176 lines
21 KiB
XML
176 lines
21 KiB
XML
<document id="08C7EEC289E4574F88A825B667E7E434" ID-CLB-Dataset="113070" ID-DOI="10.1093/zoolinnean/zlab043" ID-GBIF-Dataset="e070d52f-d008-4f08-988a-a8b17c27418f" ID-ISSN="0024-4082" ID-Zenodo-Dep="6959516" ID-ZooBank="1C89556A-12CB-43B7-9B49-E02EFF1543D3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1653991014928" checkinUser="plazi" docAuthor="Medrano, Miguel, Kury, Adriano Brilhante & Mendes, Amanda Cruz" docDate="2022" docId="03CA878CFFBAFFCAFCB0FC6EFD982369" docLanguage="en" docName="zlab043.pdf" docOrigin="Zoological Journal of the Linnean Society 195" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Libitiinae Medrano, Kury & Mendes, 2022, SUBFAM. NOV." docType="treatment" docUuid="8EB3A791-F22F-4C24-B9DF-498E41DFF74C" docUuidSource="ZooBank" docVersion="8" lastPageNumber="601" masterDocId="FFF3FFF4FFB5FFDAFFFEFFB7FFA72171" masterDocTitle="Morphology-based cladistics splinters the century-old dichotomy of the pied harvestmen (Arachnida: Gonyleptoidea: Cosmetidae)" masterLastPageNumber="672" masterPageNumber="585" pageNumber="600" updateTime="1699338404186" updateUser="ExternalLinkService">
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<mods:title id="5DCC09B8C8962A23714A66482E977733">Morphology-based cladistics splinters the century-old dichotomy of the pied harvestmen (Arachnida: Gonyleptoidea: Cosmetidae)</mods:title>
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<mods:namePart id="9D7681CF471D495E38E5BBCC45EF3483">Medrano, Miguel</mods:namePart>
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<mods:namePart id="D6B02963336B9A27AB0415F290A33A3E">Kury, Adriano Brilhante</mods:namePart>
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</mods:name>
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<mods:name id="432C58C26C51035D2F27067DFB6E7C60" type="personal">
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<mods:namePart id="0304481BE8894AC6D632C58D63A78557">Mendes, Amanda Cruz</mods:namePart>
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<mods:title id="B565F10C7545AC68C15CF97E2F8D4FF6">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="EDAFD6FE8142EB9DE1A342CD8B9E8085">2022</mods:date>
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<mods:number id="F0BA240FAB83650FE86E5BB2F2FE37F7">2021-08-30</mods:number>
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<mods:number id="666C7D8398AE6511E27010582C84FC32">195</mods:number>
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<treatment id="03CA878CFFBAFFCAFCB0FC6EFD982369" ID-DOI="http://doi.org/10.5281/zenodo.6963827" ID-GBIF-Taxon="198579180" ID-Zenodo-Dep="6963827" LSID="urn:lsid:zoobank.org:act:8EB3A791-F22F-4C24-B9DF-498E41DFF74C" httpUri="http://treatment.plazi.org/id/03CA878CFFBAFFCAFCB0FC6EFD982369" lastPageId="16" lastPageNumber="601" pageId="15" pageNumber="600">
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<subSubSection id="C3796511FFBAFFD5FCB0FC6EFB2C257E" pageId="15" pageNumber="600" type="nomenclature">
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<paragraph id="8BDC369AFFBAFFD5FCB0FC6EFB2C257E" blockId="15.[846,1423,985,1039]" pageId="15" pageNumber="600">
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<heading id="D09481F6FFBAFFD5FCB0FC6EFB2C257E" centered="true" fontSize="9" level="2" pageId="15" pageNumber="600" reason="3">
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<taxonomicName id="4C634D19FFBAFFD5FCB0FC6EFA862283" authorityName="Medrano, Kury, Mendes" authorityYear="2022" box="[846,1313,985,1010]" class="Arachnida" family="Cosmetidae" kingdom="Animalia" order="Opiliones" pageId="15" pageNumber="600" phylum="Arthropoda" rank="subFamily" status="subfam. nov." subFamily="Libitiinae">
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<emphasis id="B917EA88FFBAFFD5FCB0FC6EFC742280" bold="true" box="[846,979,985,1009]" pageId="15" pageNumber="600">LIBITIINAE</emphasis>
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MEDRANO, KURY & MENDES
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</taxonomicName>
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<emphasis id="B917EA88FFBAFFD5FAD6FC6BFB2C257E" bold="true" pageId="15" pageNumber="600">
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<taxonomicNameLabel id="A22457F3FFBAFFD5FAD6FC6BFB2C257E" pageId="15" pageNumber="600" rank="subFamily">SUBFAM. NOV.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3796511FFBAFFD5FCC5FB97FA3B2525" pageId="15" pageNumber="600" type="description">
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<paragraph id="8BDC369AFFBAFFD5FCC5FB97FA3B2525" blockId="15.[827,1443,1056,1108]" pageId="15" pageNumber="600">
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<emphasis id="B917EA88FFBAFFD5FCC5FB97FB2F2547" box="[827,1160,1056,1078]" italics="true" pageId="15" pageNumber="600">Zoobank registration:</emphasis>
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urn: lsid: zoobank. org:act:
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<uuid id="FFC50C4FFFBAFFD5FC74FB88FA3B2525" box="[906,1436,1087,1108]" pageId="15" pageNumber="600">8EB3A791-F22F-4C24-B9DF-498E41DFF74C</uuid>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3796511FFBAFFD5FCC5FBC9FA87270F" pageId="15" pageNumber="600" type="discussion">
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<paragraph id="8BDC369AFFBAFFD5FCC5FBC9FA87270F" blockId="15.[827,1444,1149,1662]" pageId="15" pageNumber="600">
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<emphasis id="B917EA88FFBAFFD5FCC5FBC9FC1525E2" box="[827,946,1150,1171]" italics="true" pageId="15" pageNumber="600">Comment:</emphasis>
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The monophylum herein labelled ‘clade A’ and described as the new subfamily
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<taxonomicName id="4C634D19FFBAFFD5FADAFB2BFA3925C3" box="[1316,1438,1180,1202]" class="Arachnida" family="Cosmetidae" kingdom="Animalia" order="Opiliones" pageId="15" pageNumber="600" phylum="Arthropoda" rank="subFamily" subFamily="Libitiinae">Libitiinae</taxonomicName>
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, includes four terminals of our analysis, which may be extrapolated to ten species.
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<taxonomicName id="4C634D19FFBAFFD5FB3EFB6EFBA5247C" authority="Sorensen, 1932" authorityName="Sorensen" authorityYear="1932" class="Arachnida" family="Cosmetidae" genus="Libitia" kingdom="Animalia" order="Opiliones" pageId="15" pageNumber="600" phylum="Arthropoda" rank="species" species="bipunctata">
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<emphasis id="B917EA88FFBAFFD5FB3EFB6EFA04259F" box="[1216,1443,1241,1262]" italics="true" pageId="15" pageNumber="600">Libitia bipunctata</emphasis>
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Sørensen, 1932
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</taxonomicName>
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and *
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<taxonomicName id="4C634D19FFBAFFD5FBA7FB4FFCDC245D" authority="(Gervais, 1844)" baseAuthorityName="Gervais" baseAuthorityYear="1844" class="Arachnida" family="Cosmetidae" genus="Libitia" kingdom="Animalia" order="Opiliones" pageId="15" pageNumber="600" phylum="Arthropoda" rank="species" species="cordata">
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<emphasis id="B917EA88FFBAFFD5FBA7FB4FFA87247C" box="[1113,1312,1272,1293]" italics="true" pageId="15" pageNumber="600">Libitia cordata</emphasis>
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(
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<bibRefCitation id="EFF24B6BFFBAFFD5FAC6FB4FFCD4245D" author="Gervais P" pageId="15" pageNumber="600" pagination="94 - 131" refId="ref51474" refString="Gervais P. 1844. Aceres Phryneides, Scorpionides, Solpugides, Phalangides et Acarides; Diceres Epizoiques, Aphanipteres et Thysanoures. In: Walckenaer CA, ed. Org. Histoire naturelle des Insectes Apteres, Tome 3, viii + 476 pp. Paris: Librairie Encyclopedique de Roret, 94 - 131." type="book chapter" year="1844">Gervais, 1844</bibRefCitation>
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)
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</taxonomicName>
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here represent the genus
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<taxonomicName id="4C634D19FFBAFFD5FB4DFAA1FA3B245D" authority="Simon, 1879" authorityName="Simon" authorityYear="1879" box="[1203,1436,1302,1324]" class="Arachnida" family="Cosmetidae" genus="Libitia" kingdom="Animalia" order="Opiliones" pageId="15" pageNumber="600" phylum="Arthropoda" rank="genus">
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<emphasis id="B917EA88FFBAFFD5FB4DFAA1FAA4245A" box="[1203,1283,1302,1323]" italics="true" pageId="15" pageNumber="600">Libitia</emphasis>
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<bibRefCitation id="EFF24B6BFFBAFFD5FAF2FAA1FA3B245D" author="Simon E" box="[1292,1436,1302,1324]" pageId="15" pageNumber="600" pagination="183 - 241" refId="ref55071" refString="Simon E. 1879. Essai d'une classification des Opiliones Mecostethi. Remarques synonymiques et descriptions d'especes nouvelles. Premiere partie. Annales de la Societe Entomologique de Belgique 22: 183 - 241." type="journal article" year="1879">Simon, 1879</bibRefCitation>
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</taxonomicName>
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, which has two further species, all four of them from
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<collectingCountry id="F374760AFFBAFFD5FCC5FAE3FC15241B" box="[827,946,1364,1386]" name="Colombia" pageId="15" pageNumber="600">Colombia</collectingCountry>
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(
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<bibRefCitation id="EFF24B6BFFBAFFD5FC37FAE3FB7C2418" author="Medrano M & Azara LN & Kury AB" box="[969,1243,1364,1386]" pageId="15" pageNumber="600" pagination="1 - 25" refId="ref53087" refString="Medrano M, Azara LN, Kury AB. 2020. The short-legged Andean cosmetids revisited: the genus Libitia Simon, 1879 with description of two new species (Opiliones, Cosmetidae). European Journal of Taxonomy 634: 1 - 25." type="journal article" year="2020">
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Medrano
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<emphasis id="B917EA88FFBAFFD5FBB9FAE3FB2D2418" box="[1095,1162,1364,1385]" italics="true" pageId="15" pageNumber="600">et al.</emphasis>
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, 2020
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</bibRefCitation>
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). *
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<taxonomicName id="4C634D19FFBAFFD5FAF6FAE3FB3524F9" authority="Mello-Leitao, 1943" authorityName="Mello-Leitao" authorityYear="1943" class="Arachnida" family="Cosmetidae" genus="Ambatoiella" kingdom="Animalia" order="Opiliones" pageId="15" pageNumber="600" phylum="Arthropoda" rank="species" species="vigilans">
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<emphasis id="B917EA88FFBAFFD5FAF6FAE3FC3924F6" italics="true" pageId="15" pageNumber="600">Ambatoiella vigilans</emphasis>
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<bibRefCitation id="EFF24B6BFFBAFFD5FC57FAC5FB3524F9" author="de Mello-Leitao CF" box="[937,1170,1394,1416]" pageId="15" pageNumber="600" pagination="1 - 8" refId="ref53478" refString="de Mello-Leitao CF. 1943. Aracnidos recogidos en el Ecuador y el Peru por la Senora H. E. Frizell Don. Comunicaciones zoologicas del Museo de Historia natural de Montevideo 1: 1 - 8." type="journal article" year="1943">Mello-Leitão, 1943</bibRefCitation>
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</taxonomicName>
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stands for the genus
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<taxonomicName id="4C634D19FFBAFFD5FCC5FA26FB0924D7" authority="Mello-Leitao, 1943" authorityName="Mello-Leitao" authorityYear="1943" box="[827,1198,1425,1447]" class="Arachnida" family="Cosmetidae" genus="Ambatoiella" kingdom="Animalia" order="Opiliones" pageId="15" pageNumber="600" phylum="Arthropoda" rank="genus">
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<emphasis id="B917EA88FFBAFFD5FCC5FA26FC6D24D7" box="[827,970,1425,1446]" italics="true" pageId="15" pageNumber="600">Ambatoiella</emphasis>
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<bibRefCitation id="EFF24B6BFFBAFFD5FC2CFA26FB0924D7" author="de Mello-Leitao CF" box="[978,1198,1425,1447]" pageId="15" pageNumber="600" pagination="1 - 8" refId="ref53478" refString="de Mello-Leitao CF. 1943. Aracnidos recogidos en el Ecuador y el Peru por la Senora H. E. Frizell Don. Comunicaciones zoologicas del Museo de Historia natural de Montevideo 1: 1 - 8." type="journal article" year="1943">Mello-Leitão, 1943</bibRefCitation>
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</taxonomicName>
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, which has a second species, both occurring in
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<collectingCountry id="F374760AFFBAFFD5FB97FA07FB6E24B7" box="[1129,1225,1456,1478]" name="Ecuador" pageId="15" pageNumber="600">Ecuador</collectingCountry>
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(
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<bibRefCitation id="EFF24B6BFFBAFFD5FB24FA07FCD52495" author="Medrano M & Kury AB" pageId="15" pageNumber="600" pagination="1 - 55" refId="ref52961" refString="Medrano M, Kury AB. 2017. Taxonomic revision of the Andean genus Eulibitia Roewer, 1912 (Arachnida, Opiliones, Cosmetidae), with the description of five new species. European Journal of Taxonomy 357: 1 - 55." type="journal article" year="2017">Medrano & Kury, 2017</bibRefCitation>
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). *
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<taxonomicName id="4C634D19FFBAFFD5FC6AFA79FACF2495" authority="Caporiacco, 1951" authorityName="Caporiacco" authorityYear="1951" box="[916,1384,1486,1508]" class="Arachnida" family="Cosmetidae" genus="Oligovonones" kingdom="Animalia" order="Opiliones" pageId="15" pageNumber="600" phylum="Arthropoda" rank="species" species="brunneus">
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<emphasis id="B917EA88FFBAFFD5FC6AFA79FB382492" box="[916,1183,1486,1507]" italics="true" pageId="15" pageNumber="600">Oligovonones brunneus</emphasis>
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<bibRefCitation id="EFF24B6BFFBAFFD5FB58FA79FACF2495" author="di Caporiacco L" box="[1190,1384,1486,1508]" pageId="15" pageNumber="600" pagination="1 - 46" refId="ref51143" refString="di Caporiacco L. 1951. Studi sugli Aracnidi del Venezuela raccolti dalla Sezione di Biologia (Universita Centrale del Venezuela). I Parte: Scorpiones, Opiliones, Solifuga y Chernetes. Acta Biologica Venezuelica 1: 1 - 46." type="journal article" year="1951">Caporiacco, 1951</bibRefCitation>
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</taxonomicName>
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here represents the genus
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<taxonomicName id="4C634D19FFBAFFD5FBC9FA5AFA3A2772" authority="Caporiacco, 1951" authorityName="di Caporiacco" authorityYear="1951" box="[1079,1437,1517,1539]" class="Arachnida" family="Cosmetidae" genus="Oligovonones" kingdom="Animalia" order="Opiliones" pageId="15" pageNumber="600" phylum="Arthropoda" rank="genus">
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<emphasis id="B917EA88FFBAFFD5FBC9FA5AFB762773" box="[1079,1233,1517,1538]" italics="true" pageId="15" pageNumber="600">Oligovonones</emphasis>
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<bibRefCitation id="EFF24B6BFFBAFFD5FB26FA5AFA3A2772" author="di Caporiacco L" box="[1240,1437,1517,1539]" pageId="15" pageNumber="600" pagination="1 - 46" refId="ref51143" refString="di Caporiacco L. 1951. Studi sugli Aracnidi del Venezuela raccolti dalla Sezione di Biologia (Universita Centrale del Venezuela). I Parte: Scorpiones, Opiliones, Solifuga y Chernetes. Acta Biologica Venezuelica 1: 1 - 46." type="journal article" year="1951">Caporiacco, 1951</bibRefCitation>
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</taxonomicName>
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, which has further three species, all found in
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<collectingCountry id="F374760AFFBAFFD5FAD0F9BBFA052753" box="[1326,1442,1548,1570]" name="Venezuela" pageId="15" pageNumber="600">Venezuela</collectingCountry>
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||
(
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<bibRefCitation id="EFF24B6BFFBAFFD5FCBAF99DFBFA2731" author="Gonzalez-Sponga MA" box="[836,1117,1578,1600]" pageId="15" pageNumber="600" refId="ref51725" refString="Gonzalez-Sponga MA. 1992. Aracnidos de Venezuela. Opiliones Laniatores II. Familia Cosmetidae. Caracas: Academia de Ciencias Fisicas, Matematicas y Naturales." type="book" year="1992">González-Sponga, 1992</bibRefCitation>
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). All three genera in their current configurations may, for the present purposes, be treated as probable monophyletic units.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3796511FFBAFFCAFCC5F910FEAF2046" lastPageId="16" lastPageNumber="601" pageId="15" pageNumber="600" type="diagnosis">
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||
<paragraph id="8BDC369AFFBAFFCAFCC5F910FEAF2046" blockId="15.[827,1443,1702,1878]" lastBlockId="16.[145,760,197,311]" lastPageId="16" lastPageNumber="601" pageId="15" pageNumber="600">
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<emphasis id="B917EA88FFBAFFD5FCC5F910FC1127CD" box="[827,950,1703,1724]" italics="true" pageId="15" pageNumber="600">Diagnosis:</emphasis>
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Small animals with short legs. DS alphatype with large coda. Fe IV curved and shorter than DS length. DS flat in lateral view, areas of mesotergum without any armature. Coxa IV entirely visible in dorsal view, convex and short (0.3× DS length in dorsal view). Leg I with five tarsomeres; leg III with five to six tarsomeres. Penis with two pairs of MS-A. Yellow spots mostly reduced to small dots in laterals of scutal grooves, extending medially in
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<taxonomicName id="4C634D19FFA5FFCAFF4EFE96FEA72047" authorityName="Simon" authorityYear="1879" box="[176,256,289,310]" class="Arachnida" family="Cosmetidae" genus="Libitia" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="601" phylum="Arthropoda" rank="genus">
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<emphasis id="B917EA88FFA5FFCAFF4EFE96FEA72047" box="[176,256,289,310]" italics="true" pageId="16" pageNumber="601">Libitia</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3796511FFA5FFCAFF6FFEEFFDA4201C" box="[145,515,344,366]" pageId="16" pageNumber="601" type="type_taxon">
|
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<paragraph id="8BDC369AFFA5FFCAFF6FFEEFFDA4201C" blockId="16.[145,515,344,366]" box="[145,515,344,366]" pageId="16" pageNumber="601">
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||
<emphasis id="B917EA88FFA5FFCAFF6FFEEFFEB3201C" box="[145,276,344,365]" italics="true" pageId="16" pageNumber="601">
|
||
<typeStatus id="54D88838FFA5FFCAFF6FFEEFFF61201C" box="[145,198,344,365]" pageId="16" pageNumber="601">Type</typeStatus>
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genus:
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||
</emphasis>
|
||
<taxonomicName id="4C634D19FFA5FFCAFEE5FEEFFE5B201C" ID-CoL="7PMS7" authority="Simon, 1879" authorityName="Simon" authorityYear="1879" box="[283,508,344,366]" class="Arachnida" family="Cosmetidae" genus="Libitia" kingdom="Animalia" order="Opiliones" pageId="16" pageNumber="601" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B917EA88FFA5FFCAFEE5FEEFFECE201C" box="[283,361,344,365]" italics="true" pageId="16" pageNumber="601">Libitia</emphasis>
|
||
<bibRefCitation id="EFF24B6BFFA5FFCAFE91FEEFFE5B201C" author="Simon E" box="[367,508,344,366]" pageId="16" pageNumber="601" pagination="183 - 241" refId="ref55071" refString="Simon E. 1879. Essai d'une classification des Opiliones Mecostethi. Remarques synonymiques et descriptions d'especes nouvelles. Premiere partie. Annales de la Societe Entomologique de Belgique 22: 183 - 241." type="journal article" year="1879">Simon, 1879</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3796511FFA5FFCAFF6FFE39FEFF2090" pageId="16" pageNumber="601" type="synonymic_list">
|
||
<paragraph id="8BDC369AFFA5FFCAFF6FFE39FEFF2090" blockId="16.[145,761,398,482]" pageId="16" pageNumber="601">
|
||
<emphasis id="B917EA88FFA5FFCAFF6FFE39FD9620D5" box="[145,561,398,420]" italics="true" pageId="16" pageNumber="601">Included genera: Ambatoiella</emphasis>
|
||
Mello - Leitão, 1943;
|
||
<emphasis id="B917EA88FFA5FFCAFF1BFE1AFEE720B2" box="[229,320,429,451]" italics="true" pageId="16" pageNumber="601">Libitia</emphasis>
|
||
Simon, 1879; and
|
||
<emphasis id="B917EA88FFA5FFCAFDB4FE1AFD5E20B2" box="[586,761,429,451]" italics="true" pageId="16" pageNumber="601">Oligovonones</emphasis>
|
||
<bibRefCitation id="EFF24B6BFFA5FFCAFF6FFE7BFEF62093" author="di Caporiacco L" box="[145,337,460,482]" pageId="16" pageNumber="601" pagination="1 - 46" refId="ref51143" refString="di Caporiacco L. 1951. Studi sugli Aracnidi del Venezuela raccolti dalla Sezione di Biologia (Universita Centrale del Venezuela). I Parte: Scorpiones, Opiliones, Solifuga y Chernetes. Acta Biologica Venezuelica 1: 1 - 46." type="journal article" year="1951">Caporiacco, 1951</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3796511FFA5FFCAFF6FFDB5FD982369" box="[145,575,514,536]" pageId="16" pageNumber="601" type="distribution">
|
||
<paragraph id="8BDC369AFFA5FFCAFF6FFDB5FD982369" blockId="16.[145,575,514,536]" box="[145,575,514,536]" pageId="16" pageNumber="601">
|
||
<emphasis id="B917EA88FFA5FFCAFF6FFDB5FE842366" box="[145,291,514,535]" italics="true" pageId="16" pageNumber="601">Distribution:</emphasis>
|
||
Northern Andes (
|
||
<figureCitation id="13582A1FFFA5FFCAFE0EFDB5FD882369" box="[496,559,514,536]" captionStart="Figure 7" captionStartId="12.[145,224,1450,1472]" captionTargetBox="[145,1425,196,1410]" captionTargetId="figure-24@12.[145,1425,195,1410]" captionTargetPageId="12" captionText="Figure 7. Strict consensus of trees obtained with K = 6.2878 and 7.4809, indicating larger groups as defined herein, with asserted distributions juxtaposed. * indicates type species of the genera.Abbreviations: BY = Bayesian analyses, EW = equal weighting, IW = implied weighting, RBS = relative Bremer support, BT = bootstrap, GC = difference in frequencies GC (Group present/Contradicted), PP = posterior probabilities of clades." figureDoi="http://doi.org/10.5281/zenodo.6959536" httpUri="https://zenodo.org/record/6959536/files/figure.png" pageId="16" pageNumber="601">Fig. 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |