloration." figureDoi="http://doi.org/10.5281/zenodo.7385090" httpUri="https://zenodo.org/record/7385090/files/figure.png" pageId="7" pageNumber="268">Fig. 1</figureCitation>
). The dorsal carapace and pereopods are generally tan or pale brown with patches and scattered spots of dark purple that are sometimes surrounded by lighter areas of purple. The setae covering them are similarly tan. The thoracic sternum, pleon, and ventral and internal surfaces are creamy white. The fingers of the chelae are brown with pale tips.
</paragraph>
</subSubSection>
<subSubSection id="C357650AFFE3FF9FFF714998C0A9FEF4" lastPageId="9" lastPageNumber="270" pageId="7" pageNumber="268" type="etymology">
<paragraph id="8BF23681FFE3FF91FF714998C312F8E7" blockId="7.[151,1437,1596,2019]" pageId="7" pageNumber="268">
<emphasis id="B939EA93FFE3FF91FF714998C309F8BF" bold="true" box="[199,334,1776,1802]" pageId="7" pageNumber="268">Etymology.</emphasis>
The species is named in honor of Prof. S. Bijoy Nandan, Dean, Faculty of Marine Sciences, 
<collectingCountry id="F35A7611FFE3FF91FAFD4998C7DBF8BF" box="[1355,1436,1776,1802]" name="India" pageId="7" pageNumber="268">Cochin</collectingCountry>
University of Science and Technology, for his significant contributions in the field of marine biology and wetland ecology in 
<collectingCountry id="F35A7611FFE3FF91FEA24850C316F8E7" box="[276,337,1848,1874]" name="India" pageId="7" pageNumber="268">India</collectingCountry>
.
</paragraph>
<paragraph id="8BF23681FFE3FF9FFF714834C0A9FEF4" blockId="7.[151,1437,1596,2019]" lastBlockId="9.[151,1437,151,1401]" lastPageId="9" lastPageNumber="270" pageId="7" pageNumber="268">
<emphasis id="B939EA93FFE3FF91FF714834C384F8C3" bold="true" box="[199,451,1884,1910]" pageId="7" pageNumber="268">Habitat and ecology.</emphasis>
<taxonomicName id="4C4D4D02FFE3FF91FE7B4835C0DAF8C3" authority="Hari &amp; Hershey &amp; Mendoza, 2022" authorityName="Hari &amp; Hershey &amp; Mendoza" authorityYear="2022" box="[461,669,1884,1911]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="268" phylum="Arthropoda" rank="species" species="bijoyi" status="sp. nov.">
<emphasis id="B939EA93FFE3FF91FE7B4835C0DAF8C3" box="[461,669,1884,1911]" italics="true" pageId="7" pageNumber="268">Aniptumnus bijoyi</emphasis>
</taxonomicName>
<emphasis id="B939EA93FFE3FF91FD114834C143F8C3" bold="true" box="[679,772,1884,1910]" pageId="7" pageNumber="268">
<taxonomicNameLabel id="A20A57E8FFE3FF91FD114834C143F8C3" box="[679,772,1884,1910]" pageId="7" pageNumber="268" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was found in larger numbers on encrusting communities such as clumps of mussels (
<taxonomicName id="4C4D4D02FFE3FF91FE7848E8C0C6F82F" baseAuthorityName="Hanley" baseAuthorityYear="1843" box="[462,641,1920,1946]" class="Bivalvia" family="Mytilidae" genus="Mytella" kingdom="Animalia" order="Mytilida" pageId="7" pageNumber="268" phylum="Mollusca" rank="species" species="strigata">
<emphasis id="B939EA93FFE3FF91FE7848E8C0C6F82F" box="[462,641,1920,1946]" italics="true" pageId="7" pageNumber="268">Mytella strigata</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C4D4D02FFE3FF91FD0148E9C106F82F" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[695,833,1920,1946]" class="Insecta" family="Lasiocampidae" genus="Perna" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="268" phylum="Arthropoda" rank="species" species="virdis">
<emphasis id="B939EA93FFE3FF91FD0148E9C106F82F" box="[695,833,1920,1946]" italics="true" pageId="7" pageNumber="268">Perna virdis</emphasis>
</taxonomicName>
), submerged structures, and roots of decaying aquatic weeds (
<taxonomicName id="4C4D4D02FFE3FF91FF5948CCC383F80B" box="[239,452,1956,1982]" class="Liliopsida" family="Pontederiaceae" genus="Eichornia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Commelinales" pageId="7" pageNumber="268" phylum="Tracheophyta" rank="species" species="crasipes">
<emphasis id="B939EA93FFE3FF91FF5948CCC383F80B" box="[239,452,1956,1982]" italics="true" pageId="7" pageNumber="268">Eichornia crasipes</emphasis>
</taxonomicName>
) in the School of Marine Science Jetty. Their occurrence was also noticed in the other parts of the 
<collectingCountry id="F35A7611FFE3FF91FEAD48A0C32BF857" box="[283,364,1992,2018]" name="India" pageId="7" pageNumber="268">Cochin</collectingCountry>
Estuary. The species is usually abundant during the pre-monsoon period (January–May) in sites with a salinity range of 10–28 ppt and depth range of 
<quantity id="4CB59B64FFEDFF9FFCB54FFFC109FF04" box="[771,846,151,177]" metricMagnitude="0" metricUnit="m" metricValue="3.5" metricValueMax="6.0" metricValueMin="1.0" pageId="9" pageNumber="270" unit="m" value="3.5" valueMax="6.0" valueMin="1.0">
1– 
<specimenCount id="9D4BFD08FFEDFF9FFCA84FFFC109FF04" box="[798,846,151,177]" pageId="9" pageNumber="270" type="generic">6 m</specimenCount>
</quantity>
. The habitat is muddy to clayey-silty, the pH of the sediment generally ranged from 7.5–8, and the average redox potential from -135 to -275mV. The total carbon of the sediment ranged from 
<quantity id="4CB59B64FFEDFF9FFE744FB6C05AFF4C" box="[450,541,222,249]" metricMagnitude="-2" metricUnit="kg" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="9" pageNumber="270" unit="g" value="32.5" valueMax="35.0" valueMin="30.0">30–35 g</quantity>
/kg and the total organic carbon ranged from 1.9–2.5 %. 
