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<document id="767ECEA43537B62878F23E25CF9CF5EE" ID-CLB-Dataset="29367" ID-DOI="10.11646/zootaxa.4486.4.1" ID-GBIF-Dataset="de2f897b-3d16-4bcd-99a7-fb46a61b5bd1" ID-ISSN="1175-5326" ID-Zenodo-Dep="1442168" ID-ZooBank="7652EE8A-437B-4A61-ACF6-4A3BF6553808" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538406769457" checkinUser="plazi" docAuthor="Vondráček, Dominik, Hadjiconstantis, Michael &amp; Šípek, Petr" docDate="2018" docId="03CA879D6224661A43A7FA99FE6FFF03" docLanguage="en" docName="zootaxa.4486.4.1.pdf" docOrigin="Zootaxa 4486 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oxythyrea tripolitana Reitter 1891" docType="treatment" docVersion="7" lastPageNumber="421" masterDocId="FFF3FFE56230660F4330FF8FFF93FFFB" masterDocTitle="Immature stages of the genus Oxythyrea (Coleoptera: Scarabaeidae: Cetoniinae) with a key to third instar larvae, and notes on the biology of the genus" masterLastPageNumber="434" masterPageNumber="401" pageNumber="421" updateTime="1698562072066" updateUser="ExternalLinkService">
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<mods:title id="9AC3AC12583CA6EE1FDE4540A5350A5D">Immature stages of the genus Oxythyrea (Coleoptera: Scarabaeidae: Cetoniinae) with a key to third instar larvae, and notes on the biology of the genus</mods:title>
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<mods:namePart id="CB5C78ABE4D0652CDF7668F5C8B86080">Vondráček, Dominik</mods:namePart>
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<mods:namePart id="EF0EEDD644955C6C9175D405C8CBA668">Hadjiconstantis, Michael</mods:namePart>
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<mods:namePart id="14C4ACBC51AD40B2A7E29D2E2D2BAB5F">Šípek, Petr</mods:namePart>
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<mods:title id="1C7965F4C2B4A3ED3A9986EA68499A7B">Zootaxa</mods:title>
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<treatment id="03CA879D6224661A43A7FA99FE6FFF03" ID-DOI="http://doi.org/10.5281/zenodo.5970925" ID-GBIF-Taxon="148425486" ID-Zenodo-Dep="5970925" LSID="urn:lsid:plazi:treatment:03CA879D6224661A43A7FA99FE6FFF03" httpUri="http://treatment.plazi.org/id/03CA879D6224661A43A7FA99FE6FFF03" lastPageId="21" lastPageNumber="421" pageId="20" pageNumber="421">
<subSubSection id="C37965006224661B43A7FA99FC8AFAA9" pageId="20" pageNumber="421" type="nomenclature">
<paragraph id="8BDC368B6224661B43A7FA99FDD3FACB" blockId="20.[151,793,1302,1362]" box="[151,576,1302,1328]" pageId="20" pageNumber="421">
<heading id="D09481E76224661B43A7FA99FDD3FACB" bold="true" box="[151,576,1302,1328]" fontSize="11" level="1" pageId="20" pageNumber="421" reason="1">
<taxonomicName id="4C634D086224661B43A7FA99FDD3FACB" ID-CoL="92YTZ" authority="Reitter, 1891" authorityName="Reitter" authorityYear="1891" box="[151,576,1302,1328]" class="Insecta" family="Scarabaeidae" genus="Oxythyrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="421" phylum="Arthropoda" rank="species" species="tripolitana">
<emphasis id="B917EA996224661B43A7FA99FDD3FACB" bold="true" box="[151,576,1302,1328]" pageId="20" pageNumber="421">
<emphasis id="B917EA996224661B43A7FA99FE09FACB" bold="true" box="[151,410,1302,1328]" italics="true" pageId="20" pageNumber="421">Oxythyrea tripolitana</emphasis>
Reitter, 1891
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BDC368B6224661B43A7FAB6FC8AFAA9" blockId="20.[151,793,1302,1362]" box="[151,793,1337,1362]" pageId="20" pageNumber="421">
<emphasis id="B917EA996224661B43A7FAB6FF33FAA9" bold="true" box="[151,160,1337,1362]" pageId="20" pageNumber="421">(</emphasis>
<figureCitation id="13582A0E6224661B4390FAB6FF6BFAA9" box="[160,248,1337,1362]" captionStart="FIGURE 3" captionStartId="6.[151,250,1961,1983]" captionTargetBox="[186,1402,493,1929]" captionTargetId="figure@6.[166,1402,480,1930]" captionTargetPageId="6" captionText="FIGURE 3ai. Habitus of third instar Oxythyrea larvae. a, O. abigail; b, O. albopicta; c, O. cinctella; d, O. dulcis; e, O. funesta; f, O. noemi; g, O. pantherina; h, O. subcalva; i, O. tripolitana." httpUri="https://zenodo.org/record/1442174/files/figure.png" pageId="20" pageNumber="421">
Figs.
