222 lines
29 KiB
XML
222 lines
29 KiB
XML
<document id="ADF562980D336DDA760A8C03060516FA" ID-DOI="10.5281/zenodo.5188356" ID-ISSN="1942-1354" ID-Zenodo-Dep="5188356" ID-ZooBank="AC2597CC-301F-4E91-9711-5C17399C9AA2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628346141153" checkinUser="felipe" docAuthor="King, Jonas G. & Lago, Paul K." docDate="2012" docId="03E1561898280B2DFF19FF3B4F841FB9" docLanguage="en" docName="InsectaMundi.0275.1-53.pdf" docOrigin="Insecta Mundi 2012 (275)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Heterocerus insolens Miller 1994" docType="treatment" docVersion="4" lastPageNumber="18" masterDocId="FFD82E60983B0B3EFFD9FFF94D431A43" masterDocTitle="The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Mississippi and Alabama, with discussion and keys to the species occurring in the southeastern United States" masterLastPageNumber="53" masterPageNumber="1" pageNumber="18" updateTime="1699045744038" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
|
||
<mods:mods id="44CC45CED797F7B9BA3F9D56DD7ACE90" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="4480008EB743B9FAD36876588A144A4B">
|
||
<mods:title id="2E46F6000C1BE5D8DFF58909217584DE">The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Mississippi and Alabama, with discussion and keys to the species occurring in the southeastern United States</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="1752891B565A562DE00E2C772090A9F7" type="personal">
|
||
<mods:role id="C921AC644AAB556F2E4844CED40AAC83">
|
||
<mods:roleTerm id="9FF3A0CC91EDBF2F3C58A793F5E911A9">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="93EC166B1D3CE2AFFF08990018D5DD51">King, Jonas G.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="20C6B81CB025EC62EFB7B98B865DB103" type="personal">
|
||
<mods:role id="AEFC892FE411C4642D2ABD0CAB059A6B">
|
||
<mods:roleTerm id="B8D062CC714D4044108A4F250808C2EB">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="24AD4CB0FB3E3EA97EBA9ABBE8182DEE">Lago, Paul K.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="35AE6354F6A4C3AA597304BAC18BF7E4">text</mods:typeOfResource>
|
||
<mods:relatedItem id="02577BBE81FE74E7B6CACC7145B01430" type="host">
|
||
<mods:titleInfo id="98C9BB793176E4BBF1146E3A9A4B5C5F">
|
||
<mods:title id="D6F6E980E0DBF4123447D7421F74DF72">Insecta Mundi</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="84B2530D979532B8E949B39B7141C41B">
|
||
<mods:date id="6C86BDA56BA98E3442D461F60A11BD25">2012</mods:date>
|
||
<mods:detail id="07BE05CC1A1ACF803AD662AA07590382" type="pubDate">
|
||
<mods:number id="D111B835BC58602F828769E8183F05A3">2012-12-28</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="78CD29629B828FD84282973F8A6F8010" type="volume">
|
||
<mods:number id="047A4E04A39C1ACD0BA8B6D0BEEC7F7C">2012</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="5B4BDC91621D037E04AADAD70DD9E0B6" type="issue">
|
||
<mods:number id="DFEC0ED3CC5FE80F335D0E6C456A3F3B">275</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="7F5FB07F43D3DDFABC4F5C937D85E749" unit="page">
|
||
<mods:start id="0C1006066F1585C51647E911EEBD4CB7">1</mods:start>
|
||
<mods:end id="D3537D26883679327A886294D8B0BBF1">53</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="1AE5070BB1D76E604E1DA81C48F876E0">journal article</mods:classification>
|
||
<mods:identifier id="9B2F71E7BDF16AC8DB7A15C4C12D339A" type="DOI">10.5281/zenodo.5188356</mods:identifier>
|
||
<mods:identifier id="1A5B1660925D291677348F4B4FAFFF1A" type="ISSN">1942-1354</mods:identifier>
