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<document id="C2437F3703BF4507E55F0F816FF6FA8D" ID-CLB-Dataset="24900" ID-DOI="10.1093/isd/ixz015" ID-GBIF-Dataset="56172c62-3e7c-4ab7-83d4-1dd9f5c57eb5" ID-Zenodo-Dep="3542130" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1573763013583" checkinUser="plazi" docAuthor="Hita-Garcia, Francisco, Lieberman, Ziv, Audisio, Tracy L., Liu, Cong &amp; Economo, Evan P." docDate="2019" docId="03D9AC4AE54BFF9AFCC4FB1DBFD105F2" docLanguage="en" docName="insectSystematicsDiversity.ixz015.pdf" docOrigin="Insect Systematics and Diversity 5" docStyle="DocumentStyle:20C34595BA01015AC07A3B955E96DF74.8:InsectSystDivers.2017-.journal_article" docStyleId="20C34595BA01015AC07A3B955E96DF74" docStyleName="InsectSystDivers.2017-.journal_article" docStyleVersion="8" docTitle="Discothyrea athene Hita-Garcia &amp; Lieberman &amp; Audisio &amp; Liu &amp; Economo 2019, sp. n." docType="treatment" docVersion="6" lastPageNumber="34" masterDocId="FFE0D432E555FFBBFFF6FF95BB120352" masterDocTitle="Revision of the Highly Specialized Ant Genus Discothyrea (Hymenoptera: Formicidae) in the Afrotropics with X-Ray Microtomography and 3 D Cybertaxonomy" masterLastPageNumber="84" masterPageNumber="1" pageNumber="31" updateTime="1698765580234" updateUser="ExternalLinkService">
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<mods:title id="103E81EE3A00E756AA8A8863B86A3E8D">Revision of the Highly Specialized Ant Genus Discothyrea (Hymenoptera: Formicidae) in the Afrotropics with X-Ray Microtomography and 3 D Cybertaxonomy</mods:title>
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Hita Garcia &amp; Lieberman
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(
<figureCitation id="134B01D9E54BFFA5FCCCFB3EB8870790" box="[826,917,1195,1218]" captionStart="Fig" captionStartId="5.[146,173,1283,1300]" captionTargetBox="[153,1435,148,1247]" captionTargetId="figure@5.[152,1435,147,1248]" captionTargetPageId="5" captionText="Fig. 4. Maps showing the known distribution ranges of Afrotropical Discothyrea species." figureDoi="http://doi.org/10.5281/zenodo.3542140" httpUri="https://zenodo.org/record/3542140/files/figure.png" pageId="30" pageNumber="31">Figs. 4D</figureCitation>
,
<figureCitation id="134B01D9E54BFFA5FC54FB3EB8D20790" box="[930,960,1195,1218]" captionStart="Fig" captionStartId="10.[146,173,1454,1471]" captionTargetBox="[155,1434,149,1418]" captionTargetId="figure@10.[154,1434,147,1419]" captionTargetPageId="10" captionText="Fig. 6. Diagnostic plate showing still images from surface volume renderings of the head in full-face view (the remainder of the body virtually removed). All Afrotropical species are shown, species names are embedded in each particular image for better comparison and orientation.Specimen codes are given inTable 1." figureDoi="http://doi.org/10.5281/zenodo.3542144" httpUri="https://zenodo.org/record/3542144/files/figure.png" pageId="30" pageNumber="31">6D</figureCitation>
,
<figureCitation id="134B01D9E54BFFA5FC38FB3EB8FF0790" box="[974,1005,1195,1218]" captionStart="Fig" captionStartId="11.[146,173,1457,1474]" captionTargetBox="[153,1435,149,1421]" captionTargetId="figure@11.[152,1435,147,1422]" captionTargetPageId="11" captionText="Fig. 7. Diagnostic plate showing still images from surface volume renderings of the head in profile (the remainder of the body virtually removed).All Afrotropical species are shown, species names are embedded in each particular image for better comparison and orientation. Specimen codes are given inTable 1." figureDoi="http://doi.org/10.5281/zenodo.3542148" httpUri="https://zenodo.org/record/3542148/files/figure.png" pageId="30" pageNumber="31">7D</figureCitation>
,
<figureCitation id="134B01D9E54BFFA5FC0CFB3EBF0B0790" box="[1018,1049,1195,1218]" captionStart="Fig" captionStartId="12.[146,172,1457,1474]" captionTargetBox="[153,1435,149,1421]" captionTargetId="figure@12.[152,1435,147,1422]" captionTargetPageId="12" captionText="Fig. 8. Diagnostic plate showing still images from surface volume renderings of the right mandible in dorsal view (the remainder of the body virtually removed).All Afrotropical species are shown,species names are embedded in each particular image for better comparison and orientation.Specimen codes are given inTable 1." figureDoi="http://doi.org/10.5281/zenodo.3542150" httpUri="https://zenodo.org/record/3542150/files/figure.png" pageId="30" pageNumber="31">8D</figureCitation>
,
<figureCitation id="134B01D9E54BFFA5FBD0FB3EBF570790" box="[1062,1093,1195,1218]" captionStart="Fig" captionStartId="13.[146,173,1457,1474]" captionTargetBox="[153,1435,149,1421]" captionTargetId="figure@13.[152,1435,147,1422]" captionTargetPageId="13" captionText="Fig. 9. Diagnostic plate showing still images from surface volume renderings of the mesosoma in profile (the remainder of the body virtually removed). All Afrotropical species are shown, species names are embedded in each particular image for better comparison and orientation.Specimen codes are given inTable 1." figureDoi="http://doi.org/10.5281/zenodo.3542152" httpUri="https://zenodo.org/record/3542152/files/figure.png" pageId="30" pageNumber="31">9D</figureCitation>
,
<figureCitation id="134B01D9E54BFFA5FBA4FB3EBF6E0790" box="[1106,1148,1195,1218]" captionStart="Fig" captionStartId="14.[146,173,1457,1474]" captionTargetBox="[153,1435,149,1421]" captionTargetId="figure@14.[152,1435,147,1422]" captionTargetPageId="14" captionText="Fig. 10. Diagnostic plate showing still images from surface volume renderings of the body in dorsal view with focus on mesosoma.All Afrotropical species are shown, species names are embedded in each particular image for better comparison and orientation. Specimen codes are given inTable 1." figureDoi="http://doi.org/10.5281/zenodo.3542154" httpUri="https://zenodo.org/record/3542154/files/figure.png" pageId="30" pageNumber="31">10D</figureCitation>
,
<figureCitation id="134B01D9E54BFFA5FB7CFB3EBFA20790" box="[1162,1200,1195,1218]" captionStart="Fig" captionStartId="15.[146,173,1454,1471]" captionTargetBox="[155,1434,149,1418]" captionTargetId="figure@15.[154,1434,147,1419]" captionTargetPageId="15" captionText="Fig. 11. Diagnostic plate showing still images from surface volume renderings of the posterior propodeum in posterior view (the remainder of the body virtually removed). All Afrotropical species are shown, species names are embedded in each particular image for better comparison and orientation. Specimen codes are given inTable 1." figureDoi="http://doi.org/10.5281/zenodo.3542156" httpUri="https://zenodo.org/record/3542156/files/figure.png" pageId="30" pageNumber="31">11D</figureCitation>
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<figureCitation id="134B01D9E54BFFA5FB36FB3EBFF90790" box="[1216,1259,1195,1218]" captionStart="Fig" captionStartId="16.[146,173,1454,1471]" captionTargetBox="[155,1434,149,1418]" captionTargetId="figure@16.[154,1434,147,1419]" captionTargetPageId="16" captionText="Fig. 12. Diagnostic plate showing still images from surface volume renderings of the petiole and gaster in profile (the remainder of the body virtually removed).All Afrotropical species are shown, species names are embedded in each particular image for better comparison and orientation.Specimen codes are given inTable 1." figureDoi="http://doi.org/10.5281/zenodo.3542158" httpUri="https://zenodo.org/record/3542158/files/figure.png" pageId="30" pageNumber="31">12D</figureCitation>
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<figureCitation id="134B01D9E54BFFA5FB0FFB3EBE360790" box="[1273,1316,1195,1218]" captionStart="Fig" captionStartId="18.[146,173,1454,1471]" captionTargetBox="[155,1434,149,1419]" captionTargetId="figure@18.[154,1434,147,1419]" captionTargetPageId="18" captionText="Fig. 14. Diagnostic plate showing differences in body pilosity. (A) D. mixta (CASENT0235473),(B) D. oculata (CASENT0235467),(C) D. aisnetu (CASENT0235475), (D) D.athene (CASENT0235476),(E) D.chimera (CASENT0235471),(F) D.damato (CASENT0247362),(G) D.dryad (CASENT0247371),(H) D.gaia (CASENT0247040), (I) D. gryphon (CASENT0247367), (J) D. hawkesi (CASENT0235470), (K) D. kalypso (CASENT0235468), (L) D. maia (CASENT0790541), (M) D. michelae (CASENT0235469), (N) D. patrizii (CASENT0235472), (O) D. penthos (CASENT0247383), (P) D. poweri (SAM-ENT-0011509), (Q) D. schulzei (CASENT0247370), (R) D. traegaordhi (CASENT0790122), (S) D. venus (CASENT0247017), (T) D. wakanda (CASENT0790326).With the exception of F, L, R,T, all other images are from https://www.antweb.org—photographers Michele Esposito and Will Ericson." figureDoi="http://doi.org/10.5281/zenodo.3542162" httpUri="https://zenodo.org/record/3542162/files/figure.png" pageId="30" pageNumber="31">14D</figureCitation>
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<figureCitation id="134B01D9E54BFFA5FAC7FB3EBE5E0790" box="[1329,1356,1195,1218]" captionStart="Fig" captionStartId="31.[146,173,1061,1078]" captionTargetBox="[147,768,147,1026]" captionTargetId="figure@31.[147,768,147,1026]" captionTargetPageId="31" captionText="Fig. 25. Stacked digital color images of D. athene sp. n. (CASENT0235476 - from https://www.antweb.org, photographer Will Ericson). (A) body in profile, (B) body in dorsal view, (C) head in full-face view." figureDoi="http://doi.org/10.5281/zenodo.3542184" httpUri="https://zenodo.org/record/3542184/files/figure.png" pageId="30" pageNumber="31">25</figureCitation>
,
<figureCitation id="134B01D9E54BFFA5FAACFB3EBE670790" box="[1370,1397,1195,1218]" captionStart="Fig" captionStartId="32.[146,173,1782,1799]" captionTargetBox="[202,1386,147,1747]" captionTargetId="figure@32.[202,1386,147,1747]" captionTargetPageId="32" captionText="Fig. 26. Still images from shaded surface display volume renderings of D. athene sp. n. holotype (CASENT0764088) showing virtually segmented body parts. (A) Body in dorsal view, (B) body in profile, (C) head in full-face view, (D) head in dorsal view, (E) head in anterodorsal view, (F) head in anterior view, (G) head in profile, (H) head in ventral view, (I) posterior propodeum in posterior view, (J) petiole in profile, (K) petiole and gaster in profile, (L) petiole in dorsal view, (M) petiole in anterior view, (N) petiole oblique anterior view, (O) gaster in dorsal view, (P) petiole in ventral view, (Q) abdominal sternite 3 in ventral view, (R) mesotibia and mesotarsus in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3542186" httpUri="https://zenodo.org/record/3542186/files/figure.png" pageId="30" pageNumber="31">26</figureCitation>
; Supp Video S4 [online only])
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:
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<materialsCitation id="3B181701E54BFFA5FB52FAF6B8F7069C" ID-GBIF-Occurrence="2451560204" collectingDate="2012-08-06" collectingDateMax="2012-08-16" collectingDateMin="2012-08-06" collectionCode="CASC, MHNG, SAMC, ZMFK" collectorName="F. Hita Garcia" country="Uganda" elevation="1510" latitude="0.56437" location="Kanyawara Biological Station" longLatPrecision="5" longitude="30.3605" municipality="National Park" pageId="30" pageNumber="31" specimenCode="CASENT0764102, CASENT0764100, CASENT0764101, CASENT0764087" specimenCount="1" stateProvince="Kibale" typeStatus="paratype">
<emphasis id="B904C14EE54BFFA5FB52FAF6BE0C0624" bold="true" box="[1188,1310,1379,1398]" pageId="30" pageNumber="31">
<typeStatus id="54CBA3FEE54BFFA5FB52FAF6BE0C0624" box="[1188,1310,1379,1398]" pageId="30" pageNumber="31" type="paratype">PARATYPES</typeStatus>
</emphasis>
, four pinned workers with same data as holotype (
<collectionCode id="ED618599E54BFFA5FB5FFA15BFF706C1" box="[1193,1253,1408,1427]" pageId="30" pageNumber="31">CASC</collectionCode>
:
<specimenCode id="DBD6B527E54BFFA5FB05FA15BE8C06C1" box="[1267,1438,1408,1427]" collectionCode="CASC" pageId="30" pageNumber="31">CASENT0764102</specimenCode>
;
<collectionCode id="ED618599E54BFFA5FCC4FA0BB86D06E3" box="[818,895,1438,1457]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="30" pageNumber="31" type="Museum">MHNG</collectionCode>
:
<specimenCode id="DBD6B527E54BFFA5FC79FA08BF2906E2" box="[911,1083,1437,1456]" collectionCode="MHNG" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="30" pageNumber="31" type="Museum">CASENT0764100</specimenCode>
;
<collectionCode id="ED618599E54BFFA5FBBAFA0BBF9F06E3" box="[1100,1165,1438,1457]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="30" pageNumber="31" type="Museum">SAMC</collectionCode>
:
<specimenCode id="DBD6B527E54BFFA5FB69FA08BE5806E2" box="[1183,1354,1437,1456]" collectionCode="SAMC" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="30" pageNumber="31" type="Museum">CASENT0764101</specimenCode>
;
<collectionCode id="ED618599E54BFFA5FAAAFA0BBE8D06E3" box="[1372,1439,1438,1457]" pageId="30" pageNumber="31">ZMFK</collectionCode>
:
<specimenCode id="DBD6B527E54BFFA5FCC4FA2EB8CD069C" box="[818,991,1467,1486]" collectionCode="ZMFK" pageId="30" pageNumber="31">CASENT0764087</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BCF1D5CE54BFFA5FCC4FA6EBEB105AB" blockId="30.[818,1443,1531,1785]" pageId="30" pageNumber="31">
<emphasis id="B904C14EE54BFFA5FCC4FA6EB885055C" box="[818,919,1531,1550]" italics="true" pageId="30" pageNumber="31">Cybertype.</emphasis>
Volumetric raw data (in DICOM format), 3D rotation video, still images of surface volume rendering, and 3D surface (in PLY format) of the physical
<typeStatus id="54CBA3FEE54BFFA5FBB2F9A0BF87051A" box="[1092,1173,1589,1608]" pageId="30" pageNumber="31" type="holotype">holotype</typeStatus>
(CASENT0764088) in addition to stacked digital color images illustrating head in full-face view, profile and dorsal views of the body.The data are deposited at Dryad (Hita Garcia et al. 2019, http://doi.org/10.5061/dryad.3qm4183) and can be freely accessed as virtual representation of the type. In addition to the cybertype data at Dryad, we also provide a freely accessible 3D surface model of the
<typeStatus id="54CBA3FEE54BFFA5FB85F970BFD605AA" box="[1139,1220,1765,1784]" pageId="30" pageNumber="31" type="holotype">holotype</typeStatus>
at Sketchfab (Model 4).
</paragraph>
<paragraph id="8BCF1D5CE54BFFA5FCC4F8BDB8C90469" blockId="30.[818,1443,1832,1910]" box="[818,987,1832,1851]" pageId="30" pageNumber="31">
<heading id="D087AA30E54BFFA5FCC4F8BDB8C90469" bold="true" box="[818,987,1832,1851]" fontSize="8" level="3" pageId="30" pageNumber="31" reason="6">
<emphasis id="B904C14EE54BFFA5FCC4F8BDB8C90469" bold="true" box="[818,987,1832,1851]" pageId="30" pageNumber="31">Non-type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF1D5CE54BFFA5FCC4F8D3BEB10424" blockId="30.[818,1443,1832,1910]" pageId="30" pageNumber="31">
<materialsCitation id="3B181701E54BFFA5FCC4F8D3BEB10424" ID-GBIF-Occurrence="2451560245" collectingDate="2009-06" collectorName="G. Fischer" country="Kenya" elevation="136" latitude="-3.314367" location="Coast Province" longLatPrecision="1" longitude="39.94073" municipality="Arabuko Sokoke Forest" pageId="30" pageNumber="31" specimenCount="1">
<emphasis id="B904C14EE54BFFA5FCC4F8D3B890040B" box="[818,898,1862,1881]" italics="true" pageId="30" pageNumber="31">
<collectingCountry id="F3675DCCE54BFFA5FCC4F8D3B86C040B" box="[818,894,1862,1881]" name="Kenya" pageId="30" pageNumber="31">KENYA</collectingCountry>
:
</emphasis>
<location id="8EAF4B87E54BFFA5FC64F8D3BF34040A" LSID="urn:lsid:plazi:treatment:03D9AC4AE54BFF9AFCC4FB1DBFD105F2:8EAF4B87E54BFFA5FC64F8D3BF34040A" box="[914,1062,1861,1881]" country="Kenya" latitude="-3.314367" longLatPrecision="1" longitude="39.94073" municipality="Arabuko Sokoke Forest" name="Coast Province" pageId="30" pageNumber="31">Coast Province</location>
,
<collectingMunicipality id="6BAB8726E54BFFA5FBCFF8D0BE34040B" box="[1081,1318,1861,1881]" pageId="30" pageNumber="31">Arabuko Sokoke Forest</collectingMunicipality>
,
<geoCoordinate id="EE447B9BE54BFFA5FACFF8D0BE8C040A" box="[1337,1438,1861,1880]" degrees="3.314367" direction="south" orientation="latitude" pageId="30" pageNumber="31" precision="1" value="-3.314367">3.314367</geoCoordinate>
,
<geoCoordinate id="EE447B9BE54BFFA5FCC4F8F6B8850424" box="[818,919,1891,1910]" degrees="39.940731" direction="east" orientation="longitude" pageId="30" pageNumber="31" precision="1" value="39.94073">39.940731</geoCoordinate>
,
<quantity id="4C88B0B9E54BFFA5FC52F8F6B8F10424" box="[932,995,1891,1910]" metricMagnitude="2" metricUnit="m" metricValue="1.3599999999999999" pageId="30" pageNumber="31" unit="m" value="136.0">
<elevation id="005DFA6FE54BFFA5FC52F8F6B8F10424" box="[932,995,1891,1910]" metricMagnitude="2" metricUnit="m" metricValue="1.3599999999999999" pageId="30" pageNumber="31" unit="m" value="136.0">136 m</elevation>
</quantity>
, mixed forest,
<collectingMethod id="5231654BE54BFFA5FB84F8F6BFCA0424" box="[1138,1240,1891,1910]" pageId="30" pageNumber="31">pitfall trap</collectingMethod>
,
<date id="FFCE3B9CE54BFFA5FB12F8F6BE3D0424" box="[1252,1327,1891,1910]" pageId="30" pageNumber="31" value="2009-06">
<collectingDate id="EF8AC274E54BFFA5FB12F8F6BE3D0424" box="[1252,1327,1891,1910]" pageId="30" pageNumber="31" value="2009-06">VI.2009</collectingDate>
</date>
(
<emphasis id="B904C14EE54BFFA5FAC9F8F6BEB10424" box="[1343,1443,1891,1910]" italics="true" pageId="30" pageNumber="31">
<collectorName id="2685788AE54BFFA5FAC9F8F6BE8D0424" box="[1343,1439,1891,1910]" pageId="30" pageNumber="31">G. Fischer</collectorName>
,
</emphasis>
</materialsCitation>
</paragraph>
<caption id="DF0F4DD4E54AFFA4FF64FBB0B975073B" ID-DOI="http://doi.org/10.5281/zenodo.3542184" ID-Zenodo-Dep="3542184" httpUri="https://zenodo.org/record/3542184/files/figure.png" pageId="31" pageNumber="32" startId="31.[146,173,1061,1078]" targetBox="[147,768,147,1026]" targetPageId="31">
<paragraph id="8BCF1D5CE54AFFA4FF64FBB0B975073B" blockId="31.[146,770,1061,1129]" pageId="31" pageNumber="32">
<emphasis id="B904C14EE54AFFA4FF64FBB0BBC20764" bold="true" box="[146,208,1061,1078]" pageId="31" pageNumber="32">Fig. 25.</emphasis>
Stacked digital color images of
<taxonomicName id="4C7066DFE54AFFA4FE13FBB0B9240764" authority="Hita-Garcia &amp; Lieberman &amp; Audisio &amp; Liu &amp; Economo, 2019" authorityName="Hita-Garcia &amp; Lieberman &amp; Audisio &amp; Liu &amp; Economo" authorityYear="2019" box="[485,566,1061,1078]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="athene" status="sp. nov.">
<emphasis id="B904C14EE54AFFA4FE13FBB0B9240764" box="[485,566,1061,1078]" italics="true" pageId="31" pageNumber="32">D. athene</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2377C35E54AFFA4FDCBFBB0B97F0764" box="[573,621,1061,1078]" pageId="31" pageNumber="32" rank="species">sp. n.</taxonomicNameLabel>
(CASENT0235476 - from https://www.antweb.org, photographer Will Ericson). (A) body in profile, (B) body in dorsal view, (C) head in full-face view.
</paragraph>
</caption>
<paragraph id="8BCF1D5CE54AFFA4FF64FB08BF440253" blockId="31.[146,771,1180,1933]" lastBlockId="31.[818,1442,149,257]" pageId="31" pageNumber="32">
<emphasis id="B904C14EE54AFFA4FF64FB08BA0007E2" box="[146,274,1181,1200]" italics="true" pageId="31" pageNumber="32">F. Hita Garcia</emphasis>
); Coast Province, Arabuko Sokoke Forest,
<geoCoordinate id="EE447B9BE54AFFA4FD6FFB08B9EC07E2" box="[665,766,1181,1200]" degrees="3.321139" direction="south" orientation="latitude" pageId="31" pageNumber="32" precision="1" value="-3.321139">3.321139</geoCoordinate>
,
<geoCoordinate id="EE447B9BE54AFFA4FF64FB2FBBEA079F" box="[146,248,1210,1229]" degrees="39.929194" direction="east" orientation="longitude" pageId="31" pageNumber="32" precision="1" value="39.929195">39.929194</geoCoordinate>
, ca.
<quantity id="4C88B0B9E54AFFA4FED7FB2FBA40079F" box="[289,338,1210,1229]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="31" pageNumber="32" unit="m" value="50.0">50 m</quantity>
, mixed forest, hand collected,
<date id="FFCE3B9CE54AFFA4FD9DFB2FB9A4079F" box="[619,694,1210,1229]" pageId="31" pageNumber="32" value="2009-06">VI.2009</date>
(
<emphasis id="B904C14EE54AFFA4FD32FB2FBA2707B8" italics="true" pageId="31" pageNumber="32">F. Hita Garcia, G. Fischer</emphasis>
);
<collectingCountry id="F3675DCCE54AFFA4FEB0FB42BAF707B8" box="[326,485,1239,1258]" name="Mozambique" pageId="31" pageNumber="32">MOZAMBIQUE</collectingCountry>
:
<materialsCitation id="3B181701E54AFFA4FE1BFB42BAC90610" ID-GBIF-Occurrence="2451560241" collectingDate="2013-05-03" collectorName="Winkler &amp; L. E. Alonso" country="Rwanda" county="Gorongosa" elevation="115" latitude="-19.03299" location="Camp" longLatPrecision="1" longitude="34.671757" municipality="Gorongosa National Park" pageId="31" pageNumber="32" specimenCount="1" stateProvince="Sofala">
<collectingRegion id="49B4D3BEE54AFFA4FE1BFB42B93507B8" box="[493,551,1239,1258]" country="Mozambique" name="Sofala" pageId="31" pageNumber="32">Sofala</collectingRegion>
,
<collectingCounty id="62AE65D0E54AFFA4FDD8FB42B98707B8" box="[558,661,1239,1258]" pageId="31" pageNumber="32">Gorongosa</collectingCounty>
,
<collectingMunicipality id="6BAB8726E54AFFA4FD6BFB42BA040655" pageId="31" pageNumber="32">Gorongosa National Park</collectingMunicipality>
,
<location id="8EAF4B87E54AFFA4FED7FB60BA4A065A" LSID="urn:lsid:plazi:treatment:03D9AC4AE54BFF9AFCC4FB1DBFD105F2:8EAF4B87E54AFFA4FED7FB60BA4A065A" box="[289,344,1269,1288]" country="Rwanda" county="Gorongosa" latitude="-19.03299" longLatPrecision="1" longitude="34.671757" municipality="Gorongosa National Park" name="Camp" pageId="31" pageNumber="32" stateProvince="Sofala">Camp</location>
#3,
<geoCoordinate id="EE447B9BE54AFFA4FE72FB60BAE7065A" box="[388,501,1269,1288]" degrees="19.032989" direction="south" orientation="latitude" pageId="31" pageNumber="32" precision="1" value="-19.03299">19.032989</geoCoordinate>
,
<geoCoordinate id="EE447B9BE54AFFA4FDF6FB61B9740655" box="[512,614,1268,1287]" degrees="34.671758" direction="east" orientation="longitude" pageId="31" pageNumber="32" precision="1" value="34.671757">34.671758</geoCoordinate>
,
<quantity id="4C88B0B9E54AFFA4FD86FB61B9BC065A" box="[624,686,1268,1288]" metricMagnitude="2" metricUnit="m" metricValue="1.15" pageId="31" pageNumber="32" unit="m" value="115.0">
<elevation id="005DFA6FE54AFFA4FD86FB61B9BC065A" box="[624,686,1268,1288]" metricMagnitude="2" metricUnit="m" metricValue="1.15" pageId="31" pageNumber="32" unit="m" value="115.0">115 m</elevation>
</quantity>
, riparian forest along creek, darker and closed, atop creek bed,
<collectorName id="2685788AE54AFFA4FD43FA87B9EC0677" box="[693,766,1298,1317]" pageId="31" pageNumber="32">Winkler</collectorName>
,
<date id="FFCE3B9CE54AFFA4FF64FABABBF60610" box="[146,228,1327,1346]" pageId="31" pageNumber="32" value="2013-05-03">
<collectingDate id="EF8AC274E54AFFA4FF64FABABBF60610" box="[146,228,1327,1346]" pageId="31" pageNumber="32" value="2013-05-03">3.V.2013</collectingDate>
</date>
(
<emphasis id="B904C14EE54AFFA4FF04FAA5BA770610" box="[242,357,1327,1346]" italics="true" pageId="31" pageNumber="32">
<collectorName id="2685788AE54AFFA4FF04FAA5BA770610" box="[242,357,1327,1346]" pageId="31" pageNumber="32">L. E. Alonso</collectorName>
</emphasis>
);
<collectingCountry id="F3675DCCE54AFFA4FE81FABABAC90610" box="[375,475,1327,1346]" name="Rwanda" pageId="31" pageNumber="32">RWANDA</collectingCountry>
</materialsCitation>
:
<materialsCitation id="3B181701E54AFFA4FE15FABAB956062F" ID-GBIF-Occurrence="2451560198" collectingDate="1973-07-10" collectingDateMax="1976-09" collectingDateMin="1973-07-10" collectorName="Werner &amp; P. Werner" country="Tanzania" elevation="1800" latitude="-2.39361" location="Rangiro" longLatPrecision="1" longitude="29.18278" municipality="Rangiro" pageId="31" pageNumber="32" specimenCount="1">
<collectingMunicipality id="6BAB8726E54AFFA4FE15FABAB93D0610" box="[483,559,1327,1346]" pageId="31" pageNumber="32">Rangiro</collectingMunicipality>
, [
<geoCoordinate id="EE447B9BE54AFFA4FDC8FABAB9860610" box="[574,660,1327,1346]" degrees="2.39361" direction="south" orientation="latitude" pageId="31" pageNumber="32" precision="1" value="-2.39361">-2.39361</geoCoordinate>
,
<geoCoordinate id="EE447B9BE54AFFA4FD6BFABAB9EB0610" box="[669,761,1327,1346]" degrees="29.18278" direction="east" orientation="longitude" pageId="31" pageNumber="32" precision="1" value="29.18278">29.18278</geoCoordinate>
],
<quantity id="4C88B0B9E54AFFA4FF64FAD8BBCB0632" box="[146,217,1357,1376]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="31" pageNumber="32" unit="m" value="1800.0">
<elevation id="005DFA6FE54AFFA4FF64FAD8BBCB0632" box="[146,217,1357,1376]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="31" pageNumber="32" unit="m" value="1800.0">1800 m</elevation>
</quantity>
,
<date id="FFCE3B9CE54AFFA4FF14FAD8BA400632" box="[226,338,1357,1376]" pageId="31" pageNumber="32" value="1973-07-10">
<collectingDate id="EF8AC274E54AFFA4FF14FAD8BA400632" box="[226,338,1357,1376]" pageId="31" pageNumber="32" value="1973-07-10">10.VII.1973</collectingDate>
</date>
(
<emphasis id="B904C14EE54AFFA4FEA8FAD8BAA6060D" box="[350,436,1357,1375]" italics="true" pageId="31" pageNumber="32">
P.
<collectorName id="2685788AE54AFFA4FE87FAD8BAA6060D" box="[369,436,1357,1375]" pageId="31" pageNumber="32">Werner</collectorName>
</emphasis>
);
<location id="8EAF4B87E54AFFA4FE30FAD9B900060D" LSID="urn:lsid:plazi:treatment:03D9AC4AE54BFF9AFCC4FB1DBFD105F2:8EAF4B87E54AFFA4FE30FAD9B900060D" box="[454,530,1356,1375]" country="Tanzania" latitude="-2.39361" longLatPrecision="1" longitude="29.18278" municipality="Rangiro" name="Rangiro" pageId="31" pageNumber="32">Rangiro</location>
, [
<geoCoordinate id="EE447B9BE54AFFA4FDD6FAD9B964060D" box="[544,630,1356,1375]" degrees="2.39361" direction="south" orientation="latitude" pageId="31" pageNumber="32" precision="1" value="-2.39361">-2.39361</geoCoordinate>
,
<geoCoordinate id="EE447B9BE54AFFA4FD89FAD8B9C80632" box="[639,730,1357,1376]" degrees="29.18278" direction="east" orientation="longitude" pageId="31" pageNumber="32" precision="1" value="29.18278">29.18278</geoCoordinate>
], ca.
<quantity id="4C88B0B9E54AFFA4FF64FAFFBBCA062F" box="[146,216,1386,1405]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="31" pageNumber="32" unit="m" value="1800.0">
<elevation id="005DFA6FE54AFFA4FF64FAFFBBCA062F" box="[146,216,1386,1405]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="31" pageNumber="32" unit="m" value="1800.0">1800 m</elevation>
</quantity>
, litter,
<date id="FFCE3B9CE54AFFA4FEE7FAFFBA4C062F" box="[273,350,1386,1405]" pageId="31" pageNumber="32" value="1976-09">
<collectingDate id="EF8AC274E54AFFA4FEE7FAFFBA4C062F" box="[273,350,1386,1405]" pageId="31" pageNumber="32" value="1976-09">IX.1976</collectingDate>
</date>
(
<emphasis id="B904C14EE54AFFA4FE9EFAFFBAAC062E" box="[360,446,1386,1404]" italics="true" pageId="31" pageNumber="32">
<collectorName id="2685788AE54AFFA4FE9EFAFFBAAC062E" box="[360,446,1386,1404]" pageId="31" pageNumber="32">P.Werner</collectorName>
</emphasis>
);
<collectingCountry id="F3675DCCE54AFFA4FE3BFAFFB956062F" box="[461,580,1386,1405]" name="Tanzania" pageId="31" pageNumber="32">TANZANIA</collectingCountry>
</materialsCitation>
:
<materialsCitation id="3B181701E54AFFA4FDBDFAFFB95206E5" ID-GBIF-Occurrence="2451560168" collectingDate="1996-05-16" collectorName="Winkler &amp; H. G. Robertson" country="Tanzania" elevation="550" latitude="-4.283333" location="Gonja Forest Reserve" longLatPrecision="1" longitude="38.03333" pageId="31" pageNumber="32" specimenCount="1" stateProvince="Kilimanjaro">
<collectingRegion id="49B4D3BEE54AFFA4FDBDFAFFB9EC062F" box="[587,766,1386,1405]" country="Tanzania" name="Kilimanjaro" pageId="31" pageNumber="32">Kilimanjaro Region</collectingRegion>
,
<location id="8EAF4B87E54AFFA4FF64FA12BA4506C8" LSID="urn:lsid:plazi:treatment:03D9AC4AE54BFF9AFCC4FB1DBFD105F2:8EAF4B87E54AFFA4FF64FA12BA4506C8" box="[146,343,1415,1434]" country="Tanzania" latitude="-4.283333" longLatPrecision="1" longitude="38.03333" name="Gonja Forest Reserve" pageId="31" pageNumber="32" stateProvince="Kilimanjaro">Gonja Forest Reserve</location>
,
<geoCoordinate id="EE447B9BE54AFFA4FE96FA12BAD706C8" box="[352,453,1415,1434]" degrees="4.283333" direction="south" orientation="latitude" pageId="31" pageNumber="32" precision="1" value="-4.283333">4.283333</geoCoordinate>
,
<geoCoordinate id="EE447B9BE54AFFA4FE39FA12B93B06C8" box="[463,553,1415,1434]" degrees="38.03333" direction="east" orientation="longitude" pageId="31" pageNumber="32" precision="1" value="38.03333">38.03333</geoCoordinate>
,
<quantity id="4C88B0B9E54AFFA4FDC5FA12B97D06C8" box="[563,623,1415,1434]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="31" pageNumber="32" unit="m" value="550.0">
<elevation id="005DFA6FE54AFFA4FDC5FA12B97D06C8" box="[563,623,1415,1434]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="31" pageNumber="32" unit="m" value="550.0">550 m</elevation>
</quantity>
, lowland forest, leaf litter,
<collectorName id="2685788AE54AFFA4FF1CFA31BA2106E5" box="[234,307,1444,1463]" pageId="31" pageNumber="32">Winkler</collectorName>
,
<date id="FFCE3B9CE54AFFA4FECDFA31BA8806E5" box="[315,410,1444,1463]" pageId="31" pageNumber="32" value="1996-05-16">
<collectingDate id="EF8AC274E54AFFA4FECDFA31BA8806E5" box="[315,410,1444,1463]" pageId="31" pageNumber="32" value="1996-05-16">16.V.1996</collectingDate>
</date>
(
<emphasis id="B904C14EE54AFFA4FE50FA30B92A06E5" box="[422,568,1444,1464]" italics="true" pageId="31" pageNumber="32">
<collectorName id="2685788AE54AFFA4FE50FA30B92A06E5" box="[422,568,1444,1464]" pageId="31" pageNumber="32">H.G. Robertson</collectorName>
</emphasis>
)
</materialsCitation>
;
<materialsCitation id="3B181701E54AFFA4FDBCFA31B927055D" ID-GBIF-Occurrence="2451560265" collectingDate="1995-12-02" collectorName="H. G. Robertson" country="Tanzania" county="Mkomazi Game Reserve" elevation="900" location="Winkler" municipality="Ibaya" pageId="31" pageNumber="32" specimenCount="1" stateProvince="Kilimanjaro">
<collectingRegion id="49B4D3BEE54AFFA4FDBCFA31B9EC06E5" box="[586,766,1444,1463]" country="Tanzania" name="Kilimanjaro" pageId="31" pageNumber="32">Kilimanjaro Region</collectingRegion>
,
<collectingCounty id="62AE65D0E54AFFA4FF64FA57BA6F0687" box="[146,381,1474,1493]" pageId="31" pageNumber="32">Mkomazi Game Reserve</collectingCounty>
, valley behind
<collectingMunicipality id="6BAB8726E54AFFA4FDE4FA57B9540687" box="[530,582,1474,1493]" pageId="31" pageNumber="32">Ibaya</collectingMunicipality>
, 3.966667, 37.80,
<quantity id="4C88B0B9E54AFFA4FF64FA4ABBC306A0" box="[146,209,1503,1522]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="31" pageNumber="32" unit="m" value="900.0">
<elevation id="005DFA6FE54AFFA4FF64FA4ABBC306A0" box="[146,209,1503,1522]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="31" pageNumber="32" unit="m" value="900.0">900 m</elevation>
</quantity>
, closed woodland next to stream in steep-sided valley, leaf litter,
<location id="8EAF4B87E54AFFA4FF3FFA69BA00055D" LSID="urn:lsid:plazi:treatment:03D9AC4AE54BFF9AFCC4FB1DBFD105F2:8EAF4B87E54AFFA4FF3FFA69BA00055D" box="[201,274,1532,1551]" country="Tanzania" county="Mkomazi Game Reserve" municipality="Ibaya" name="Winkler" pageId="31" pageNumber="32" stateProvince="Kilimanjaro">Winkler</location>
,
<date id="FFCE3B9CE54AFFA4FED6FA69BA97055D" box="[288,389,1532,1551]" pageId="31" pageNumber="32" value="1995-12-02">
<collectingDate id="EF8AC274E54AFFA4FED6FA69BA97055D" box="[288,389,1532,1551]" pageId="31" pageNumber="32" value="1995-12-02">2.XII.1995</collectingDate>
</date>
(
<emphasis id="B904C14EE54AFFA4FE60FA68B93F055D" box="[406,557,1532,1552]" italics="true" pageId="31" pageNumber="32">
<collectorName id="2685788AE54AFFA4FE60FA68B93F055D" box="[406,557,1532,1552]" pageId="31" pageNumber="32">H.G. Robertson</collectorName>
</emphasis>
)
</materialsCitation>
;
<materialsCitation id="3B181701E54AFFA4FDB3FA69B93E0535" ID-GBIF-Occurrence="2451560216" collectingDate="1996-05-12" collectorName="Winkler &amp; H. G. Robertson" country="Tanzania" elevation="1600" latitude="-3.883333" location="Maji Kununua" longLatPrecision="1" longitude="37.816666" municipality="Mkomazi Game Reserve" pageId="31" pageNumber="32" specimenCount="1" stateProvince="Kilimanjaro">
<collectingRegion id="49B4D3BEE54AFFA4FDB3FA69B9EC055D" box="[581,766,1532,1551]" country="Tanzania" name="Kilimanjaro" pageId="31" pageNumber="32">Kilimanjaro Region</collectingRegion>
,
<collectingMunicipality id="6BAB8726E54AFFA4FF64F98FBA92057F" box="[146,384,1562,1581]" pageId="31" pageNumber="32">Mkomazi Game Reserve</collectingMunicipality>
, at foot of
<location id="8EAF4B87E54AFFA4FE0AF98FB99B057F" LSID="urn:lsid:plazi:treatment:03D9AC4AE54BFF9AFCC4FB1DBFD105F2:8EAF4B87E54AFFA4FE0AF98FB99B057F" box="[508,649,1562,1581]" country="Tanzania" latitude="-3.883333" longLatPrecision="1" longitude="37.816666" municipality="Mkomazi Game Reserve" name="Maji Kununua" pageId="31" pageNumber="32" stateProvince="Kilimanjaro">Maji Kununua</location>
,
<geoCoordinate id="EE447B9BE54AFFA4FD6FF98FB9EC057F" box="[665,766,1562,1581]" degrees="3.883333" direction="south" orientation="latitude" pageId="31" pageNumber="32" precision="1" value="-3.883333">3.883333</geoCoordinate>
,
<geoCoordinate id="EE447B9BE54AFFA4FF64F9A2BBEA0518" box="[146,248,1591,1610]" degrees="37.816667" direction="east" orientation="longitude" pageId="31" pageNumber="32" precision="1" value="37.816666">37.816667</geoCoordinate>
,
<quantity id="4C88B0B9E54AFFA4FEF1F9A2BA440518" box="[263,342,1591,1610]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="31" pageNumber="32" unit="m" value="1600.0">
<elevation id="005DFA6FE54AFFA4FEF1F9A2BA440518" box="[263,342,1591,1610]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="31" pageNumber="32" unit="m" value="1600.0">1600 m</elevation>
</quantity>
,
<taxonomicName id="4C7066DFE54AFFA4FE90F9A2BAC60518" box="[358,468,1591,1610]" class="Magnoliopsida" family="Combretaceae" genus="Combretum" kingdom="Plantae" order="Myrtales" pageId="31" pageNumber="32" phylum="Tracheophyta" rank="genus">
<emphasis id="B904C14EE54AFFA4FE90F9A2BAC60518" box="[358,468,1591,1610]" italics="true" pageId="31" pageNumber="32">Combretum</emphasis>
</taxonomicName>
thicket at base of valley, leaf litter,
<collectorName id="2685788AE54AFFA4FF3EF9C1BA030535" box="[200,273,1620,1639]" pageId="31" pageNumber="32">Winkler</collectorName>
,
<date id="FFCE3B9CE54AFFA4FEE8F9C1BA6F0535" box="[286,381,1620,1639]" pageId="31" pageNumber="32" value="1996-05-12">
<collectingDate id="EF8AC274E54AFFA4FEE8F9C1BA6F0535" box="[286,381,1620,1639]" pageId="31" pageNumber="32" value="1996-05-12">12.V.1996</collectingDate>
</date>
(
<emphasis id="B904C14EE54AFFA4FE7BF9C0B9360535" box="[397,548,1620,1640]" italics="true" pageId="31" pageNumber="32">
<collectorName id="2685788AE54AFFA4FE7BF9C0B9360535" box="[397,548,1620,1640]" pageId="31" pageNumber="32">H.G. Robertson</collectorName>
</emphasis>
)
</materialsCitation>
;
<materialsCitation id="3B181701E54AFFA4FDCDF9C1B9EF05F0" ID-GBIF-Occurrence="2451560172" collectingDate="2008-02-20" collectingDateMax="2008-02-23" collectingDateMin="2008-02-20" collectorName="Winkler &amp; Hawkes &amp; Y. Mlacha &amp; F. Ninga" country="Tanzania" elevation="870" latitude="-10.11569" location="Rondo Forest Reserve" longLatPrecision="1" longitude="39.18858" pageId="31" pageNumber="32" specimenCount="1" stateProvince="Lindi">
<collectingRegion id="49B4D3BEE54AFFA4FDCDF9C1B9A40535" box="[571,694,1620,1639]" country="Tanzania" name="Lindi" pageId="31" pageNumber="32">Lindi Region</collectingRegion>
,
<location id="8EAF4B87E54AFFA4FD35F9C1BA0905D7" LSID="urn:lsid:plazi:treatment:03D9AC4AE54BFF9AFCC4FB1DBFD105F2:8EAF4B87E54AFFA4FD35F9C1BA0905D7" country="Tanzania" latitude="-10.11569" longLatPrecision="1" longitude="39.18858" name="Rondo Forest Reserve" pageId="31" pageNumber="32" stateProvince="Lindi">Rondo Forest Reserve</location>
,
<geoCoordinate id="EE447B9BE54AFFA4FEDEF9E7BA9F05D7" box="[296,397,1650,1669]" degrees="10.11569" direction="south" orientation="latitude" pageId="31" pageNumber="32" precision="1" value="-10.11569">10.11569</geoCoordinate>
,
<geoCoordinate id="EE447B9BE54AFFA4FE6CF9E7BAE605D7" box="[410,500,1650,1669]" degrees="39.18858" direction="east" orientation="longitude" pageId="31" pageNumber="32" precision="1" value="39.18858">39.18858</geoCoordinate>
,
<quantity id="4C88B0B9E54AFFA4FDF7F9E7B95205D7" box="[513,576,1650,1669]" metricMagnitude="2" metricUnit="m" metricValue="8.7" pageId="31" pageNumber="32" unit="m" value="870.0">
<elevation id="005DFA6FE54AFFA4FDF7F9E7B95205D7" box="[513,576,1650,1669]" metricMagnitude="2" metricUnit="m" metricValue="8.7" pageId="31" pageNumber="32" unit="m" value="870.0">870 m</elevation>
</quantity>
, primary forest, leaf litter,
<collectorName id="2685788AE54AFFA4FF3CF91ABA0105F0" box="[202,275,1679,1698]" pageId="31" pageNumber="32">Winkler</collectorName>
,
<date id="FFCE3B9CE54AFFA4FED4F91ABAB805F0" box="[290,426,1679,1698]" pageId="31" pageNumber="32" value="2008-02-20" valueMax="2008-02-23" valueMin="2008-02-20">
<collectingDate id="EF8AC274E54AFFA4FED4F91ABAB805F0" box="[290,426,1679,1698]" pageId="31" pageNumber="32" value="2008-02-20" valueMax="2008-02-23" valueMin="2008-02-20">20.23.II.2008</collectingDate>
</date>
(
<emphasis id="B904C14EE54AFFA4FE4BF905B9E705F0" box="[445,757,1679,1698]" italics="true" pageId="31" pageNumber="32">
P.
<collectorName id="2685788AE54AFFA4FE21F91AB93005F0" box="[471,546,1679,1698]" pageId="31" pageNumber="32">Hawkes</collectorName>
,
<collectorName id="2685788AE54AFFA4FDC7F905B98605F0" box="[561,660,1679,1698]" pageId="31" pageNumber="32">Y. Mlacha</collectorName>
,
<collectorName id="2685788AE54AFFA4FD54F905B9E705F0" box="[674,757,1680,1698]" pageId="31" pageNumber="32">F. Ninga</collectorName>
</emphasis>
)
</materialsCitation>
;
<materialsCitation id="3B181701E54AFFA4FF64F938BA4C05A8" ID-GBIF-Occurrence="2451560232" collectingDate="2005-08-23" collectingDateMax="2005-08-26" collectingDateMin="2005-08-23" collectorName="Winkler &amp; Hawkes &amp; J. Makwati &amp; R. Mtana" country="Tanzania" elevation="820" latitude="-5.94905" location="Kanga Forest Reserve" longLatPrecision="1" longitude="37.68539" pageId="31" pageNumber="32" specimenCount="1" stateProvince="Morogoro">
<collectingRegion id="49B4D3BEE54AFFA4FF64F938BA2805ED" box="[146,314,1708,1728]" country="Tanzania" name="Morogoro" pageId="31" pageNumber="32">Morogoro Region</collectingRegion>
,
<location id="8EAF4B87E54AFFA4FEB2F938B91E0592" LSID="urn:lsid:plazi:treatment:03D9AC4AE54BFF9AFCC4FB1DBFD105F2:8EAF4B87E54AFFA4FEB2F938B91E0592" box="[324,524,1709,1728]" country="Tanzania" latitude="-5.94905" longLatPrecision="1" longitude="37.68539" name="Kanga Forest Reserve" pageId="31" pageNumber="32" stateProvince="Morogoro">Kanga Forest Reserve</location>
,
<geoCoordinate id="EE447B9BE54AFFA4FDE1F939B96205ED" box="[535,624,1708,1727]" degrees="5.94905" direction="south" orientation="latitude" pageId="31" pageNumber="32" precision="1" value="-5.94905">5.94905</geoCoordinate>
,
<geoCoordinate id="EE447B9BE54AFFA4FD8CF939B9C605ED" box="[634,724,1708,1727]" degrees="37.68539" direction="east" orientation="longitude" pageId="31" pageNumber="32" precision="1" value="37.68539">37.68539</geoCoordinate>
,
<quantity id="4C88B0B9E54AFFA4FD28F938BBB7058F" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="31" pageNumber="32" unit="m" value="820.0">
<elevation id="005DFA6FE54AFFA4FD28F938BBB7058F" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="31" pageNumber="32" unit="m" value="820.0">820 m</elevation>
</quantity>
, primary forest, leaf litter,
<collectorName id="2685788AE54AFFA4FE6AF95FBAF7058F" box="[412,485,1738,1757]" pageId="31" pageNumber="32">Winkler</collectorName>
,
<date id="FFCE3B9CE54AFFA4FE06F95FB99C058F" box="[496,654,1738,1757]" pageId="31" pageNumber="32" value="2005-08-23" valueMax="2005-08-26" valueMin="2005-08-23">
<collectingDate id="EF8AC274E54AFFA4FE06F95FB99C058F" box="[496,654,1738,1757]" pageId="31" pageNumber="32" value="2005-08-23" valueMax="2005-08-26" valueMin="2005-08-23">23.26.VIII.2005</collectingDate>
</date>
(
<emphasis id="B904C14EE54AFFA4FD6AF95FBA4405A8" italics="true" pageId="31" pageNumber="32">
P.
<collectorName id="2685788AE54AFFA4FD45F95FB9ED058F" box="[691,767,1738,1757]" pageId="31" pageNumber="32">Hawkes</collectorName>
,
<collectorName id="2685788AE54AFFA4FF64F97DBBE505A8" box="[146,247,1767,1786]" pageId="31" pageNumber="32">J. Makwati</collectorName>
,
<collectorName id="2685788AE54AFFA4FEF6F97DBA4405A8" box="[256,342,1768,1786]" pageId="31" pageNumber="32">R. Mtana</collectorName>
</emphasis>
)
</materialsCitation>
;
<materialsCitation id="3B181701E54AFFA4FE9EF972B95A0467" ID-GBIF-Occurrence="2451560274" collectingDate="2007-10-12" collectingDateMax="2007-10-15" collectingDateMin="2007-10-12" collectorName="Winkler &amp; Hawkes &amp; M. Bhoke &amp; U. Richard" country="Tanzania" elevation="700" latitude="-6.89388" location="Mkungwe Forest Reserve" longLatPrecision="1" longitude="37.90414" pageId="31" pageNumber="32" specimenCount="1" stateProvince="Morogoro">
<collectingRegion id="49B4D3BEE54AFFA4FE9EF972B91C05A8" box="[360,526,1767,1786]" country="Tanzania" name="Morogoro" pageId="31" pageNumber="32">Morogoro Region</collectingRegion>
,
<location id="8EAF4B87E54AFFA4FDE1F972B9EC05A8" LSID="urn:lsid:plazi:treatment:03D9AC4AE54BFF9AFCC4FB1DBFD105F2:8EAF4B87E54AFFA4FDE1F972B9EC05A8" box="[535,766,1767,1786]" country="Tanzania" latitude="-6.89388" longLatPrecision="1" longitude="37.90414" name="Mkungwe Forest Reserve" pageId="31" pageNumber="32" stateProvince="Morogoro">Mkungwe Forest Reserve</location>
,
<geoCoordinate id="EE447B9BE54AFFA4FF64F891BBF90445" box="[146,235,1796,1815]" degrees="6.89388" direction="south" orientation="latitude" pageId="31" pageNumber="32" precision="1" value="-6.89388">6.89388</geoCoordinate>
,
<geoCoordinate id="EE447B9BE54AFFA4FF03F890BA5D044A" box="[245,335,1797,1816]" degrees="37.90414" direction="east" orientation="longitude" pageId="31" pageNumber="32" precision="1" value="37.90414">37.90414</geoCoordinate>
,
<quantity id="4C88B0B9E54AFFA4FEAFF890BA87044A" box="[345,405,1797,1816]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="31" pageNumber="32" unit="m" value="700.0">
<elevation id="005DFA6FE54AFFA4FEAFF890BA87044A" box="[345,405,1797,1816]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="31" pageNumber="32" unit="m" value="700.0">700 m</elevation>
</quantity>
, primary forest, leaf litter,
<collectorName id="2685788AE54AFFA4FD71F891B9C20445" box="[647,720,1796,1815]" pageId="31" pageNumber="32">Winkler</collectorName>
,
<date id="FFCE3B9CE54AFFA4FD2CF890BBE70467" pageId="31" pageNumber="32" value="2007-10-12" valueMax="2007-10-15" valueMin="2007-10-12">
<collectingDate id="EF8AC274E54AFFA4FD2CF890BBE70467" pageId="31" pageNumber="32" value="2007-10-12" valueMax="2007-10-15" valueMin="2007-10-12">12. 15.X.2007</collectingDate>
</date>
(
<emphasis id="B904C14EE54AFFA4FEF5F8B7B9520467" box="[259,576,1826,1845]" italics="true" pageId="31" pageNumber="32">
P.
<collectorName id="2685788AE54AFFA4FEECF8B7BA770467" box="[282,357,1826,1845]" pageId="31" pageNumber="32">Hawkes</collectorName>
,
<collectorName id="2685788AE54AFFA4FE87F8B7BADD0467" box="[369,463,1826,1845]" pageId="31" pageNumber="32">M. Bhoke</collectorName>
,
<collectorName id="2685788AE54AFFA4FE2CF8B7B9520467" box="[474,576,1826,1845]" pageId="31" pageNumber="32">U. Richard</collectorName>
</emphasis>
)
</materialsCitation>
;
<materialsCitation id="3B181701E54AFFA4FDA3F8B7B9EF043D" ID-GBIF-Occurrence="2451560189" collectingDate="2007-10-17" collectingDateMax="2007-10-20" collectingDateMin="2007-10-17" collectorName="Winkler &amp; Hawkes &amp; M. Bhoke &amp; U. Richard" country="Tanzania" elevation="1150" latitude="-8.94497" location="Sali Forest Reserve" longLatPrecision="5" longitude="36.6726" pageId="31" pageNumber="32" specimenCount="1" stateProvince="Morogoro">
<collectingRegion id="49B4D3BEE54AFFA4FDA3F8B7B9EC0467" box="[597,766,1826,1845]" country="Tanzania" name="Morogoro" pageId="31" pageNumber="32">Morogoro Region</collectingRegion>
,
<location id="8EAF4B87E54AFFA4FF64F8AABA2D0400" LSID="urn:lsid:plazi:treatment:03D9AC4AE54BFF9AFCC4FB1DBFD105F2:8EAF4B87E54AFFA4FF64F8AABA2D0400" box="[146,319,1855,1874]" country="Tanzania" latitude="-8.94497" longLatPrecision="5" longitude="36.6726" name="Sali Forest Reserve" pageId="31" pageNumber="32" stateProvince="Morogoro">Sali Forest Reserve</location>
,
<geoCoordinate id="EE447B9BE54AFFA4FEBEF8AABAB30400" box="[328,417,1855,1874]" degrees="8.94497" direction="south" orientation="latitude" pageId="31" pageNumber="32" precision="1" value="-8.94497">8.94497</geoCoordinate>
,
<geoCoordinate id="EE447B9BE54AFFA4FE5DF8AABAEB0400" box="[427,505,1855,1874]" degrees="36.6726" direction="east" orientation="longitude" pageId="31" pageNumber="32" precision="5" value="36.6726">36.6726</geoCoordinate>
,
<quantity id="4C88B0B9E54AFFA4FDF4F8AAB9580400" box="[514,586,1855,1874]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="31" pageNumber="32" unit="m" value="1150.0">
<elevation id="005DFA6FE54AFFA4FDF4F8AAB9580400" box="[514,586,1855,1874]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="31" pageNumber="32" unit="m" value="1150.0">1150 m</elevation>
</quantity>
, primary forest, leaf litter,
<collectorName id="2685788AE54AFFA4FF31F8C9BA02043D" box="[199,272,1884,1903]" pageId="31" pageNumber="32">Winkler</collectorName>
,
<date id="FFCE3B9CE54AFFA4FEEAF8C8BAB40422" box="[284,422,1885,1904]" pageId="31" pageNumber="32" value="2007-10-17" valueMax="2007-10-20" valueMin="2007-10-17">
<collectingDate id="EF8AC274E54AFFA4FEEAF8C8BAB40422" box="[284,422,1885,1904]" pageId="31" pageNumber="32" value="2007-10-17" valueMax="2007-10-20" valueMin="2007-10-17">17.20.X.2007</collectingDate>
</date>
(
<emphasis id="B904C14EE54AFFA4FE43F8C8B9E6043D" box="[437,756,1884,1903]" italics="true" pageId="31" pageNumber="32">
P.
<collectorName id="2685788AE54AFFA4FE3BF8C9B90A043D" box="[461,536,1884,1903]" pageId="31" pageNumber="32">Hawkes</collectorName>
,
<collectorName id="2685788AE54AFFA4FDD2F8C8B990043D" box="[548,642,1884,1903]" pageId="31" pageNumber="32">M. Bhoke</collectorName>
,
<collectorName id="2685788AE54AFFA4FD78F8C8B9E6043D" box="[654,756,1884,1903]" pageId="31" pageNumber="32">U. Richard</collectorName>
</emphasis>
)
</materialsCitation>
;
<materialsCitation id="3B181701E54AFFA4FF64F8EFBF9D0394" ID-GBIF-Occurrence="2451560263" collectingDate="2005-08-27" collectingDateMax="2005-08-30" collectingDateMin="2005-08-27" collectorName="Winkler, F &amp; Hawkes &amp; J. Makwati &amp; R. Mtana" country="Uganda" elevation="1015" latitude="-5.57934" location="Kilindi Forest Reserve" longLatPrecision="1" longitude="37.57971" pageId="31" pageNumber="32" specimenCount="1" stateProvince="Tanga">
<collectingRegion id="49B4D3BEE54AFFA4FF64F8EFBA0004DF" box="[146,274,1914,1933]" country="Tanzania" name="Tanga" pageId="31" pageNumber="32">Tanga Region</collectingRegion>
,
<location id="8EAF4B87E54AFFA4FEEDF8EFBAF404DF" LSID="urn:lsid:plazi:treatment:03D9AC4AE54BFF9AFCC4FB1DBFD105F2:8EAF4B87E54AFFA4FEEDF8EFBAF404DF" box="[283,486,1914,1933]" country="Uganda" latitude="-5.57934" longLatPrecision="1" longitude="37.57971" name="Kilindi Forest Reserve" pageId="31" pageNumber="32" stateProvince="Tanga">Kilindi Forest Reserve</location>
,
<geoCoordinate id="EE447B9BE54AFFA4FE19F8EFB95B04DF" box="[495,585,1914,1933]" degrees="5.57934" direction="south" orientation="latitude" pageId="31" pageNumber="32" precision="1" value="-5.57934">5.57934</geoCoordinate>
,
<geoCoordinate id="EE447B9BE54AFFA4FDA4F8EFB9BE04DF" box="[594,684,1914,1933]" degrees="37.57971" direction="east" orientation="longitude" pageId="31" pageNumber="32" precision="1" value="37.57971">37.57971</geoCoordinate>
,
<quantity id="4C88B0B9E54AFFA4FD40F8EFB9EC04DF" box="[694,766,1914,1933]" metricMagnitude="3" metricUnit="m" metricValue="1.015" pageId="31" pageNumber="32" unit="m" value="1015.0">
<elevation id="005DFA6FE54AFFA4FD40F8EFB9EC04DF" box="[694,766,1914,1933]" metricMagnitude="3" metricUnit="m" metricValue="1.015" pageId="31" pageNumber="32" unit="m" value="1015.0">1015 m</elevation>
</quantity>
, primary forest, leaf litter,
<collectorName id="2685788AE54AFFA4FBE3FF00BF6703FB" box="[1045,1141,149,169]" pageId="31" pageNumber="32">Winkler, F</collectorName>
21,
<date id="FFCE3B9CE54AFFA4FB62FF00BE2003FA" box="[1172,1330,149,168]" pageId="31" pageNumber="32" value="2005-08-27" valueMax="2005-08-30" valueMin="2005-08-27">
<collectingDate id="EF8AC274E54AFFA4FB62FF00BE2003FA" box="[1172,1330,149,168]" pageId="31" pageNumber="32" value="2005-08-27" valueMax="2005-08-30" valueMin="2005-08-27">27.30.VIII.2005</collectingDate>
</date>
(
<emphasis id="B904C14EE54AFFA4FAC8FF03BF000397" italics="true" pageId="31" pageNumber="32">
P.
<collectorName id="2685788AE54AFFA4FAA5FF00BE8D03FA" box="[1363,1439,149,168]" pageId="31" pageNumber="32">Hawkes</collectorName>
,
<collectorName id="2685788AE54AFFA4FCC4FF26B8B30394" box="[818,929,179,198]" pageId="31" pageNumber="32">J. Makwati</collectorName>
,
<collectorName id="2685788AE54AFFA4FC45FF26BF000397" box="[947,1042,179,197]" pageId="31" pageNumber="32">R. Mtana</collectorName>
</emphasis>
);
<collectingCountry id="F3675DCCE54AFFA4FBDBFF26BF9D0394" box="[1069,1167,179,198]" name="Uganda" pageId="31" pageNumber="32">UGANDA</collectingCountry>
</materialsCitation>
:
<materialsCitation id="3B181701E54AFFA4FB57FF26BF400253" ID-GBIF-Occurrence="2451560170" collectingDate="2012-08-17" collectorName="J. Longino" country="Uganda" elevation="690" latitude="0.85617" location="Semliki National Park" longLatPrecision="1" longitude="30.16583" municipality="Rwankasenyi" pageId="31" pageNumber="32" specimenCount="1" stateProvince="Bundibugyo">
<collectingRegion id="49B4D3BEE54AFFA4FB57FF26BE030394" box="[1185,1297,179,198]" country="Uganda" name="Bundibugyo" pageId="31" pageNumber="32">Bundibugyo</collectingRegion>
,
<collectingMunicipality id="6BAB8726E54AFFA4FAD4FF26BE8C0394" box="[1314,1438,179,198]" pageId="31" pageNumber="32">Rwankasenyi</collectingMunicipality>
,
<location id="8EAF4B87E54AFFA4FCC4FF45B8E903B1" LSID="urn:lsid:plazi:treatment:03D9AC4AE54BFF9AFCC4FB1DBFD105F2:8EAF4B87E54AFFA4FCC4FF45B8E903B1" box="[818,1019,208,227]" country="Uganda" latitude="0.85617" longLatPrecision="1" longitude="30.16583" municipality="Rwankasenyi" name="Semliki National Park" pageId="31" pageNumber="32" stateProvince="Bundibugyo">Semliki National Park</location>
,
<geoCoordinate id="EE447B9BE54AFFA4FBF5FF45BF4303B1" box="[1027,1105,208,227]" degrees="0.85617" direction="north" orientation="latitude" pageId="31" pageNumber="32" precision="1" value="0.85617">0.85617</geoCoordinate>
,
<geoCoordinate id="EE447B9BE54AFFA4FBAFFF45BFA103B1" box="[1113,1203,208,227]" degrees="30.16583" direction="east" orientation="longitude" pageId="31" pageNumber="32" precision="1" value="30.16583">30.16583</geoCoordinate>
,
<quantity id="4C88B0B9E54AFFA4FB4DFF45BFE403B6" box="[1211,1270,208,228]" metricMagnitude="2" metricUnit="m" metricValue="6.9" pageId="31" pageNumber="32" unit="m" value="690.0">
<elevation id="005DFA6FE54AFFA4FB4DFF45BFE403B6" box="[1211,1270,208,228]" metricMagnitude="2" metricUnit="m" metricValue="6.9" pageId="31" pageNumber="32" unit="m" value="690.0">690 m</elevation>
</quantity>
, swamp forest, leaf litter,
<date id="FFCE3B9CE54AFFA4FC93FF7BB8C90253" box="[869,987,238,257]" pageId="31" pageNumber="32" value="2012-08-17">
<collectingDate id="EF8AC274E54AFFA4FC93FF7BB8C90253" box="[869,987,238,257]" pageId="31" pageNumber="32" value="2012-08-17">17.VIII.2012</collectingDate>
</date>
(
<emphasis id="B904C14EE54AFFA4FC1FFF7BBF5B0252" box="[1001,1097,238,256]" italics="true" pageId="31" pageNumber="32">
<collectorName id="2685788AE54AFFA4FC1FFF7BBF5B0252" box="[1001,1097,238,256]" pageId="31" pageNumber="32">J. Longino</collectorName>
</emphasis>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36A4ED7E54AFFA4FCC4FEB8BF0F011A" pageId="31" pageNumber="32" type="diagnosis">
<paragraph id="8BCF1D5CE54AFFA4FCC4FEB8B89E0212" blockId="31.[818,1442,301,585]" box="[818,908,301,320]" pageId="31" pageNumber="32">
<heading id="D087AA30E54AFFA4FCC4FEB8B89E0212" bold="true" box="[818,908,301,320]" fontSize="8" level="3" pageId="31" pageNumber="32" reason="6">
<emphasis id="B904C14EE54AFFA4FCC4FEB8B89E0212" bold="true" box="[818,908,301,320]" pageId="31" pageNumber="32">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF1D5CE54AFFA4FCC4FEDEBF0F011A" blockId="31.[818,1442,301,585]" pageId="31" pageNumber="32">
The following character combination distinguishes
<taxonomicName id="4C7066DFE54AFFA4FAC5FEDEB87C0229" authorityName="Hita-Garcia &amp; Lieberman &amp; Audisio &amp; Liu &amp; Economo" authorityYear="2019" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="athene">
<emphasis id="B904C14EE54AFFA4FAC5FEDEB87C0229" italics="true" pageId="31" pageNumber="32">Discothyrea athene</emphasis>
</taxonomicName>
from the remainder of the complex: standing pilosity absent from mesosoma and abdominal terga; propodeum denticulate; eyes present, relatively large (OI 59); in dorsal view mesosoma relatively broad and robust (DMI 5967, DMI2 92100); mesotibia without apicoventral spur; petiole attenuated dorsally (DPeI 300500, LPeI 286500); sculpture distinct, declivitous face of propodeum foveolate; generally smaller species (WL 0.300.53); color usually orange, variably infuscated.
</paragraph>
</subSubSection>
<subSubSection id="C36A4ED7E54AFF9AFCC4FDEBBFFB0394" lastPageId="33" lastPageNumber="34" pageId="31" pageNumber="32" type="description">
<paragraph id="8BCF1D5CE54AFFA4FCC4FDEBBFD201C2" blockId="31.[818,1442,637,833]" box="[818,1216,637,657]" pageId="31" pageNumber="32">
<heading id="D087AA30E54AFFA4FCC4FDEBBFD201C2" bold="true" box="[818,1216,637,657]" fontSize="8" level="3" pageId="31" pageNumber="32" reason="6">
<emphasis id="B904C14EE54AFFA4FCC4FDEBBFD201C2" bold="true" box="[818,1216,637,657]" pageId="31" pageNumber="32">
Worker Measurements and Indices (
<emphasis id="B904C14EE54AFFA4FB8BFDEBBF9B01C3" bold="true" box="[1149,1161,638,657]" italics="true" pageId="31" pageNumber="32">n</emphasis>
= 17)
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF1D5CE54AFFA4FCC4FD0EBF830012" blockId="31.[818,1442,637,833]" pageId="31" pageNumber="32">
EL 0.030.04; HL 0.390.49; HW 0.330.42; SL 0.190.26;
<collectingCountry id="F3675DCCE54AFFA4FA72FD0EBEB001FC" box="[1412,1442,667,686]" name="Philippines" pageId="31" pageNumber="32">PH</collectingCountry>
0.190.27; PW 0.230.32; PrH 0.230.30; DML 0.230.34; WL 0.380.53; HFL 0.210.30; PeL 0.040.07; PeW 0.150.21; PeH 0.150.20; LT3 0.270.39; LT40.250.33; OI 59; CI 8089; SI 4853; LMI 4751; DMI 5967; DMI2 92100; ASI 85103; HFI 5360; DPeI 300500; LPeI 286500.
</paragraph>
<paragraph id="8BCF1D5CE54AFFA4FCC4FCEDB8F500D9" blockId="31.[818,1443,888,1934]" box="[818,999,888,907]" pageId="31" pageNumber="32">
<heading id="D087AA30E54AFFA4FCC4FCEDB8F500D9" bold="true" box="[818,999,888,907]" fontSize="8" level="3" pageId="31" pageNumber="32" reason="6">
<emphasis id="B904C14EE54AFFA4FCC4FCEDB8F500D9" bold="true" box="[818,999,888,907]" pageId="31" pageNumber="32">Worker Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF1D5CE54AFFA4FCC4FC00B8C1057C" blockId="31.[818,1443,888,1934]" pageId="31" pageNumber="32">
<emphasis id="B904C14EE54AFFA4FCC4FC00B87600FA" box="[818,868,917,936]" italics="true" pageId="31" pageNumber="32">Head</emphasis>
fairly broad (CI 8089) and round, posterior head margin gently convex; posterodorsal corners of head broadly rounded; in frontal view, sides of head convex and tapering somewhat anteriorly; eyes present, relatively large (OI 59) and round, comprising several ommatidia, placed about a third of the way between anterolateral corner of gena and posterior head margin; eyes visible in frontal view; frontal lamella broadly triangular in profile, apex rounded; lamella with translucent basal region, variably developed, from an indistinct patch to an elliptical and well-defined fenestra; medial clypeus convex, lateral clypeus curving gently between antennal sockets and anterolateral corners of head, bearing short, curved setae.
<emphasis id="B904C14EE54AFFA4FC44FB4DBF1207B9" box="[946,1024,1240,1259]" italics="true" pageId="31" pageNumber="32">Antenna</emphasis>
with short scape (SI 4853), scape strongly incrassate, slightly bent; pedicel subglobose, broader than long; true antennomere count nine; apparent antennomere count seven to nine; flagellomeres basad apical club highly compressed, taken together only about as long as apical club.
<emphasis id="B904C14EE54AFFA4FB8DFAD8BFE00632" box="[1147,1266,1357,1376]" italics="true" pageId="31" pageNumber="32">Ventral head</emphasis>
with postoccipital ridge weakly developed, without anteromedian carina; hypostoma medially rounded, arms somewhat narrowed, similar in width across their length; palpal formula not examined.
<emphasis id="B904C14EE54AFFA4FB4EFA30BE0206EA" box="[1208,1296,1445,1464]" italics="true" pageId="31" pageNumber="32">Mandible</emphasis>
edentate except for small, sharp prebasal denticle; basal angle broadly rounded; longitudinal carina on ectal face confluent with masticatory margin for most of its length, leaving small depressed triangular area anterad prebasal denticle.
</paragraph>
<paragraph id="8BCF1D5CE54AFFA4FCA4F9ACBE950401" blockId="31.[818,1443,888,1934]" pageId="31" pageNumber="32">
<emphasis id="B904C14EE54AFFA4FCA4F9ACB8A60519" box="[850,948,1593,1611]" italics="true" pageId="31" pageNumber="32">Mesosoma</emphasis>
gently convex, pronotum slightly lower than propodeum. In dorsal view mesosoma broad, robust (DMI 5967; DMI2 92100), somewhat narrowed posteriorly, pronotum wider than propodeum; pronotal humeri moderately rounded; posterior propodeal margin concave, distinctly concave between dentae; posterodorsal corners of propodeum angulate to dentate, teeth blunt but distinct; declivitous face of propodeum distinctly concave in profile and oblique posterior view; propodeal spiracle relatively large, subcircular, directed posterolaterally, often conspicuous due to shiny, unsculptured spiracular opening; propodeal lobes short, rounded.
</paragraph>
<paragraph id="8BCF1D5CE54AFF9AFCA4F8CBBA070196" blockId="31.[818,1443,888,1934]" lastBlockId="33.[146,772,615,708]" lastPageId="33" lastPageNumber="34" pageId="31" pageNumber="32">
<emphasis id="B904C14EE54AFFA4FCA4F8CBB86F0422" box="[850,893,1886,1904]" italics="true" pageId="31" pageNumber="32">Legs</emphasis>
short to moderate in length (HFI 5360) and slender; mesotibia without apicoventral spur, with small but distinct
<emphasis id="B904C14EE574FF9AFF64FDF2BBCB012A" bold="true" box="[146,217,615,632]" pageId="33" pageNumber="34">Model 4.</emphasis>
3D surface model of
<taxonomicName id="4C7066DFE574FF9AFE67FDF2BAF3012A" authority="Hita-Garcia &amp; Lieberman &amp; Audisio &amp; Liu &amp; Economo, 2019" authorityName="Hita-Garcia &amp; Lieberman &amp; Audisio &amp; Liu &amp; Economo" authorityYear="2019" box="[401,481,615,632]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="athene" status="sp. nov.">
<emphasis id="B904C14EE574FF9AFE67FDF2BAF3012A" box="[401,481,615,632]" italics="true" pageId="33" pageNumber="34">D. athene</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2377C35E574FF9AFE11FDF2B907012A" box="[487,533,615,632]" pageId="33" pageNumber="34" rank="species">sp. n.</taxonomicNameLabel>
<typeStatus id="54CBA3FEE574FF9AFDEDFDF2B971012A" box="[539,611,615,632]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
(CASENT0764088). An interactive version of this model is available in the HTML version of this article online and at https://sketchfab.com/3d-models/8486543f757b484d895 2a7dd01c8d733.
</paragraph>
<caption id="DF0F4DD4E575FF9BFF64F963BAE4043F" ID-DOI="http://doi.org/10.5281/zenodo.3542186" ID-Zenodo-Dep="3542186" httpUri="https://zenodo.org/record/3542186/files/figure.png" pageId="32" pageNumber="33" startId="32.[146,173,1782,1799]" targetBox="[202,1386,147,1747]" targetPageId="32">
<paragraph id="8BCF1D5CE575FF9BFF64F963BAE4043F" blockId="32.[146,1444,1782,1901]" pageId="32" pageNumber="33">
<emphasis id="B904C14EE575FF9BFF64F963BBDC0455" bold="true" box="[146,206,1782,1799]" pageId="32" pageNumber="33">Fig. 26.</emphasis>
Still images from shaded surface display volume renderings of
<taxonomicName id="4C7066DFE575FF9BFD15F963B8200455" authority="Hita-Garcia &amp; Lieberman &amp; Audisio &amp; Liu &amp; Economo, 2019" authorityName="Hita-Garcia &amp; Lieberman &amp; Audisio &amp; Liu &amp; Economo" authorityYear="2019" box="[739,818,1782,1799]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="athene" status="sp. nov.">
<emphasis id="B904C14EE575FF9BFD15F963B8200455" box="[739,818,1782,1799]" italics="true" pageId="32" pageNumber="33">D. athene</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2377C35E575FF9BFCC1F963B8770455" box="[823,869,1782,1799]" pageId="32" pageNumber="33" rank="species">sp. n.</taxonomicNameLabel>
holotype (CASENT0764088) showing virtually segmented body parts. (A) Body in dorsal view, (B) body in profile, (C) head in full-face view, (D) head in dorsal view, (E) head in anterodorsal view, (F) head in anterior view, (G) head in profile, (H) head in ventral view, (I) posterior propodeum in posterior view, (J) petiole in profile, (K) petiole and gaster in profile, (L) petiole in dorsal view, (M) petiole in anterior view, (N) petiole oblique anterior view, (O) gaster in dorsal view, (P) petiole in ventral view, (Q) abdominal sternite 3 in ventral view, (R) mesotibia and mesotarsus in anterior view.
</paragraph>
</caption>
<paragraph id="8BCF1D5CE574FF9AFF64FD60BAA50077" blockId="33.[146,771,757,1950]" pageId="33" pageNumber="34">seta inserted in apical pit; mesobasitarsus short, about as long as tarsomeres II-IV taken together.</paragraph>
<paragraph id="8BCF1D5CE574FF9AFF44FCA5BA64073A" blockId="33.[146,771,757,1950]" pageId="33" pageNumber="34">
<emphasis id="B904C14EE574FF9AFF44FCA5BA3F0011" box="[178,301,816,835]" italics="true" pageId="33" pageNumber="34">Petiolar node</emphasis>
moderately to strongly attenuated dorsally, about 3.7 times higher than broad (LPeI 286500). In profile, anterior face of node convex, apex broadly peaked, posterior face sloping posteroventrally. In dorsal view, petiole about 35 times wider than long (DPeI 300500); sides divergent posteriorly, posterior margin slightly concave, anterior margin nearly straight. In anterior view, petiolar outline very wide, pentagonal, edges and apex slightly rounded but distinct; in oblique anterior view, anterior face flat; in ventral view approximately rectangular, sides diverging posteriorly; subpetiolar process large, lobate, apex rounded; petiolar spiracles reniform in ventral view.
</paragraph>
<paragraph id="8BCF1D5CE574FF9AFF44FBE7BA7E06DF" blockId="33.[146,771,757,1950]" pageId="33" pageNumber="34">
<emphasis id="B904C14EE574FF9AFF44FBE7BA9307D7" box="[178,385,1138,1157]" italics="true" pageId="33" pageNumber="34">Abdominal segment 3</emphasis>
roughly campaniform, tergite prolonged anteriorly pasted anterior sternal margin, tergite widest just anterad end of segment, sternite sloping to convex in profile;
<collectingCountry id="F3675DCCE574FF9AFD86FB38B99F0792" box="[624,653,1197,1216]" name="American Samoa" pageId="33" pageNumber="34">AS</collectingCountry>
3 with scarce trace of medial carina; prora present as a very fine carina, shallowly concave in ventral view; AT4 about 0.85 the length of AT3 to slightly longer, usually approximately equal in length (ASI 85103); AT4 almost perfectly hemidemispherical;
<collectingCountry id="F3675DCCE574FF9AFE2EFAB7BAE10667" box="[472,499,1314,1333]" name="American Samoa" pageId="33" pageNumber="34">AS</collectingCountry>
4 with anterior lip short but fairly broad, covering more than the one-third of the width of
<collectingCountry id="F3675DCCE574FF9AFD21FAD5B9E70601" box="[727,757,1344,1363]" name="American Samoa" pageId="33" pageNumber="34">AS</collectingCountry>
3, anterior margin straight; successive abdominal segments short, telescopic, often concealed.
</paragraph>
<paragraph id="8BCF1D5CE574FF9AFF44FA0DBBE605E1" blockId="33.[146,771,757,1950]" pageId="33" pageNumber="34">
<emphasis id="B904C14EE574FF9AFF44FA0DBA1E06F9" box="[178,268,1432,1451]" italics="true" pageId="33" pageNumber="34">Sculpture</emphasis>
on head, dorsal mesosoma, and petiole regularly punctate-reticulate or foveolate-reticulate; on ventral head surface and lateral mesosoma becoming foveolate to punctate; lateral mesosoma with interspaces of foveolate forming rugulae, especially on lower surfaces; declivitous face of propodeum shallowly foveolate-reticulate to laterally rugulose; mandibles with fairly coarse piligerous punctae; frontal lamella and medial clypeus roughly granulate; AT3 shallowly punctulate-reticulate; AT4 with minute, dense piligerous punctulae, clearly shinier than AT3.
</paragraph>
<paragraph id="8BCF1D5CE574FF9AFF44F928BA8E04CF" blockId="33.[146,771,757,1950]" pageId="33" pageNumber="34">
<emphasis id="B904C14EE574FF9AFF44F928BBEF0582" box="[178,253,1725,1744]" italics="true" pageId="33" pageNumber="34">Setation</emphasis>
mostly consisting of very fine, appressed white pubescence, similarly distributed on dorsal surfaces, dilute to absent on lateral surfaces of head and mesosoma; abdominal sternite 3 with longer, curved decumbent white setae; abdominal segments five through seven with longer, flexuous standing setae; appendages with only very fine, appressed white pubescence; ectal face of mandible with curved, appressed to decumbent setae; row of straight, stout setae on masticatory margin.
</paragraph>
<paragraph id="8BCF1D5CE574FF9AFCA4FF00BFFB0394" blockId="33.[818,1442,149,198]" pageId="33" pageNumber="34">
<emphasis id="B904C14EE574FF9AFCA4FF00B89503FA" box="[850,903,149,168]" italics="true" pageId="33" pageNumber="34">Color</emphasis>
unicolorous luteous-orange to brown, usually brownish orange, upper surfaces often slightly infuscated.
</paragraph>
</subSubSection>
<subSubSection id="C36A4ED7E574FF9AFCC4FF7BB8AD02BE" pageId="33" pageNumber="34" type="etymology">
<paragraph id="8BCF1D5CE574FF9AFCC4FF7BB8870253" blockId="33.[818,1443,238,492]" box="[818,917,238,257]" pageId="33" pageNumber="34">
<heading id="D087AA30E574FF9AFCC4FF7BB8870253" bold="true" box="[818,917,238,257]" fontSize="8" level="3" pageId="33" pageNumber="34" reason="6">
<emphasis id="B904C14EE574FF9AFCC4FF7BB8870253" bold="true" box="[818,917,238,257]" pageId="33" pageNumber="34">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF1D5CE574FF9AFCC4FE9EB8AD02BE" blockId="33.[818,1443,238,492]" pageId="33" pageNumber="34">
The Greek goddess Athene was fabled to have competed against a mortal woman named Arachne, whom the goddess punished for her hubris by transforming her into the first spider. Although the trophic biology of
<taxonomicName id="4C7066DFE574FF9AFC6FFEF6B8E60224" authorityName="Hita-Garcia &amp; Lieberman &amp; Audisio &amp; Liu &amp; Economo" authorityYear="2019" box="[921,1012,355,374]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="athene">
<emphasis id="B904C14EE574FF9AFC6FFEF6B8E60224" box="[921,1012,355,374]" italics="true" pageId="33" pageNumber="34">D. athene</emphasis>
</taxonomicName>
specifically is unknown, the species is named in reference to the known arachnophagic habits of its congeners. Additionally, Athene is archetypically portrayed bearing a helmet and a spear, reflecting the notably thick cuticle and prominent sting of
<taxonomicName id="4C7066DFE574FF9AFCBDFE4DB8A802B9" authorityName="Roger" authorityYear="1863" box="[843,954,472,491]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
<emphasis id="B904C14EE574FF9AFCBDFE4DB8A802B9" box="[843,954,472,491]" italics="true" pageId="33" pageNumber="34">Discothyrea</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C36A4ED7E574FF9AFCC4FD86BEB20184" pageId="33" pageNumber="34" type="distribution">
<paragraph id="8BCF1D5CE574FF9AFCC4FD86BF0A0174" blockId="33.[818,1442,531,726]" box="[818,1048,531,550]" pageId="33" pageNumber="34">
<heading id="D087AA30E574FF9AFCC4FD86BF0A0174" bold="true" box="[818,1048,531,550]" fontSize="8" level="3" pageId="33" pageNumber="34" reason="6">
<emphasis id="B904C14EE574FF9AFCC4FD86BF0A0174" bold="true" box="[818,1048,531,550]" pageId="33" pageNumber="34">Distribution and Biology</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF1D5CE574FF9AFCC4FDA5BEB20184" blockId="33.[818,1442,531,726]" pageId="33" pageNumber="34">
<taxonomicName id="4C7066DFE574FF9AFCC4FDA5B8F60111" authorityName="Hita-Garcia &amp; Lieberman &amp; Audisio &amp; Liu &amp; Economo" authorityYear="2019" box="[818,996,560,579]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="athene">
<emphasis id="B904C14EE574FF9AFCC4FDA5B8F60111" box="[818,996,560,579]" italics="true" pageId="33" pageNumber="34">Discothyrea athene</emphasis>
</taxonomicName>
is known from various localities in East Africa including
<collectingCountry id="F3675DCCE574FF9AFC64FDD8B8F30132" box="[914,993,589,608]" name="Rwanda" pageId="33" pageNumber="34">Rwanda</collectingCountry>
,
<collectingCountry id="F3675DCCE574FF9AFC06FDD8BF2B0132" box="[1008,1081,589,608]" name="Uganda" pageId="33" pageNumber="34">Uganda</collectingCountry>
,
<collectingCountry id="F3675DCCE574FF9AFBB1FDDBBF910133" box="[1095,1155,590,609]" name="Kenya" pageId="33" pageNumber="34">Kenya</collectingCountry>
,
<collectingCountry id="F3675DCCE574FF9AFB67FDD8BFF40132" box="[1169,1254,589,608]" name="Tanzania" pageId="33" pageNumber="34">Tanzania</collectingCountry>
, and
<collectingCountry id="F3675DCCE574FF9AFAD4FDD8BE8C0132" box="[1314,1438,589,608]" name="Mozambique" pageId="33" pageNumber="34">Mozambique</collectingCountry>
, where it occurs in forest and woodland, usually but not always below
<quantity id="4C88B0B9E574FF9AFC85FD1DB8AC01CE" box="[883,958,648,668]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="33" pageNumber="34" unit="m" value="1000.0">1000 m</quantity>
(
<figureCitation id="134B01D9E574FF9AFC38FD1DBF0501C9" box="[974,1047,648,667]" captionStart="Fig" captionStartId="5.[146,173,1283,1300]" captionTargetBox="[153,1435,148,1247]" captionTargetId="figure@5.[152,1435,147,1248]" captionTargetPageId="5" captionText="Fig. 4. Maps showing the known distribution ranges of Afrotropical Discothyrea species." figureDoi="http://doi.org/10.5281/zenodo.3542140" httpUri="https://zenodo.org/record/3542140/files/figure.png" pageId="33" pageNumber="34">Fig. 4D</figureCitation>
). It has been collected in riparian forest, swamp forest, and near streams throughout its range, suggesting a possible preference for mesic habitats. The species lives in leaf litter.
</paragraph>
</subSubSection>
<subSubSection id="C36A4ED7E574FF9AFCC4FC95BFD105F2" pageId="33" pageNumber="34" type="discussion">
<paragraph id="8BCF1D5CE574FF9AFCC4FC95B8840041" blockId="33.[818,1443,768,1345]" box="[818,918,768,787]" pageId="33" pageNumber="34">
<heading id="D087AA30E574FF9AFCC4FC95B8840041" bold="true" box="[818,918,768,787]" fontSize="8" level="3" pageId="33" pageNumber="34" reason="6">
<emphasis id="B904C14EE574FF9AFCC4FC95B8840041" bold="true" box="[818,918,768,787]" pageId="33" pageNumber="34">Comments</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF1D5CE574FF9AFCC4FC88BE750749" blockId="33.[818,1443,768,1345]" pageId="33" pageNumber="34">
This species is relatively unspecialized in character states but can only be confused with the potentially sympatric species from East Africa possessing a robust and stocky mesosoma (LMI2 92103), such as
<taxonomicName id="4C7066DFE574FF9AFC8CFCE3B8CF00DA" authorityName="Hita-Garcia &amp; Lieberman &amp; Audisio &amp; Liu &amp; Economo" authorityYear="2019" box="[890,989,885,904]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="damato">
<emphasis id="B904C14EE574FF9AFC8CFCE3B8CF00DA" box="[890,989,885,904]" italics="true" pageId="33" pageNumber="34">D. damato</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7066DFE574FF9AFC1EFCE3BF4B00DA" authorityName="Hita-Garcia &amp; Lieberman &amp; Audisio &amp; Liu &amp; Economo" authorityYear="2019" box="[1000,1113,885,904]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="wakanda">
<emphasis id="B904C14EE574FF9AFC1EFCE3BF4B00DA" box="[1000,1113,885,904]" italics="true" pageId="33" pageNumber="34">D. wakanda</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7066DFE574FF9AFB92FCE3BFDB00DA" authorityName="Hita-Garcia &amp; Lieberman &amp; Audisio &amp; Liu &amp; Economo" authorityYear="2019" box="[1124,1225,885,904]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="schulzei">
<emphasis id="B904C14EE574FF9AFB92FCE3BFDB00DA" box="[1124,1225,885,904]" italics="true" pageId="33" pageNumber="34">D. schulzei</emphasis>
</taxonomicName>
, or
<taxonomicName id="4C7066DFE574FF9AFB19FCE3BE5200DA" authorityName="Hita-Garcia &amp; Lieberman &amp; Audisio &amp; Liu &amp; Economo" authorityYear="2019" box="[1263,1344,886,904]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="venus">
<emphasis id="B904C14EE574FF9AFB19FCE3BE5200DA" box="[1263,1344,886,904]" italics="true" pageId="33" pageNumber="34">D. venus</emphasis>
</taxonomicName>
. However, the latter is unique by having AT4 around 1.6 to 1.8 times longer than AT3 (ASI 158183), and
<taxonomicName id="4C7066DFE574FF9AFBBEFC24BFBF0091" authorityName="Hita-Garcia &amp; Lieberman &amp; Audisio &amp; Liu &amp; Economo" authorityYear="2019" box="[1096,1197,944,963]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="schulzei">
<emphasis id="B904C14EE574FF9AFBBEFC24BFBF0091" box="[1096,1197,944,963]" italics="true" pageId="33" pageNumber="34">D. schulzei</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7066DFE574FF9AFB2AFC24BE5F0091" authorityName="Hita-Garcia &amp; Lieberman &amp; Audisio &amp; Liu &amp; Economo" authorityYear="2019" box="[1244,1357,944,963]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="wakanda">
<emphasis id="B904C14EE574FF9AFB2AFC24BE5F0091" box="[1244,1357,944,963]" italics="true" pageId="33" pageNumber="34">D. wakanda</emphasis>
</taxonomicName>
have noticeable standing pilosity on the mesosomal and abdominal dorsa. The closest species is
<taxonomicName id="4C7066DFE574FF9AFC0AFC7EBF7200AC" authorityName="Hita-Garcia &amp; Lieberman &amp; Audisio &amp; Liu &amp; Economo" authorityYear="2019" box="[1020,1120,1003,1022]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="damato">
<emphasis id="B904C14EE574FF9AFC0AFC7EBF7200AC" box="[1020,1120,1003,1022]" italics="true" pageId="33" pageNumber="34">D. damato</emphasis>
</taxonomicName>
, but both can be differentiated by the eye size, the shape of petiole, and abdominal proportions.
</paragraph>
<paragraph id="8BCF1D5CE574FF9AFCA4FBB0BFF60613" blockId="33.[818,1443,768,1345]" pageId="33" pageNumber="34">
Admittedly, the diversity of habitats and the elevational range of
<taxonomicName id="4C7066DFE574FF9AFCB8FBD6B8B80704" authorityName="Hita-Garcia &amp; Lieberman &amp; Audisio &amp; Liu &amp; Economo" authorityYear="2019" box="[846,938,1091,1110]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="athene">
<emphasis id="B904C14EE574FF9AFCB8FBD6B8B80704" box="[846,938,1091,1110]" italics="true" pageId="33" pageNumber="34">D. athene</emphasis>
</taxonomicName>
are rather unusual for species of the
<taxonomicName id="4C7066DFE574FF9AFAEEFBD6BEB30704" authorityName="Santschi" authorityYear="1913" box="[1304,1441,1091,1110]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="traegaordhi">
<emphasis id="B904C14EE574FF9AFAEEFBD6BEB30704" box="[1304,1441,1091,1110]" italics="true" pageId="33" pageNumber="34">D. traegaordhi</emphasis>
</taxonomicName>
complex. Also, considering that there is some pronounced variation in size between series, it possible that the material considered as
<taxonomicName id="4C7066DFE574FF9AFCBDFB0EB8B407FC" authorityName="Hita-Garcia &amp; Lieberman &amp; Audisio &amp; Liu &amp; Economo" authorityYear="2019" box="[843,934,1179,1198]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="athene">
<emphasis id="B904C14EE574FF9AFCBDFB0EB8B407FC" box="[843,934,1179,1198]" italics="true" pageId="33" pageNumber="34">D. athene</emphasis>
</taxonomicName>
might represent more than one species. Unfortunately, the number of specimens from each locality is often limited, thus precluding a more in-depth analysis. Additional material from East Africa and molecular data could provide stronger evidence for heterospecificity, but based on the studied specimens we prefer to regard
<taxonomicName id="4C7066DFE574FF9AFC85FABBB8DE0612" authorityName="Hita-Garcia &amp; Lieberman &amp; Audisio &amp; Liu &amp; Economo" authorityYear="2019" box="[883,972,1325,1344]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="athene">
<emphasis id="B904C14EE574FF9AFC85FABBB8DE0612" box="[883,972,1325,1344]" italics="true" pageId="33" pageNumber="34">D. athene</emphasis>
</taxonomicName>
as one species for the present.
</paragraph>
<paragraph id="8BCF1D5CE574FF9AFCC4FAFDB89A0629" blockId="33.[818,1442,1384,1697]" box="[818,904,1384,1403]" pageId="33" pageNumber="34">
<heading id="D087AA30E574FF9AFCC4FAFDB89A0629" bold="true" box="[818,904,1384,1403]" fontSize="8" level="3" pageId="33" pageNumber="34" reason="6">
<emphasis id="B904C14EE574FF9AFCC4FAFDB89A0629" bold="true" box="[818,904,1384,1403]" pageId="33" pageNumber="34">Variation</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF1D5CE574FF9AFCC4FA10BFD105F2" blockId="33.[818,1442,1384,1697]" pageId="33" pageNumber="34">
<taxonomicName id="4C7066DFE574FF9AFCC4FA10B8CC06CA" authorityName="Hita-Garcia &amp; Lieberman &amp; Audisio &amp; Liu &amp; Economo" authorityYear="2019" box="[818,990,1413,1432]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="athene">
<emphasis id="B904C14EE574FF9AFCC4FA10B8CC06CA" box="[818,990,1413,1432]" italics="true" pageId="33" pageNumber="34">Discothyrea athene</emphasis>
</taxonomicName>
varies geographically in size, with specimens from
<collectingCountry id="F3675DCCE574FF9AFCC4FA36B86506E4" box="[818,887,1443,1462]" name="Uganda" pageId="33" pageNumber="34">Uganda</collectingCountry>
and
<collectingCountry id="F3675DCCE574FF9AFC57FA36B8FE06E4" box="[929,1004,1443,1462]" name="Rwanda" pageId="33" pageNumber="34">Rwanda</collectingCountry>
being generally larger (HW 0.380.42; WL 0.45 0.53) than those from
<collectingCountry id="F3675DCCE574FF9AFC0CFA54BF260686" box="[1018,1076,1473,1492]" name="Kenya" pageId="33" pageNumber="34">Kenya</collectingCountry>
,
<collectingCountry id="F3675DCCE574FF9AFBCBFA55BF820681" box="[1085,1168,1472,1491]" name="Tanzania" pageId="33" pageNumber="34">Tanzania</collectingCountry>
, and
<collectingCountry id="F3675DCCE574FF9AFB49FA55BE240681" box="[1215,1334,1472,1491]" name="Mozambique" pageId="33" pageNumber="34">Mozambique</collectingCountry>
(HW 0.33 0.39; WL 0.380.50). This variation in size is not related to elevation. There is also some variation in color which may correspond to distribution: most individuals are similarly orange, while those from Rangiro are notably darker. The degree of attenuation of the petiolar node is also quite variable (DPeI 300500; LPeI 286500) but does not seem to be correlated to geography since specimens from the same series display pronounced differences.
</paragraph>
</subSubSection>
</treatment>
</document>