<date id="FFDD104B6224661B43EEFAB5FF6BFAA9" box="[222,248,1338,1362]" pageId="20" pageNumber="421">3I</date>
</figureCitation>
;
<figureCitation id="13582A0E6224661B4234FAB5FED3FAAA" box="[260,320,1338,1362]" captionStart="FIGURE 4" captionStartId="7.[151,250,1906,1928]" captionTargetBox="[192,1405,223,1831]" captionTargetId="figure@7.[184,1408,193,1886]" captionTargetPageId="7" captionText="FIGURE 4ar. Cranium of third instar Oxythyrea larvae (ai) and antenna of third instar Oxythyrea larvae, dorsal aspect (jr). a, O. abigail; b, O. albopicta; c, O. cinctella; d, O. dulcis; e, O. funesta; f, O. noemi; g, O. pantherina; h, O. subcalva; i, O. tripolitana; j, O. abigail; k, O. albopicta; l, O. cinctella; m, O. dulcis; n, O. funesta; o, O. noemi; p, O. pantherina; q, O. subcalva; r, O. tripolitana. Scale bars = 1 mm (ai); 0.5 mm (jr)." httpUri="https://zenodo.org/record/1442176/files/figure.png" pageId="20" pageNumber="421">
<date id="FFDD104B6224661B4234FAB5FE8DFAA9" box="[260,286,1338,1362]" pageId="20" pageNumber="421">4I</date>
, R
</figureCitation>
;
<figureCitation id="13582A0E6224661B427BFAB6FE14FAAA" box="[331,391,1337,1362]" captionStart="FIGURE 5" captionStartId="8.[151,250,1821,1843]" captionTargetBox="[169,1436,274,1741]" captionTargetId="figure@8.[151,1436,246,1777]" captionTargetPageId="8" captionText="FIGURE 5ar. Epipharynx of third instar Oxythyrea larvae (ai) and antenna of third instar Oxythyrea larvae, ventral aspect (jr). a, O. abigail; b, O. albopicta; c, O. cinctella; d, O. dulcis; e, O. funesta; f, O. noemi; g, O. pantherina; h, O. subcalva; i, O. tripolitana; j, O. abigail; k, O. albopicta; l, O. cinctella; m, O. dulcis; n, O. funesta; o, O. noemi; p, O. pantherina; q, O. subcalva; r, O. tripolitana. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/1442178/files/figure.png" pageId="20" pageNumber="421">
<date id="FFDD104B6224661B427BFAB6FEF6FAA9" box="[331,357,1337,1362]" pageId="20" pageNumber="421">5I</date>
, R
</figureCitation>
;
<figureCitation id="13582A0E6224661B42A2FAB6FE5DFAA9" box="[402,462,1337,1362]" captionStart="FIGURE 6" captionStartId="9.[151,250,1908,1930]" captionTargetBox="[162,1426,193,1888]" captionTargetId="figure@9.[162,1426,193,1888]" captionTargetPageId="9" captionText="FIGURE 6aq. Maxillo-labial complex of third instar Oxythyrea larvae, dorsal view (ai) and maxillary stridulatory teeth of third instar Oxythyrea larvae (jq). a, O. abigail; b, O. albopicta; c, O. cinctella; d, O. dulcis; e, O. funesta; f, O. noemi; g, O. pantherina; h, O. subcalva; i, O. tripolitana; j, O. abigail; k, O. albopicta; l, O. cinctella; m, O. dulcis; n, O. funesta; o, O. noemi; p, O. subcalva; q, O. tripolitana. Scale bars = 0.5 mm (ai)." httpUri="https://zenodo.org/record/1442180/files/figure.png" pageId="20" pageNumber="421">
<date id="FFDD104B6224661B42A2FAB6FE38FAA9" box="[402,427,1337,1362]" pageId="20" pageNumber="421">6I</date>
, Q
</figureCitation>
;
<figureCitation id="13582A0E6224661B42EAFAB5FD86FAAA" box="[474,533,1338,1362]" captionStart="FIGURE 8" captionStartId="11.[151,250,1908,1930]" captionTargetBox="[208,1371,223,1864]" captionTargetId="figure@11.[178,1402,193,1884]" captionTargetPageId="11" captionText="FIGURE 8al. Left mandible of third instar Oxythyrea larvae, dorsal, medial and ventral view (ac), right mandible of third instar Oxythyrea larvae: ventral, medial and dorsal view (df), maxillo-labial complex of third instar Oxythyrea larvae, ventral view (gi) and metathoracic leg (jl). a, O. pantherina; b, O. subcalva; c, O. tripolitana; d, O. abigail; e, O. albopicta; f, O. cinctella; g, O. pantherina; h, O. subcalva; i, O. tripolitana; j, O. abigail; k, O. albopicta; l, O. cinctella. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/1442186/files/figure.png" pageId="20" pageNumber="421">8C, I</figureCitation>
;
<figureCitation id="13582A0E6224661B4111FAB5FDC4FAAA" box="[545,599,1338,1362]" captionStart="FIGURE 9" captionStartId="12.[151,250,1928,1950]" captionTargetBox="[202,1398,209,1902]" captionTargetId="figure@12.[175,1419,193,1911]" captionTargetPageId="12" captionText="FIGURE 9al. Right mandible of third instar Oxythyrea larvae, ventral, medial and dorsal view (af) and metathoracic leg of third instar Oxythyrea larvae (gl). a, O. dulcis; b, O. funesta; c, O. noemi; d, O. pantherina; e, O. subcalva; f, O. tripolitana; g, O. dulcis; h, O. funesta; i, O. noemi; j, O. pantherina; k, O. subcalva; l, O. tripolitana. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/1442188/files/figure.png" pageId="20" pageNumber="421">9F, I</figureCitation>
;
<figureCitation id="13582A0E6224661B4153FAB5FD3FFAAA" box="[611,684,1338,1362]" captionStart="FIGURE 10" captionStartId="13.[151,250,1900,1922]" captionTargetBox="[224,1362,193,1870]" captionTargetId="figure@13.[224,1362,193,1870]" captionTargetPageId="13" captionText="FIGURE 10ar. Last abdominal segment of third instar Oxythyrea larvae, ventral view (ai) and last abdominal segment of third instar Oxythyrea larvae, detail of raster (jr). a, O. abigail; b, O. albopicta; c, O. cinctella; d, O. dulcis; e, O. funesta; f, O. noemi; g, O. pantherina; h, O. subcalva; i, O. tripolitana; j, O. abigail; k, O. albopicta; l, O. cinctella; m, O. dulcis; n, O. funesta; o, O. noemi; p, O. pantherina; q, O. subcalva; r, O. tripolitana. Scale bars = 1 mm (ai)." httpUri="https://zenodo.org/record/1442190/files/figure.png" pageId="20" pageNumber="421">
<date id="FFDD104B6224661B4153FAB5FD19FAA9" box="[611,650,1338,1362]" pageId="20" pageNumber="421" value="2010-01">10I</date>
, R
</figureCitation>
;
<figureCitation id="13582A0E6224661B4187FAB5FD4CFAA9" box="[695,735,1338,1362]" captionStart="FIGURE 11" captionStartId="14.[151,250,1903,1925]" captionTargetBox="[161,1436,249,1852]" captionTargetId="figure@14.[151,1436,193,1881]" captionTargetPageId="14" captionText="FIGURE 11am. Last abdominal segment of third instar Oxythyrea larvae, lateral view (ai) and last abdominal segment of third instar Oxythyrea larvae, posterior view (jm). a, O. abigail; b, O. albopicta; c, O. cinctella; d, O. dulcis; e, O. funesta; f, O. noemi; g, O. pantherina; h, O. subcalva; i, O. tripolitana; j, O. albopicta; k, O. cinctella; l, O. dulcis; m, O. funesta. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/1442192/files/figure.png" pageId="20" pageNumber="421">
<date id="FFDD104B6224661B4187FAB5FD4CFAA9" box="[695,735,1338,1362]" pageId="20" pageNumber="421" value="2011-01">11I</date>
</figureCitation>
;
<figureCitation id="13582A0E6224661B41DAFAB5FC82FAA9" box="[746,785,1338,1362]" captionStart="FIGURE 12" captionStartId="15.[151,250,1504,1526]" captionTargetBox="[171,1407,184,1445]" captionTargetId="figure@15.[151,1436,167,1483]" captionTargetPageId="15" captionText="FIGURE 12ao. Thoracic spiracles of third instar Oxythyrea larvae, entire view (ai) and detail of sieve plate (jo). a, O. abigail; b, O. albopicta; c, O. cinctella; d, O. dulcis; e, O. funesta; f, O. noemi; g, O. pantherina; h, O. subcalva; i, O. tripolitana; j, O. abigail; k, O. albopicta; l, O. cinctella; m, O. dulcis; n, O. funesta; o, O. subcalva. Scale bars = 0.1 mm (ai)." httpUri="https://zenodo.org/record/1442194/files/figure.png" pageId="20" pageNumber="421">
<date id="FFDD104B6224661B41DAFAB5FC82FAA9" box="[746,785,1338,1362]" pageId="20" pageNumber="421" value="2012-01">12I</date>
</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37965006224661A43A7FA0EFE6FFF03" lastPageId="21" lastPageNumber="422" pageId="20" pageNumber="421" type="materials_examined">
<paragraph id="8BDC368B6224661B43A7FA0EFEA9F9FD" blockId="20.[151,1437,1409,2010]" pageId="20" pageNumber="421">
<emphasis id="B917EA996224661B43A7FA0EFEF6FA61" bold="true" box="[151,357,1409,1434]" pageId="20" pageNumber="421">Material studied.</emphasis>
3 L3 reared ex adults collected at:
<collectingCountry id="F374761B6224661B41C3FA0EFCDAFA61" box="[755,841,1409,1434]" name="Tunisia" pageId="20" pageNumber="421">Tunisia</collectingCountry>
,
<collectingCountry id="F374761B6224661B4065FA0EFC00FA61" box="[853,915,1409,1434]" name="Tunisia" pageId="20" pageNumber="421">Jerba</collectingCountry>
, Rass Rmel, sand dunes, N°33.883, E°10.933,
<date id="FFDD104B6224661B43A7FA2AFE97FA45" box="[151,260,1445,1470]" pageId="20" pageNumber="421" value="2006-06-07">7.vi.2006</date>
, J. Batelka, J. Straka lgt.; 12 L3 reared ex adults collected at:
<collectingCountry id="F374761B6224661B4088FA2AFB9DFA45" box="[952,1038,1445,1470]" name="Tunisia" pageId="20" pageNumber="421">Tunisia</collectingCountry>
,
<collectingCountry id="F374761B6224661B4729FA2AFBFBFA45" box="[1049,1128,1445,1470]" name="Tunisia" pageId="20" pageNumber="421">Djerba</collectingCountry>
,
<collectingCountry id="F374761B6224661B4744FA2AFB53FA45" box="[1140,1216,1445,1470]" name="Tunisia" pageId="20" pageNumber="421">Djerba</collectingCountry>
island,
<quantity id="4C9B9B6E6224661B4628FA29FAC2FA45" box="[1304,1361,1445,1470]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="20" pageNumber="421" unit="km" value="9.0">9 km</quantity>
NE of Houmt Souk, Rass Rmel sand spit, ca
<quantity id="4C9B9B6E6224661B417CFA45FDE8FA1A" box="[588,635,1482,1506]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="421" unit="m" value="1.0">1 m</quantity>
, N°33.883, E°10.933,
<date id="FFDD104B6224661B404CFA46FBBAFA19" box="[892,1065,1481,1506]" pageId="20" pageNumber="421" value="2014-03-25" valueMax="2014-03-26" valueMin="2014-03-25">25.26.iii.2014</date>
, O. Hillert, D. Král, P. Šípek, D. Vondráček lgt.
</paragraph>
<paragraph id="8BDC368B6224661A43F7F99EFE6FFF03" blockId="20.[151,1437,1409,2010]" lastBlockId="21.[151,1436,151,248]" lastPageId="21" lastPageNumber="422" pageId="20" pageNumber="421">
Body length
<quantity id="4C9B9B6E6224661B4251F99DFE71F9D2" box="[353,482,1554,1578]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" metricValueMax="2.4" metricValueMin="2.0" pageId="20" pageNumber="421" unit="mm" value="22.0" valueMax="24.0" valueMin="20.0">2024 mm</quantity>
. Cranium: width
<quantity id="4C9B9B6E6224661B4182F99EFCDDF9D2" box="[690,846,1553,1578]" metricMagnitude="-3" metricUnit="m" metricValue="2.64" metricValueMax="2.68" metricValueMin="2.6" pageId="20" pageNumber="421" unit="mm" value="2.64" valueMax="2.68" valueMin="2.6">2.62.68 mm</quantity>
, height
<quantity id="4C9B9B6E6224661B4080F99EFBC8F9D2" box="[944,1115,1553,1578]" metricMagnitude="-3" metricUnit="m" metricValue="1.5699999999999998" metricValueMax="1.67" metricValueMin="1.47" pageId="20" pageNumber="421" unit="mm" value="1.5699999999999998" valueMax="1.67" valueMin="1.47">1.471.67 mm</quantity>
. Cranium brown with faint sculpture; slightly elongate in shape, with its dorsal margin more or less convex at its entire length (length of dorsal straight portion of cranial between ¼ and ⅓ of maximum width of cranium). Chaetotaxy as in
<tableCitation id="C6E103306224661B4782F9D6FA96F989" box="[1202,1285,1625,1650]" captionStart="TABLE 1" captionStartId="5.[147,235,556,579]" captionText="TABLE 1. Cranial chaetotaxy of last instar Oxythyrea larvae" pageId="20" pageNumber="421">Table 1</tableCitation>
. Epipharynx: Zygum with 1215 stout conical setae and 57 similarly shaped setae at its proximal base. Chaetoparia with 6873 and 3247 setae on right and left side respectively. Acanthoparia with 68 falcate setae. Stridulatory area of mandibles with 1317 transverse ridges, ridges in apical third only slightly narrower than remaining ridges. If apical scissorial tooth of left mandible falcate, than its size not twice large as size of remaining. Transverse proximal row of setae and sensilla on ligula composed of 78 stout lanceolate setae and campaniform sensilla combined (mostly only lanceolate setae present). Setae of thoracic legs stout (spine-like) and short to slender hairlike. Respiratory plate of spiracle with 1317 holes across diameter. Raster monostichous, sometimes with few scattered irregular pali around the main row; with 1823 pali on each side, septula elliptic more than three times longer than wide. Tegites variable in size from half of the length of palus to four times as long as palus (longer tegites sparse). Tegilla densely setose; average distance between two setae equals from one to two times the length of palus. Tegilla fused into single tegillum, campus absent. Dorsal parts of last abdominal segment densely setose, covered with numerous short and medium long (up to three times length of palus) setae, without ring of prominent long setae or bald circular field.
</paragraph>
</subSubSection>
</treatment>
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