|
||
<mods:identifier id="0385B3420085653815B30900AC3DDA6F" type="Zenodo-Dep">5188356</mods:identifier>
|
||
<mods:identifier id="3494603C76C21A3C6AFA01F74E7C0512" type="ZooBank">AC2597CC-301F-4E91-9711-5C17399C9AA2</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03E1561898280B2DFF19FF3B4F841FB9" ID-DOI="http://doi.org/10.5281/zenodo.5187841" ID-Zenodo-Dep="5187841" LSID="urn:lsid:plazi:treatment:03E1561898280B2DFF19FF3B4F841FB9" httpUri="http://treatment.plazi.org/id/03E1561898280B2DFF19FF3B4F841FB9" lastPageNumber="18" pageId="19" pageNumber="18">
|
||
<subSubSection id="C352B48598280B2DFF19FF3B4F121A99" box="[192,593,194,218]" pageId="19" pageNumber="18" type="nomenclature">
|
||
<paragraph id="8BF7E70E98280B2DFF19FF3B4F121A99" blockId="19.[192,593,194,250]" box="[192,593,194,218]" pageId="19" pageNumber="18">
|
||
<heading id="D0BF506298280B2DFF19FF3B4F121A99" bold="true" box="[192,593,194,218]" fontSize="10" level="2" pageId="19" pageNumber="18" reason="3">
|
||
<emphasis id="B93C3B1C98280B2DFF19FF3B4F121A99" bold="true" box="[192,593,194,218]" pageId="19" pageNumber="18">
|
||
4.
|
||
<taxonomicName id="4C489C8D98280B2DFF04FF3A4F121A99" authority="Miller" authorityName="Miller" authorityYear="1994" box="[221,593,194,218]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="insolens">
|
||
<emphasis id="B93C3B1C98280B2DFF04FF3A4CBB1A99" bold="true" box="[221,504,194,218]" italics="true" pageId="19" pageNumber="18">Heterocerus insolens</emphasis>
|
||
Miller
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C352B48598280B2DFF19FF1B4CC11AB9" box="[192,386,226,250]" pageId="19" pageNumber="18" type="description">
|
||
<paragraph id="8BF7E70E98280B2DFF19FF1B4CC11AB9" blockId="19.[192,593,194,250]" box="[192,386,226,250]" pageId="19" pageNumber="18">
|
||
(
|
||
<figureCitation id="1373FB8B98280B2DFF10FF1B4C621AB9" box="[201,289,226,250]" captionStart="Figures 18-20" captionStartId="16.[192,286,1329,1351]" captionTargetBox="[217,1350,191,1276]" captionTargetId="figure-640@16.[746,784,656,734]" captionTargetPageId="16" captionText="Figures 18-20. Habiti of large Heterocerus species with post-mesocoxal lines. 18) H. insolens. 19) H. sinuosus. 20) H. mollinus. Specimens 20a and 20c are hypermandibulate. Scale line = 1mm." figureDoi="http://doi.org/10.5281/zenodo.5188370" httpUri="https://zenodo.org/record/5188370/files/figure.png" pageId="19" pageNumber="18">Fig. 18</figureCitation>
|
||
,
|
||
<figureCitation id="1373FB8B98280B2DFEF7FF1B4C0D1AB9" box="[302,334,226,250]" captionStart="Figures 32-44" captionStartId="38.[192,286,1682,1704]" captionTargetBox="[192,1357,192,1649]" captionTargetId="figure-419@38.[688,779,856,946]" captionTargetPageId="38" captionText="Figures 32-44. Male genitalia of southeastern Heterocerus species. The ventral view is shown unless otherwise stated. 32-39) Species that lack post-mesocoxal lines. 32) H. fatuus. 33) H. glicki. 34) H. intermuralis. 35) H. angustatus; left - ventral view, right - dorsal view. 36) H. longilobulus. 37) H. pallidus; left - ventral view, right - dorsal view. 38) H. gnatho; left - ventral view, right - dorsal view. 39) H. sandersoni; left - ventral view, center - dorsal view, left - lateral view. 40-44) Species with post-mesocoxal lines. 40) H. sinuosus; left - dorsal view, right - ventral aspect. 41) H. mollinus. 42) H. insolens; left - dorsal view, right - ventral aspect. 43) H. parrotus. 44) H. brunneus." figureDoi="http://doi.org/10.5281/zenodo.5188382" httpUri="https://zenodo.org/record/5188382/files/figure.png" pageId="19" pageNumber="18">42</figureCitation>
