464 lines
59 KiB
XML
464 lines
59 KiB
XML
<document id="1F0D944C907D9123D2EC5E91FC91EAC4" ID-CLB-Dataset="51220" ID-DOI="10.11646/zootaxa.4061.3.1" ID-GBIF-Dataset="518241f5-6ca9-4376-825e-c4c5de256b28" ID-ISSN="1175-5326" ID-Zenodo-Dep="200905" ID-ZooBank="719E677E-45E4-40A1-BE1D-5030CFB421CD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459676408665" checkinUser="plazi" docAuthor="Stanley, Edward L., Ceríaco, Luis M. P., Bandeira, Suzana, Valerio, Hilaria, Bates, Michael F. & Branch, William R." docDate="2016" docId="03D78791FF88424ABC8402C4BEA40CDE" docLanguage="en" docName="zootaxa.4061.3.1.pdf" docOrigin="Zootaxa 4061 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cordylus namakuiyus Stanley, Ceríaco, Bandeira, Valerio, Bates & Branch, 2016, sp. nov." docType="treatment" docVersion="10" lastPageNumber="217" masterDocId="FFEEFFE9FF83425ABC13021DBF070F33" masterDocTitle="A review of Cordylus machadoi (Squamata: Cordylidae) in southwestern Angola, with the description of a new species from the Pro-Namib desert" masterLastPageNumber="226" masterPageNumber="201" pageNumber="212" updateTime="1698206681825" updateUser="ExternalLinkService">
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<mods:title id="B63849F92D4324A830F0B187DDF6103E">A review of Cordylus machadoi (Squamata: Cordylidae) in southwestern Angola, with the description of a new species from the Pro-Namib desert</mods:title>
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<mods:namePart id="889686B541AA15E9658522DFD2820053">Stanley, Edward L.</mods:namePart>
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<mods:namePart id="CA73504EEBA2CD6B3B91D3601A4E7C46">Ceríaco, Luis M. P.</mods:namePart>
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<mods:namePart id="E763293B3B5BED80B72D64CEFE28A9F6">Bandeira, Suzana</mods:namePart>
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<mods:namePart id="01265530564D1D6C0D86854538B19D4F">Valerio, Hilaria</mods:namePart>
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<mods:namePart id="C309B26AAB2B600B13D8CBA83EE73706">Bates, Michael F.</mods:namePart>
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<mods:namePart id="EA3CA40662C2636DB368B51595B08C60">Branch, William R.</mods:namePart>
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<mods:title id="032A8D508DED497FE27E4356B46057F0">Zootaxa</mods:title>
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<mods:date id="4533DC8B530FB24DD6F61BFCECA31D66">2016</mods:date>
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<mods:number id="DBDA5E18907869A9A25EBC53F448F35D">4061</mods:number>
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<mods:identifier id="ED9EF4CD76241A14275A3E669F29B54B" type="DOI">10.11646/zootaxa.4061.3.1</mods:identifier>
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<treatment id="03D78791FF88424ABC8402C4BEA40CDE" ID-DOI="http://doi.org/10.5281/zenodo.3506980" ID-GBIF-Taxon="119332804" ID-Zenodo-Dep="3506980" LSID="urn:lsid:plazi:treatment:03D78791FF88424ABC8402C4BEA40CDE" httpUri="http://treatment.plazi.org/id/03D78791FF88424ABC8402C4BEA40CDE" lastPageId="16" lastPageNumber="217" pageId="11" pageNumber="212">
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<subSubSection id="C364650CFF884251BC8402C4BE740E27" pageId="11" pageNumber="212" type="nomenclature">
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<paragraph id="8BC13687FF884251BC8402C4BD030FC0" blockId="11.[151,516,217,277]" box="[151,516,217,243]" pageId="11" pageNumber="212">
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<heading id="D08981EBFF884251BC8402C4BD030FC0" bold="true" box="[151,516,217,243]" fontSize="11" level="1" pageId="11" pageNumber="212" reason="1">
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<emphasis id="B90AEA95FF884251BC8402C4BD030FC0" bold="true" box="[151,516,217,243]" pageId="11" pageNumber="212">
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<taxonomicName id="4C7E4D04FF884251BC8402C4BE980FC0" box="[151,415,217,243]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="212" phylum="Chordata" rank="species" species="namakuiyus" status="sp. nov.">
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<emphasis id="B90AEA95FF884251BC8402C4BE980FC0" bold="true" box="[151,415,217,243]" italics="true" pageId="11" pageNumber="212">Cordylus namakuiyus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23957EEFF884251BDB402C4BD030FC0" box="[423,516,217,243]" pageId="11" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC13687FF884251BC8402E1BE740E27" blockId="11.[151,516,217,277]" box="[151,371,252,277]" pageId="11" pageNumber="212">
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(
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<figureCitation id="13452A02FF884251BC8C02E1BE0C0E26" box="[159,267,252,277]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="2.[151,249,1395,1417]" captionStartId-1="8.[151,249,1861,1883]" captionStartId-2="9.[151,249,1120,1142]" captionStartId-3="12.[151,249,1924,1946]" captionStartId-4="13.[151,249,1634,1656]" captionStartId-5="15.[151,249,1882,1904]" captionTargetBox-0="[163,1436,375,1369]" captionTargetBox-1="[170,1414,201,1825]" captionTargetBox-2="[161,1413,208,1091]" captionTargetBox-3="[229,1353,231,1873]" captionTargetBox-4="[150,1436,194,1612]" captionTargetBox-5="[152,1436,193,1850]" captionTargetId-0="figure@2.[151,1436,375,1373]" captionTargetId-1="figure@8.[162,1427,193,1838]" captionTargetId-2="figure@9.[151,1436,193,1099]" captionTargetId-3="figure@12.[217,1370,193,1903]" captionTargetId-4="figure@13.[150,1436,193,1613]" captionTargetId-5="figure@15.[151,1436,193,1861]" captionTargetPageId-0="2" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="12" captionTargetPageId-4="13" captionTargetPageId-5="15" captionText-0="FIGURE 1. Head and body diagrams of Cordylus sp. (AMNH 47301, “ Angola ”) from an unpublished species description by Charles M. Bogert, illustrated by Alma W. Froderstrom, reproduced with the permission of the Department of Herpetology at the American Museum of Natural History" captionText-1="FIGURE 2. Map showing localities for Cordylus in Angola and Namibia. Cordylus namakuiyus sp. nov. 1) Iona national park: PEM R 18005, 2) Pico Azevedo: CAS 254754, 254755, 256529 – 31, 3) Caraculo: CAS 254912, 256529, 4) Iona National Park: TM 40430. C. machadoi: 5) Humpata environs: PEM R 18006 – 8, 19782, 19784, 6) Nasecute do Tchiviuguira: PEM R 18009, 7) NMN 7002, 8) TM 40131 – 3, 9) TM 40095, 40096. C. cf. machadoi: 10) Otjihipa Mountains: NCN – FN 377, 378, 11) Baynes Mountains: TM 57561, NCN-FP 402, 403, 12) Sera de Neve (sight record, P. Vaz Pinto), 13) C. angolensis, Caconda: type locality, and 14) Mombolo: AMNH 47331 – 35." captionText-2="FIGURE 3. Maximum likelihood phylogeny of 11 - gene dataset of Cordylus. Bootstrap support for MP and ML shown above branches, Bayesian posterior probabilities below. The new species, Cordylus namakuiyus sp. nov., is highlighted for reference. The numbered circles to the right of the Angolan specimens correspond to the numbered localities in Figure 2." captionText-3="FIGURE 4. Dorsal, lateral, and ventral views of A) Cordylus namakuiyus sp. nov. (CAS 254912, holotype) and B) C. machadoi (PEM R 18006)." captionText-4="FIGURE 5. High-resolution Computed Tomography reconstructions showing the ventral aspects of (top) Cordylus namakuiyus sp. nov. (CAS 254912, holotype) and (bottom) C. machadoi PEM R 18006. Osteoderms are shown in green. A – F tomograms showing cross-sections of the throat (A + D), torso (B + E), and hind limb (C + F)." captionText-5="FIGURE 6. PCA plots of A) 32 meristic and continuous and B) 22 meristic characters from 73 specimens of Cordylus. Green dots = C. namakuiyus sp. nov. (N = 10), red dots = C. machadoi (N = 13), yellow dots = AMNH C. cf. namakuiyus collected on the 1925 Vernay expedition (N = 48), purple star = C. cf. machadoi (TM 57561) from northern Namibia. Polygons with solid lines show morphospace of adult specimens, those with dotted lines represent morphospace of adults + juveniles." httpUri-0="https://zenodo.org/record/200906/files/figure.png" httpUri-1="https://zenodo.org/record/267461/files/figure.png" httpUri-2="https://zenodo.org/record/267462/files/figure.png" httpUri-3="https://zenodo.org/record/267463/files/figure.png" httpUri-4="https://zenodo.org/record/200910/files/figure.png" httpUri-5="https://zenodo.org/record/200911/files/figure.png" pageId="11" pageNumber="212">Figs. 1–6</figureCitation>
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,
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<tableCitation id="C6FC033CFF884251BD0502E1BE6D0E27" box="[278,362,252,277]" captionStart="TABLE 2 a" captionStartId="5.[125,203,568,588]" captionTargetBox="[126,1918,645,1655]" captionTargetPageId="5" captionText="TABLE 2 a. Variation in continuous measurements (mean and range) for specimens of Cordylus namakuiyus sp. nov., C. machadoi, C. cf. namakuiyus (AMNH R 47274 – 47321), and C. cf. machadoi from northern Namibia (TM 57561). All measurements following snout – vent length (SVL) are size-corrected (represented as percentage of SVL)." httpUri="http://table.plazi.org/id/DF01660FFF86425FBC6E0025B9180D55" pageId="11" pageNumber="212" tableUuid="DF01660FFF86425FBC6E0025B9180D55">Table 2</tableCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C364650CFF884251BC840323BBCE0C40" pageId="11" pageNumber="212" type="materials_examined">
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<paragraph id="8BC13687FF884251BC840323BA5B0EAC" blockId="11.[151,1437,318,2035]" pageId="11" pageNumber="212">
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<emphasis id="B90AEA95FF884251BC840323BE0C0E64" bold="true" box="[151,267,318,343]" pageId="11" pageNumber="212">
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<typeStatus id="54C58825FF884251BC840323BE010E64" box="[151,262,318,343]" pageId="11" pageNumber="212" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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254912, an adult female collected from a large rock outcrop near Caraculo, on the road from Lubango and Namibe, Namibe province,
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<collectingCountry id="F3697617FF884251BE78037FBDB90E48" box="[619,702,354,379]" name="Angola" pageId="11" pageNumber="212">Angola</collectingCountry>
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[
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,
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<geoCoordinate id="EE4A5040FF884251BF61037EBB1E0E48" box="[882,1049,355,379]" direction="east" orientation="longitude" pageId="11" pageNumber="212" precision="1" value="12.642028">12°38'31.30"E</geoCoordinate>
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;
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elevation], collected by E. L. Stanley, S. de Sá, S. Bandeira, H. Valerio, A. L. Kuhn, J. V. Vindum and L. Ceríaco, on
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<date id="FFC01047FF884251B89E039BBA5F0EAC" box="[1165,1368,390,415]" pageId="11" pageNumber="212" value="2013-12-06">6 December 2013</date>
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.
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</paragraph>
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<paragraph id="8BC13687FF884251BCD403B7BB6A0DA8" blockId="11.[151,1437,318,2035]" pageId="11" pageNumber="212">
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<typeStatus id="54C58825FF884251BCD403B7BE460EF0" box="[199,321,426,451]" pageId="11" pageNumber="212" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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Eight specimens:
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<collectionCode id="ED6FAE42FF884251BE0603B6BD4C0EF0" LSID="urn:lsid:biocol.org:col:34700" box="[533,587,427,451]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="11" pageNumber="212">CAS</collectionCode>
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254913–14, two fetuses taken from the
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<typeStatus id="54C58825FF884251B80003B7BB7F0EF0" box="[1043,1144,426,451]" pageId="11" pageNumber="212" type="holotype">holotype</typeStatus>
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, with the same data;
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<collectionCode id="ED6FAE42FF884251B97503B6BA9B0EF0" LSID="urn:lsid:biocol.org:col:34700" box="[1382,1436,427,451]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="11" pageNumber="212">CAS</collectionCode>
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256529, one juvenile collected at the same locality as the
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<typeStatus id="54C58825FF884251BF3F03D3BC890ED4" box="[812,910,462,487]" pageId="11" pageNumber="212" type="holotype">holotype</typeStatus>
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[
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,
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<geoCoordinate id="EE4A5040FF884251B82603D3BBC90ED4" box="[1077,1230,462,487]" direction="east" orientation="longitude" pageId="11" pageNumber="212" precision="1" value="12.642388">12°38'32.6"E</geoCoordinate>
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;
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<quantity id="4C869B62FF884251B8CF03D3BA230ED5" box="[1244,1316,462,487]" metricMagnitude="2" metricUnit="m" metricValue="5.09" pageId="11" pageNumber="212" unit="m" value="509.0">509 m</quantity>
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elevation] by the same collectors;
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<collectionCode id="ED6FAE42FF884251BD8C03EEBED20D38" LSID="urn:lsid:biocol.org:col:34700" box="[415,469,499,523]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="11" pageNumber="212">CAS</collectionCode>
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||
254754, adult female,
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||
<collectionCode id="ED6FAE42FF884251BECA03EEBC080D38" LSID="urn:lsid:biocol.org:col:34700" box="[729,783,499,523]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="11" pageNumber="212">CAS</collectionCode>
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254755, adult male,
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<collectionCode id="ED6FAE42FF884251BFEF03EEBB350D38" LSID="urn:lsid:biocol.org:col:34700" box="[1020,1074,499,523]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="11" pageNumber="212">CAS</collectionCode>
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256530, juvenile,
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<collectionCode id="ED6FAE42FF884251B91703EEBA3D0D38" LSID="urn:lsid:biocol.org:col:34700" box="[1284,1338,499,523]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="11" pageNumber="212">CAS</collectionCode>
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256531, adult female, all collected on the outskirts of Pico Azevedo [
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<geoCoordinate id="EE4A5040FF884251BF57000BBCDA0D1C" box="[836,989,534,559]" direction="south" orientation="latitude" pageId="11" pageNumber="212" precision="1" value="-15.475888">15°28'33.2"S</geoCoordinate>
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,
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<geoCoordinate id="EE4A5040FF884251BFFA000BBB850D1C" box="[1001,1154,534,559]" direction="east" orientation="longitude" pageId="11" pageNumber="212" precision="1" value="12.462694">12°27'45.7"E</geoCoordinate>
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;
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<quantity id="4C869B62FF884251B89C000ABBD10D1D" box="[1167,1238,535,559]" metricMagnitude="2" metricUnit="m" metricValue="4.21" pageId="11" pageNumber="212" unit="m" value="421.0">421 m</quantity>
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elevation] by the same collectors of the
|
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<typeStatus id="54C58825FF884251BD860027BEF00D60" box="[405,503,570,595]" pageId="11" pageNumber="212" type="holotype">holotype</typeStatus>
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on
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||
<date id="FFC01047FF884251BE300026BDE90D60" box="[547,750,570,595]" pageId="11" pageNumber="212" value="2013-12-07">7 December 2013</date>
|
||
;
|
||
<collectionCode id="ED6FAE42FF884251BEE90026BC330D61" box="[762,820,571,594]" httpUri="http://grbio.org/cool/xqn4-34sv" name="Port Elizabeth Museum" pageId="11" pageNumber="212">PEM</collectionCode>
|
||
R18005, adult female, collected in a low rock outcrop bordering the road between Namibe and Omahua lodge [
|
||
<geoCoordinate id="EE4A5040FF884251BF370043BCBB0D44" box="[804,956,606,631]" direction="south" orientation="latitude" pageId="11" pageNumber="212" precision="1" value="-15.996806">15°59'48.5"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE4A5040FF884251BFD90043BB630D44" box="[970,1124,606,631]" direction="east" orientation="longitude" pageId="11" pageNumber="212" precision="1" value="12.406833">12°24'24.6"E</geoCoordinate>
|
||
;
|
||
<quantity id="4C869B62FF884251B8620042BBBD0D45" box="[1137,1210,607,631]" metricMagnitude="2" metricUnit="m" metricValue="3.08" pageId="11" pageNumber="212" unit="m" value="308.0">308 m</quantity>
|
||
elevation], Namibe Province,
|
||
<collectingCountry id="F3697617FF884251BD1B009FBE5A0DA8" box="[264,349,642,667]" name="Angola" pageId="11" pageNumber="212">Angola</collectingCountry>
|
||
, by W. R. Branch, K. A. Tolley and G. J. Measey on
|
||
<date id="FFC01047FF884251BFBF009EBB6E0DA8" box="[940,1129,643,667]" pageId="11" pageNumber="212" value="2009-01-20">20 January 2009</date>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC13687FF884251BCD400BBBBCE0C40" blockId="11.[151,1437,318,2035]" pageId="11" pageNumber="212">
|
||
<emphasis id="B90AEA95FF884251BCD400BBBEB90D8C" bold="true" box="[199,446,678,703]" pageId="11" pageNumber="212">Additional material.</emphasis>
|
||
Forty-nine specimens. One specimen: TM 40430, adult female from
|
||
<quantity id="4C869B62FF884251B8E200BABA3A0D8C" box="[1265,1341,678,703]" metricMagnitude="4" metricUnit="m" metricValue="4.9" pageId="11" pageNumber="212" unit="km" value="49.0">49 km</quantity>
|
||
ESE of Tombua, Namibe Province,
|
||
<collectingCountry id="F3697617FF884251BDC200D7BD230DD0" box="[465,548,714,739]" name="Angola" pageId="11" pageNumber="212">Angola</collectingCountry>
|
||
[
|
||
<geoCoordinate id="EE4A5040FF884251BE2100D7BD890DD0" box="[562,654,714,739]" direction="south" orientation="latitude" pageId="11" pageNumber="212" precision="925" value="-15.883333">15°53'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE4A5040FF884251BE8900D7BDF00DD0" box="[666,759,714,739]" direction="east" orientation="longitude" pageId="11" pageNumber="212" precision="925" value="12.266666">12°16'E</geoCoordinate>
|
||
], collected by W. D. Haacke on
|
||
<date id="FFC01047FF884251B87300D6BA090DD0" box="[1120,1294,714,739]" pageId="11" pageNumber="212" value="1971-03-31">31 March 1971</date>
|
||
; and a series of
|
||
<specimenCount id="9D78FD0EFF884251BCA500F2BE490C34" box="[182,334,750,775]" pageId="11" pageNumber="212" type="generic">48 specimens</specimenCount>
|
||
collected by A. S. Vernay, H. Lang and R. Boulton between
|
||
<date id="FFC01047FF884251BFED00F3BA3A0C34" box="[1022,1341,750,775]" pageId="11" pageNumber="212" value="1925-04-25" valueMax="1925-08-04" valueMin="1925-04-25">25 April and 4 August 1925</date>
|
||
, locality “
|
||
<collectingCountry id="F3697617FF884251BCB2010FBFFC0C18" box="[161,251,786,811]" name="Angola" pageId="11" pageNumber="212">Angola</collectingCountry>
|
||
”: Adult females:
|
||
<collectionCode id="ED6FAE42FF884251BDD9010EBD180C18" LSID="urn:lsid:biocol.org:col:34925" box="[458,543,787,811]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="11" pageNumber="212">AMNH</collectionCode>
|
||
R47274, 47276, 47278, 47279, 47282–84, 47286, 47287, 47289-95, 47297, 47298, 47299, 47304, 47305, and 47315. Adult males:
|
||
<collectionCode id="ED6FAE42FF884251BF30012ABC7F0C7C" LSID="urn:lsid:biocol.org:col:34925" box="[803,888,823,847]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="11" pageNumber="212">AMNH</collectionCode>
|
||
R47275, 47277, 47280, 47281, 47285, 47288, 47296, 47300–03, 47306–09, and 47311. Juveniles:
|
||
<collectionCode id="ED6FAE42FF884251BEF30146BC310C40" LSID="urn:lsid:biocol.org:col:34925" box="[736,822,859,883]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="11" pageNumber="212">AMNH</collectionCode>
|
||
R47310, 47312–14, and 47316–23.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C364650CFF884251BCD40163BB5D0AE4" pageId="11" pageNumber="212" type="diagnosis">
|
||
<paragraph id="8BC13687FF884251BCD40163BB5D0AE4" blockId="11.[151,1437,318,2035]" pageId="11" pageNumber="212">
|
||
<emphasis id="B90AEA95FF884251BCD40163BE460CA4" bold="true" box="[199,321,894,919]" pageId="11" pageNumber="212">Diagnosis.</emphasis>
|
||
A relatively large species of
|
||
<taxonomicName id="4C7E4D04FF884251BEA20163BC100CA5" box="[689,791,894,918]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="212" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90AEA95FF884251BEA20163BC100CA5" box="[689,791,894,918]" italics="true" pageId="11" pageNumber="212">Cordylus</emphasis>
|
||
</taxonomicName>
|
||
, identified to genus by the following combination of characters: four-limbed, rupicolous, viviparous, strongly depressed triangular head and body, osteoderms present, rhomboidal, imbricate and keeled dorsal scales present, occipitals non-spinose, and spinose caudal and limb scales enlarged (
|
||
<bibRefCitation id="EFEF4B76FF884251BD1B01F7BE980B30" author="Branch" box="[264,415,1002,1027]" pageId="11" pageNumber="218" refString="Branch, W. R. (1998) Field guide to snakes and other reptiles of southern Africa. Ralph Curtis Books Publishing: Sanibel Island, Florida, 399 pp." type="book" year="1998">Branch 1998</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEF4B76FF884251BDBD01F7BDE10B30" author="Broadley" box="[430,742,1002,1027]" pageId="11" pageNumber="218" refString="Broadley, D. G. & Branch, W. R. (2002) A review of the small east African Cordylus (Sauria: Cordylidae), with the description of a new species. African Journal of Herpetology, 51, 9 - 34. http: // dx. doi. org / 10.1080 / 21564574.2002.9635459" type="journal article" year="2002">Broadley and Branch 2002</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEF4B76FF884251BEE601F7BCDE0B30" author="Stanley" box="[757,985,1002,1027]" pageId="11" pageNumber="219" refString="Stanley, E. L., Bauer, A. M., Jackman, T. R., Branch, W. R. & Mouton, P. leF. N. (2011) Between a rock and a hard polytomy: Rapid radiation in the rupicolous girdled lizards (Squamata: Cordylidae). Molecular Phylogenetics and Evolution, 58, 53 - 70. http: // dx. doi. org / 10.1016 / j. ympev. 2010.08.024" type="journal article" year="2011">
|
||
Stanley
|
||
<emphasis id="B90AEA95FF884251BF4101F6BC930B30" box="[850,916,1003,1027]" italics="true" pageId="11" pageNumber="212">et al.,</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C7E4D04FF884251BFFD01F6BBE30B30" box="[1006,1252,1003,1027]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="212" phylum="Chordata" rank="species" species="namakuiyus" status="sp. nov.">
|
||
<emphasis id="B90AEA95FF884251BFFD01F6BBE30B30" box="[1006,1252,1003,1027]" italics="true" pageId="11" pageNumber="212">Cordylus namakuiyus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23957EEFF884251B8FE01F7BA4E0B30" box="[1261,1353,1002,1027]" pageId="11" pageNumber="212" rank="species">
|
||
<emphasis id="B90AEA95FF884251B8FE01F7BA4E0B30" bold="true" box="[1261,1353,1002,1027]" pageId="11" pageNumber="212">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from all other species in the genus except for
|
||
<taxonomicName id="4C7E4D04FF884251BEB00612BC170B14" box="[675,784,1039,1063]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="212" phylum="Chordata" rank="species" species="vittifer">
|
||
<emphasis id="B90AEA95FF884251BEB00612BC170B14" box="[675,784,1039,1063]" italics="true" pageId="11" pageNumber="212">C. vittifer</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7E4D04FF884251BF5A0612BCDE0B15" box="[841,985,1038,1062]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="212" phylum="Chordata" rank="species" species="machadoi">
|
||
<emphasis id="B90AEA95FF884251BF5A0612BCDE0B15" box="[841,985,1038,1062]" italics="true" pageId="11" pageNumber="212">C. machadoi</emphasis>
|
||
</taxonomicName>
|
||
, by the presence of a transverse row of elongated dorsal scales immediately posterior to occipitals (vs. absence); from
|
||
<taxonomicName id="4C7E4D04FF884251B842062EBBC30B78" box="[1105,1220,1075,1099]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="212" phylum="Chordata" rank="species" species="vittifer">
|
||
<emphasis id="B90AEA95FF884251B842062EBBC30B78" box="[1105,1220,1075,1099]" italics="true" pageId="11" pageNumber="212">C. vittifer</emphasis>
|
||
</taxonomicName>
|
||
by possessing an incomplete row of pre-occipital scales between posterior parietal and occipital scales (absent in
|
||
<taxonomicName id="4C7E4D04FF884251B8DE064ABA3E0B5C" box="[1229,1337,1111,1135]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="212" phylum="Chordata" rank="species" species="vittifer">
|
||
<emphasis id="B90AEA95FF884251B8DE064ABA3E0B5C" box="[1229,1337,1111,1135]" italics="true" pageId="11" pageNumber="212">C. vittifer</emphasis>
|
||
</taxonomicName>
|
||
), and by having infralabials that are moderately-deeply corrugated (vs. usually smooth); from
|
||
<taxonomicName id="4C7E4D04FF884251B8660666BA000BA0" box="[1141,1287,1147,1171]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="212" phylum="Chordata" rank="species" species="machadoi">
|
||
<emphasis id="B90AEA95FF884251B8660666BA000BA0" box="[1141,1287,1147,1171]" italics="true" pageId="11" pageNumber="212">C. machadoi</emphasis>
|
||
</taxonomicName>
|
||
by having a large, keyhole-shaped interparietal in broad contact with frontoparietals, thereby separating anterior parietals (vs. small, diamond-shaped interparietal not in contact with frontoparietal, thereby never completely separating anterior parietals), temporal scales that are strongly keeled (vs. weakly keeled), fewer transverse gular rows (14–17 vs. 17– 24), light brown dorsal body coloration (vs. darker brown), and by the absence of dark speckles on throat and ventral body surfaces (vs. presence), presence of osteoderms on throat and ventral surfaces (vs. absence); from
|
||
<taxonomicName id="4C7E4D04FF884251B9910732BE080A58" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="212" phylum="Chordata" rank="species" species="angolensis">
|
||
<emphasis id="B90AEA95FF884251B9910732BE080A58" italics="true" pageId="11" pageNumber="212">C. angolensis</emphasis>
|
||
</taxonomicName>
|
||
(based on original description;
|
||
<bibRefCitation id="EFEF4B76FF884251BE66074EBC0D0A58" author="Bocage" box="[629,778,1362,1387]" pageId="11" pageNumber="218" refString="Bocage, J. (1895) Herpetologie d'Angola et du Congo. Imprimerie Nationale IXX: Lisbon, Lisbonne, 203 pp., 220 pls." type="book" year="1895">Bocage 1895</bibRefCitation>
|
||
) by having fewer ventral transverse scale rows (21–24 vs. 27), light brown dorsal body coloration (vs. brown with blackish speckles over paler dorsal ground coloration), and by the absence of longitudinal series of whitish speckles along dorsal surface (vs. presence of two longitudinal series of small whitish speckles along dorsum), and presence of a loreal (vs. absence).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C364650CFF88424ABCD407FFBA4F0E2E" lastPageId="16" lastPageNumber="217" pageId="11" pageNumber="212" type="description">
|
||
<paragraph id="8BC13687FF884251BCD407FFBDA208C0" blockId="11.[151,1437,318,2035]" pageId="11" pageNumber="212">
|
||
<emphasis id="B90AEA95FF884251BCD407FFBEED0AC8" bold="true" box="[199,490,1506,1531]" pageId="11" pageNumber="212">
|
||
Description of
|
||
<typeStatus id="54C58825FF884251BD6F07FFBEE20AC8" box="[380,485,1506,1531]" pageId="11" pageNumber="212" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
SVL
|
||
<quantity id="4C869B62FF884251BE2107FEBD930AC9" box="[562,660,1507,1531]" metricMagnitude="-1" metricUnit="m" metricValue="1.01" pageId="11" pageNumber="212" unit="mm" value="101.0">101 mm</quantity>
|
||
. Head and body depressed. Head 1.2
|
||
<emphasis id="B90AEA95FF884251B85807FEBB5C0AC9" box="[1099,1115,1507,1530]" italics="true" pageId="11" pageNumber="212">×</emphasis>
|
||
as long as broad. Nasals in median broad contact; frontonasal lozenge-shaped, as broad as long, separated from frontal by enlarged prefrontals (in median contact, forming a suture), separated from rostral by nasals, separated from loreal by prefrontals; frontal in contact with first and second supraoculars, followed by a pair of large frontoparietals in broad, median contact; a distinctive keyhole-shaped interparietal in broad contact with the frontoparietals, separating anterior parietals; parietal eye visible; a row of 10 rugose occipitals. A row of 11 elongated nuchal scales. Four supraoculars and three supraciliaries. Nasals large, with nostril pierced centrally on upper margin. Loreal in contact with preocular, nasal and first two supralabials; three suboculars, well separated from the lip by the fourth and fifth supralabials. Rostral twice as broad as deep; supralabials five; infralabials five; chin shields five. Mental twice as broad as long; gulars smooth, enlarged and forming transverse rows posteriorly, with 16 gulars between the posterior extent of the jaws. Dorsal scales rectangular, rugose, obtusely keeled, mucronate and serrate posteriorly; dorsals and laterals in 25 transverse and 24 longitudinal rows; ventrals squarish, smooth, in 21 transverse and 15 longitudinal rows. Scales on limbs above large, strongly keeled and spinose; subdigital lamellae under fourth toe 14; femoral pores five; generation glands absent. Tail with whorls of large, elongate, strongly keeled, spinose and serrated scales, spines directed posteriorly and largest superolaterally.
|
||
</paragraph>
|
||
<caption id="DF01660FFF8F4256BC840599BEA4088A" httpUri="https://zenodo.org/record/267463/files/figure.png" pageId="12" pageNumber="213" targetBox="[229,1353,231,1873]" targetPageId="12">
|
||
<paragraph id="8BC13687FF8F4256BC840599BEA4088A" blockId="12.[151,1436,1924,1977]" pageId="12" pageNumber="213">
|
||
<emphasis id="B90AEA95FF8F4256BC840599BE1008A9" bold="true" box="[151,279,1924,1946]" pageId="12" pageNumber="213">FIGURE 4.</emphasis>
|
||
Dorsal, lateral, and ventral views of
|
||
<emphasis id="B90AEA95FF8F4256BEA30599BDCD08A9" bold="true" box="[688,714,1924,1946]" pageId="12" pageNumber="213">A)</emphasis>
|
||
<taxonomicName id="4C7E4D04FF8F4256BEC50599BCB108AA" box="[726,950,1924,1945]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="213" phylum="Chordata" rank="species" species="namakuiyus" status="sp. nov.">
|
||
<emphasis id="B90AEA95FF8F4256BEC50599BCB108AA" box="[726,950,1924,1945]" italics="true" pageId="12" pageNumber="213">Cordylus namakuiyus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23957EEFF8F4256BFD20599BB1108A9" box="[961,1046,1924,1946]" pageId="12" pageNumber="213" rank="species">
|
||
<emphasis id="B90AEA95FF8F4256BFD20599BB1108A9" bold="true" box="[961,1046,1924,1946]" pageId="12" pageNumber="213">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(CAS 254912, holotype) and
|
||
<emphasis id="B90AEA95FF8F4256B9730599BA7D08A9" bold="true" box="[1376,1402,1924,1946]" pageId="12" pageNumber="213">B)</emphasis>
|
||
<taxonomicName id="4C7E4D04FF8F4256B9970598BFFC088B" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="213" phylum="Chordata" rank="species" species="machadoi">
|
||
<emphasis id="B90AEA95FF8F4256B9970598BFFC088B" italics="true" pageId="12" pageNumber="213">C. machadoi</emphasis>
|
||
</taxonomicName>
|
||
(PEM R18006).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF01660FFF8E4257BC84047FBCB00986" httpUri="https://zenodo.org/record/200910/files/figure.png" pageId="13" pageNumber="214" targetBox="[150,1436,194,1612]" targetPageId="13">
|
||
<paragraph id="8BC13687FF8E4257BC84047FBCB00986" blockId="13.[151,1436,1634,1717]" pageId="13" pageNumber="214">
|
||
<emphasis id="B90AEA95FF8E4257BC84047FBE16094B" bold="true" box="[151,273,1634,1656]" pageId="13" pageNumber="214">FIGURE 5.</emphasis>
|
||
High-resolution Computed Tomography reconstructions showing the ventral aspects of (top)
|
||
<taxonomicName id="4C7E4D04FF8E4257B8D2047FBA9B0944" box="[1217,1436,1634,1655]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="214" phylum="Chordata" rank="species" species="namakuiyus" status="sp. nov.">
|
||
<emphasis id="B90AEA95FF8E4257B8D2047FBA9B0944" box="[1217,1436,1634,1655]" italics="true" pageId="13" pageNumber="214">Cordylus namakuiyus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23957EEFF8E4257BC84049DBFEF09A5" box="[151,232,1664,1686]" pageId="13" pageNumber="214" rank="species">
|
||
<emphasis id="B90AEA95FF8E4257BC84049DBFEF09A5" bold="true" box="[151,232,1664,1686]" pageId="13" pageNumber="214">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(CAS 254912, holotype) and (bottom)
|
||
<taxonomicName id="4C7E4D04FF8E4257BE93049CBC0409A5" box="[640,771,1665,1686]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="214" phylum="Chordata" rank="species" species="machadoi">
|
||
<emphasis id="B90AEA95FF8E4257BE93049CBC0409A5" box="[640,771,1665,1686]" italics="true" pageId="13" pageNumber="214">C. machadoi</emphasis>
|
||
</taxonomicName>
|
||
PEM R18006. Osteoderms are shown in green. A–F tomograms showing cross-sections of the throat (A+D), torso (B+E), and hind limb (C+F).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC13687FF8E4254BCD404C1BE910EC6" blockId="13.[151,1436,1756,2033]" lastBlockId="14.[151,1437,151,2013]" lastPageId="14" lastPageNumber="215" pageId="13" pageNumber="214">
|
||
<emphasis id="B90AEA95FF8E4257BCD404C1BEA609C6" bold="true" box="[199,417,1756,1781]" pageId="13" pageNumber="214">Cranial osteology.</emphasis>
|
||
The parietal is pentagonal with a well-developed, unbifurcated medio-posterior process that extends past the postero-lateral processes. The premaxilla is unpaired and contains seven pleurodont teeth and five foramina, with a dorsal process that extends posteriorly to be clasped by the nasals, which themselves insert into an unpaired frontal. The maxilla is typically scinciform, with a deeply grooved crista dentalis and 16 pleurodont teeth. No palpebral is present, though the prefrontal has a small, flattened, laterally projecting tubercle that supports the anterior-most superorbital osteoderm in much the same way. The jugal is triangular in crosssection and asymmetrically T-shaped, with a tapering anterior process and a broad, truncated posterior process that extends along and past the posterior edge of the maxilla. The lacrimal bone is small, flattened and extends along the postero-ventral boundary of the maxilla and prefrontal. Edentate pterygoids extend back to connect with the quadrates, becoming C-shaped in cross-section posterior to the epipterygoid condyle. The squamosal is curved and blade-like, circular in cross-section anteriorly, becoming flattened posteriorly, where it articulates with the cephalic condyle of the quadrate and the braincase. Supratemporals are flattened, reniform and fused to the paroccipital processes. The supraoccipital has a strong sagittal crest that extends posteriorly to contact the ventral surface of the medio-posterior process of the parietal. The prootic bears an extended alar process and a well-developed, rhomboid christa prootica, and a very weak supratrigeminal process. Basipterygoid processes are well developed and flattened. The lower jaw possesses a large adductor fossa, a highly flattened and medially extended retroarticular process, a medially open meckelian canal and a dentary with a strong subdental shelf, 21 mandibular teeth, and nine dentary foramina.
|
||
</paragraph>
|
||
<paragraph id="8BC13687FF8D4254BCD403E2BB020CDE" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="215">
|
||
<emphasis id="B90AEA95FF8D4254BCD403E2BECC0D2B" bold="true" box="[199,459,511,536]" pageId="14" pageNumber="215">Postcranial osteology.</emphasis>
|
||
The
|
||
<typeStatus id="54C58825FF8D4254BE1403E2BD6E0D2B" box="[519,617,511,536]" pageId="14" pageNumber="215" type="holotype">holotype</typeStatus>
|
||
possesses 25 presacral and 17 postsacral vertebrae with the last
|
||
<quantity id="4C869B62FF8D4254B95F001DBA9B0D24" box="[1356,1436,512,536]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="14" pageNumber="215" unit="mm" value="21.0">21 mm</quantity>
|
||
(roughly 20%) of the tail made up of regenerated cartilage. There are five cervical ribs, three sternal ribs, two xiphisternal ribs, five long asternal ribs with ossified costal cartilage, and eight short asternal ribs. The first three cervical ribs are distally flattened with bifid cartilaginous projections. Sacral vertebrae are fused asymmetrically, with the left diapophysis formed from the fused sacral ribs, while the right diapophysis is formed by the fusing of the second sacral and first caudal ribs. The next two pairs of caudal ribs are well developed and angled anteriorly, with subsequent ribs becoming increasingly smaller and posteriorly angled. Pelvic girdle is well developed and flattened. No iliac tubercle is present. There is a well-developed, ventrally angled pubic tubercle directly anterior to the obturator foramen. Both hypoischium and hyperischium are well developed. Pubic bones are well separated by triangular prepubic cartilage. The sternal plate is broad and lacks a fontanelle. Interclavicle cruciform, clavicles rod-like and flattened dorsally. The epicoracoid is narrow and curved, connecting the scapular ray to the primary and secondary coracoid rays, but not to the anterior process of the scapular. Limb bones well developed. Digits display the typical squamate phalangeal arrangement of 3–5–4–3–2. A large sesamoid is found in the palm of the hands. The fifth metatarsal possesses an elongated medial process at midbody.
|
||
</paragraph>
|
||
<paragraph id="8BC13687FF8D4254BCD401EABE760A66" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="215">
|
||
<emphasis id="B90AEA95FF8D4254BCD401EABE5A0B23" bold="true" box="[199,349,1015,1040]" pageId="14" pageNumber="215">Osteoderms.</emphasis>
|
||
Scales of the dorsal and temporal regions of the skull and the ventrolateral aspects of the jaws are underlain with rugose osteoderms. These osteoderms fuse to the proximal parietal, frontal and postorbital bones, although the mesokinetic and metakinetic joints appear unobstructed and flexible. The entire body is covered in robust osteoderms, with only the femoral pores and the axillary, inguinal and cloacal regions unarmoured. Large, rectangular, imbricate osteoderms protect the dorsal and lateral aspect of the trunk, becoming more keeled and mucronate laterally. Unkeeled, imbricate osteoderms cover the entire venter, from the gulars to the cloaca, rectangular on the throat and abdomen, becoming rhomboid around the sternum and cloaca. Several abdominal osteoderms appear to be fractured. Caudal osteoderms are thick, sharply spined and arranged in imbricate transverse whorls. Limbs covered by imbricate circular/rhomboid osteoderms, keeled and mucronate dorsally, plate-like ventrally.
|
||
</paragraph>
|
||
<paragraph id="8BC13687FF8D4254BCD40742BD5D0963" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="215">
|
||
<emphasis id="B90AEA95FF8D4254BCD40742BEF50A4B" bold="true" box="[199,498,1375,1400]" pageId="14" pageNumber="215">
|
||
Colouration of
|
||
<typeStatus id="54C58825FF8D4254BD970742BEEA0A4B" box="[388,493,1375,1400]" pageId="14" pageNumber="215" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Dorsal coloration is medially brown, blending to a straw-yellow laterally. Head yellow-brown; supra and infralabials yellowish; a dark-brown bar extends from the posterior aspect of the eye to the temporals. The base of the tail is medially brown, surrounded by a straw-yellow coloration that extends towards the tip and eventually dominates the tail; the dorsal aspect of the limbs is brown. Laterally a dark-brown line, similar to the eye-mask coloration, extends from the neck towards the insertion of the forelimbs. The venter is yellowish-white, cream on the tail; the underside of the hind limbs, cloacal region and tail are cream, grading into yellowish on the anterior half of the tail.
|
||
</paragraph>
|
||
<paragraph id="8BC13687FF8D424ABCD40441BA4F0E2E" blockId="14.[151,1437,151,2013]" lastBlockId="16.[151,1437,151,1005]" lastPageId="16" lastPageNumber="217" pageId="14" pageNumber="215">
|
||
<emphasis id="B90AEA95FF8D4254BCD40441BE3C0946" bold="true" box="[199,315,1628,1653]" pageId="14" pageNumber="215">Variation.</emphasis>
|
||
Variation in scalation and body measurements of
|
||
<taxonomicName id="4C7E4D04FF8D4254BF690441BB250947" box="[890,1058,1628,1652]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="215" phylum="Chordata" rank="species" species="namakuiyus" status="sp. nov.">
|
||
<emphasis id="B90AEA95FF8D4254BF690441BB250947" box="[890,1058,1628,1652]" italics="true" pageId="14" pageNumber="215">C. namakuiyus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23957EEFF8D4254B8380441BB810946" box="[1067,1158,1628,1653]" pageId="14" pageNumber="215" rank="species">
|
||
<emphasis id="B90AEA95FF8D4254B8380441BB810946" bold="true" box="[1067,1158,1628,1653]" pageId="14" pageNumber="215">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
are reported in
|
||
<tableCitation id="C6FC033CFF8D4254B92E0441BA9B0947" box="[1341,1436,1628,1653]" captionStart="TABLE 2 a" captionStartId="5.[125,203,568,588]" captionTargetBox="[126,1918,645,1655]" captionTargetPageId="5" captionText="TABLE 2 a. Variation in continuous measurements (mean and range) for specimens of Cordylus namakuiyus sp. nov., C. machadoi, C. cf. namakuiyus (AMNH R 47274 – 47321), and C. cf. machadoi from northern Namibia (TM 57561). All measurements following snout – vent length (SVL) are size-corrected (represented as percentage of SVL)." httpUri="http://table.plazi.org/id/DF01660FFF86425FBC6E0025B9180D55" pageId="14" pageNumber="215" tableUuid="DF01660FFF86425FBC6E0025B9180D55">Tables 2</tableCitation>
|
||
and 3. The sole male
|
||
<typeStatus id="54C58825FF8D4254BD80049CBEF309AA" box="[403,500,1665,1689]" pageId="14" pageNumber="215" type="paratype">paratype</typeStatus>
|
||
CAS 254755 has 12 generation glands on the ventral surface of each thigh. One female
|
||
<typeStatus id="54C58825FF8D4254BCE004B8BE54098E" box="[243,339,1701,1725]" pageId="14" pageNumber="215" type="paratype">paratype</typeStatus>
|
||
(CAS 254754) differs from the
|
||
<typeStatus id="54C58825FF8D4254BEE004B9BC52098E" box="[755,853,1700,1725]" pageId="14" pageNumber="215" type="holotype">holotype</typeStatus>
|
||
in having the loreal and preocular fused. The medioposterior parietal process is slightly bifurcated distally in CAS 254754, CAS 265529 and PEM R18005, and does not extend back beyond the posterolateral processes in CAS 25630. Five premaxillary foramina are present in all
|
||
<typeStatus id="54C58825FF8D4254BCA8050CBFEC081A" box="[187,235,1809,1833]" pageId="14" pageNumber="215">type</typeStatus>
|
||
specimens except CAS 254754 (two) and CAS 256531 (four). Specimens CAS 254755 and PEM R18005 have 15 maxillary teeth, and CAS 256531 has 17. All
|
||
<typeStatus id="54C58825FF8D4254BF1D0528BC7E087E" box="[782,889,1845,1869]" pageId="14" pageNumber="215" type="paratype">paratypes</typeStatus>
|
||
have 21 mandibular teeth, except CAS 254754 (20), CAS 256530 (18), and CAS 254755 and CAS 256529 (16). The supratemporals are not fused to the paroccipital processes in CAS 254754, CAS 256529, and CAS 256530. Five of the
|
||
<typeStatus id="54C58825FF8D4254B8970560BBE808A6" box="[1156,1263,1917,1941]" pageId="14" pageNumber="215" type="paratype">paratypes</typeStatus>
|
||
have full, unautotomised tails: CAS 254755 has 29 postsacral vertebrae, CAS 25474, CAS 256529 each have 27, and the two fetuses, CAS 254913 and CAS 254914, have 25 and 26 respectively. One specimen, CAS 256531, has 26 presacral vertebrae. CAS 254755, CAS 256529 and CAS 256530 each possess four pairs of long asternal ribs, while CAS 256529 and PEM R18005 each have seven short asternal ribs. None of the
|
||
<typeStatus id="54C58825FF93424AB81802A0BB710FE6" box="[1035,1142,189,213]" pageId="16" pageNumber="217" type="paratype">paratypes</typeStatus>
|
||
exhibit the asymmetrical sacral fusion of the
|
||
<typeStatus id="54C58825FF93424ABD6102C2BED00FCB" box="[370,471,223,248]" pageId="16" pageNumber="217" type="holotype">holotype</typeStatus>
|
||
, but rather display the normal condition of having the sacral diapophyses formed from the sacral ribs. The hyperischium of CAS 254755 extends anteriorly to connect with the prepubic cartilage.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF01660FFF8C4255BC840547BA1608FF" httpUri="https://zenodo.org/record/200911/files/figure.png" pageId="15" pageNumber="216" targetBox="[152,1436,193,1850]" targetPageId="15">
|
||
<paragraph id="8BC13687FF8C4255BC840547BA1608FF" blockId="15.[151,1436,1882,1996]" pageId="15" pageNumber="216">
|
||
<emphasis id="B90AEA95FF8C4255BC840547BE140843" bold="true" box="[151,275,1882,1904]" pageId="15" pageNumber="216">FIGURE 6.</emphasis>
|
||
PCA plots of A) 32 meristic and continuous and B) 22 meristic characters from 73 specimens of
|
||
<taxonomicName id="4C7E4D04FF8C4255B8E60546BA560843" box="[1269,1361,1883,1904]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="216" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90AEA95FF8C4255B8E60546BA560843" box="[1269,1361,1883,1904]" italics="true" pageId="15" pageNumber="216">Cordylus</emphasis>
|
||
</taxonomicName>
|
||
. Green dots =
|
||
<taxonomicName id="4C7E4D04FF8C4255BCC80567BE7608BD" box="[219,369,1913,1935]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="216" phylum="Chordata" rank="species" species="namakuiyus" status="sp. nov.">
|
||
<emphasis id="B90AEA95FF8C4255BCC80567BE7608BD" box="[219,369,1913,1935]" italics="true" pageId="15" pageNumber="216">C. namakuiyus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23957EEFF8C4255BD640564BEC008BC" box="[375,455,1913,1935]" pageId="15" pageNumber="216" rank="species">
|
||
<emphasis id="B90AEA95FF8C4255BD640564BEC008BC" bold="true" box="[375,455,1913,1935]" pageId="15" pageNumber="216">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(N = 10), red dots =
|
||
<taxonomicName id="4C7E4D04FF8C4255BE890567BC1C08BD" box="[666,795,1913,1935]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="216" phylum="Chordata" rank="species" species="machadoi">
|
||
<emphasis id="B90AEA95FF8C4255BE890567BC1C08BD" box="[666,795,1913,1935]" italics="true" pageId="15" pageNumber="216">C. machadoi</emphasis>
|
||
</taxonomicName>
|
||
(N = 13), yellow dots = AMNH
|
||
<taxonomicName id="4C7E4D04FF8C4255B8760567BA1B08BD" box="[1125,1308,1913,1935]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="216" phylum="Chordata" rank="species" species="namakuiyus">
|
||
<emphasis id="B90AEA95FF8C4255B8760567BB7A08BC" box="[1125,1149,1914,1935]" italics="true" pageId="15" pageNumber="216">C.</emphasis>
|
||
cf.
|
||
<emphasis id="B90AEA95FF8C4255B8B00564BA1B08BD" box="[1187,1308,1913,1934]" italics="true" pageId="15" pageNumber="216">namakuiyus</emphasis>
|
||
</taxonomicName>
|
||
collected on the 1925 Vernay expedition (N = 48), purple star =
|
||
<taxonomicName id="4C7E4D04FF8C4255BEBB0585BC4C089E" box="[680,843,1943,1965]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="216" phylum="Chordata" rank="species" species="machadoi">
|
||
<emphasis id="B90AEA95FF8C4255BEBB0585BDC7089E" box="[680,704,1944,1965]" italics="true" pageId="15" pageNumber="216">C.</emphasis>
|
||
cf.
|
||
<emphasis id="B90AEA95FF8C4255BEF40585BC4C089E" box="[743,843,1944,1965]" italics="true" pageId="15" pageNumber="216">machadoi</emphasis>
|
||
</taxonomicName>
|
||
(TM 57561) from northern Namibia. Polygons with solid lines show morphospace of adult specimens, those with dotted lines represent morphospace of adults + juveniles.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C364650CFF93424ABCD4033ABD7D0EBB" pageId="16" pageNumber="217" type="etymology">
|
||
<paragraph id="8BC13687FF93424ABCD4033ABD7D0EBB" blockId="16.[151,1437,151,1005]" pageId="16" pageNumber="217">
|
||
<emphasis id="B90AEA95FF93424ABCD4033ABE490E73" bold="true" box="[199,334,295,320]" pageId="16" pageNumber="217">Etymology.</emphasis>
|
||
The specific epithet “
|
||
<taxonomicName id="4C7E4D04FF93424ABE430335BDD10E73" box="[592,726,296,320]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="217" phylum="Chordata" rank="species" species="namakuiyus">
|
||
<emphasis id="B90AEA95FF93424ABE430335BDD10E73" box="[592,726,296,320]" italics="true" pageId="16" pageNumber="217">namakuiyus</emphasis>
|
||
</taxonomicName>
|
||
” is the masculine latinised form of namakuiya, which means “thorny” in the Herero language, referring to the sharp spines on the limbs and tail of this species. Suggested common name: Kaokoveld Girdled Lizard.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C364650CFF93424ABCD40389BC1B0DB6" pageId="16" pageNumber="217" type="distribution">
|
||
<paragraph id="8BC13687FF93424ABCD40389BCE80EE3" blockId="16.[151,1437,151,1005]" pageId="16" pageNumber="217">
|
||
<emphasis id="B90AEA95FF93424ABCD40389BE670E9E" bold="true" box="[199,352,404,429]" pageId="16" pageNumber="217">Distribution.</emphasis>
|
||
Currently, the new species is known only from the south and central parts of Namibe Province,
|
||
<collectingCountry id="F3697617FF93424ABC8403AABFEB0EE3" box="[151,236,439,464]" name="Angola" pageId="16" pageNumber="217">Angola</collectingCountry>
|
||
, southwest of the Leba escarpment, at elevations of
|
||
<quantity id="4C869B62FF93424ABF2703AABCB40EFC" box="[820,947,439,464]" metricMagnitude="2" metricUnit="m" metricValue="3.62" metricValueMax="5.09" metricValueMin="2.15" pageId="16" pageNumber="217" unit="m" value="362.0" valueMax="509.0" valueMin="215.0">215–509 m</quantity>
|
||
a.s.l.
|
||
</paragraph>
|
||
<paragraph id="8BC13687FF93424ABCD403C1BC1B0DB6" blockId="16.[151,1437,151,1005]" pageId="16" pageNumber="217">
|
||
<emphasis id="B90AEA95FF93424ABCD403C1BE460EC6" bold="true" box="[199,321,476,501]" pageId="16" pageNumber="217">Localities.</emphasis>
|
||
<collectingCountry id="F3697617FF93424ABD4303C1BEA00EC6" box="[336,423,476,501]" name="Angola" pageId="16" pageNumber="217">Angola</collectingCountry>
|
||
: AMNH R47274–47321. Namibe Province, outskirts of Caraculo: CAS 254912–14 [
|
||
<geoCoordinate id="EE4A5040FF93424ABC8D03E2BE2F0D2B" box="[158,296,511,536]" direction="south" orientation="latitude" pageId="16" pageNumber="217" precision="1" value="-15.0165">15°0'59.4"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE4A5040FF93424ABD2F001DBED20D2B" box="[316,469,512,536]" direction="east" orientation="longitude" pageId="16" pageNumber="217" precision="1" value="12.642028">12°38'31.3"E</geoCoordinate>
|
||
;
|
||
<quantity id="4C869B62FF93424ABDFA03E2BD3F0D24" box="[489,568,511,536]" metricMagnitude="2" metricUnit="m" metricValue="5.029999999999999" pageId="16" pageNumber="217" unit="m" value="503.0">503 m</quantity>
|
||
elevation], CAS 256529 [
|
||
<geoCoordinate id="EE4A5040FF93424ABF6F03E2BB010D2B" box="[892,1030,511,536]" direction="south" orientation="latitude" pageId="16" pageNumber="217" precision="1" value="-15.015917">15°0'57.3"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE4A5040FF93424AB80903E2BBB40D2B" box="[1050,1203,511,536]" direction="east" orientation="longitude" pageId="16" pageNumber="217" precision="1" value="12.642388">12°38'32.6"E</geoCoordinate>
|
||
;
|
||
<quantity id="4C869B62FF93424AB8D503E2BA120D24" box="[1222,1301,511,536]" metricMagnitude="2" metricUnit="m" metricValue="5.09" pageId="16" pageNumber="217" unit="m" value="509.0">509 m</quantity>
|
||
elevation]; outskirts of Pico Azevedo: CAS 254754–55, 256530, 256531) [
|
||
<geoCoordinate id="EE4A5040FF93424ABF6D0039BB110D0E" box="[894,1046,548,573]" direction="south" orientation="latitude" pageId="16" pageNumber="217" precision="1" value="-15.475888">15°28'33.2"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE4A5040FF93424AB8350039BBB80D0E" box="[1062,1215,548,573]" direction="east" orientation="longitude" pageId="16" pageNumber="217" precision="1" value="12.462694">12°27'45.7"E</geoCoordinate>
|
||
;
|
||
<quantity id="4C869B62FF93424AB8DC0039BA1D0D0F" box="[1231,1306,548,572]" metricMagnitude="2" metricUnit="m" metricValue="4.21" pageId="16" pageNumber="217" unit="m" value="421.0">421 m</quantity>
|
||
elevation]; Road from Namibe to Omahua lodge PEM R18005 [
|
||
<geoCoordinate id="EE4A5040FF93424ABEF4005ABC870D53" box="[743,896,583,608]" direction="south" orientation="latitude" pageId="16" pageNumber="217" precision="1" value="-15.996806">15°59'48.5"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE4A5040FF93424ABF9F005ABB220D53" box="[908,1061,583,608]" direction="east" orientation="longitude" pageId="16" pageNumber="217" precision="1" value="12.406833">12°24'24.6"E</geoCoordinate>
|
||
;
|
||
<quantity id="4C869B62FF93424AB8220055BB700D6C" box="[1073,1143,584,608]" metricMagnitude="2" metricUnit="m" metricValue="3.08" pageId="16" pageNumber="217" unit="m" value="308.0">308 m</quantity>
|
||
elevation];
|
||
<quantity id="4C869B62FF93424AB8ED0055BA420D53" box="[1278,1349,583,608]" metricMagnitude="4" metricUnit="m" metricValue="4.9" pageId="16" pageNumber="217" unit="km" value="49.0">49 km</quantity>
|
||
ESE of Tombua: TM 40430 (
|
||
<geoCoordinate id="EE4A5040FF93424ABD9B0071BEE50DB6" box="[392,482,620,645]" direction="south" orientation="latitude" pageId="16" pageNumber="217" precision="925" value="-15.883333">15°53'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE4A5040FF93424ABDFD0071BD4D0DB6" box="[494,586,620,645]" direction="east" orientation="longitude" pageId="16" pageNumber="217" precision="925" value="12.266666">12°16'E</geoCoordinate>
|
||
;
|
||
<quantity id="4C869B62FF93424ABE450071BD9A0DB7" box="[598,669,620,645]" metricMagnitude="2" metricUnit="m" metricValue="2.15" pageId="16" pageNumber="217" unit="m" value="215.0">215 m</quantity>
|
||
elevation).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C364650CFF93424ABCD40092BEA40CDE" pageId="16" pageNumber="217" type="description">
|
||
<paragraph id="8BC13687FF93424ABCD40092BEA40CDE" blockId="16.[151,1437,151,1005]" pageId="16" pageNumber="217">
|
||
<emphasis id="B90AEA95FF93424ABCD40092BD580D9B" bold="true" box="[199,607,655,680]" pageId="16" pageNumber="217">Habitat and natural history notes.</emphasis>
|
||
This species is found in gently sloping crevices of granite outcrops in the arid Kaokoveld. When approached, specimens retreat into their fissure as far as possible, wedging themselves head-first and protecting their head and flanks with their spiny tails.
|
||
<collectingCountry id="F3697617FF93424ABFB400C5BCC10DC3" box="[935,966,728,752]" name="American Samoa" pageId="16" pageNumber="217">As</collectingCountry>
|
||
with all other cordylines,
|
||
<taxonomicName id="4C7E4D04FF93424AB8E700C5BA9B0DC3" box="[1268,1436,728,752]" class="Reptilia" family="Cordylidae" genus="Cordylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="217" phylum="Chordata" rank="species" species="namakuiyus" status="sp. nov.">
|
||
<emphasis id="B90AEA95FF93424AB8E700C5BA9B0DC3" box="[1268,1436,728,752]" italics="true" pageId="16" pageNumber="217">C. namakuiyus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23957EEFF93424ABC8400E1BFF70C26" box="[151,240,764,789]" pageId="16" pageNumber="217" rank="species">
|
||
<emphasis id="B90AEA95FF93424ABC8400E1BFF70C26" bold="true" box="[151,240,764,789]" pageId="16" pageNumber="217">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is viviparous; the
|
||
<typeStatus id="54C58825FF93424ABDAE00E1BD180C26" box="[445,543,764,789]" pageId="16" pageNumber="217" type="holotype">holotype</typeStatus>
|
||
contained two large fetuses (SVL 49.5% of the
|
||
<typeStatus id="54C58825FF93424AB82500E1BB9A0C26" box="[1078,1181,764,789]" pageId="16" pageNumber="217" type="holotype">holotype</typeStatus>
|
||
). The fetuses appear to be approaching full-term, suggesting a spring/early summer parturition period. Although these fetuses lack any body osteoderms, the slightly larger neonates (SVL 55.6–56.4% of the
|
||
<typeStatus id="54C58825FF93424ABFE20159BB520C6E" box="[1009,1109,836,861]" pageId="16" pageNumber="217" type="holotype">holotype</typeStatus>
|
||
) possess significant dermal ossification, suggesting that the young rapidly accumulate osteodermal armour following parturition. One of the Pico Azevedo specimens was found occupying the same rock crack as an adult
|
||
<taxonomicName id="4C7E4D04FF93424AB84C0191BE0D0CFB" authority="Schmidt" authorityName="Schmidt" class="Reptilia" family="Gekkonidae" genus="Chondrodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="217" phylum="Chordata" rank="species" species="pulitzerae">
|
||
<emphasis id="B90AEA95FF93424AB84C0191BA9B0C97" box="[1119,1436,908,932]" italics="true" pageId="16" pageNumber="217">Chondrodactylus pulitzerae</emphasis>
|
||
(Schmidt)
|
||
</taxonomicName>
|
||
, and PEM R18005 from Iona National Park inhabited the same crack as an adult
|
||
<taxonomicName id="4C7E4D04FF93424AB8CF01ADBE980CDE" authority="Loveridge" authorityName="Loveridge" class="Reptilia" family="Gekkonidae" genus="Chondrodactylus" kingdom="Animalia" order="Squamata" pageId="16" pageNumber="217" phylum="Chordata" rank="species" species="fitzsimonsi">
|
||
<emphasis id="B90AEA95FF93424AB8CF01ADBE080CDF" italics="true" pageId="16" pageNumber="217">Chondrodactylus fitzsimonsi</emphasis>
|
||
(Loveridge)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |