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<document id="9FF1224A18DC41E8168C8FC928DA0CDF" ID-DOI="10.11646/zootaxa.3670.3.2" ID-GBIF-Dataset="214a6898-e56a-4a5a-948f-7a197b245bcb" ID-ISSN="1175-5326" ID-Zenodo-Dep="223299" ID-ZooBank="3F2904A2-A999-484A-850B-4E2E13A9DBD7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460634060144" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2013" docId="03DC5A34FF8D5031DBEF406816D4FA41" docLanguage="en" docName="zt03670p328.pdf" docOrigin="Zootaxa 3670 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Plaumanniola sanctaecatharinae Costa Lima" docType="treatment" docVersion="7" lastPageNumber="322" masterDocId="FFE5224CFF8E5034DB7845541762FFA5" masterDocTitle="A new species of the putatively myrmecophilous genus Plaumanniola Costa Lima, with notes on the systematic position of Plaumanniolini (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="328" masterPageNumber="317" pageNumber="320" updateTime="1698334982260" updateUser="plazi">
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<mods:title id="6A5321800A66025816A9793B3F511B9A">A new species of the putatively myrmecophilous genus Plaumanniola Costa Lima, with notes on the systematic position of Plaumanniolini (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
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<mods:namePart id="9E0A7277283154FAD2FDB2BB9706BC9D">Jałoszyński, Paweł</mods:namePart>
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<mods:date id="5B641D86E6D3B336790EA3DECEC679AD">2013</mods:date>
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<treatment id="03DC5A34FF8D5031DBEF406816D4FA41" ID-DOI="http://doi.org/10.5281/zenodo.6148788" ID-GBIF-Taxon="119570269" ID-Zenodo-Dep="6148788" LSID="urn:lsid:plazi:treatment:03DC5A34FF8D5031DBEF406816D4FA41" httpUri="http://treatment.plazi.org/id/03DC5A34FF8D5031DBEF406816D4FA41" lastPageId="5" lastPageNumber="322" pageId="3" pageNumber="320">
<subSubSection id="C36FB8A9FF8D5037DBEF406816E3FAD3" pageId="3" pageNumber="320" type="nomenclature">
<paragraph id="8BCAEB22FF8D5037DBEF406815A3FAF0" blockId="3.[151,705,1339,1398]" box="[151,705,1339,1366]" pageId="3" pageNumber="320">
<heading id="D0825C4EFF8D5037DBEF406815A3FAF0" bold="true" box="[151,705,1339,1366]" fontSize="11" level="1" pageId="3" pageNumber="320" reason="1">
<taxonomicName id="4C7590A1FF8D5037DBEF406815A3FAF0" ID-CoL="4K24B" authority="Costa Lima" authorityName="Costa Lima" box="[151,705,1339,1366]" class="Insecta" family="Staphylinidae" genus="Plaumanniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="320" phylum="Arthropoda" rank="species" species="sanctaecatharinae">
<emphasis id="B9013730FF8D5037DBEF406815A3FAF0" bold="true" box="[151,705,1339,1366]" pageId="3" pageNumber="320">
<emphasis id="B9013730FF8D5037DBEF4068154AFAF3" bold="true" box="[151,552,1340,1366]" italics="true" pageId="3" pageNumber="320">Plaumanniola sanctaecatharinae</emphasis>
Costa Lima
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BCAEB22FF8D5037DBEF400A16E3FAD3" blockId="3.[151,705,1339,1398]" box="[151,385,1374,1398]" pageId="3" pageNumber="320">
(
<figureCitation id="134EF7A7FF8D5037DBE7400A1669FAD3" box="[159,267,1374,1398]" captionStart="FIGURES 1 3" captionStartId="4.[151,264,1464,1487]" captionTargetBox="[158,1432,707,1439]" captionTargetId="figure@4.[151,1436,697,1443]" captionTargetPageId="4" captionText="FIGURES 1 3. Plaumanniola sanctaecatharinae Costa Lima, dorsal habitus of paratype male (1) and its original set of labels (2); Plaumanniola regina sp. n., dorsal habitus of paratype male (3)." httpUri="https://zenodo.org/record/223300/files/figure.png" pageId="3" pageNumber="320">Figs. 12</figureCitation>
,
<figureCitation id="134EF7A7FF8D5037DA6F400A1645FAD3" box="[279,295,1374,1398]" captionStart="FIGURES 4 5" captionStartId="5.[151,264,888,911]" captionTargetBox="[151,1422,204,842]" captionTargetId="figure@5.[151,1436,193,867]" captionTargetPageId="5" captionText="FIGURES 4 5. Heads of paratype males in dorsal view. Plaumanniola sanctaecatharinae Costa Lima (4) and Plaumanniola regina sp. n. (5)." httpUri="https://zenodo.org/record/223301/files/figure.png" pageId="3" pageNumber="320">4</figureCitation>
,
<figureCitation id="134EF7A7FF8D5037DA4A400A1618FAD3" box="[306,378,1374,1398]" captionStart="FIGURES 12 15" captionStartId="8.[151,264,771,794]" captionTargetBox="[158,1432,196,740]" captionTargetId="figure@8.[151,1436,193,750]" captionTargetPageId="8" captionText="FIGURES 12 15. Aedeagus of Plaumanniola sanctaecatharinae Costa Lima (12 13) and Plaumanniola regina sp. n. (14 15) in ventral (12, 14) and lateral (13, 15) views." httpUri="https://zenodo.org/record/223304/files/figure.png" pageId="3" pageNumber="320">1213</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36FB8A9FF8D5037DBEF40F31425FA19" box="[151,839,1446,1469]" pageId="3" pageNumber="320" type="reference_group">
<paragraph id="8BCAEB22FF8D5037DBEF40F31425FA19" blockId="3.[151,839,1446,1469]" box="[151,839,1446,1469]" pageId="3" pageNumber="320">
<treatmentCitationGroup id="AB65CC0CFF8D5037DBEF40F31425FA19" box="[151,839,1446,1469]" pageId="3" pageNumber="320">
<taxonomicName id="4C7590A1FF8D5037DBEF40F315BAFA19" ID-CoL="4K24B" authority="Costa Lima, 1962: 416" authorityName="Costa Lima" authorityPageNumber="416" authorityYear="1962" box="[151,728,1446,1469]" class="Insecta" family="Staphylinidae" genus="Plaumanniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="320" phylum="Arthropoda" rank="species" species="sanctaecatharinae">
<emphasis id="B9013730FF8D5037DBEF40F3168AFA18" box="[151,488,1447,1469]" italics="true" pageId="3" pageNumber="320">Plaumanniola sanctaecatharinae</emphasis>
<treatmentCitation id="0AD4CD33FF8D5037DA9540F315BAFA19" author="Costa" box="[493,728,1446,1468]" page="416" pageId="3" pageNumber="320" year="1962">
<bibRefCitation id="EFE496D3FF8D5037DA9540F315C4FA19" author="Costa" box="[493,678,1446,1468]" pageId="3" pageNumber="320" refString="Costa Lima, A. (1962) Micro-Coleoptero representante da nova subfamia Plaumanniolinae (Col., Ptinidae). Revista Brasileira de Biologia, 22, 413 - 418." type="journal article" year="1962">Costa Lima, 1962</bibRefCitation>
: 416
</treatmentCitation>
</taxonomicName>
,
<figureCitation id="134EF7A7FF8D5037D99A40F21421FA19" box="[738,835,1446,1468]" captionStart-0="FIGURES 1 3" captionStart-1="FIGURES 4 5" captionStartId-0="4.[151,264,1464,1487]" captionStartId-1="5.[151,264,888,911]" captionTargetBox-0="[158,1432,707,1439]" captionTargetBox-1="[151,1422,204,842]" captionTargetId-0="figure@4.[151,1436,697,1443]" captionTargetId-1="figure@5.[151,1436,193,867]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURES 1 3. Plaumanniola sanctaecatharinae Costa Lima, dorsal habitus of paratype male (1) and its original set of labels (2); Plaumanniola regina sp. n., dorsal habitus of paratype male (3)." captionText-1="FIGURES 4 5. Heads of paratype males in dorsal view. Plaumanniola sanctaecatharinae Costa Lima (4) and Plaumanniola regina sp. n. (5)." httpUri-0="https://zenodo.org/record/223300/files/figure.png" httpUri-1="https://zenodo.org/record/223301/files/figure.png" pageId="3" pageNumber="320">Figs. 14</figureCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36FB8A9FF8D5037DBEF40BE1383F9C8" pageId="3" pageNumber="320" type="materials_examined">
<paragraph id="8BCAEB22FF8D5037DBEF40BE1383F9C8" blockId="3.[151,1437,1513,2005]" pageId="3" pageNumber="320">
<emphasis id="B9013730FF8D5037DBEF40BE1430F9A7" bold="true" box="[151,850,1513,1538]" pageId="3" pageNumber="320">
<typeStatus id="54CE5580FF8D5037DBEF40BE17B0F9A7" box="[151,210,1514,1538]" pageId="3" pageNumber="320">Type</typeStatus>
material.
<typeStatus id="54CE5580FF8D5037DA3540BE16DCF9A7" box="[333,446,1514,1538]" pageId="3" pageNumber="320" type="paratype">Paratype</typeStatus>
:
<collectingCountry id="F362ABB2FF8D5037DAB340BD1554F9A7" box="[459,566,1513,1538]" name="Brazil" pageId="3" pageNumber="320">BRAZIL</collectingCountry>
(Santa Catarina state):
</emphasis>
3, five labels (
<figureCitation id="134EF7A7FF8D5037DF7B40BD132BF9A4" box="[1027,1097,1513,1537]" captionStart="FIGURES 1 3" captionStartId="4.[151,264,1464,1487]" captionTargetBox="[158,1432,707,1439]" captionTargetId="figure@4.[151,1436,697,1443]" captionTargetPageId="4" captionText="FIGURES 1 3. Plaumanniola sanctaecatharinae Costa Lima, dorsal habitus of paratype male (1) and its original set of labels (2); Plaumanniola regina sp. n., dorsal habitus of paratype male (3)." httpUri="https://zenodo.org/record/223300/files/figure.png" pageId="3" pageNumber="320">Fig. 2</figureCitation>
): &quot;CHAPECO' / 27'07 52'36 /
<quantity id="4C8D46C7FF8D5037DBEF43591782F980" box="[151,224,1549,1573]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="320" unit="m" value="600.0">600 m</quantity>
&quot; [yellowish, printed]; &quot;F. Plaumann /
<date id="FFCBCDE2FF8D5037D9F3435A147CF980" box="[651,798,1549,1574]" pageId="3" pageNumber="320" value="1960-08">August 1960</date>
&quot; [yellowish, printed]; &quot;
<taxonomicName id="4C7590A1FF8D5037DF5B435913A2F980" box="[1059,1216,1549,1573]" class="Insecta" family="Staphylinidae" genus="Plaumanniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="328" phylum="Arthropoda" rank="genus">Plaumanniola</taxonomicName>
/
<taxonomicName id="4C7590A1FF8D5037DFAF435912FFF980" box="[1239,1437,1549,1573]" class="Insecta" family="Staphylinidae" genus="Plaumanniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="320" phylum="Arthropoda" rank="species" species="sanctaecatharinae">sanctaecatharinae</taxonomicName>
/ C.L. 1962 / Costa Lima det.&quot; [yellowish with black frame, handwritten and printed]; &quot;PARATYPO&quot; [white with reddish hue and black frame; reverse side red, printed]; and red rectangle with black line (
<collectionCode id="ED6473E7FF8D5037DFFD430113B6F9C8" box="[1157,1236,1621,1645]" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="3" pageNumber="320">MZSP</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C36FB8A9FF8D5037DBBF432D1393F910" pageId="3" pageNumber="320" type="diagnosis">
<paragraph id="8BCAEB22FF8D5037DBBF432D1393F910" blockId="3.[151,1437,1513,2005]" pageId="3" pageNumber="320">
<emphasis id="B9013730FF8D5037DBBF432D1623F937" bold="true" box="[199,321,1657,1682]" pageId="3" pageNumber="320">Diagnosis.</emphasis>
Body shorter than
<quantity id="4C8D46C7FF8D5037D96F432D1538F934" box="[535,602,1657,1681]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="320" unit="mm" value="2.0">2 mm</quantity>
; antennae with sharply delimited club composed of antennomeres VIIXI; tempora longer than small eyes; all tibiae broadest near distal third; each paramere with three setae.
</paragraph>
</subSubSection>
<subSubSection id="C36FB8A9FF8D5031DBBF4395152CFBFD" lastPageId="5" lastPageNumber="322" pageId="3" pageNumber="320" type="description">
<paragraph id="8BCAEB22FF8D5037DBBF439516F9F958" blockId="3.[151,1437,1513,2005]" pageId="3" pageNumber="320">
<emphasis id="B9013730FF8D5037DBBF43951616F97F" bold="true" box="[199,372,1729,1754]" pageId="3" pageNumber="320">Redescription.</emphasis>
Body of male (
<figureCitation id="134EF7A7FF8D5037D95743951510F97C" box="[559,626,1729,1753]" captionStart="FIGURES 1 3" captionStartId="4.[151,264,1464,1487]" captionTargetBox="[158,1432,707,1439]" captionTargetId="figure@4.[151,1436,697,1443]" captionTargetPageId="4" captionText="FIGURES 1 3. Plaumanniola sanctaecatharinae Costa Lima, dorsal habitus of paratype male (1) and its original set of labels (2); Plaumanniola regina sp. n., dorsal habitus of paratype male (3)." httpUri="https://zenodo.org/record/223300/files/figure.png" pageId="3" pageNumber="320">Fig. 1</figureCitation>
) moderately stout and convex, light brown, covered with light brown vestiture. BL
<quantity id="4C8D46C7FF8D5037DA4843B116F5F958" box="[304,407,1765,1789]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="320" unit="mm" value="1.9">1.90 mm</quantity>
.
</paragraph>
<paragraph id="8BCAEB22FF8D5037DBBF425D12FFF870" blockId="3.[151,1437,1513,2005]" pageId="3" pageNumber="320">
Head (
<figureCitation id="134EF7A7FF8D5037DA6A425D1635F884" box="[274,343,1801,1825]" captionStart="FIGURES 4 5" captionStartId="5.[151,264,888,911]" captionTargetBox="[151,1422,204,842]" captionTargetId="figure@5.[151,1436,193,867]" captionTargetPageId="5" captionText="FIGURES 4 5. Heads of paratype males in dorsal view. Plaumanniola sanctaecatharinae Costa Lima (4) and Plaumanniola regina sp. n. (5)." httpUri="https://zenodo.org/record/223301/files/figure.png" pageId="3" pageNumber="320">Fig. 4</figureCitation>
) in dorsal view subtrapezoid, strongly flattened, broadest at posterior margin of vertex, HL
<quantity id="4C8D46C7FF8D5037DE13425D17A4F8E0" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="3" pageNumber="320" unit="mm" value="0.38">0.38 mm</quantity>
, HW
<quantity id="4C8D46C7FF8D5037DA7542791615F8E0" box="[269,375,1837,1861]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="3" pageNumber="320" unit="mm" value="0.68">0.68 mm</quantity>
; vertex strongly transverse and only weakly, evenly convex; tempora distinctly longer than eyes; frons posteriorly confluent with vertex, between eyes regularly convex, between feebly marked supraantennal tubercles distinctly impressed; compound eyes small, with deeply and broadly emarginate posterior margin. Vertex and frons glossy, covered with sparse and fine setiferous punctures separated by spaces as long as 4
<date id="FFCBCDE2FF8D5037DBCC42E917A2F870" box="[180,192,1981,2005]" pageId="3" pageNumber="320">5</date>
x puncture diameters. Setae on head dorsum sparse but long, curved and suberect, directed posteriorly.
</paragraph>
<paragraph id="8BCAEB22FF8A5030DBEF45CC15A0FD21" blockId="4.[151,1437,151,644]" pageId="4" pageNumber="321">
Antennae (
<figureCitation id="134EF7A7FF8A5030DA6945C31637FF0A" box="[273,341,151,175]" captionStart="FIGURES 4 5" captionStartId="5.[151,264,888,911]" captionTargetBox="[151,1422,204,842]" captionTargetId="figure@5.[151,1436,193,867]" captionTargetPageId="5" captionText="FIGURES 4 5. Heads of paratype males in dorsal view. Plaumanniola sanctaecatharinae Costa Lima (4) and Plaumanniola regina sp. n. (5)." httpUri="https://zenodo.org/record/223301/files/figure.png" pageId="4" pageNumber="321">Fig. 4</figureCitation>
) short, with compact, slightly flattened dorso-ventrally and sharply demarcated club composed of five distal antennomeres, AnL 0.53, antennomere I about twice as long as broad; II as broad as I but much shorter, slightly broader than long; III slightly narrower than II and much shorter, strongly transverse; IV about as long and broad as III; V slightly broader and longer than IV, strongly transverse and distinctly asymmetrical, with proximal margin transverse in relation to long axis of antenna while distal margin strongly oblique and external lateral margin much longer than mesal lateral margin; VI slightly broader and longer than IV, of similar shape except that both proximal and distal margins are strongly oblique in relation to long axis of antenna and mesal lateral margin forming small projection; VII slightly shorter than VI but distinctly broader, strongly transverse,
<date id="FFCBCDE2FF8A5030DF8D44C01273FE09" box="[1269,1297,404,428]" pageId="4" pageNumber="321">4x</date>
as broad as long; VIII as broad as VII but distinctly longer, strongly transverse,
<date id="FFCBCDE2FF8A5030D8DA44E314DCFE6A" box="[930,958,439,463]" pageId="4" pageNumber="321">3x</date>
as broad as long; IX as broad as VIII but slightly longer, strongly transverse, 2.8x as broad as long; X as broad as IX and yet longer, strongly transverse and ca. 2.5x as broad as long; XI subconical with blunt apex, distinctly narrower than X, about 1.8x as long as broad. Antennomeres IVI with sparse long and erect setae distributed mainly along their mesal margin; antennomeres VIIXI moderately densely covered with long and nearly recumbent setae directed toward antennal apex and additionally with sparse strongly erect long setae.
</paragraph>
<caption id="DF0ABBAAFF8A5030DBEF40EC142EFA48" httpUri="https://zenodo.org/record/223300/files/figure.png" pageId="4" pageNumber="321" targetBox="[158,1432,707,1439]" targetPageId="4">
<paragraph id="8BCAEB22FF8A5030DBEF40EC142EFA48" blockId="4.[151,1436,1464,1517]" pageId="4" pageNumber="321">
<emphasis id="B9013730FF8A5030DBEF40EC1658FA68" bold="true" box="[151,314,1464,1487]" pageId="4" pageNumber="321">FIGURES 13.</emphasis>
<taxonomicName id="4C7590A1FF8A5030DA3940EC1472FA6B" authority="Costa Lima" authorityName="Costa Lima" box="[321,784,1464,1486]" class="Insecta" family="Staphylinidae" genus="Plaumanniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="321" phylum="Arthropoda" rank="species" species="sanctaecatharinae">
<emphasis id="B9013730FF8A5030DA3940EC15F0FA6B" box="[321,658,1464,1486]" italics="true" pageId="4" pageNumber="321">Plaumanniola sanctaecatharinae</emphasis>
Costa Lima
</taxonomicName>
, dorsal habitus of paratype male (1) and its original set of labels (2);
<taxonomicName id="4C7590A1FF8A5030DBBA408316FAFA48" box="[194,408,1495,1517]" class="Insecta" family="Staphylinidae" genus="Plaumanniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="321" phylum="Arthropoda" rank="species" species="regina" status="sp. nov.">
<emphasis id="B9013730FF8A5030DBBA408316FAFA48" box="[194,408,1495,1517]" italics="true" pageId="4" pageNumber="321">Plaumanniola regina</emphasis>
</taxonomicName>
<emphasis id="B9013730FF8A5030DAE7408316B5FA49" bold="true" box="[415,471,1495,1516]" pageId="4" pageNumber="321">
<taxonomicNameLabel id="A2328A4BFF8A5030DAE7408316B5FA49" box="[415,471,1495,1516]" pageId="4" pageNumber="321" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, dorsal habitus of paratype male (3).
</paragraph>
</caption>
<paragraph id="8BCAEB22FF8A5030DBBF43411371F945" blockId="4.[151,1436,1556,1976]" pageId="4" pageNumber="321">
Pronotum slightly more convex than head, approximately oval, broadest distinctly anterior to middle; PL
<quantity id="4C8D46C7FF8A5030DE13434017A7F9F5" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="4" pageNumber="321" unit="mm" value="0.53">0.53 mm</quantity>
, PW
<quantity id="4C8D46C7FF8A5030DA7C436C160FF9F5" box="[260,365,1592,1616]" metricMagnitude="-4" metricUnit="m" metricValue="8.299999999999999" pageId="4" pageNumber="321" unit="mm" value="0.83">0.83 mm</quantity>
. Anterior and lateral margins confluent, anterior pronotal corners not marked; anterior margin arcuate and slightly concave just behind head; lateral margins strongly rounded; posterior pronotal corners strongly obtuse and blunt but well-marked; base of pronotum in middle strongly and abruptly projecting posterad, forming short and broad subtrapezoid lobe over base of elytra demarcated laterally by obtuse angles; posterior pronotal margin nearly straight. Punctures and setae similar to those on vertex and frons.
</paragraph>
<paragraph id="8BCAEB22FF8A5030DBBF43B815F8F8D5" blockId="4.[151,1436,1556,1976]" pageId="4" pageNumber="321">
Elytra more convex than pronotum, oval, broadest between middle and anterior third; EL 1.00 mm, EW
<quantity id="4C8D46C7FF8A5030DE1343B817A7F88D" metricMagnitude="-4" metricUnit="m" metricValue="8.299999999999999" pageId="4" pageNumber="321" unit="mm" value="0.83">0.83 mm</quantity>
, EI 1.21. Punctures more distinct than those on head and pronotum, shallow and small, with moderately distinct margins, separated by spaces 2
<date id="FFCBCDE2FF8A5030D91642601518F8E9" box="[622,634,1844,1868]" pageId="4" pageNumber="321">3</date>
x as wide as puncture diameters; setae similar to those on head and pronotum but distinctly longer and more erect
</paragraph>
<paragraph id="8BCAEB22FF8A5030DBBF4229169BF81D" blockId="4.[151,1436,1556,1976]" pageId="4" pageNumber="321">Legs short and robust; protibiae slightly curved inwards in distal third; meso- and metatibiae nearly straight; all tibiae broadest near distal third.</paragraph>
<caption id="DF0ABBAAFF8B5031DBEF462C1620FC08" httpUri="https://zenodo.org/record/223301/files/figure.png" pageId="5" pageNumber="322" targetBox="[151,1422,204,842]" targetPageId="5">
<paragraph id="8BCAEB22FF8B5031DBEF462C1620FC08" blockId="5.[151,1435,888,941]" pageId="5" pageNumber="322">
<emphasis id="B9013730FF8B5031DBEF462C165EFC2B" bold="true" box="[151,316,888,911]" pageId="5" pageNumber="322">FIGURES 45.</emphasis>
Heads of paratype males in dorsal view.
<taxonomicName id="4C7590A1FF8B5031D99C462C13D1FC2B" authority="Costa Lima" authorityName="Costa Lima" box="[740,1203,888,910]" class="Insecta" family="Staphylinidae" genus="Plaumanniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="322" phylum="Arthropoda" rank="species" species="sanctaecatharinae">
<emphasis id="B9013730FF8B5031D99C462C1354FC2B" box="[740,1078,888,910]" italics="true" pageId="5" pageNumber="322">Plaumanniola sanctaecatharinae</emphasis>
Costa Lima
</taxonomicName>
(4) and
<taxonomicName id="4C7590A1FF8B5031DE74462C17BAFC08" class="Insecta" family="Staphylinidae" genus="Plaumanniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="322" phylum="Arthropoda" rank="species" species="regina" status="sp. nov.">
<emphasis id="B9013730FF8B5031DE74462C17BAFC08" italics="true" pageId="5" pageNumber="322">Plaumanniola regina</emphasis>
</taxonomicName>
<emphasis id="B9013730FF8B5031DBA646C31675FC09" bold="true" box="[222,279,919,940]" pageId="5" pageNumber="322">
<taxonomicNameLabel id="A2328A4BFF8B5031DBA646C31675FC09" box="[222,279,919,940]" pageId="5" pageNumber="322" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(5).
</paragraph>
</caption>
<paragraph id="8BCAEB22FF8B5031DBBF468017B4FB91" blockId="5.[151,1436,980,1508]" pageId="5" pageNumber="322">
Aedeagus (
<figureCitation id="134EF7A7FF8B5031DA31468016B6FC49" box="[329,468,980,1004]" captionStart="FIGURES 12 15" captionStartId="8.[151,264,771,794]" captionTargetBox="[158,1432,196,740]" captionTargetId="figure@8.[151,1436,193,750]" captionTargetPageId="8" captionText="FIGURES 12 15. Aedeagus of Plaumanniola sanctaecatharinae Costa Lima (12 13) and Plaumanniola regina sp. n. (14 15) in ventral (12, 14) and lateral (13, 15) views." httpUri="https://zenodo.org/record/223304/files/figure.png" pageId="5" pageNumber="322">Figs. 1213</figureCitation>
) pear-shaped; AeL
<quantity id="4C8D46C7FF8B5031D9C64680144BFC49" box="[702,809,980,1004]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="5" pageNumber="322" unit="mm" value="0.38">0.38 mm</quantity>
; median lobe in ventral view with long subtriangular apical part; internal armature complex and partly asymmetrical; parameres in lateral view curved, each with three setae.
</paragraph>
<paragraph id="8BCAEB22FF8B5031DBBF4114152CFBFD" blockId="5.[151,1436,980,1508]" box="[199,590,1088,1112]" pageId="5" pageNumber="322">Female. Not studied; see Remarks.</paragraph>
</subSubSection>
<subSubSection id="C36FB8A9FF8B5031DBBF4130146EFBD9" box="[199,780,1124,1149]" pageId="5" pageNumber="322" type="distribution">
<paragraph id="8BCAEB22FF8B5031DBBF4130146EFBD9" blockId="5.[151,1436,980,1508]" box="[199,780,1124,1149]" pageId="5" pageNumber="322">
<emphasis id="B9013730FF8B5031DBBF41301602FBD8" bold="true" box="[199,352,1124,1149]" pageId="5" pageNumber="322">Distribution.</emphasis>
Southern
<collectingCountry id="F362ABB2FF8B5031DAAB41301578FBD9" box="[467,538,1124,1148]" name="Brazil" pageId="5" pageNumber="322">Brazil</collectingCountry>
, state Santa Catarina.
</paragraph>
</subSubSection>
<subSubSection id="C36FB8A9FF8B5031DBBF41DD16D4FA41" pageId="5" pageNumber="322" type="discussion">
<paragraph id="8BCAEB22FF8B5031DBBF41DD16D4FA41" blockId="5.[151,1436,980,1508]" pageId="5" pageNumber="322">
<emphasis id="B9013730FF8B5031DBBF41DD1659FB04" bold="true" box="[199,315,1161,1185]" pageId="5" pageNumber="322">Remarks.</emphasis>
According to
<bibRefCitation id="EFE496D3FF8B5031DAA241DC15CDFB05" box="[474,687,1160,1184]" pageId="5" pageNumber="322" refString="Costa Lima, A. (1962) Micro-Coleoptero representante da nova subfamia Plaumanniolinae (Col., Ptinidae). Revista Brasileira de Biologia, 22, 413 - 418." type="journal article">Costa Lima (1962)</bibRefCitation>
, the
<typeStatus id="54CE5580FF8B5031D99B41DD1471FB04" box="[739,787,1161,1185]" pageId="5" pageNumber="322">type</typeStatus>
series of
<taxonomicName id="4C7590A1FF8B5031D80641DC1390FB05" box="[894,1266,1160,1184]" class="Insecta" family="Staphylinidae" genus="Plaumanniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="322" phylum="Arthropoda" rank="species" species="sanctaecatharinae">
<emphasis id="B9013730FF8B5031D80641DC1390FB05" box="[894,1266,1160,1184]" italics="true" pageId="5" pageNumber="322">Plaumanniola sanctaecatharinae</emphasis>
</taxonomicName>
was composed of a female
<typeStatus id="54CE5580FF8B5031DA6741F816E3FB61" box="[287,385,1196,1220]" pageId="5" pageNumber="322" type="holotype">holotype</typeStatus>
and 11
<typeStatus id="54CE5580FF8B5031DAA441F91529FB60" box="[476,587,1197,1221]" pageId="5" pageNumber="322" type="paratype">paratypes</typeStatus>
. The
<typeStatus id="54CE5580FF8B5031D9F241F8158EFB61" box="[650,748,1196,1220]" pageId="5" pageNumber="322" type="holotype">holotype</typeStatus>
and five
<typeStatus id="54CE5580FF8B5031D82041F914A6FB60" box="[856,964,1197,1221]" pageId="5" pageNumber="322" type="paratype">paratypes</typeStatus>
were originally in the collection of Costa Lima and six
<typeStatus id="54CE5580FF8B5031DA48418516F9FB4C" box="[304,411,1233,1257]" pageId="5" pageNumber="322" type="paratype">paratypes</typeStatus>
in the collection of Plaumann (
<bibRefCitation id="EFE496D3FF8B5031D981418414DDFB4D" author="Costa" box="[761,959,1232,1256]" pageId="5" pageNumber="322" refString="Costa Lima, A. (1962) Micro-Coleoptero representante da nova subfamia Plaumanniolinae (Col., Ptinidae). Revista Brasileira de Biologia, 22, 413 - 418." type="journal article" year="1962">Costa Lima 1962</bibRefCitation>
).
<bibRefCitation id="EFE496D3FF8B5031D8AA41851241FB4D" author="Lawrence" box="[978,1315,1232,1256]" pageId="5" pageNumber="322" refString="Lawrence, J. F. &amp; Reichardt, H. (1966) The systematic position of Plaumanniola Costa Lima (Coleoptera: Scydmaenidae). The Coleopterists' Bulletin, 20, 39 - 42." type="journal article" year="1966">Lawrence &amp; Reichardt (1966)</bibRefCitation>
stated that a single
<typeStatus id="54CE5580FF8B5031DB8C41A11636FAA8" box="[244,340,1269,1293]" pageId="5" pageNumber="322" type="paratype">paratype</typeStatus>
specimen was bought by Reichardt from Plaumann and deposited &quot;in the collection at São Paulo&quot; (i.e., at Departamento de Zoologia). Currently three
<typeStatus id="54CE5580FF8B5031D98B404D143CFA94" box="[755,862,1305,1329]" pageId="5" pageNumber="322" type="paratype">paratypes</typeStatus>
are housed at MZSP (email from S. Casari dated
<date id="FFCBCDE2FF8B5031DBEF4068167BFAF1" box="[151,281,1340,1364]" pageId="5" pageNumber="322" value="2013-03-15">15.03.2013</date>
), and I was not able to locate neither the
<typeStatus id="54CE5580FF8B5031D87A40681406FAF1" box="[770,868,1340,1364]" pageId="5" pageNumber="322" type="holotype">holotype</typeStatus>
nor any of the remaining
<typeStatus id="54CE5580FF8B5031DFE14069126AFAF0" box="[1177,1288,1341,1365]" pageId="5" pageNumber="322" type="paratype">paratypes</typeStatus>
. Illustrations given by
<bibRefCitation id="EFE496D3FF8B5031DA79403416B4FADD" box="[257,470,1376,1400]" pageId="5" pageNumber="322" refString="Costa Lima, A. (1962) Micro-Coleoptero representante da nova subfamia Plaumanniolinae (Col., Ptinidae). Revista Brasileira de Biologia, 22, 413 - 418." type="journal article">Costa Lima (1962)</bibRefCitation>
show that the general body shape and the structure of antennae of males and females do not differ, and structures of the
<typeStatus id="54CE5580FF8B5031D96740D1151DFA38" box="[543,639,1413,1437]" pageId="5" pageNumber="322" type="paratype">paratype</typeStatus>
male studied during the present work match well images presented by Costa Lima. Later &quot;redescription&quot; by
<bibRefCitation id="EFE496D3FF8B5031D93E40FD15BBFA65" author="Franz" box="[582,729,1448,1472]" pageId="5" pageNumber="322" refString="Franz, H. (1990) Neue Beitrage zur Kenntnis der sud- und mittelamerikanischen Scydmaenidae (Coleoptera). Zeitschrift der Arbeitsgemeinshaft Osterreichischer Entomologen, 42, 32 - 38." type="journal article" year="1990">Franz (1990)</bibRefCitation>
was based on misidentified specimens (belonging to the new species described below).
</paragraph>
</subSubSection>
</treatment>
</document>