|
||
,
|
||
<figureCitation id="1373FB8B98280B2DFE82FF1B4C3A1AB9" box="[347,377,226,250]" captionStart="Figures 54-57" captionStartId="39.[192,286,1864,1886]" captionTargetBox="[192,1436,1076,1850]" captionTargetId="figure-932@39.[788,832,1441,1510]" captionTargetPageId="39" captionText="Figures 54-57. Distribution by county in the United States. 54) Heterocerus pallidus. 55) Heterocerus gnatho. 56) Heterocerus longilobulus. 57) Heterocerus insolens." figureDoi="http://doi.org/10.5281/zenodo.5188388" httpUri="https://zenodo.org/record/5188388/files/figure.png" pageId="19" pageNumber="18">57</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C352B48598280B2DFF19FEDA4F381B79" box="[192,635,290,314]" pageId="19" pageNumber="18" type="reference_group">
|
||
<paragraph id="8BF7E70E98280B2DFF19FEDA4F381B79" blockId="19.[192,635,290,314]" box="[192,635,290,314]" pageId="19" pageNumber="18">
|
||
<treatmentCitationGroup id="AB58C02098280B2DFF19FEDA4F381B79" box="[192,635,290,314]" pageId="19" pageNumber="18">
|
||
<treatmentCitation id="0AE9C11F98280B2DFF19FEDA4F351B79" author="Miller, W. V." box="[192,630,290,314]" page="16" pageId="19" pageNumber="18" year="1994">
|
||
<taxonomicName id="4C489C8D98280B2DFF19FEDA4F351B79" authority="Miller 1994: 16" authorityName="Miller" authorityPageNumber="16" authorityYear="1994" box="[192,630,290,314]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="insolens">
|
||
<emphasis id="B93C3B1C98280B2DFF19FEDA4CFA1B79" box="[192,441,290,314]" italics="true" pageId="19" pageNumber="18">Heterocerus insolens</emphasis>
|
||
<bibRefCitation id="EFD99AFF98280B2DFE19FEDB4F351B79" author="Miller, W. V." box="[448,630,290,314]" pageId="19" pageNumber="18" pagination="11 - 18" refId="ref21914" refString="Miller, W. V. 1994. New species of Heteroceridae from Canada and the United States. Coleopterists Bulletin 48: 11 - 18." type="journal article" year="1994">Miller 1994:16</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C352B48598280B2DFF19FE9B4F731879" pageId="19" pageNumber="18" type="description">
|
||
<paragraph id="8BF7E70E98280B2DFF19FE9B4F731879" blockId="19.[192,1441,354,570]" pageId="19" pageNumber="18">
|
||
<emphasis id="B93C3B1C98280B2DFF19FE9B4C281B39" bold="true" box="[192,363,354,378]" pageId="19" pageNumber="18">Description.</emphasis>
|
||
Length
|
||
<quantity id="4CB04AEB98280B2DFE15FE9B4F231B39" box="[460,608,354,378]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.5" metricValueMin="4.5" pageId="19" pageNumber="18" unit="mm" value="5.0" valueMax="5.5" valueMin="4.5">4.5 - 5.5 mm</quantity>
|
||
. Chestnut brown. Apical angles of pronotum usually pale. Elytra dark brown and distinctly trifaciate with relatively large medial and distal pale markings, and small broken pale basal marks; striae absent (
|
||
<figureCitation id="1373FB8B98280B2DFD9FFE5B4FDF1BF9" box="[582,668,418,442]" captionStart="Figures 18-20" captionStartId="16.[192,286,1329,1351]" captionTargetBox="[217,1350,191,1276]" captionTargetId="figure-640@16.[746,784,656,734]" captionTargetPageId="16" captionText="Figures 18-20. Habiti of large Heterocerus species with post-mesocoxal lines. 18) H. insolens. 19) H. sinuosus. 20) H. mollinus. Specimens 20a and 20c are hypermandibulate. Scale line = 1mm." figureDoi="http://doi.org/10.5281/zenodo.5188370" httpUri="https://zenodo.org/record/5188370/files/figure.png" pageId="19" pageNumber="18">Fig. 18</figureCitation>