<taxonomicName id="4C4D4D02FFEDFF9FFB134FB7C734FF4C" authority="Hari &amp; Hershey &amp; Mendoza, 2022" authorityName="Hari &amp; Hershey &amp; Mendoza" authorityYear="2022" box="[1189,1395,223,249]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="270" phylum="Arthropoda" rank="species" species="bijoyi" status="sp. nov.">
<emphasis id="B939EA93FFEDFF9FFB134FB7C734FF4C" box="[1189,1395,223,249]" italics="true" pageId="9" pageNumber="270">Aniptumnus bijoyi</emphasis>
</taxonomicName>
<emphasis id="B939EA93FFEDFF9FFACD4FB7C28FFEA8" bold="true" pageId="9" pageNumber="270">
<taxonomicNameLabel id="A20A57E8FFEDFF9FFACD4FB7C28FFEA8" pageId="9" pageNumber="270" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was observed to occur together with the brachyuran crabs 
<taxonomicName id="4C4D4D02FFEDFF9FFCE94E6BC69FFEA8" authority="(Ruppell, 1830)" baseAuthorityName="Ruppell" baseAuthorityYear="1830" box="[863,1240,259,285]" class="Malacostraca" family="Portunidae" genus="Thranita" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="270" phylum="Arthropoda" rank="species" species="crenata">
<emphasis id="B939EA93FFEDFF9FFCE94E6BC65AFEA9" box="[863,1053,259,285]" italics="true" pageId="9" pageNumber="270">Thranita crenata</emphasis>
(Rüppell, 1830)
</taxonomicName>
, 
<taxonomicName id="4C4D4D02FFEDFF9FFB524E6BC3C9FEF4" authority="(Kemp, 1917)" baseAuthorityName="Kemp" baseAuthorityYear="1917" class="Malacostraca" family="Hymenosomatidae" genus="Neorhynchoplax" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="270" phylum="Arthropoda" rank="species" species="alcocki">
<emphasis id="B939EA93FFEDFF9FFB524E6BC2AFFEF4" italics="true" pageId="9" pageNumber="270">Neorhynchoplax alcocki</emphasis>
(Kemp, 1917)
</taxonomicName>
, and 
<taxonomicName id="4C4D4D02FFEDFF9FFE7C4E4FC0ADFEF4" authority="(Kemp, 1917)" baseAuthorityName="Kemp" baseAuthorityYear="1917" box="[458,746,295,321]" class="Malacostraca" family="Hymenosomatidae" genus="Neorhynchoplax" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="270" phylum="Arthropoda" rank="species" species="demeloi">
<emphasis id="B939EA93FFEDFF9FFE7C4E4FC004FEF4" box="[458,579,295,321]" italics="true" pageId="9" pageNumber="270">N. demeloi</emphasis>
(Kemp, 1917)
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="DF326609FFECFF9EFF214828C347F85C" ID-DOI="http://doi.org/10.5281/zenodo.7385102" ID-Zenodo-Dep="7385102" httpUri="https://zenodo.org/record/7385102/files/figure.png" pageId="8" pageNumber="269" startId="8.[151,250,1856,1880]" targetBox="[164,1422,181,1833]" targetPageId="8">
<paragraph id="8BF23681FFECFF9EFF214828C347F85C" blockId="8.[151,1437,1856,2025]" pageId="8" pageNumber="269">
<emphasis id="B939EA93FFECFF9EFF214828C34AF8EC" bold="true" box="[151,269,1856,1881]" pageId="8" pageNumber="269">FIGURE 5</emphasis>
. 
<taxonomicName id="4C4D4D02FFECFF9EFEAF4829C099F8ED" authority="(De Man, 1895)" baseAuthorityName="De Man" baseAuthorityYear="1895" box="[281,734,1856,1881]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="269" phylum="Arthropoda" rank="species" species="quadridentatus">
<emphasis id="B939EA93FFECFF9EFEAF4829C075F8EC" box="[281,562,1857,1881]" italics="true" pageId="8" pageNumber="269">Aniptumnus quadridentatus</emphasis>
(
<bibRefCitation id="EFDC4B70FFECFF9EFDF64829C091F8ED" author="De Man, J. G." box="[576,726,1856,1881]" pageId="8" pageNumber="269" pagination="485 - 609" refId="ref7241" refString="De Man, J. G. (1895) Bericht uber die von Herrn Schiffscapitan Storm zur Atjeh, an den westlichen Kusten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8 (4), 485 - 609. https: // doi. org / 10.5962 / bhl. title. 16084" type="journal article" year="1895">De Man, 1895</bibRefCitation>
)
</taxonomicName>
, male, 11.1 by 7.4 mm (ZRC 2021.0421), off Pulau Ubin, Singapore. A, carapace, dorsal view; B, third maxillipeds, external view; C, cephalothorax, frontal view; D, anterior thoracic sternum and pleon, ventral view; E, pleonal somites 1–3 and thoracic sternite 8, posteroventral view; F–I, left G1 (F, pleonal/ventral view; G, mesiosternal/mesiodorsal view; H, lateral view; I, sternal/dorsal view). Scale bars: 2.0 mm (A, C); 1.0 mm (B, D, E); 0.5 mm (F–I).
</paragraph>
</caption>
<subSubSection id="C357650AFFEDFF9FFF714E23C62CFACC" pageId="9" pageNumber="270" type="discussion">
<paragraph id="8BF23681FFEDFF9FFF714E23C606FE40" blockId="9.[151,1437,151,1401]" pageId="9" pageNumber="270">
<emphasis id="B939EA93FFEDFF9FFF714E23C37CFED0" bold="true" box="[199,315,331,357]" pageId="9" pageNumber="270">Remarks.</emphasis>
<taxonomicName id="4C4D4D02FFEDFF9FFEF24E23C054FED0" authority="Hari &amp; Hershey &amp; Mendoza, 2022" authorityName="Hari &amp; Hershey &amp; Mendoza" authorityYear="2022" box="[324,531,331,357]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="270" phylum="Arthropoda" rank="species" species="bijoyi" status="sp. nov.">
<emphasis id="B939EA93FFEDFF9FFEF24E23C054FED0" box="[324,531,331,357]" italics="true" pageId="9" pageNumber="270">Aniptumnus bijoyi</emphasis>
</taxonomicName>
<emphasis id="B939EA93FFEDFF9FFDAD4E23C030FED0" bold="true" box="[539,631,331,357]" pageId="9" pageNumber="270">
<taxonomicNameLabel id="A20A57E8FFEDFF9FFDAD4E23C030FED0" box="[539,631,331,357]" pageId="9" pageNumber="270" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is morphologically most similar to 
<taxonomicName id="4C4D4D02FFEDFF9FFBA74E23C7DBFED0" authority="(De Man, 1895)" baseAuthorityName="De Man" baseAuthorityYear="1895" box="[1041,1436,330,357]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="270" phylum="Arthropoda" rank="species" species="quadridentatus">
<emphasis id="B939EA93FFEDFF9FFBA74E23C69BFED0" box="[1041,1244,331,357]" italics="true" pageId="9" pageNumber="270">A. quadridentatus</emphasis>
(
<bibRefCitation id="EFDC4B70FFEDFF9FFB5B4E23C7D3FED0" author="De Man, J. G." box="[1261,1428,330,357]" pageId="9" pageNumber="270" pagination="485 - 609" refId="ref7241" refString="De Man, J. G. (1895) Bericht uber die von Herrn Schiffscapitan Storm zur Atjeh, an den westlichen Kusten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8 (4), 485 - 609. https: // doi. org / 10.5962 / bhl. title. 16084" type="journal article" year="1895">De Man, 1895</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="4C4D4D02FFEDFF9FFF7F4E07C3A1FE3C" authority="(Deb, 1985)" authorityName="Hari &amp; Hershey &amp; Mendoza" authorityYear="2022" baseAuthorityName="Deb" baseAuthorityYear="1985" box="[201,486,366,393]" class="Malacostraca" family="Pilumnidae" genus="Heteropanope" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="270" phylum="Arthropoda" rank="species" species="neolaevis" status="comb. nov.">
<emphasis id="B939EA93FFEDFF9FFF7F4E07C313FE3C" box="[201,340,367,393]" italics="true" pageId="9" pageNumber="270">A. neolaevis</emphasis>
(Deb, 1985)
</taxonomicName>
<taxonomicNameLabel id="A20A57E8FFEDFF9FFE594E07C02AFE3C" box="[495,621,367,393]" pageId="9" pageNumber="270" rank="species">comb. nov.</taxonomicNameLabel>
in the general form of the carapace, thoracic sternum, male pleon, and pereopods (cf. 
<bibRefCitation id="EFDC4B70FFEDFF9FFEF64EFBC3E1FE18" author="Ng, P. K. L." box="[320,422,403,429]" pageId="9" pageNumber="270" pagination="209 - 226" refId="ref7709" refString="Ng, P. K. L. (2002) The Indo-Pacific Pilumnidae. XVI. On the Identity of Pilumnus cristimanus A. Milne Edwards and the status of Parapilumnus Kossman, 1877 (Crustacea: Decapoda: Brachyura), with description of a new species from the rubble beds in Guam. Micronesica, 34 (2), 209 - 226." type="journal article" year="2002">Ng 2002</bibRefCitation>
; 
<bibRefCitation id="EFDC4B70FFEDFF9FFE044EFBC0C0FE18" author="Trivedi, J. &amp; Mitra, S. &amp; Patel, P. &amp; Maheta, N. &amp; Patel, K. &amp; Ng, P. K. L." box="[434,647,403,429]" pageId="9" pageNumber="270" pagination="1 - 18" refId="ref7973" refString="Trivedi, J., Mitra, S., Patel, P., Maheta, N., Patel, K. &amp; Ng, P. K. L. (2021) On the Indian species of Eurycarcinus A. MilneEdwards, 1867, Heteropanope Stimpson, 1858, and Pilumnopeus A. Milne-Edwards, 1867 (Decapoda: Brachyura: Pilumnidae). Nauplius, 29 (e 2021004), 1 - 18. https: // doi. org / 10.1590 / 2358 - 2936 e 2021004" type="journal article" year="2021">
Trivedi 
<emphasis id="B939EA93FFEDFF9FFDBD4EFBC003FE18" box="[523,580,403,429]" italics="true" pageId="9" pageNumber="270">et al.</emphasis>
2021
</bibRefCitation>
; 
<bibRefCitation id="EFDC4B70FFEDFF9FFD254EFBC18AFE18" author="Widyastuti, E. &amp; Rahayu, D. L." box="[659,973,403,429]" pageId="9" pageNumber="270" pagination="15 - 22" refId="ref8067" refString="Widyastuti, E. &amp; Rahayu, D. L. (2022) The new record of Heteropilumnus sasekumari (Serene, 1971) and Aniptumnus quadridentatus (De Man, 1895) (Crustacea, Decapoda, Brachyura, Pilumnidae) from mangrove habitat in Papua, Indonesia. Oseanologi dan Limnologi di Indonesia, 7 (1), 15 - 22." type="journal article" year="2022">Widyastuti &amp; Rahayu 2022</bibRefCitation>
). The new species is easily distinguished from 
<taxonomicName id="4C4D4D02FFEDFF9FFF634EDFC320FE64" baseAuthorityName="Garth &amp; Kim" baseAuthorityYear="1983" box="[213,359,439,465]" class="Malacostraca" family="Acidopsidae" genus="Parapilumnus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="270" phylum="Arthropoda" rank="species" species="nefissurus">
<emphasis id="B939EA93FFEDFF9FFF634EDFC320FE64" box="[213,359,439,465]" italics="true" pageId="9" pageNumber="270">A. nefissurus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C4D4D02FFEDFF9FFE284EDFC004FE65" authorityName="Ng &amp; Clark" authorityYear="2008" box="[414,579,439,465]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="270" phylum="Arthropoda" rank="species" species="vietnamicus">
<emphasis id="B939EA93FFEDFF9FFE284EDFC004FE65" box="[414,579,439,465]" italics="true" pageId="9" pageNumber="270">A. vietnamicus</emphasis>
</taxonomicName>
by the general form of the carapace, by its less granular major chela and by the G1 morphology (cf. 
<bibRefCitation id="EFDC4B70FFEDFF9FFE114EB3C038FE40" author="Garth, J. S. &amp; Kim, H. S." box="[423,639,475,501]" pageId="9" pageNumber="270" pagination="663 - 729" refId="ref7385" refString="Garth, J. S. &amp; Kim, H. S. (1983) Crabs of the family Xanthidae (Crustacea: Brachyura) from the Philippine Islands and adjacent waters based largely on collections of the U. S. Fish Commission steamer Albatross in 1909 - 1909. Journal of Natural History, 17, 663 - 729. https: // doi. org / 10.1080 / 00222938300770561" type="journal article" year="1983">Garth &amp; Kim 1983</bibRefCitation>
: fig. 11; 
<bibRefCitation id="EFDC4B70FFEDFF9FFD564EB3C1EFFE40" author="Ng, P. K. L. &amp; Clark, P. F." box="[736,936,475,501]" pageId="9" pageNumber="270" pagination="885 - 912" refId="ref7776" refString="Ng, P. K. L. &amp; Clark, P. F. (2008) A revision of Latopilumnus Turkay &amp; Schuhmacher, 1985, and Aniptumnus Ng, 2002 (Crustacea: Decapoda: Brachyura: Pilumnidae) with larval comparisons. Journal of Natural History, 42 (9), 885 - 912. https: // doi. org / 10.1080 / 00222930701869422" type="journal article" year="2008">Ng &amp; Clark 2008</bibRefCitation>
: figs. 19–21).
</paragraph>
<paragraph id="8BF23681FFEDFF9FFF714E97C0A8FCA0" blockId="9.[151,1437,151,1401]" pageId="9" pageNumber="270">
<taxonomicName id="4C4D4D02FFEDFF9FFF714E97C3D1FDAC" authority="Hari &amp; Hershey &amp; Mendoza, 2022" authorityName="Hari &amp; Hershey &amp; Mendoza" authorityYear="2022" box="[199,406,511,537]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="270" phylum="Arthropoda" rank="species" species="bijoyi" status="sp. nov.">
<emphasis id="B939EA93FFEDFF9FFF714E97C3D1FDAC" box="[199,406,511,537]" italics="true" pageId="9" pageNumber="270">Aniptumnus bijoyi</emphasis>
</taxonomicName>
<emphasis id="B939EA93FFEDFF9FFE294E97C3BDFDAC" bold="true" box="[415,506,511,537]" pageId="9" pageNumber="270">
<taxonomicNameLabel id="A20A57E8FFEDFF9FFE294E97C3BDFDAC" box="[415,506,511,537]" pageId="9" pageNumber="270" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can most easily be distinguished from 
<taxonomicName id="4C4D4D02FFEDFF9FFC0B4E97C60FFDAC" authorityName="Hari &amp; Hershey &amp; Mendoza" authorityYear="2022" baseAuthorityName="Deb" baseAuthorityYear="1985" box="[957,1096,511,537]" class="Malacostraca" family="Pilumnidae" genus="Heteropanope" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="270" phylum="Arthropoda" rank="species" species="neolaevis">
<emphasis id="B939EA93FFEDFF9FFC0B4E97C60FFDAC" box="[957,1096,511,537]" italics="true" pageId="9" pageNumber="270">A. neolaevis</emphasis>
</taxonomicName>
in the following features: the tuberculation on the posteroproximal margin of the P5 merus is much more prominent, with the conical tubercles being larger (
<figureCitation id="13762A04FFEDFF9FFE9B4D2FC3C4FDD4" box="[301,387,583,609]" captionStart="FIGURE 3" captionStartId="6.[151,250,1891,1915]" captionTargetBox="[151,1435,181,1867]" captionTargetId="figure-21@6.[151,1435,181,1867]" captionTargetPageId="6" captionText="FIGURE 3. Aniptumnus bijoyi sp. nov., holotype, male, 10.7 by 7.1 mm (CUSAT/SBN/HRJ/1), Kochi, Kerala, SW India. A, minor cheliped, external view; B, major cheliped, external view; C, right P5, dorsal view; D, E, left G1 (D, pleonal/ventral view; E, mesiosternal/mesiodorsal view); F, left G2, sternal/dorsal view; G, H, distal tip of left G1 (G, pleonal/ventral view; H, mesiosternal/mesiodorsal view). Scale bars: 1.0 mm (A–C); 0.5 mm (D–F); 0.25 mm (G, H)." figureDoi="http://doi.org/10.5281/zenodo.7385098" httpUri="https://zenodo.org/record/7385098/files/figure.png" pageId="9" pageNumber="270">Fig. 3C</figureCitation>
) (vs. tuberculation, while present, is much reduced, tubercles are smaller in 
<taxonomicName id="4C4D4D02FFEDFF9FFB5E4D2FC736FDD4" authorityName="Hari &amp; Hershey &amp; Mendoza" authorityYear="2022" baseAuthorityName="Deb" baseAuthorityYear="1985" box="[1256,1393,583,609]" class="Malacostraca" family="Pilumnidae" genus="Heteropanope" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="270" phylum="Arthropoda" rank="species" species="neolaevis">
<emphasis id="B939EA93FFEDFF9FFB5E4D2FC736FDD4" box="[1256,1393,583,609]" italics="true" pageId="9" pageNumber="270">A. neolaevis</emphasis>
</taxonomicName>
, cf. 
<bibRefCitation id="EFDC4B70FFEDFF9FFF214D03C321FD30" author="Trivedi, J. &amp; Mitra, S. &amp; Patel, P. &amp; Maheta, N. &amp; Patel, K. &amp; Ng, P. K. L." box="[151,358,619,645]" pageId="9" pageNumber="270" pagination="1 - 18" refId="ref7973" refString="Trivedi, J., Mitra, S., Patel, P., Maheta, N., Patel, K. &amp; Ng, P. K. L. (2021) On the Indian species of Eurycarcinus A. MilneEdwards, 1867, Heteropanope Stimpson, 1858, and Pilumnopeus A. Milne-Edwards, 1867 (Decapoda: Brachyura: Pilumnidae). Nauplius, 29 (e 2021004), 1 - 18. https: // doi. org / 10.1590 / 2358 - 2936 e 2021004" type="journal article" year="2021">
Trivedi 
<emphasis id="B939EA93FFEDFF9FFF584D03C359FD30" box="[238,286,619,645]" italics="true" pageId="9" pageNumber="270">et al</emphasis>
. 2021
</bibRefCitation>
: fig. 8A); pleonal somite 2 of the male is considerably wider, and consequently, thoracic sternite 8 is completely concealed when the pleon is folded and locked onto the thoracic sternum (
<figureCitation id="13762A04FFEDFF9FFBD14DE7C6FCFD1C" box="[1127,1211,655,681]" captionStart="FIGURE 2" captionStartId="5.[151,250,1891,1915]" captionTargetBox="[151,1435,181,1867]" captionTargetId="figure-20@5.[151,1435,181,1867]" captionTargetPageId="5" captionText="FIGURE 2. Aniptumnus bijoyi sp. nov., holotype, male, 10.7 by 7.1 mm (CUSAT/SBN/HRJ/1), Kochi, Kerala, SW India. A, carapace, dorsal view; B, left third maxilliped, external view; cephalothorax and chelipeds, frontal view; D, anterior thoracic sternum and telson, ventral view; E, thoracic sternum and sternopleonal cavity, ventral view; F, pleon, ventral view; G, pleonal somites 1–3, posteroventral view. Scale bars: 2.0 mm (A, C); 1.0 mm (B, D–G)." figureDoi="http://doi.org/10.5281/zenodo.7385093" httpUri="https://zenodo.org/record/7385093/files/figure.png" pageId="9" pageNumber="270">Fig. 2E</figureCitation>
) (vs. pleonal somite 2 much narrower, thoracic sternite 8 exposed in 
<taxonomicName id="4C4D4D02FFEDFF9FFD0F4DDBC104FD78" authorityName="Hari &amp; Hershey &amp; Mendoza" authorityYear="2022" baseAuthorityName="Deb" baseAuthorityYear="1985" box="[697,835,691,717]" class="Malacostraca" family="Pilumnidae" genus="Heteropanope" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="270" phylum="Arthropoda" rank="species" species="neolaevis">
<emphasis id="B939EA93FFEDFF9FFD0F4DDBC104FD78" box="[697,835,691,717]" italics="true" pageId="9" pageNumber="270">A. neolaevis</emphasis>
</taxonomicName>
; cf. 
<bibRefCitation id="EFDC4B70FFEDFF9FFCC04DDBC60BFD78" author="Trivedi, J. &amp; Mitra, S. &amp; Patel, P. &amp; Maheta, N. &amp; Patel, K. &amp; Ng, P. K. L." box="[886,1100,691,717]" pageId="9" pageNumber="270" pagination="1 - 18" refId="ref7973" refString="Trivedi, J., Mitra, S., Patel, P., Maheta, N., Patel, K. &amp; Ng, P. K. L. (2021) On the Indian species of Eurycarcinus A. MilneEdwards, 1867, Heteropanope Stimpson, 1858, and Pilumnopeus A. Milne-Edwards, 1867 (Decapoda: Brachyura: Pilumnidae). Nauplius, 29 (e 2021004), 1 - 18. https: // doi. org / 10.1590 / 2358 - 2936 e 2021004" type="journal article" year="2021">
Trivedi 
<emphasis id="B939EA93FFEDFF9FFC664DDBC645FD78" box="[976,1026,691,717]" italics="true" pageId="9" pageNumber="270">et al</emphasis>
. 2021
</bibRefCitation>
: fig. 8D); and the distal tip of the G1 has a short, projecting, scoop-shaped apical lobe (
<figureCitation id="13762A04FFEDFF9FFCAA4DBFC19EFD44" box="[796,985,727,753]" captionStart="FIGURE 3" captionStartId="6.[151,250,1891,1915]" captionTargetBox="[151,1435,181,1867]" captionTargetId="figure-21@6.[151,1435,181,1867]" captionTargetPageId="6" captionText="FIGURE 3. Aniptumnus bijoyi sp. nov., holotype, male, 10.7 by 7.1 mm (CUSAT/SBN/HRJ/1), Kochi, Kerala, SW India. A, minor cheliped, external view; B, major cheliped, external view; C, right P5, dorsal view; D, E, left G1 (D, pleonal/ventral view; E, mesiosternal/mesiodorsal view); F, left G2, sternal/dorsal view; G, H, distal tip of left G1 (G, pleonal/ventral view; H, mesiosternal/mesiodorsal view). Scale bars: 1.0 mm (A–C); 0.5 mm (D–F); 0.25 mm (G, H)." figureDoi="http://doi.org/10.5281/zenodo.7385098" httpUri="https://zenodo.org/record/7385098/files/figure.png" pageId="9" pageNumber="270">Fig. 3D, E, G, H</figureCitation>
) (vs. apical lobe absent in 
<taxonomicName id="4C4D4D02FFEDFF9FFABE4DBFC7D2FD44" authorityName="Hari &amp; Hershey &amp; Mendoza" authorityYear="2022" baseAuthorityName="Deb" baseAuthorityYear="1985" box="[1288,1429,727,753]" class="Malacostraca" family="Pilumnidae" genus="Heteropanope" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="270" phylum="Arthropoda" rank="species" species="neolaevis">
<emphasis id="B939EA93FFEDFF9FFABE4DBFC7D2FD44" box="[1288,1429,727,753]" italics="true" pageId="9" pageNumber="270">H. neolaevis</emphasis>
</taxonomicName>
; cf. 
<bibRefCitation id="EFDC4B70FFEDFF9FFF0D4D93C36AFCA0" author="Deb, M." box="[187,301,763,789]" pageId="9" pageNumber="270" pagination="345 - 403" refId="ref7187" refString="Deb, M. (1999) Crustacea: Decapoda: Crabs. In: Director ZSI, Kolkata (Ed.), Fauna of west Bengal. State Fauna Series 3. Zoological Survey of India, Kolkata, pp. 345 - 403. [dated 1998, published 1999]" type="journal article" year="1999">Deb 1999</bibRefCitation>
: fig. 4; 
<bibRefCitation id="EFDC4B70FFEDFF9FFE374D93C013FCA0" author="Trivedi, J. &amp; Mitra, S. &amp; Patel, P. &amp; Maheta, N. &amp; Patel, K. &amp; Ng, P. K. L." box="[385,596,763,789]" pageId="9" pageNumber="270" pagination="1 - 18" refId="ref7973" refString="Trivedi, J., Mitra, S., Patel, P., Maheta, N., Patel, K. &amp; Ng, P. K. L. (2021) On the Indian species of Eurycarcinus A. MilneEdwards, 1867, Heteropanope Stimpson, 1858, and Pilumnopeus A. Milne-Edwards, 1867 (Decapoda: Brachyura: Pilumnidae). Nauplius, 29 (e 2021004), 1 - 18. https: // doi. org / 10.1590 / 2358 - 2936 e 2021004" type="journal article" year="2021">
Trivedi 
<emphasis id="B939EA93FFEDFF9FFE6F4D93C04DFCA0" box="[473,522,763,789]" italics="true" pageId="9" pageNumber="270">et al</emphasis>
. 2021
</bibRefCitation>
: fig. 10G, H).
</paragraph>
<paragraph id="8BF23681FFEDFF9FFF714C77C62CFACC" blockId="9.[151,1437,151,1401]" pageId="9" pageNumber="270">
<taxonomicName id="4C4D4D02FFEDFF9FFF714C77C3D4FC8C" authority="Hari &amp; Hershey &amp; Mendoza, 2022" authorityName="Hari &amp; Hershey &amp; Mendoza" authorityYear="2022" box="[199,403,799,825]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="270" phylum="Arthropoda" rank="species" species="bijoyi" status="sp. nov.">
<emphasis id="B939EA93FFEDFF9FFF714C77C3D4FC8C" box="[199,403,799,825]" italics="true" pageId="9" pageNumber="270">Aniptumnus bijoyi</emphasis>
</taxonomicName>
<emphasis id="B939EA93FFEDFF9FFE2E4C77C3B7FC8C" bold="true" box="[408,496,799,825]" pageId="9" pageNumber="270">
<taxonomicNameLabel id="A20A57E8FFEDFF9FFE2E4C77C3B7FC8C" box="[408,496,799,825]" pageId="9" pageNumber="270" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from 
<taxonomicName id="4C4D4D02FFEDFF9FFD364C77C10FFC8C" baseAuthorityName="De Man" baseAuthorityYear="1895" box="[640,840,799,825]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="270" phylum="Arthropoda" rank="species" species="quadridentatus">
<emphasis id="B939EA93FFEDFF9FFD364C77C10FFC8C" box="[640,840,799,825]" italics="true" pageId="9" pageNumber="270">A. quadridentatus</emphasis>
</taxonomicName>
in the following morphological features: the carapace outline is generally wider and shorter, with anterolateral and posterolateral margins nearly identical in length, and the posterolateral margins slightly convex (
<figureCitation id="13762A04FFEDFF9FFDC94C0FC09EFC34" box="[639,729,871,897]" captionStart="FIGURE 2" captionStartId="5.[151,250,1891,1915]" captionTargetBox="[151,1435,181,1867]" captionTargetId="figure-20@5.[151,1435,181,1867]" captionTargetPageId="5" captionText="FIGURE 2. Aniptumnus bijoyi sp. nov., holotype, male, 10.7 by 7.1 mm (CUSAT/SBN/HRJ/1), Kochi, Kerala, SW India. A, carapace, dorsal view; B, left third maxilliped, external view; cephalothorax and chelipeds, frontal view; D, anterior thoracic sternum and telson, ventral view; E, thoracic sternum and sternopleonal cavity, ventral view; F, pleon, ventral view; G, pleonal somites 1–3, posteroventral view. Scale bars: 2.0 mm (A, C); 1.0 mm (B, D–G)." figureDoi="http://doi.org/10.5281/zenodo.7385093" httpUri="https://zenodo.org/record/7385093/files/figure.png" pageId="9" pageNumber="270">Fig. 2A</figureCitation>
) (vs. carapace outline narrower and longer, with posterolateral margins longer than anterolateral margins, and posterolateral margins straight in 
<taxonomicName id="4C4D4D02FFEDFF9FFB8F4CE3C742FC10" baseAuthorityName="De Man" baseAuthorityYear="1895" box="[1081,1285,907,933]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="270" phylum="Arthropoda" rank="species" species="quadridentatus">
<emphasis id="B939EA93FFEDFF9FFB8F4CE3C742FC10" box="[1081,1285,907,933]" italics="true" pageId="9" pageNumber="270">A. quadridentatus</emphasis>
</taxonomicName>
; 
<figureCitation id="13762A04FFEDFF9FFAA04CE3C736FC10" box="[1302,1393,906,933]" captionStart="FIGURE 5" captionStartId="8.[151,250,1856,1880]" captionTargetBox="[164,1422,181,1833]" captionTargetId="figure-21@8.[164,1422,181,1833]" captionTargetPageId="8" captionText="FIGURE 5. Aniptumnus quadridentatus (De Man, 1895), male, 11.1 by 7.4 mm (ZRC 2021.0421), off Pulau Ubin, Singapore. A, carapace, dorsal view; B, third maxillipeds, external view; C, cephalothorax, frontal view; D, anterior thoracic sternum and pleon, ventral view; E, pleonal somites 1–3 and thoracic sternite 8, posteroventral view; F–I, left G1 (F, pleonal/ventral view; G, mesiosternal/mesiodorsal view; H, lateral view; I, sternal/dorsal view). Scale bars: 2.0 mm (A, C); 1.0 mm (B, D, E); 0.5 mm (F–I)." figureDoi="http://doi.org/10.5281/zenodo.7385102" httpUri="https://zenodo.org/record/7385102/files/figure.png" pageId="9" pageNumber="270">Fig. 5A</figureCitation>
; cf. 
<bibRefCitation id="EFDC4B70FFEDFF9FFF214CC7C37AFC7C" author="De Man, J. G." box="[151,317,943,969]" pageId="9" pageNumber="270" pagination="725 - 790" refId="ref7316" refString="De Man, J. G. (1897) Bericht uber die von Herrn Schiffscapitan Storm zur Atjeh, an den westlichen Kusten von Malakka, Borneo und Celebes sowie in der Java-See gesammelten Decapoden und Stomatopoden. Funfter Theil. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 9 (6), 725 - 790, pls. 12 - 14" type="journal article" year="1897">De Man 1897</bibRefCitation>
: pl. 13 fig. 6; 
<bibRefCitation id="EFDC4B70FFEDFF9FFE5B4CC7C0C2FC7C" author="Takeda, M." box="[493,645,943,969]" pageId="9" pageNumber="270" pagination="87 - 93" refId="ref7918" refString="Takeda, M. (2001) On two estuarine crabs from Hainan Island, South China Sea. In: Matsuura, K. (Ed.), Marine fauna of the shallow waters around Hainan Island, South China Sea. National Science Museum Monographs, 21, 87 - 93." type="journal article" year="2001">Takeda 2001</bibRefCitation>
: figs. 2, 3A; 
<bibRefCitation id="EFDC4B70FFEDFF9FFC934CC7C620FC7C" author="Widyastuti, E. &amp; Rahayu, D. L." box="[805,1127,943,969]" pageId="9" pageNumber="270" pagination="15 - 22" refId="ref8067" refString="Widyastuti, E. &amp; Rahayu, D. L. (2022) The new record of Heteropilumnus sasekumari (Serene, 1971) and Aniptumnus quadridentatus (De Man, 1895) (Crustacea, Decapoda, Brachyura, Pilumnidae) from mangrove habitat in Papua, Indonesia. Oseanologi dan Limnologi di Indonesia, 7 (1), 15 - 22." type="journal article" year="2022">Widyastuti &amp; Rahayu 2022</bibRefCitation>
: fig. 3A); on the carapace anterolateral margin, the first tooth after the exorbital angle is much reduced, more like a flat lobe (
<figureCitation id="13762A04FFEDFF9FFABF4CBBC724FC58" box="[1289,1379,979,1005]" captionStart="FIGURE 2" captionStartId="5.[151,250,1891,1915]" captionTargetBox="[151,1435,181,1867]" captionTargetId="figure-20@5.[151,1435,181,1867]" captionTargetPageId="5" captionText="FIGURE 2. Aniptumnus bijoyi sp. nov., holotype, male, 10.7 by 7.1 mm (CUSAT/SBN/HRJ/1), Kochi, Kerala, SW India. A, carapace, dorsal view; B, left third maxilliped, external view; cephalothorax and chelipeds, frontal view; D, anterior thoracic sternum and telson, ventral view; E, thoracic sternum and sternopleonal cavity, ventral view; F, pleon, ventral view; G, pleonal somites 1–3, posteroventral view. Scale bars: 2.0 mm (A, C); 1.0 mm (B, D–G)." figureDoi="http://doi.org/10.5281/zenodo.7385093" httpUri="https://zenodo.org/record/7385093/files/figure.png" pageId="9" pageNumber="270">Fig. 2A</figureCitation>
) (vs. first tooth lobiform and low but with distinct, denticulate convex protuberance in 
<taxonomicName id="4C4D4D02FFEDFF9FFB8B4C9FC74FFBA4" baseAuthorityName="De Man" baseAuthorityYear="1895" box="[1085,1288,1015,1041]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="270" phylum="Arthropoda" rank="species" species="quadridentatus">
<emphasis id="B939EA93FFEDFF9FFB8B4C9FC74FFBA4" box="[1085,1288,1015,1041]" italics="true" pageId="9" pageNumber="270">A. quadridentatus</emphasis>
</taxonomicName>
; 
<figureCitation id="13762A04FFEDFF9FFAAE4C9FC735FBA4" box="[1304,1394,1014,1041]" captionStart="FIGURE 5" captionStartId="8.[151,250,1856,1880]" captionTargetBox="[164,1422,181,1833]" captionTargetId="figure-21@8.[164,1422,181,1833]" captionTargetPageId="8" captionText="FIGURE 5. Aniptumnus quadridentatus (De Man, 1895), male, 11.1 by 7.4 mm (ZRC 2021.0421), off Pulau Ubin, Singapore. A, carapace, dorsal view; B, third maxillipeds, external view; C, cephalothorax, frontal view; D, anterior thoracic sternum and pleon, ventral view; E, pleonal somites 1–3 and thoracic sternite 8, posteroventral view; F–I, left G1 (F, pleonal/ventral view; G, mesiosternal/mesiodorsal view; H, lateral view; I, sternal/dorsal view). Scale bars: 2.0 mm (A, C); 1.0 mm (B, D, E); 0.5 mm (F–I)." figureDoi="http://doi.org/10.5281/zenodo.7385102" httpUri="https://zenodo.org/record/7385102/files/figure.png" pageId="9" pageNumber="270">Fig. 5A</figureCitation>
; cf. 
<bibRefCitation id="EFDC4B70FFEDFF9FFF214B73C36AFB80" author="Takeda, M." box="[151,301,1051,1077]" pageId="9" pageNumber="270" pagination="87 - 93" refId="ref7918" refString="Takeda, M. (2001) On two estuarine crabs from Hainan Island, South China Sea. In: Matsuura, K. (Ed.), Marine fauna of the shallow waters around Hainan Island, South China Sea. National Science Museum Monographs, 21, 87 - 93." type="journal article" year="2001">Takeda 2001</bibRefCitation>
: fig. 3B; 
<bibRefCitation id="EFDC4B70FFEDFF9FFE2F4B73C046FB80" author="Ng, P. K. L." box="[409,513,1051,1077]" pageId="9" pageNumber="270" pagination="209 - 226" refId="ref7709" refString="Ng, P. K. L. (2002) The Indo-Pacific Pilumnidae. XVI. On the Identity of Pilumnus cristimanus A. Milne Edwards and the status of Parapilumnus Kossman, 1877 (Crustacea: Decapoda: Brachyura), with description of a new species from the rubble beds in Guam. Micronesica, 34 (2), 209 - 226." type="journal article" year="2002">Ng 2002</bibRefCitation>
: fig. 1A); the dorsal surface of the carapace is widely convex transversely in the frontal view (
<figureCitation id="13762A04FFEDFF9FFE844B57C3CDFBEC" box="[306,394,1087,1113]" captionStart="FIGURE 2" captionStartId="5.[151,250,1891,1915]" captionTargetBox="[151,1435,181,1867]" captionTargetId="figure-20@5.[151,1435,181,1867]" captionTargetPageId="5" captionText="FIGURE 2. Aniptumnus bijoyi sp. nov., holotype, male, 10.7 by 7.1 mm (CUSAT/SBN/HRJ/1), Kochi, Kerala, SW India. A, carapace, dorsal view; B, left third maxilliped, external view; cephalothorax and chelipeds, frontal view; D, anterior thoracic sternum and telson, ventral view; E, thoracic sternum and sternopleonal cavity, ventral view; F, pleon, ventral view; G, pleonal somites 1–3, posteroventral view. Scale bars: 2.0 mm (A, C); 1.0 mm (B, D–G)." figureDoi="http://doi.org/10.5281/zenodo.7385093" httpUri="https://zenodo.org/record/7385093/files/figure.png" pageId="9" pageNumber="270">Fig. 2C</figureCitation>
) (vs. dorsal surface more acutely convex transversely in frontal view in 
<taxonomicName id="4C4D4D02FFEDFF9FFB7C4B57C7D2FBEC" baseAuthorityName="De Man" baseAuthorityYear="1895" box="[1226,1429,1087,1113]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="270" phylum="Arthropoda" rank="species" species="quadridentatus">
<emphasis id="B939EA93FFEDFF9FFB7C4B57C7D2FBEC" box="[1226,1429,1087,1113]" italics="true" pageId="9" pageNumber="270">A. quadridentatus</emphasis>
</taxonomicName>
; 
<figureCitation id="13762A04FFEDFF9FFF214B0BC2A8FBC8" box="[151,239,1122,1149]" captionStart="FIGURE 5" captionStartId="8.[151,250,1856,1880]" captionTargetBox="[164,1422,181,1833]" captionTargetId="figure-21@8.[164,1422,181,1833]" captionTargetPageId="8" captionText="FIGURE 5. Aniptumnus quadridentatus (De Man, 1895), male, 11.1 by 7.4 mm (ZRC 2021.0421), off Pulau Ubin, Singapore. A, carapace, dorsal view; B, third maxillipeds, external view; C, cephalothorax, frontal view; D, anterior thoracic sternum and pleon, ventral view; E, pleonal somites 1–3 and thoracic sternite 8, posteroventral view; F–I, left G1 (F, pleonal/ventral view; G, mesiosternal/mesiodorsal view; H, lateral view; I, sternal/dorsal view). Scale bars: 2.0 mm (A, C); 1.0 mm (B, D, E); 0.5 mm (F–I)." figureDoi="http://doi.org/10.5281/zenodo.7385102" httpUri="https://zenodo.org/record/7385102/files/figure.png" pageId="9" pageNumber="270">Fig. 5C</figureCitation>
; cf. 
<bibRefCitation id="EFDC4B70FFEDFF9FFE974B0BC01CFBC8" author="Widyastuti, E. &amp; Rahayu, D. L." box="[289,603,1123,1149]" pageId="9" pageNumber="270" pagination="15 - 22" refId="ref8067" refString="Widyastuti, E. &amp; Rahayu, D. L. (2022) The new record of Heteropilumnus sasekumari (Serene, 1971) and Aniptumnus quadridentatus (De Man, 1895) (Crustacea, Decapoda, Brachyura, Pilumnidae) from mangrove habitat in Papua, Indonesia. Oseanologi dan Limnologi di Indonesia, 7 (1), 15 - 22." type="journal article" year="2022">Widyastuti &amp; Rahayu 2022</bibRefCitation>
: fig. 3B); there is no notch on the orbit at the junction of the supraorbital and infraorbital margins (
<figureCitation id="13762A04FFEDFF9FFE034BEFC04DFB14" box="[437,522,1159,1185]" captionStart="FIGURE 2" captionStartId="5.[151,250,1891,1915]" captionTargetBox="[151,1435,181,1867]" captionTargetId="figure-20@5.[151,1435,181,1867]" captionTargetPageId="5" captionText="FIGURE 2. Aniptumnus bijoyi sp. nov., holotype, male, 10.7 by 7.1 mm (CUSAT/SBN/HRJ/1), Kochi, Kerala, SW India. A, carapace, dorsal view; B, left third maxilliped, external view; cephalothorax and chelipeds, frontal view; D, anterior thoracic sternum and telson, ventral view; E, thoracic sternum and sternopleonal cavity, ventral view; F, pleon, ventral view; G, pleonal somites 1–3, posteroventral view. Scale bars: 2.0 mm (A, C); 1.0 mm (B, D–G)." figureDoi="http://doi.org/10.5281/zenodo.7385093" httpUri="https://zenodo.org/record/7385093/files/figure.png" pageId="9" pageNumber="270">Fig. 2C</figureCitation>
) (vs. present in 
<taxonomicName id="4C4D4D02FFEDFF9FFD094BEFC1CFFB14" baseAuthorityName="De Man" baseAuthorityYear="1895" box="[703,904,1159,1185]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="270" phylum="Arthropoda" rank="species" species="quadridentatus">
<emphasis id="B939EA93FFEDFF9FFD094BEFC1CFFB14" box="[703,904,1159,1185]" italics="true" pageId="9" pageNumber="270">A. quadridentatus</emphasis>
</taxonomicName>
; 
<figureCitation id="13762A04FFEDFF9FFC204BEFC1AAFB14" box="[918,1005,1158,1185]" captionStart="FIGURE 5" captionStartId="8.[151,250,1856,1880]" captionTargetBox="[164,1422,181,1833]" captionTargetId="figure-21@8.[164,1422,181,1833]" captionTargetPageId="8" captionText="FIGURE 5. Aniptumnus quadridentatus (De Man, 1895), male, 11.1 by 7.4 mm (ZRC 2021.0421), off Pulau Ubin, Singapore. A, carapace, dorsal view; B, third maxillipeds, external view; C, cephalothorax, frontal view; D, anterior thoracic sternum and pleon, ventral view; E, pleonal somites 1–3 and thoracic sternite 8, posteroventral view; F–I, left G1 (F, pleonal/ventral view; G, mesiosternal/mesiodorsal view; H, lateral view; I, sternal/dorsal view). Scale bars: 2.0 mm (A, C); 1.0 mm (B, D, E); 0.5 mm (F–I)." figureDoi="http://doi.org/10.5281/zenodo.7385102" httpUri="https://zenodo.org/record/7385102/files/figure.png" pageId="9" pageNumber="270">Fig. 5C</figureCitation>
); the distolateral angle of mxp3 merus is more rounded and projecting (
<figureCitation id="13762A04FFEDFF9FFDBD4BC3C025FB70" box="[523,610,1195,1221]" captionStart="FIGURE 2" captionStartId="5.[151,250,1891,1915]" captionTargetBox="[151,1435,181,1867]" captionTargetId="figure-20@5.[151,1435,181,1867]" captionTargetPageId="5" captionText="FIGURE 2. Aniptumnus bijoyi sp. nov., holotype, male, 10.7 by 7.1 mm (CUSAT/SBN/HRJ/1), Kochi, Kerala, SW India. A, carapace, dorsal view; B, left third maxilliped, external view; cephalothorax and chelipeds, frontal view; D, anterior thoracic sternum and telson, ventral view; E, thoracic sternum and sternopleonal cavity, ventral view; F, pleon, ventral view; G, pleonal somites 1–3, posteroventral view. Scale bars: 2.0 mm (A, C); 1.0 mm (B, D–G)." figureDoi="http://doi.org/10.5281/zenodo.7385093" httpUri="https://zenodo.org/record/7385093/files/figure.png" pageId="9" pageNumber="270">Fig. 2B</figureCitation>
) (vs. less projecting in 
<taxonomicName id="4C4D4D02FFEDFF9FFCD84BC3C67EFB70" baseAuthorityName="De Man" baseAuthorityYear="1895" box="[878,1081,1195,1221]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="270" phylum="Arthropoda" rank="species" species="quadridentatus">
<emphasis id="B939EA93FFEDFF9FFCD84BC3C67EFB70" box="[878,1081,1195,1221]" italics="true" pageId="9" pageNumber="270">A. quadridentatus</emphasis>
</taxonomicName>
; 
<figureCitation id="13762A04FFEDFF9FFBFE4BC3C6E6FB70" box="[1096,1185,1194,1221]" captionStart="FIGURE 5" captionStartId="8.[151,250,1856,1880]" captionTargetBox="[164,1422,181,1833]" captionTargetId="figure-21@8.[164,1422,181,1833]" captionTargetPageId="8" captionText="FIGURE 5. Aniptumnus quadridentatus (De Man, 1895), male, 11.1 by 7.4 mm (ZRC 2021.0421), off Pulau Ubin, Singapore. A, carapace, dorsal view; B, third maxillipeds, external view; C, cephalothorax, frontal view; D, anterior thoracic sternum and pleon, ventral view; E, pleonal somites 1–3 and thoracic sternite 8, posteroventral view; F–I, left G1 (F, pleonal/ventral view; G, mesiosternal/mesiodorsal view; H, lateral view; I, sternal/dorsal view). Scale bars: 2.0 mm (A, C); 1.0 mm (B, D, E); 0.5 mm (F–I)." figureDoi="http://doi.org/10.5281/zenodo.7385102" httpUri="https://zenodo.org/record/7385102/files/figure.png" pageId="9" pageNumber="270">Fig. 5B</figureCitation>
; cf. 
<bibRefCitation id="EFDC4B70FFEDFF9FFB654BC3C77EFB70" author="Ng, P. K. L." box="[1235,1337,1195,1221]" pageId="9" pageNumber="270" pagination="209 - 226" refId="ref7709" refString="Ng, P. K. L. (2002) The Indo-Pacific Pilumnidae. XVI. On the Identity of Pilumnus cristimanus A. Milne Edwards and the status of Parapilumnus Kossman, 1877 (Crustacea: Decapoda: Brachyura), with description of a new species from the rubble beds in Guam. Micronesica, 34 (2), 209 - 226." type="journal article" year="2002">Ng 2002</bibRefCitation>
: fig. 1B; 
<bibRefCitation id="EFDC4B70FFEDFF9FFF214BA7C38CFB5C" author="Widyastuti, E. &amp; Rahayu, D. L." box="[151,459,1231,1257]" pageId="9" pageNumber="270" pagination="15 - 22" refId="ref8067" refString="Widyastuti, E. &amp; Rahayu, D. L. (2022) The new record of Heteropilumnus sasekumari (Serene, 1971) and Aniptumnus quadridentatus (De Man, 1895) (Crustacea, Decapoda, Brachyura, Pilumnidae) from mangrove habitat in Papua, Indonesia. Oseanologi dan Limnologi di Indonesia, 7 (1), 15 - 22." type="journal article" year="2022">Widyastuti &amp; Rahayu 2022</bibRefCitation>
: fig. 3C); in the male, thoracic sternite 8 not visible when the pleon is folded and locked onto the thoracic sternum (
<figureCitation id="13762A04FFEDFF9FFE7A4B9BC062FAB8" box="[460,549,1267,1293]" captionStart="FIGURE 2" captionStartId="5.[151,250,1891,1915]" captionTargetBox="[151,1435,181,1867]" captionTargetId="figure-20@5.[151,1435,181,1867]" captionTargetPageId="5" captionText="FIGURE 2. Aniptumnus bijoyi sp. nov., holotype, male, 10.7 by 7.1 mm (CUSAT/SBN/HRJ/1), Kochi, Kerala, SW India. A, carapace, dorsal view; B, left third maxilliped, external view; cephalothorax and chelipeds, frontal view; D, anterior thoracic sternum and telson, ventral view; E, thoracic sternum and sternopleonal cavity, ventral view; F, pleon, ventral view; G, pleonal somites 1–3, posteroventral view. Scale bars: 2.0 mm (A, C); 1.0 mm (B, D–G)." figureDoi="http://doi.org/10.5281/zenodo.7385093" httpUri="https://zenodo.org/record/7385093/files/figure.png" pageId="9" pageNumber="270">Fig. 2G</figureCitation>
) (vs. sternite 8, sometimes exposed, small but distinct, positioned in between the lateral margin of the pleonal somite 2 and the mesial margin of the coxo-sternal condyle of the P5 coxa in 
<taxonomicName id="4C4D4D02FFEDFF9FFA324A7FC306FAE0" baseAuthorityName="De Man" baseAuthorityYear="1895" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="270" phylum="Arthropoda" rank="species" species="quadridentatus">
<emphasis id="B939EA93FFEDFF9FFA324A7FC306FAE0" italics="true" pageId="9" pageNumber="270">A. quadridentatus</emphasis>
</taxonomicName>
; 
<figureCitation id="13762A04FFEDFF9FFEE64A53C3E1FAE0" box="[336,422,1338,1365]" captionStart="FIGURE 5" captionStartId="8.[151,250,1856,1880]" captionTargetBox="[164,1422,181,1833]" captionTargetId="figure-21@8.[164,1422,181,1833]" captionTargetPageId="8" captionText="FIGURE 5. Aniptumnus quadridentatus (De Man, 1895), male, 11.1 by 7.4 mm (ZRC 2021.0421), off Pulau Ubin, Singapore. A, carapace, dorsal view; B, third maxillipeds, external view; C, cephalothorax, frontal view; D, anterior thoracic sternum and pleon, ventral view; E, pleonal somites 1–3 and thoracic sternite 8, posteroventral view; F–I, left G1 (F, pleonal/ventral view; G, mesiosternal/mesiodorsal view; H, lateral view; I, sternal/dorsal view). Scale bars: 2.0 mm (A, C); 1.0 mm (B, D, E); 0.5 mm (F–I)." figureDoi="http://doi.org/10.5281/zenodo.7385102" httpUri="https://zenodo.org/record/7385102/files/figure.png" pageId="9" pageNumber="270">Fig. 5E</figureCitation>
); and the distal tip of the G1 has a narrower apical lobe (
<figureCitation id="13762A04FFEDFF9FFB934A53C6A5FAE0" box="[1061,1250,1339,1365]" captionStart="FIGURE 3" captionStartId="6.[151,250,1891,1915]" captionTargetBox="[151,1435,181,1867]" captionTargetId="figure-21@6.[151,1435,181,1867]" captionTargetPageId="6" captionText="FIGURE 3. Aniptumnus bijoyi sp. nov., holotype, male, 10.7 by 7.1 mm (CUSAT/SBN/HRJ/1), Kochi, Kerala, SW India. A, minor cheliped, external view; B, major cheliped, external view; C, right P5, dorsal view; D, E, left G1 (D, pleonal/ventral view; E, mesiosternal/mesiodorsal view); F, left G2, sternal/dorsal view; G, H, distal tip of left G1 (G, pleonal/ventral view; H, mesiosternal/mesiodorsal view). Scale bars: 1.0 mm (A–C); 0.5 mm (D–F); 0.25 mm (G, H)." figureDoi="http://doi.org/10.5281/zenodo.7385098" httpUri="https://zenodo.org/record/7385098/files/figure.png" pageId="9" pageNumber="270">Fig. 3D, E, G, H</figureCitation>
) (vs. apical lobe wider in 
<taxonomicName id="4C4D4D02FFEDFF9FFF4D4A37C383FACC" baseAuthorityName="De Man" baseAuthorityYear="1895" box="[251,452,1375,1401]" class="Malacostraca" family="Pilumnidae" genus="Aniptumnus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="270" phylum="Arthropoda" rank="species" species="quadridentatus">
<emphasis id="B939EA93FFEDFF9FFF4D4A37C383FACC" box="[251,452,1375,1401]" italics="true" pageId="9" pageNumber="270">A. quadridentatus</emphasis>
</taxonomicName>
; 
<figureCitation id="13762A04FFEDFF9FFE644A37C079FACC" box="[466,574,1374,1401]" captionStart="FIGURE 5" captionStartId="8.[151,250,1856,1880]" captionTargetBox="[164,1422,181,1833]" captionTargetId="figure-21@8.[164,1422,181,1833]" captionTargetPageId="8" captionText="FIGURE 5. Aniptumnus quadridentatus (De Man, 1895), male, 11.1 by 7.4 mm (ZRC 2021.0421), off Pulau Ubin, Singapore. A, carapace, dorsal view; B, third maxillipeds, external view; C, cephalothorax, frontal view; D, anterior thoracic sternum and pleon, ventral view; E, pleonal somites 1–3 and thoracic sternite 8, posteroventral view; F–I, left G1 (F, pleonal/ventral view; G, mesiosternal/mesiodorsal view; H, lateral view; I, sternal/dorsal view). Scale bars: 2.0 mm (A, C); 1.0 mm (B, D, E); 0.5 mm (F–I)." figureDoi="http://doi.org/10.5281/zenodo.7385102" httpUri="https://zenodo.org/record/7385102/files/figure.png" pageId="9" pageNumber="270">Fig. 5F–I</figureCitation>
; cf. 
<bibRefCitation id="EFDC4B70FFEDFF9FFDDA4A37C147FACC" author="Takeda, M." box="[620,768,1375,1401]" pageId="9" pageNumber="270" pagination="87 - 93" refId="ref7918" refString="Takeda, M. (2001) On two estuarine crabs from Hainan Island, South China Sea. In: Matsuura, K. (Ed.), Marine fauna of the shallow waters around Hainan Island, South China Sea. National Science Museum Monographs, 21, 87 - 93." type="journal article" year="2001">Takeda 2001</bibRefCitation>
: fig. 3H, I; 
<bibRefCitation id="EFDC4B70FFEDFF9FFC364A37C1A2FACC" author="Ng, P. K. L." box="[896,997,1375,1401]" pageId="9" pageNumber="270" pagination="209 - 226" refId="ref7709" refString="Ng, P. K. L. (2002) The Indo-Pacific Pilumnidae. XVI. On the Identity of Pilumnus cristimanus A. Milne Edwards and the status of Parapilumnus Kossman, 1877 (Crustacea: Decapoda: Brachyura), with description of a new species from the rubble beds in Guam. Micronesica, 34 (2), 209 - 226." type="journal article" year="2002">Ng 2002</bibRefCitation>
: fig. 1F, G).
</paragraph>
</subSubSection>
</treatment>
</document>