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<document id="7920C053DD60503184170E8153A07052" ID-DOI="10.11646/zootaxa.3688.1.1" ID-GBIF-Dataset="151481a0-71e1-4402-ab7e-8262caddf539" ID-ISSN="1175-5326" ID-Zenodo-Dep="284201" ID-ZooBank="DB84C679-C2AA-41CC-816E-83E2F6DCA391" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460649331738" checkinUser="plazi" docAuthor="Haddad, Charles R." docDate="2013" docId="03DB8445FFA4FFF2FF2AFCE0302BFACB" docLanguage="en" docName="zt03688p079.pdf" docOrigin="Zootaxa 3688 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Wasaka imitatrix Haddad, 2013, sp. nov." docType="treatment" docVersion="7" lastPageNumber="65" masterDocId="FFE2FC3DFF9AFFB2FFBDFFE6345DFFF0" masterDocTitle="Taxonomic notes on the spider genus Messapus Simon, 1898 (Araneae, Corinnidae), with the description of the new genera Copuetta and Wasaka and the first cladistic analysis of Afrotropical Castianeirinae" masterLastPageNumber="79" masterPageNumber="1" pageNumber="63" updateTime="1698337697679" updateUser="plazi">
<mods:mods id="4D8EB9B3F539315F70B1DCDEA871C82D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D94FBC0402B4F6DA59B62385718728D7">
<mods:title id="D2EBE6661D5B7F4681010B5AC3B81529">Taxonomic notes on the spider genus Messapus Simon, 1898 (Araneae, Corinnidae), with the description of the new genera Copuetta and Wasaka and the first cladistic analysis of Afrotropical Castianeirinae</mods:title>
</mods:titleInfo>
<mods:name id="2BF3C1EE9BEBC048C0D1DB86CA19A7A2" type="personal">
<mods:role id="40B2289490A8CA2AF9E96A681FB23998">
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<mods:namePart id="0E59B7EEF1B6116FD9F77ED50A23270B">Haddad, Charles R.</mods:namePart>
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<mods:title id="17A56D13EF3056E0C8F382DF8155E675">Zootaxa</mods:title>
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<mods:part id="87656D5DC014ECA3877F40074D2F5B09">
<mods:date id="39FCC0A6A18AE4ED3AE921DAA03B46CF">2013</mods:date>
<mods:detail id="596A4659A60A6FCD9151945B7475A535" type="volume">
<mods:number id="BB06C65DFBB8B01D50BE740EA8E607F6">3688</mods:number>
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<mods:classification id="4E75BDB68840D7814CB8EA4701582E61">journal article</mods:classification>
<mods:identifier id="F476D28105380224CFB16DBCB3871356" type="DOI">10.11646/zootaxa.3688.1.1</mods:identifier>
<mods:identifier id="3EF9CB33A12FEB6045F1CA320EA5526E" type="GBIF-Dataset">151481a0-71e1-4402-ab7e-8262caddf539</mods:identifier>
<mods:identifier id="58795719C97AEA735DED16F16C93E79A" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="C19FB578A375CF200905B9603C2F4728" type="Zenodo-Dep">284201</mods:identifier>
<mods:identifier id="B28B67671B8AFED410D72BFFEFED9C0C" type="ZooBank">DB84C679-C2AA-41CC-816E-83E2F6DCA391</mods:identifier>
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<treatment id="03DB8445FFA4FFF2FF2AFCE0302BFACB" ID-DOI="http://doi.org/10.5281/zenodo.6153967" ID-GBIF-Taxon="127664904" ID-Zenodo-Dep="6153967" LSID="urn:lsid:plazi:treatment:03DB8445FFA4FFF2FF2AFCE0302BFACB" httpUri="http://treatment.plazi.org/id/03DB8445FFA4FFF2FF2AFCE0302BFACB" lastPageId="64" lastPageNumber="65" pageId="62" pageNumber="63">
<subSubSection id="C36866D8FFA4FF8CFF2AFCE03585FCB1" pageId="62" pageNumber="63" type="nomenclature">
<paragraph id="8BCD3553FFA4FF8CFF2AFCE03595FCD0" blockId="62.[151,472,774,833]" box="[151,456,774,800]" pageId="62" pageNumber="63">
<heading id="D085823FFFA4FF8CFF2AFCE03595FCD0" bold="true" box="[151,456,774,800]" fontSize="11" level="1" pageId="62" pageNumber="63" reason="1">
<emphasis id="B906E941FFA4FF8CFF2AFCE03595FCD0" bold="true" box="[151,456,774,800]" pageId="62" pageNumber="63">
<taxonomicName id="4C724ED0FFA4FF8CFF2AFCE03539FCD0" ID-CoL="5BV2T" box="[151,356,774,800]" class="Arachnida" family="Corinnidae" genus="Wasaka" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="imitatrix" status="sp. nov.">
<emphasis id="B906E941FFA4FF8CFF2AFCE03539FCD0" bold="true" box="[151,356,774,800]" italics="true" pageId="62" pageNumber="63">Wasaka imitatrix</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A235543AFFA4FF8CFED6FCE03595FCD0" box="[363,456,774,800]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCD3553FFA4FF8CFF2AFCCF3585FCB1" blockId="62.[151,472,774,833]" box="[151,472,809,833]" pageId="62" pageNumber="63">
<figureCitation id="134929D6FFA4FF8CFF2AFCCF356FFCB1" box="[151,306,809,833]" captionStart="FIGURES 154 – 161" captionStartId="61.[151,264,1108,1131]" captionTargetBox="[153,1436,259,1087]" captionTargetId="figure@61.[153,1438,259,1087]" captionTargetPageId="61" captionText="FIGURES 154 – 161. Digital microscope photographs of Wasaka imitatrix sp. nov. (154, 155), W. montana sp. nov. (156, 157), W. occulta sp. nov. (158, 159) and W. ventralis sp. nov. (160, 161): 154, 156, 158, 160. Male habitus, dorsal view. 155, 157, 159, 161. Female habitus, dorsal view. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/284230/files/figure.png" pageId="62" pageNumber="63">Figs 154, 155</figureCitation>
,
<figureCitation id="134929D6FFA4FF8CFE83FCCF3537FCB1" box="[318,362,809,833]" captionStart="FIGURES 162 – 165" captionStartId="62.[151,264,641,664]" captionTargetBox="[151,1436,208,617]" captionTargetId="figure@62.[151,1436,208,617]" captionTargetPageId="62" captionText="FIGURES 162 – 165. Digital microscope photographs of emboli of Wasaka gen. nov. species in ventral view: 162. W. imitatrix sp. nov.. 163. W. montana sp. nov.. 164. W. occulta sp. nov.. 165. W. ventralis sp. nov.. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/284231/files/figure.png" pageId="62" pageNumber="63">162</figureCitation>
,
<figureCitation id="134929D6FFA4FF8CFECBFCCF3585FCB1" box="[374,472,809,833]" captionStart="FIGURES 166 – 169" captionStartId="63.[151,264,1723,1746]" captionTargetBox="[164,1422,810,1687]" captionTargetId="figure@63.[151,1436,803,1702]" captionTargetPageId="63" captionText="FIGURES 166 – 169. Genitalic morphology of Wasaka imitatrix sp. nov.: 166. Male palp, ventral view. 167. Same, retrolateral view. 168. Female epigyne, ventral view. 169. Same, dorsal view. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/284232/files/figure.png" pageId="62" pageNumber="63">166–169</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36866D8FFA4FF8CFF2AFC9435A6FC5D" pageId="62" pageNumber="63" type="etymology">
<paragraph id="8BCD3553FFA4FF8CFF2AFC9435A6FC5D" blockId="62.[151,1437,881,2033]" pageId="62" pageNumber="63">
<emphasis id="B906E941FFA4FF8CFF2AFC94357CFC7A" bold="true" box="[151,289,882,906]" pageId="62" pageNumber="63">Etymology:</emphasis>
The species name is Latin for “resembling”, referring to the similarity in genitalic morphology of this species and
<taxonomicName id="4C724ED0FFA4FF8CFEA3FC7035CEFC5D" box="[286,403,917,941]" class="Arachnida" family="Corinnidae" genus="Wasaka" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="occulta" status="sp. nov.">
<emphasis id="B906E941FFA4FF8CFEA3FC7035CEFC5D" box="[286,403,917,941]" italics="true" pageId="62" pageNumber="63">W. occulta</emphasis>
</taxonomicName>
<emphasis id="B906E941FFA4FF8CFE27FC7335A9FC5D" bold="true" box="[410,500,917,941]" pageId="62" pageNumber="63">
<taxonomicNameLabel id="A235543AFFA4FF8CFE27FC7335A9FC5D" box="[410,500,917,941]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C36866D8FFA4FF8CFF7AFC5F3798FB01" pageId="62" pageNumber="63" type="diagnosis">
<paragraph id="8BCD3553FFA4FF8CFF7AFC5F3798FB01" blockId="62.[151,1437,881,2033]" pageId="62" pageNumber="63">
<emphasis id="B906E941FFA4FF8CFF7AFC5F351EFC22" bold="true" box="[199,323,953,978]" pageId="62" pageNumber="63">Diagnosis:</emphasis>
This species is most closely related to
<taxonomicName id="4C724ED0FFA4FF8CFD43FC5C3729FC21" box="[766,884,953,977]" class="Arachnida" family="Corinnidae" genus="Wasaka" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="occulta" status="sp. nov.">
<emphasis id="B906E941FFA4FF8CFD43FC5C3729FC21" box="[766,884,953,977]" italics="true" pageId="62" pageNumber="63">W. occulta</emphasis>
</taxonomicName>
<emphasis id="B906E941FFA4FF8CFCC6FC5C3788FC22" bold="true" box="[891,981,954,978]" pageId="62" pageNumber="63">
<taxonomicNameLabel id="A235543AFFA4FF8CFCC6FC5C3788FC22" box="[891,981,954,978]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, from which the males can be separated by the more longitudinally compressed coils and more transversely orientated embolus tip (compare
<figureCitation id="134929D6FFA4FF8CFAB7FC3B3498FBE9" captionStart="FIGURES 162 – 165" captionStartId="62.[151,264,641,664]" captionTargetBox="[151,1436,208,617]" captionTargetId="figure@62.[151,1436,208,617]" captionTargetPageId="62" captionText="FIGURES 162 – 165. Digital microscope photographs of emboli of Wasaka gen. nov. species in ventral view: 162. W. imitatrix sp. nov.. 163. W. montana sp. nov.. 164. W. occulta sp. nov.. 165. W. ventralis sp. nov.. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/284231/files/figure.png" pageId="62" pageNumber="63">Figs 162 and 164</figureCitation>
). Females can be separated from
<taxonomicName id="4C724ED0FFA4FF8CFDF1FBE43698FBE9" box="[588,709,1025,1049]" class="Arachnida" family="Corinnidae" genus="Wasaka" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="occulta" status="sp. nov.">
<emphasis id="B906E941FFA4FF8CFDF1FBE43698FBE9" box="[588,709,1025,1049]" italics="true" pageId="62" pageNumber="63">W. occulta</emphasis>
</taxonomicName>
<emphasis id="B906E941FFA4FF8CFD72FBE43776FBEA" bold="true" box="[719,811,1026,1050]" pageId="62" pageNumber="63">
<taxonomicNameLabel id="A235543AFFA4FF8CFD72FBE43776FBEA" box="[719,811,1026,1050]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the longer coiled epigynal ridges that nearly meet medially (clearly separated in
<taxonomicName id="4C724ED0FFA4FF8CFE47FBC03629FBCD" box="[506,628,1061,1085]" class="Arachnida" family="Corinnidae" genus="Wasaka" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="occulta" status="sp. nov.">
<emphasis id="B906E941FFA4FF8CFE47FBC03629FBCD" box="[506,628,1061,1085]" italics="true" pageId="62" pageNumber="63">W. occulta</emphasis>
</taxonomicName>
<emphasis id="B906E941FFA4FF8CFDC3FBC33681FBCD" bold="true" box="[638,732,1061,1085]" pageId="62" pageNumber="63">
<taxonomicNameLabel id="A235543AFFA4FF8CFDC3FBC33681FBCD" box="[638,732,1061,1085]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) and the less strongly curved copulatory openings (strongly curved in
<taxonomicName id="4C724ED0FFA4FF8CFEADFBAC35D7FB91" box="[272,394,1097,1121]" class="Arachnida" family="Corinnidae" genus="Wasaka" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="occulta" status="sp. nov.">
<emphasis id="B906E941FFA4FF8CFEADFBAC35D7FB91" box="[272,394,1097,1121]" italics="true" pageId="62" pageNumber="63">W. occulta</emphasis>
</taxonomicName>
<emphasis id="B906E941FFA4FF8CFE29FBAC35AFFB92" bold="true" box="[404,498,1098,1122]" pageId="62" pageNumber="63">
<taxonomicNameLabel id="A235543AFFA4FF8CFE29FBAC35AFFB92" box="[404,498,1098,1122]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) (compare
<figureCitation id="134929D6FFA4FF8CFDC1FBAF36BFFB91" box="[636,738,1097,1121]" captionStart="FIGURES 166 – 169" captionStartId="63.[151,264,1723,1746]" captionTargetBox="[164,1422,810,1687]" captionTargetId="figure@63.[151,1436,803,1702]" captionTargetPageId="63" captionText="FIGURES 166 – 169. Genitalic morphology of Wasaka imitatrix sp. nov.: 166. Male palp, ventral view. 167. Same, retrolateral view. 168. Female epigyne, ventral view. 169. Same, dorsal view. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/284232/files/figure.png" pageId="62" pageNumber="63">Figs 168</figureCitation>
and
<figureCitation id="134929D6FFA4FF8CFC9CFBAF3712FB91" box="[801,847,1097,1121]" captionStart="FIGURES 174 – 177" captionStartId="68.[151,264,1194,1217]" captionTargetBox="[166,1426,198,1161]" captionTargetId="figure@68.[150,1436,193,1173]" captionTargetPageId="68" captionText="FIGURES 174 – 177. Genitalic morphology of Wasaka occulta sp. nov.: 174. Male palp, ventral view. 175. Same, retrolateral view. 176. Female epigyne, ventral view. 177. Same, dorsal view. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/284234/files/figure.png" pageId="62" pageNumber="63">176</figureCitation>
). In terms of internal structure of the epigyne,
<taxonomicName id="4C724ED0FFA4FF8CFA3DFBAC34AAFB75" class="Arachnida" family="Corinnidae" genus="Wasaka" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="imitatrix" status="sp. nov.">
<emphasis id="B906E941FFA4FF8CFA3DFBAC34AAFB75" italics="true" pageId="62" pageNumber="63">W. imitatrix</emphasis>
</taxonomicName>
<emphasis id="B906E941FFA4FF8CFEBDFB8B3506FB75" bold="true" box="[256,347,1133,1157]" pageId="62" pageNumber="63">
<taxonomicNameLabel id="A235543AFFA4FF8CFEBDFB8B3506FB75" box="[256,347,1133,1157]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
females have a distinct separation between ST II and ST I, ST II that are as long as broad, and distinct entrance ducts that are clearly separated from the ST I and II (
<figureCitation id="134929D6FFA4FF8CFC16FB773050FB59" box="[939,1037,1169,1193]" captionStart="FIGURES 166 – 169" captionStartId="63.[151,264,1723,1746]" captionTargetBox="[164,1422,810,1687]" captionTargetId="figure@63.[151,1436,803,1702]" captionTargetPageId="63" captionText="FIGURES 166 – 169. Genitalic morphology of Wasaka imitatrix sp. nov.: 166. Male palp, ventral view. 167. Same, retrolateral view. 168. Female epigyne, ventral view. 169. Same, dorsal view. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/284232/files/figure.png" pageId="62" pageNumber="63">Fig. 169</figureCitation>
), while
<taxonomicName id="4C724ED0FFA4FF8CFBDAFB743087FB59" box="[1127,1242,1169,1193]" class="Arachnida" family="Corinnidae" genus="Wasaka" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="occulta" status="sp. nov.">
<emphasis id="B906E941FFA4FF8CFBDAFB743087FB59" box="[1127,1242,1169,1193]" italics="true" pageId="62" pageNumber="63">W. occulta</emphasis>
</taxonomicName>
<emphasis id="B906E941FFA4FF8CFB59FB743163FB5A" bold="true" box="[1252,1342,1170,1194]" pageId="62" pageNumber="63">
<taxonomicNameLabel id="A235543AFFA4FF8CFB59FB743163FB5A" box="[1252,1342,1170,1194]" pageId="62" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
females have a broad fusion between ST I and II, ST II that are nearly twice as long as broad, and very short entrance ducts that are continuous with ST II and partly hidden beneath ST I (
<figureCitation id="134929D6FFA4FF8CFCEBFB3F37E5FB01" box="[854,952,1241,1265]" captionStart="FIGURES 174 – 177" captionStartId="68.[151,264,1194,1217]" captionTargetBox="[166,1426,198,1161]" captionTargetId="figure@68.[150,1436,193,1173]" captionTargetPageId="68" captionText="FIGURES 174 – 177. Genitalic morphology of Wasaka occulta sp. nov.: 174. Male palp, ventral view. 175. Same, retrolateral view. 176. Female epigyne, ventral view. 177. Same, dorsal view. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/284234/files/figure.png" pageId="62" pageNumber="63">Fig. 177</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36866D8FFA4FF8CFF7AFB1B308DFAC9" pageId="62" pageNumber="63" type="discussion">
<paragraph id="8BCD3553FFA4FF8CFF7AFB1B308DFAC9" blockId="62.[151,1437,881,2033]" pageId="62" pageNumber="63">
<emphasis id="B906E941FFA4FF8CFF7AFB1B356FFAE5" bold="true" box="[199,306,1277,1301]" pageId="62" pageNumber="63">Remark:</emphasis>
The
<typeStatus id="54C98BF1FFA4FF8CFECEFB18358EFAE6" box="[371,467,1278,1302]" pageId="62" pageNumber="63" type="paratype">paratype</typeStatus>
female is darker than the
<typeStatus id="54C98BF1FFA4FF8CFCB0FB1B3732FAE5" box="[781,879,1277,1301]" pageId="62" pageNumber="63" type="holotype">holotype</typeStatus>
and has more distinct mottling on the legs and markings on the chelicerae, endites, labium and sternum, similar to those described for the male.
</paragraph>
</subSubSection>
<subSubSection id="C36866D8FFA4FFF2FF7AFAA335D3FC07" lastPageId="64" lastPageNumber="65" pageId="62" pageNumber="63" type="description">
<paragraph id="8BCD3553FFA4FF8CFF7AFAA3378AFA55" blockId="62.[151,1437,881,2033]" pageId="62" pageNumber="63">
<emphasis id="B906E941FFA4FF8CFF7AFAA3376AFAAD" bold="true" box="[199,823,1349,1374]" pageId="62" pageNumber="63">
Male (
<typeStatus id="54C98BF1FFA4FF8CFEA9FAA335DFFAAE" box="[276,386,1349,1374]" pageId="62" pageNumber="63" type="paratype">paratype</typeStatus>
,
<taxonomicName id="4C724ED0FFA4FF8CFE2FFAA33659FAAD" box="[402,516,1349,1373]" class="Arachnida" family="Corinnidae" genus="Copuetta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="62" pageNumber="73" phylum="Arthropoda" rank="species" species="uzungwa">Uzungwa</taxonomicName>
, USNM). Measurements:
</emphasis>
CL 4.38, CW 3.34, AL 4.72, AW 2.95, TL 8.90, FL 0.51, SL 1.86, SW 1.78, AME–AME 0.10, AME–ALE 0.02, ALE–ALE 0.63, PME–PME 0.22, PME–PLE 0.16, PLE–PLE 0.98,
<collectingCountry id="F36575C3FFA4FF8CFEF3FA6835C7FA55" box="[334,410,1422,1445]" name="Peru" pageId="62" pageNumber="63">PERW</collectingCountry>
1.29, MOQAW 0.60, MOQPW 0.69, MOQL 0.73.
</paragraph>
<paragraph id="8BCD3553FFA4FF8CFF7AFA573775FA1D" blockId="62.[151,1437,881,2033]" pageId="62" pageNumber="63">Length of leg segments: I 4.43 + 1.70 + 4.20 + 4.10 + 2.20 = 16.63; II missing; III 4.10 + 1.52 + 3.40 + 3.70 + 1.80 = 14.52; IV 4.85 + 1.60 + 4.15 + 5.20 + 2.08 = 17.88.</paragraph>
<paragraph id="8BCD3553FFA4FF8DFF7AFA1F3016FD05" blockId="62.[151,1437,881,2033]" lastBlockId="63.[151,1436,151,757]" lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63">
General appearance as in
<figureCitation id="134929D6FFA4FF8CFE43FA1F3639F9E1" box="[510,612,1529,1553]" captionStart="FIGURES 154 – 161" captionStartId="61.[151,264,1108,1131]" captionTargetBox="[153,1436,259,1087]" captionTargetId="figure@61.[153,1438,259,1087]" captionTargetPageId="61" captionText="FIGURES 154 – 161. Digital microscope photographs of Wasaka imitatrix sp. nov. (154, 155), W. montana sp. nov. (156, 157), W. occulta sp. nov. (158, 159) and W. ventralis sp. nov. (160, 161): 154, 156, 158, 160. Male habitus, dorsal view. 155, 157, 159, 161. Female habitus, dorsal view. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/284230/files/figure.png" pageId="62" pageNumber="63">Fig. 154</figureCitation>
. Carapace bright yellow-orange, eye region grey; paired mediolateral markings comprising large black blotches corresponding to each coxal pair, fused between coxae III and IV; narrow black mottled marking along midline from PME to fovea; orange-brown striae with faint black mottling radiating from fovea, directed between coxae; lateral margins of carapace with narrow mottled black fringe from cephalic region to posterior margin; cephalic region with small black spot beneath each ALE and lateral of each PLE; markings covered with black feathery setae, areas between them in white feathery and short straight setae. AME separated by distance equal to 2⁄5 their diameter; AME separated from ALE by distance less than 1⁄5 AME diameter; clypeus height slightly less than 1 1⁄5 AME diameter; PME separated by distance equal to ½ their diameter; PME separated from PLE by distance equal to ¾ PME diameter; CW:
<collectingCountry id="F36575C3FFA4FF8CFBAAF8C2303AF8CC" box="[1047,1127,1828,1852]" name="Peru" pageId="62" pageNumber="63">PERW</collectingCountry>
= 2.59:1. Chelicerae bright orange-brown with black mottling; promargin with three adjacent teeth, distal tooth smallest, median tooth largest; retromargin with two teeth separated by basal width of proximal tooth, distal tooth larger, close to fang base. Endites yellow-brown with black mottling, cream prolaterally distally; labium red-brown proximally with black mottling, fading to orange and cream distally; sternum yellow with black mottling, orange-brown around margins, with large black spots at each coxa. Legs (both leg II missing) with cream femora, remaining segments yellowbrown, with black mottling and markings and black spots at spine bases; markings and mottling densely covered in black feathery setae, with yellowish feathery setae between them; femora I, III and IV with black distal band dorsally; patellae I, III and IV with dorsal line proximally, dark mottling laterally and ventrally; tibiae and metatarsi I and III with broken bands proximally, medially and distally; tibiae and metatarsi IV with proximal and median bands fused, distal end with narrow band; tarsi I, III and IV yellow with black mottling. Leg spination (both leg II missing): femora: I pl 3 do 3 rl 3, III pl 3 do 3 rl 3, IV pl 2-3 do 3 rl 2; all femora with scattered erect ventral setae; patellae: I, III and IV with fine proximal and distal do setae; tibiae: I pl 2 rl 2 plv 3 rlv 3, III pl 2 do 1 rl 2 plv 2 rlv 2 vt 1, IV pl 2-3 do 1 rl 2 plv 2 rlv 2 vt 2; metatarsi: I pl 2 rl 2 plv 2 rlv 2, III pl 3 rl 3 plv 2 rlv 2 vt 3, IV pl 3 rl 3 plv 2 rlv 2 vt 3. Palpal spination: femora: pl 1 do 2, with rlv 5 erect setae; patellae: pl 1 do 2; tibiae: pl 1 do 1 plv 1; tarsi: pl 1 plv 2. Abdomen with small anterior dorsal scutum extending only to 1⁄10 abdomen length, dorsum mottled grey behind scutum, with several narrow cream chevrons along midline; markings densely covered in black and yellow-brown feathery setae, with scattered brown short straight setae between them; spinnerets surrounded by narrow black ring; venter mottled grey with cream spots, covered in yellow-brown feathery setae and brown short straight setae; epigastric scutum, ventral sclerite and inframamillary sclerite yellow. Male palp bright yellow with dorsal and lateral mottled black spots, cymbium dark yellow-brown; tegulum orange, with black ducts; embolus with broad, nearly transverse oblique base, proximal coil strongly compressed and narrowly separated from base, embolus tip triangular, nearly transverse, directed retrolaterally (
<figureCitation id="134929D6FFA5FF8DFCD5FD3B3790FD05" box="[872,973,733,757]" captionStart="FIGURES 162 – 165" captionStartId="62.[151,264,641,664]" captionTargetBox="[151,1436,208,617]" captionTargetId="figure@62.[151,1436,208,617]" captionTargetPageId="62" captionText="FIGURES 162 – 165. Digital microscope photographs of emboli of Wasaka gen. nov. species in ventral view: 162. W. imitatrix sp. nov.. 163. W. montana sp. nov.. 164. W. occulta sp. nov.. 165. W. ventralis sp. nov.. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/284231/files/figure.png" pageId="63" pageNumber="64">Figs 162</figureCitation>
,
<figureCitation id="134929D6FFA5FF8DFC64FD3B3063FD05" box="[985,1086,733,757]" captionStart="FIGURES 166 – 169" captionStartId="63.[151,264,1723,1746]" captionTargetBox="[164,1422,810,1687]" captionTargetId="figure@63.[151,1436,803,1702]" captionTargetPageId="63" captionText="FIGURES 166 – 169. Genitalic morphology of Wasaka imitatrix sp. nov.: 166. Male palp, ventral view. 167. Same, retrolateral view. 168. Female epigyne, ventral view. 169. Same, dorsal view. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/284232/files/figure.png" pageId="63" pageNumber="64">166, 167</figureCitation>
).
</paragraph>
<caption id="DF0D65DBFFA5FF8DFF2AF95D3045F900" httpUri="https://zenodo.org/record/284232/files/figure.png" pageId="63" pageNumber="64" targetBox="[164,1422,810,1687]" targetPageId="63">
<paragraph id="8BCD3553FFA5FF8DFF2AF95D3045F900" blockId="63.[151,1436,1723,1776]" pageId="63" pageNumber="64">
<emphasis id="B906E941FFA5FF8DFF2AF95D3530F921" bold="true" box="[151,365,1723,1746]" pageId="63" pageNumber="64">FIGURES 166–169.</emphasis>
Genitalic morphology of
<taxonomicName id="4C724ED0FFA5FF8DFDC9F95A377DF922" box="[628,800,1724,1746]" class="Arachnida" family="Corinnidae" genus="Wasaka" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="imitatrix" status="sp. nov.">
<emphasis id="B906E941FFA5FF8DFDC9F95A377DF922" box="[628,800,1724,1746]" italics="true" pageId="63" pageNumber="64">Wasaka imitatrix</emphasis>
</taxonomicName>
<emphasis id="B906E941FFA5FF8DFC9AF95A3725F921" bold="true" box="[807,888,1724,1745]" pageId="63" pageNumber="64">
<taxonomicNameLabel id="A235543AFFA5FF8DFC9AF95A3725F921" box="[807,888,1724,1745]" pageId="63" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 166. Male palp, ventral view. 167. Same, retrolateral view. 168. Female epigyne, ventral view. 169. Same, dorsal view. Scale bars = 0.25 mm.
</paragraph>
</caption>
<paragraph id="8BCD3553FFA5FF8DFF7AF8FB314BF88D" blockId="63.[151,1437,1821,2025]" pageId="63" pageNumber="64">
<emphasis id="B906E941FFA5FF8DFF7AF8FB3055F8C5" bold="true" box="[199,1032,1821,1846]" pageId="63" pageNumber="64">
Female (
<typeStatus id="54C98BF1FFA5FF8DFE8DF8FB35C7F8C6" box="[304,410,1821,1846]" pageId="63" pageNumber="64" type="holotype">holotype</typeStatus>
, Amani, CAS, CASENT 9033285). Measurements:
</emphasis>
CL 3.38, CW 2.58, AL 3.75, AW 2.25, TL 7.00 (7.00–9.45), FL 0.31, SL 1.49, SW 1.38, AME–AME 0.11, AME–ALE 0.02, ALE–ALE 0.51, PME– PME 0.16, PME–PLE 0.10, PLE–PLE 0.76,
<collectingCountry id="F36575C3FFA5FF8DFD30F8803684F88D" box="[653,729,1894,1917]" name="Peru" pageId="63" pageNumber="64">PERW</collectingCountry>
1.00, MOQAW 0.48, MOQPW 0.55, MOQL 0.56.
</paragraph>
<paragraph id="8BCD3553FFA5FF8DFF7AF86F30DBF835" blockId="63.[151,1437,1821,2025]" pageId="63" pageNumber="64">Length of leg segments: I 3.15 + 1.24 + 2.80 + 2.65 + 1.45 = 11.29; II 3.05 + 1.18 + 2.63 + 2.58 + 1.37 = 10.81; III 3.00 + 1.20 + 2.40 + 2.67 + 1.30 = 10.57; IV 3.73 + 1.30 + 2.95 + 3.90 + 1.25 = 13.13.</paragraph>
<paragraph id="8BCD3553FFA5FFF2FF7AF83735D3FC07" blockId="63.[151,1437,1821,2025]" lastBlockId="64.[151,1437,151,1339]" lastPageId="64" lastPageNumber="65" pageId="63" pageNumber="64">
General appearance as in
<figureCitation id="134929D6FFA5FF8DFE55F837361AF819" box="[488,583,2001,2025]" captionStart="FIGURES 154 – 161" captionStartId="61.[151,264,1108,1131]" captionTargetBox="[153,1436,259,1087]" captionTargetId="figure@61.[153,1438,259,1087]" captionTargetPageId="61" captionText="FIGURES 154 – 161. Digital microscope photographs of Wasaka imitatrix sp. nov. (154, 155), W. montana sp. nov. (156, 157), W. occulta sp. nov. (158, 159) and W. ventralis sp. nov. (160, 161): 154, 156, 158, 160. Male habitus, dorsal view. 155, 157, 159, 161. Female habitus, dorsal view. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/284230/files/figure.png" pageId="63" pageNumber="64">Fig. 155</figureCitation>
, female more robustly built than male. Carapace cream, markings and setae as for male but slightly smaller. AME separated by distance slightly less than ½ their diameter; AME separated from ALE by distance approximately 1⁄10 AME diameter; clypeus height slightly less than AME diameter; PME separated by distance equal to ¾ their diameter; PME separated from PLE by distance equal to ½ PME diameter; CW:
<collectingCountry id="F36575C3FFDAFFF2FF7AFEEF354AFED1" box="[199,279,265,289]" name="Peru" pageId="64" pageNumber="65">PERW</collectingCountry>
= 2.58:1. Chelicerae cream, dentition as for male. Endites, labium and sternum cream, without markings. Legs cream, with black mottling and markings and black spots at spine bases; markings and mottling covered in black feathery setae, with yellowish feathery setae between them; femora I–IV with faint black mottled band distally; patellae I–IV with black mottling laterally, ventrally and distally; tibiae I–III with black mottling between spine pairs, IV with broad median black band, absent from proximal and distal ends; metatarsi I–IV with faint proximal, median and distal bands; tarsi I–IV cream. Leg spination: femora: I pl 3 do 3 rl 2, II pl 3 do 3 rl 3, III pl 3 do 3 rl 3, IV pl 2 do 3 rl 1; all femora with scattered erect ventral setae; patellae: I–IV with fine proximal and distal do setae; tibiae: I pl 2 rl 2 plv 3 rlv 3, II pl 2 rl 2 plv 3 rlv 3, III pl 2 do 1 rl 2 plv 2 rlv 2 vt 1, IV pl 2 do 1 rl 2 plv 2 rlv 2 vt 2; metatarsi: I pl 2 rl 2 plv 2 rlv 2, II pl 2 rl 2 plv 2 rlv 2, III pl 3 rl 3 plv 2 rlv 2 vt 3, IV pl 3 rl 3 plv 2 rlv 2 vt 3. Palpal spination: femora: pl 2 do 1 rl 1, with rlv 8 erect setae; patellae: pl 1 do 2; tibiae: pl 1 do 2 plv 1; tarsi: pl 1 rl 1 plv 3 rlv 1. Abdomen with tiny creamy-yellow dorsal scutum on anterior surface; dorsum cream, densely covered in golden feathery setae with scattered patches of black feathery setae, giving mottled appearance, posterior 1⁄5 of abdomen covered in black feathery setae with scattered yellowish feathery setae; dorsum cream along midline, with a few short narrow chevrons, paired black spots medially just posterior to midpoint; spinnerets surrounded by black ring dorsally and laterally, absent ventrally; venter cream, covered in white feathery and short straight setae, with many long erect straight setae; epigastric scutum weakly sclerotised, creamy-yellow; inframamillary sclerite yellow-orange. Epigyne with large strongly sclerotised 6-shaped ridges laterally in posterior half of epigyne, nearly meeting along midline, with curved copulatory openings (
<figureCitation id="134929D6FFDAFFF2FA96FC9531D2FC7B" box="[1323,1423,883,907]" captionStart="FIGURES 166 – 169" captionStartId="63.[151,264,1723,1746]" captionTargetBox="[164,1422,810,1687]" captionTargetId="figure@63.[151,1436,803,1702]" captionTargetPageId="63" captionText="FIGURES 166 – 169. Genitalic morphology of Wasaka imitatrix sp. nov.: 166. Male palp, ventral view. 167. Same, retrolateral view. 168. Female epigyne, ventral view. 169. Same, dorsal view. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/284232/files/figure.png" pageId="64" pageNumber="65">Fig. 168</figureCitation>
); copulatory ducts short, slightly longer than broad, directed anteriorly, entering anterior ST II posterolaterally; ST II more than twice as broad as ST I, narrowly joined, clearly separate structures; ST I narrow and elongate, situated posteriorly (
<figureCitation id="134929D6FFDAFFF2FEA2FC3935DFFC07" box="[287,386,991,1015]" captionStart="FIGURES 166 – 169" captionStartId="63.[151,264,1723,1746]" captionTargetBox="[164,1422,810,1687]" captionTargetId="figure@63.[151,1436,803,1702]" captionTargetPageId="63" captionText="FIGURES 166 – 169. Genitalic morphology of Wasaka imitatrix sp. nov.: 166. Male palp, ventral view. 167. Same, retrolateral view. 168. Female epigyne, ventral view. 169. Same, dorsal view. Scale bars = 0.25 mm." httpUri="https://zenodo.org/record/284232/files/figure.png" pageId="64" pageNumber="65">Fig. 169</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36866D8FFDAFFF2FF7AFBE536DCFB03" pageId="64" pageNumber="65" type="materials_examined">
<paragraph id="8BCD3553FFDAFFF2FF7AFBE53557FB93" blockId="64.[151,1437,151,1339]" pageId="64" pageNumber="65">
<emphasis id="B906E941FFDAFFF2FF7AFBE53521FBEB" bold="true" box="[199,380,1026,1051]" pageId="64" pageNumber="65">
<typeStatus id="54C98BF1FFDAFFF2FF7AFBE5355FFBEB" box="[199,258,1027,1051]" pageId="64" pageNumber="65">Type</typeStatus>
material:
</emphasis>
<typeStatus id="54C98BF1FFDAFFF2FE35FBE535ADFBEB" box="[392,496,1027,1051]" pageId="64" pageNumber="65" type="holotype">Holotype</typeStatus>
Ƥ:
<emphasis id="B906E941FFDAFFF2FD9BFBE4369EFBEB" bold="true" box="[550,707,1026,1051]" pageId="64" pageNumber="65">
<collectingCountry id="F36575C3FFDAFFF2FD9BFBE436E3FBEB" box="[550,702,1026,1051]" name="Tanzania" pageId="64" pageNumber="65">TANZANIA</collectingCountry>
:
</emphasis>
<emphasis id="B906E941FFDAFFF2FD72FBE23732FBEB" box="[719,879,1028,1051]" italics="true" pageId="64" pageNumber="65">Tanga Region</emphasis>
: East Usambara Mountains, Amani,
<geoCoordinate id="EE465394FFDAFFF2FA95FBE531C5FBEB" box="[1320,1432,1027,1051]" direction="south" orientation="latitude" pageId="64" pageNumber="65" precision="92" value="-5.095">05°05.7'S</geoCoordinate>
,
<geoCoordinate id="EE465394FFDAFFF2FF2AFBC134AFFBCF" box="[151,242,1063,1087]" direction="east" orientation="longitude" pageId="64" pageNumber="65" precision="925" value="38.633335">38°38'E</geoCoordinate>
,
<quantity id="4C8A98B6FFDAFFF2FEB8FBC13519FBCF" box="[261,324,1063,1087]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="64" pageNumber="65" unit="m" value="950.0">950m</quantity>
a.s.l., leg. C.E. Griswold, N. Scharff &amp; D. Ubick,
<date id="FFCC1393FFDAFFF2FC05FBC13033FBCF" box="[952,1134,1063,1087]" pageId="64" pageNumber="65" value="1995-10-27" valueMax="1995-11-09" valueMin="1995-10-27">27.X–9.XI.1995</date>
(forest) (
<collectionCode id="ED63AD96FFDAFFF2FB5EFBC13140FBCF" LSID="urn:lsid:biocol.org:col:34700" box="[1251,1309,1063,1087]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="64" pageNumber="65">CAS</collectionCode>
,
<collectionCode id="ED63AD96FFDAFFF2FA92FBC131C1FBCF" box="[1327,1436,1063,1087]" pageId="64" pageNumber="65">CASENT</collectionCode>
9033285).
</paragraph>
<paragraph id="8BCD3553FFDAFFF2FF7AFB963646FB3F" blockId="64.[151,1437,151,1339]" pageId="64" pageNumber="65">
<typeStatus id="54C98BF1FFDAFFF2FF7AFB963565FB78" box="[199,312,1136,1160]" pageId="64" pageNumber="65" type="paratype">Paratypes</typeStatus>
:
<emphasis id="B906E941FFDAFFF2FEFFFB883582FB77" bold="true" box="[322,479,1134,1159]" pageId="64" pageNumber="65">
<collectingCountry id="F36575C3FFDAFFF2FEFFFB883587FB77" box="[322,474,1134,1159]" name="Tanzania" pageId="64" pageNumber="65">TANZANIA</collectingCountry>
:
</emphasis>
Iringa District,
<taxonomicName id="4C724ED0FFDAFFF2FD2CFB9636A1FB78" box="[657,764,1136,1160]" class="Arachnida" family="Corinnidae" genus="Copuetta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="64" pageNumber="73" phylum="Arthropoda" rank="species" species="uzungwa">Uzungwa</taxonomicName>
Scarp Forest Reserve,
<quantity id="4C8A98B6FFDAFFF2FC43FB893060FB77" box="[1022,1085,1135,1159]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="64" pageNumber="65" unit="km" value="11.0">11km</quantity>
SE Masisiwe, Kihanga stream,
<geoCoordinate id="EE465394FFDAFFF2FF2AFB753573FB5B" box="[151,302,1171,1195]" direction="south" orientation="latitude" pageId="64" pageNumber="65" precision="1" value="-8.36825">08°22'05.7&quot;S</geoCoordinate>
,
<geoCoordinate id="EE465394FFDAFFF2FE82FB75358AFB5B" box="[319,471,1171,1195]" direction="east" orientation="longitude" pageId="64" pageNumber="65" precision="1" value="35.97822">35°58'41.6&quot;E</geoCoordinate>
,
<quantity id="4C8A98B6FFDAFFF2FE55FB75366BFB5B" box="[488,566,1171,1195]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="64" pageNumber="65" unit="m" value="1800.0">1800m</quantity>
a.s.l., leg.
<collectionCode id="ED63AD96FFDAFFF2FD7CFB75374FFB5B" box="[705,786,1171,1195]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="64" pageNumber="65">ZMUC</collectionCode>
&amp;
<collectionCode id="ED63AD96FFDAFFF2FCFCFB7537CFFB5B" LSID="urn:lsid:biocol.org:col:34871" box="[833,914,1171,1195]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="64" pageNumber="65">USNM</collectionCode>
expedition,
<date id="FFCC1393FFDAFFF2FB94FB753098FB5B" box="[1065,1221,1171,1195]" pageId="64" pageNumber="65" value="1997-05-17" valueMax="1997-05-27" valueMin="1997-05-17">17–27.V.1997</date>
(canopy), 13 1Ƥ (
<collectionCode id="ED63AD96FFDAFFF2FF22FB5134ABFB3F" LSID="urn:lsid:biocol.org:col:34871" box="[159,246,1207,1231]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="64" pageNumber="65">USNM</collectionCode>
); same data, 2Ƥ (
<collectionCode id="ED63AD96FFDAFFF2FE05FB513652FB3F" box="[440,527,1207,1231]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="64" pageNumber="65">ZMUC</collectionCode>
).
</paragraph>
<paragraph id="8BCD3553FFDAFFF2FF7AFB3C36DCFB03" blockId="64.[151,1437,151,1339]" box="[199,641,1242,1267]" pageId="64" pageNumber="65">
<emphasis id="B906E941FFDAFFF2FF7AFB3C366BFB03" bold="true" box="[199,566,1242,1267]" pageId="64" pageNumber="65">Additional material examined:</emphasis>
None.
</paragraph>
</subSubSection>
<subSubSection id="C36866D8FFDAFFF2FF7AFB183071FAE7" box="[199,1068,1278,1303]" pageId="64" pageNumber="65" type="distribution">
<paragraph id="8BCD3553FFDAFFF2FF7AFB183071FAE7" blockId="64.[151,1437,151,1339]" box="[199,1068,1278,1303]" pageId="64" pageNumber="65">
<emphasis id="B906E941FFDAFFF2FF7AFB18353FFAE7" bold="true" box="[199,354,1278,1303]" pageId="64" pageNumber="65">Distribution:</emphasis>
Known only from two localities in eastern
<collectingCountry id="F36575C3FFDAFFF2FCF4FB1937F3FAE7" box="[841,942,1279,1303]" name="Tanzania" pageId="64" pageNumber="65">Tanzania</collectingCountry>
(
<figureCitation id="134929D6FFDAFFF2FC00FB193042FAE7" box="[957,1055,1279,1303]" captionStart="FIGURE 182" captionStartId="71.[151,250,1279,1302]" captionTargetBox="[151,1436,373,1258]" captionTargetId="figure@71.[151,1436,373,1258]" captionTargetPageId="71" captionText="FIGURE 182. Distribution of Wasaka imitatrix sp. nov. (white circles), W. montana sp. nov. (black circles), W. occulta sp. nov. (black triangle) and W. ventralis sp. nov. (white triangles) in tropical Africa." httpUri="https://zenodo.org/record/284236/files/figure.png" pageId="64" pageNumber="65">Fig. 182</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36866D8FFDAFFF2FF7AFAC4302BFACB" box="[199,1142,1314,1339]" pageId="64" pageNumber="65" type="biology_ecology">
<paragraph id="8BCD3553FFDAFFF2FF7AFAC4302BFACB" blockId="64.[151,1437,151,1339]" box="[199,1142,1314,1339]" pageId="64" pageNumber="65">
<emphasis id="B906E941FFDAFFF2FF7AFAC43577FACB" bold="true" box="[199,298,1314,1339]" pageId="64" pageNumber="65">Biology:</emphasis>
Collected in forests but details of the sampling methodology are unknown.
</paragraph>
</subSubSection>
</treatment>
</document>
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