|
||
). Post-metathoracic coxal lines absent; post-mesothoracic coxal lines prominent. Male genitalia (
|
||
<figureCitation id="1373FB8B98280B2DFD8FFE3B4FED1B99" box="[598,686,450,474]" captionStart="Figures 32-44" captionStartId="38.[192,286,1682,1704]" captionTargetBox="[192,1357,192,1649]" captionTargetId="figure-419@38.[688,779,856,946]" captionTargetPageId="38" captionText="Figures 32-44. Male genitalia of southeastern Heterocerus species. The ventral view is shown unless otherwise stated. 32-39) Species that lack post-mesocoxal lines. 32) H. fatuus. 33) H. glicki. 34) H. intermuralis. 35) H. angustatus; left - ventral view, right - dorsal view. 36) H. longilobulus. 37) H. pallidus; left - ventral view, right - dorsal view. 38) H. gnatho; left - ventral view, right - dorsal view. 39) H. sandersoni; left - ventral view, center - dorsal view, left - lateral view. 40-44) Species with post-mesocoxal lines. 40) H. sinuosus; left - dorsal view, right - ventral aspect. 41) H. mollinus. 42) H. insolens; left - dorsal view, right - ventral aspect. 43) H. parrotus. 44) H. brunneus." figureDoi="http://doi.org/10.5281/zenodo.5188382" httpUri="https://zenodo.org/record/5188382/files/figure.png" pageId="19" pageNumber="18">Fig. 42</figureCitation>
|
||
) of the typical
|
||
<taxonomicName id="4C489C8D98280B2DFCB0FE3A4EBA1B99" authorityName="Fabricius" authorityYear="1792" box="[873,1017,451,474]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93C3B1C98280B2DFCB0FE3A4EBA1B99" box="[873,1017,451,474]" italics="true" pageId="19" pageNumber="18">Heterocerus</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="54F359AC98280B2DFBD8FE3A497A1B98" box="[1025,1081,451,475]" pageId="19" pageNumber="18">type</typeStatus>
|
||
, phallobase constricted to its narrowest point at anterior third and then slightly broadened towards outer edges of parameres; medial edges of parameres approximately 30
|
||
<superScript id="7C3D4A4698280B2DFD19FDF84F8B184C" attach="left" box="[704,712,513,527]" fontSize="6" pageId="19" pageNumber="18">o</superScript>
|
||
divergent; dorsal plate ovoid, with no constrictions. Hypermandibulate males rare.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C352B48598280B2DFF19FD9B4FEE1E98" pageId="19" pageNumber="18" type="diagnosis">
|
||
<paragraph id="8BF7E70E98280B2DFF19FD9B4EBD19B9" blockId="19.[192,1441,610,1018]" pageId="19" pageNumber="18">
|
||
<emphasis id="B93C3B1C98280B2DFF19FD9B4C0E1839" bold="true" box="[192,333,610,634]" pageId="19" pageNumber="18">Diagnosis.</emphasis>
|
||
The presence of post-mesothoracic coxal lines and distinctive elytral coloration distinguishes this species from all other southeastern species except
|
||
<taxonomicName id="4C489C8D98280B2DFC81FD7A4EAA18D9" authorityName="Kiesenwetter" authorityYear="1851" box="[856,1001,642,666]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="mollinus">
|
||
<emphasis id="B93C3B1C98280B2DFC81FD7A4EAA18D9" box="[856,1001,642,666]" italics="true" pageId="19" pageNumber="18">H. mollinus</emphasis>
|
||
</taxonomicName>
|
||
, the most common North American
|
||
<taxonomicName id="4C489C8D98280B2DFF19FD5A4C1318F9" authorityName="Fabricius" authorityYear="1792" box="[192,336,675,698]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93C3B1C98280B2DFF19FD5A4C1318F9" box="[192,336,675,698]" italics="true" pageId="19" pageNumber="18">Heterocerus</emphasis>
|
||
</taxonomicName>
|
||
. In most specimens of
|
||
<emphasis id="B93C3B1C98280B2DFDB3FD5A4FB918F9" box="[618,762,674,698]" italics="true" pageId="19" pageNumber="18">
|
||
<taxonomicName id="4C489C8D98280B2DFDB3FD5A4FB618F9" authorityName="Miller" authorityYear="1994" box="[618,757,674,698]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="insolens">H. insolens</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
the two distal pale marks on the elytra are much more extensive than is typical of
|
||
<taxonomicName id="4C489C8D98280B2DFDDDFD3A4FD71899" authorityName="Kiesenwetter" authorityYear="1851" box="[516,660,706,730]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="mollinus">
|
||
<emphasis id="B93C3B1C98280B2DFDDDFD3A4FD71899" box="[516,660,706,730]" italics="true" pageId="19" pageNumber="18">H. mollinus</emphasis>
|
||
</taxonomicName>
|
||
(compare
|
||
<figureCitation id="1373FB8B98280B2DFCC8FD3B4EFF1899" box="[785,956,706,730]" captionStart="Figures 18-20" captionStartId="16.[192,286,1329,1351]" captionTargetBox="[217,1350,191,1276]" captionTargetId="figure-640@16.[746,784,656,734]" captionTargetPageId="16" captionText="Figures 18-20. Habiti of large Heterocerus species with post-mesocoxal lines. 18) H. insolens. 19) H. sinuosus. 20) H. mollinus. Specimens 20a and 20c are hypermandibulate. Scale line = 1mm." figureDoi="http://doi.org/10.5281/zenodo.5188370" httpUri="https://zenodo.org/record/5188370/files/figure.png" pageId="19" pageNumber="18">Fig. 18 and 20</figureCitation>
|
||
). Some males of
|
||
<taxonomicName id="4C489C8D98280B2DFB58FD3A48451899" authorityName="Miller" authorityYear="1994" box="[1153,1286,706,730]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="insolens">
|
||
<emphasis id="B93C3B1C98280B2DFB58FD3A48451899" box="[1153,1286,706,730]" italics="true" pageId="19" pageNumber="18">H. insolens</emphasis>
|
||
</taxonomicName>
|
||
are reported to have a small lateral projection on the mandibles, which are absent on
|
||
<taxonomicName id="4C489C8D98280B2DFBE8FD1A482818B9" authority="(Miller 1994)" baseAuthorityName="Miller" baseAuthorityYear="1994" box="[1073,1387,738,762]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="mollinus">
|
||
<emphasis id="B93C3B1C98280B2DFBE8FD1A498218B9" box="[1073,1217,738,762]" italics="true" pageId="19" pageNumber="18">H. mollinus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD99AFF98280B2DFB09FD1B482118B9" author="Miller, W. V." box="[1232,1378,738,762]" pageId="19" pageNumber="18" pagination="11 - 18" refId="ref21914" refString="Miller, W. V. 1994. New species of Heteroceridae from Canada and the United States. Coleopterists Bulletin 48: 11 - 18." type="journal article" year="1994">Miller 1994</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, but we have not observed this character. The abruptly constricted anterior end of the phallobase and the divergent parameres immediately distinguish
|
||
<taxonomicName id="4C489C8D98280B2DFD24FCDA4EC41979" authorityName="Miller" authorityYear="1994" box="[765,903,802,826]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="insolens">
|
||
<emphasis id="B93C3B1C98280B2DFD24FCDA4EC41979" box="[765,903,802,826]" italics="true" pageId="19" pageNumber="18">H. insolens</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C489C8D98280B2DFC0DFCDA49241979" baseAuthorityName="Miller" baseAuthorityYear="1994" box="[980,1127,802,826]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="mollinus">
|
||
<emphasis id="B93C3B1C98280B2DFC0DFCDA49241979" box="[980,1127,802,826]" italics="true" pageId="19" pageNumber="18">H. mollinus</emphasis>
|
||
</taxonomicName>
|
||
, in which the phallobase slowly tapers anteriorly and the internal edges of the parameres only slightly diverge (compare
|
||
<figureCitation id="1373FB8B98280B2DFA95FCBB4C571939" captionStart="Figures 32-44" captionStartId="38.[192,286,1682,1704]" captionTargetBox="[192,1357,192,1649]" captionTargetId="figure-419@38.[688,779,856,946]" captionTargetPageId="38" captionText="Figures 32-44. Male genitalia of southeastern Heterocerus species. The ventral view is shown unless otherwise stated. 32-39) Species that lack post-mesocoxal lines. 32) H. fatuus. 33) H. glicki. 34) H. intermuralis. 35) H. angustatus; left - ventral view, right - dorsal view. 36) H. longilobulus. 37) H. pallidus; left - ventral view, right - dorsal view. 38) H. gnatho; left - ventral view, right - dorsal view. 39) H. sandersoni; left - ventral view, center - dorsal view, left - lateral view. 40-44) Species with post-mesocoxal lines. 40) H. sinuosus; left - dorsal view, right - ventral aspect. 41) H. mollinus. 42) H. insolens; left - dorsal view, right - ventral aspect. 43) H. parrotus. 44) H. brunneus." figureDoi="http://doi.org/10.5281/zenodo.5188382" httpUri="https://zenodo.org/record/5188382/files/figure.png" pageId="19" pageNumber="18">Fig. 41 and 42</figureCitation>
|
||
). Another southeastern species with post-mesothoracic coxal lines,
|
||
<emphasis id="B93C3B1C98280B2DFBEDFC9A498B1939" box="[1076,1224,867,890]" italics="true" pageId="19" pageNumber="18">
|
||
<taxonomicName id="4C489C8D98280B2DFBEDFC9A49801939" authorityName="King & Lago" authorityYear="2012" baseAuthorityName="King & Lago" baseAuthorityYear="2012" box="[1076,1219,867,890]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="sinuosus">H. sinuosus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
has the pronotum consistently darker than the head and elytra, which immediately distinguishes it from
|
||
<taxonomicName id="4C489C8D98280B2DFB05FC7A482019D9" authorityName="Miller" authorityYear="1994" box="[1244,1379,898,922]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="insolens">
|
||
<emphasis id="B93C3B1C98280B2DFB05FC7A482019D9" box="[1244,1379,898,922]" italics="true" pageId="19" pageNumber="18">H. insolens</emphasis>
|
||
</taxonomicName>
|
||
. The external features and genital morphology of this species are also similar to those of
|
||
<taxonomicName id="4C489C8D98280B2DFB31FC5A48C319F9" authorityName="Melsheimer" authorityYear="1844" box="[1256,1408,930,954]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="brunneus">
|
||
<emphasis id="B93C3B1C98280B2DFB31FC5A48C319F9" box="[1256,1408,930,954]" italics="true" pageId="19" pageNumber="18">H. brunneus</emphasis>
|
||
</taxonomicName>
|
||
, a western species with several dubious southeastern records (see
|
||
<bibRefCitation id="EFD99AFF98280B2DFC13FC3B49361999" author="Pacheco, F." box="[970,1141,962,986]" pageId="19" pageNumber="18" pagination="1 - 155" refId="ref22030" refString="Pacheco, F. 1964. Sistematica, filogenia y distribucion de los Heteroceridos de America (Coleoptera; Heteroceridae). Monografias del Colegio de Post-graduados, No. 1, Escuela Nacional de Agricultura, Chapingo, Mexico, Vol. 1: 1 - 155." type="journal article" year="1964">Pacheco 1964</bibRefCitation>
|
||
). Characters of the male genitalia must be used to separate these two species (see couplet 19).
|
||
</paragraph>
|
||
<paragraph id="8BF7E70E98280B2DFF19FBDB4CE51ED9" blockId="19.[192,1440,1058,1243]" pageId="19" pageNumber="18">
|
||
<emphasis id="B93C3B1C98280B2DFF19FBDB4C4C1E79" bold="true" box="[192,271,1058,1082]" pageId="19" pageNumber="18">Notes</emphasis>
|
||
. During this project,
|
||
<taxonomicName id="4C489C8D98280B2DFDC4FBDA4FE51E79" authorityName="Miller" authorityYear="1994" box="[541,678,1058,1082]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="insolens">
|
||
<emphasis id="B93C3B1C98280B2DFDC4FBDA4FE51E79" box="[541,678,1058,1082]" italics="true" pageId="19" pageNumber="18">H. insolens</emphasis>
|
||
</taxonomicName>
|
||
was collected on two occasions. In both instances the beetles were taken from cypress-dominated overflow areas along the Tallahatchie and Yazoo rivers. Previously collected
|
||
<taxonomicName id="4C489C8D98280B2DFEEDFB9A4CFD1E39" authorityName="Miller" authorityYear="1994" box="[308,446,1122,1146]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="insolens">
|
||
<emphasis id="B93C3B1C98280B2DFEEDFB9A4CFD1E39" box="[308,446,1122,1146]" italics="true" pageId="19" pageNumber="18">H. insolens</emphasis>
|
||
</taxonomicName>
|
||
came from a variety of habitats, including small streams above the fall line in northern
|
||
<collectingRegion id="498C29EC98280B2DFEEAFB7B4CE01ED9" box="[307,419,1154,1178]" country="United States of America" name="Alabama" pageId="19" pageNumber="18">Alabama</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF7E70E98280B2DFF29FB5B4FEE1E98" blockId="19.[192,1440,1058,1243]" pageId="19" pageNumber="18">
|
||
Several of Miller’s
|
||
<typeStatus id="54F359AC98280B2DFE08FB5A4F0A1EF8" box="[465,585,1187,1211]" pageId="19" pageNumber="18" type="paratype">paratypes</typeStatus>
|
||
were found to be misidentified specimens of
|
||
<taxonomicName id="4C489C8D98280B2DFBB8FB5A49AC1EF9" baseAuthorityName="Miller" baseAuthorityYear="1994" box="[1121,1263,1186,1210]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="mollinus">
|
||
<emphasis id="B93C3B1C98280B2DFBB8FB5A49AC1EF9" box="[1121,1263,1186,1210]" italics="true" pageId="19" pageNumber="18">H. mollinus</emphasis>
|
||
</taxonomicName>
|
||
. Genitalia had not been extracted from these
|
||
<typeStatus id="54F359AC98280B2DFDF5FB3A4FEA1E98" box="[556,681,1219,1243]" pageId="19" pageNumber="18" type="paratype">paratypes</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C352B48598280B2DFF19FAFB4F841FB9" pageId="19" pageNumber="18" type="distribution">
|
||
<paragraph id="8BF7E70E98280B2DFF19FAFB4C581FF9" blockId="19.[192,1440,1282,1466]" pageId="19" pageNumber="18">
|
||
<emphasis id="B93C3B1C98280B2DFF19FAFB4C2F1F59" bold="true" box="[192,364,1282,1306]" pageId="19" pageNumber="18">Distribution</emphasis>
|
||
.
|
||
<taxonomicName id="4C489C8D98280B2DFEA3FAFA4F311F59" authorityName="Miller" authorityYear="1994" box="[378,626,1282,1306]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="insolens">
|
||
<emphasis id="B93C3B1C98280B2DFEA3FAFA4F311F59" box="[378,626,1282,1306]" italics="true" pageId="19" pageNumber="18">Heterocerus insolens</emphasis>
|
||
</taxonomicName>
|
||
has been collected in four different physiographic regions above the fall line in northern
|
||
<collectingRegion id="498C29EC98280B2DFE10FADB4F751F79" box="[457,566,1314,1338]" country="United States of America" name="Alabama" pageId="19" pageNumber="18">Alabama</collectingRegion>
|
||
and it is also common throughout the Alluvial and Coastal plains of
|
||
<collectingRegion id="498C29EC98280B2DFF19FABB4C0B1F19" box="[192,328,1346,1370]" country="United States of America" name="Mississippi" pageId="19" pageNumber="18">Mississippi</collectingRegion>
|
||
and
|
||
<collectingRegion id="498C29EC98280B2DFE5FFABB4CB71F19" box="[390,500,1346,1370]" country="United States of America" name="Alabama" pageId="19" pageNumber="18">Alabama</collectingRegion>
|
||
(
|
||
<figureCitation id="1373FB8B98280B2DFDDCFABB4F1C1F19" box="[517,607,1346,1370]" captionStart="Figures 54-57" captionStartId="39.[192,286,1864,1886]" captionTargetBox="[192,1436,1076,1850]" captionTargetId="figure-932@39.[788,832,1441,1510]" captionTargetPageId="39" captionText="Figures 54-57. Distribution by county in the United States. 54) Heterocerus pallidus. 55) Heterocerus gnatho. 56) Heterocerus longilobulus. 57) Heterocerus insolens." figureDoi="http://doi.org/10.5281/zenodo.5188388" httpUri="https://zenodo.org/record/5188388/files/figure.png" pageId="19" pageNumber="18">Fig. 57</figureCitation>
|
||
).
|
||
<bibRefCitation id="EFD99AFF98280B2DFDADFABB4E551F19" author="Miller, W. V." box="[628,790,1346,1370]" pageId="19" pageNumber="18" pagination="11 - 18" refId="ref21914" refString="Miller, W. V. 1994. New species of Heteroceridae from Canada and the United States. Coleopterists Bulletin 48: 11 - 18." type="journal article" year="1994">Miller (1994)</bibRefCitation>
|
||
provided single records from
|
||
<collectingRegion id="498C29EC98280B2DFB50FABB48401F19" box="[1161,1283,1346,1370]" country="United States of America" name="Louisiana" pageId="19" pageNumber="18">Louisiana</collectingRegion>
|
||
and
|
||
<collectingRegion id="498C29EC98280B2DFA98FABB48DF1F19" box="[1345,1436,1346,1370]" country="United States of America" name="Illinois" pageId="19" pageNumber="18">Illinois</collectingRegion>
|
||
. Since this distribution indicates adaptation to a variety of habitats, and because the species was described fairly recently, it is probable that
|
||
<taxonomicName id="4C489C8D98280B2DFD18FA7A4E0A1FD9" authorityName="Miller" authorityYear="1994" box="[705,841,1410,1434]" class="Insecta" family="Heteroceridae" genus="Heterocerus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="insolens">
|
||
<emphasis id="B93C3B1C98280B2DFD18FA7A4E0A1FD9" box="[705,841,1410,1434]" italics="true" pageId="19" pageNumber="18">H. insolens</emphasis>
|
||
</taxonomicName>
|
||
has a much wider distribution than is currently known.
|
||
</paragraph>
|
||
<paragraph id="8BF7E70E98280B2DFF19FA1B4F841FB9" blockId="19.[192,711,1506,1530]" box="[192,711,1506,1530]" pageId="19" pageNumber="18">
|
||
<emphasis id="B93C3B1C98280B2DFF19FA1B4CA01FB9" bold="true" box="[192,483,1506,1530]" pageId="19" pageNumber="18">Specimens examined.</emphasis>
|
||
98 (See Appendix).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |