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<document id="9E649FC25EAAC9BB22FCBB812B061C31" ID-DOI="10.11646/zootaxa.5433.4.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10954420" ID-ZooBank="7ADF6A54-E31C-453A-AA25-44DEDBCF2752" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1712752719575" checkinUser="plazi" docAuthor="Souza, Leonardo Santos De, Araújo, Tarciso Almeida De &amp; Pimenta, Alexandre Dias" docDate="2024" docId="03CA878ABA79DE2AFF4FFAED754BF82B" docLanguage="en" docName="zootaxa.5433.4.1.pdf" docOrigin="Zootaxa 5433 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5433.4.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Aclis macrostoma Barros, Lima &amp; Francisco 2007" docType="treatment" docVersion="2" lastPageNumber="474" masterDocId="FFF3FFF2BA6EDE3DFFD8FFD77042FF9F" masterDocTitle="Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena" masterLastPageNumber="499" masterPageNumber="451" pageNumber="474" updateTime="1712753971016" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="11BBBBB3553CBA41300C353C8A1441CB">Taxonomic review of Aclis Lóven, 1846 (Gastropoda, Eulimidae) from Brazil with notes on other congeners from the western Atlantic and Saint Helena</mods:title>
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<figureCitation id="13582A19BA79DE2AFD16FAA87364FA08" box="[718,806,1407,1431]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStart-2="FIGURE 9" captionStartId-0="15.[151,246,1943,1967]" captionStartId-1="16.[151,250,1943,1967]" captionStartId-2="17.[151,250,1539,1563]" captionTargetBox-0="[162,1425,181,1918]" captionTargetBox-1="[162,1425,181,1919]" captionTargetBox-2="[151,1435,273,1515]" captionTargetId-0="figure-17@15.[162,1425,181,1918]" captionTargetId-1="figure-23@16.[162,1425,181,1919]" captionTargetId-2="figure-67@17.[151,1435,273,1515]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="17" captionText-0="FIGURE 7. Aclis pulchra sp. nov. A–C, F–I. Holotype, MNRJ 16825. D–E, J. paratype, MNRJ 26601. A, C–D. Shell in frontal view. B, E. Shell in lateral view. F–G. Detail of teleoconch surface; square in F indicates detail in G. H. Detail of body whorl in frontal view. I. Detail of apical whorls. J. Protoconch in apical view. Scale bars: A–E. 1 mm; F–G. 50 μm; H. 500 μm; I–J. 200 μm." captionText-1="FIGURE 8. Aclis pulchra sp. nov. A. Paratype, MNRJ 16323. B, F, H. Paratype, MNRJ 17713. C. Paratype, MNRJ 30872. D, G, I. paratype, MORG 39469. E. Paratype, MNRJ 26601. A–D. Shell in frontal view. E–G. Detail of apical whorls. H–I. Detail of body whorl in frontal view. Scale bars: A–D. 1 mm; E–G. 200 μm; H–I. 500 μm." captionText-2="FIGURE 9. Aclis cf. pulchra: MNRJ 34072. A, C. Shell in frontal view. B. Shell in lateral view. D. Detail of teleoconch surface. E. Detail of apical whorls. F–G. Detail of body whorl in frontal view. Scale bars: A–C. 1 mm; D. 20 μm; E. 200 μm; F–G. 500 μm." figureDoi-0="http://doi.org/10.5281/zenodo.10954440" figureDoi-1="http://doi.org/10.5281/zenodo.10954443" figureDoi-2="http://doi.org/10.5281/zenodo.10954447" httpUri-0="https://zenodo.org/record/10954440/files/figure.png" httpUri-1="https://zenodo.org/record/10954443/files/figure.png" httpUri-2="https://zenodo.org/record/10954447/files/figure.png" pageId="23" pageNumber="474">Fig. 7–9</figureCitation>
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, Fig. 490);
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,
<quantity id="4C9B9B79BA79DE2AFE65F9DA7245F9B7" box="[445,519,1549,1576]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="23" pageNumber="474" unit="m" value="500.0">
<elevation id="004ED1AFBA79DE2AFE65F9DA7245F9B7" box="[445,519,1549,1576]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="23" pageNumber="474" unit="m" value="500.0">500 m</elevation>
</quantity>
;
<date id="FFDD105CBA79DE2AFDCAF9DA72D2F9B8" box="[530,656,1549,1575]" pageId="23" pageNumber="474" value="2001-11-26">
<collectingDate id="EF99E9B4BA79DE2AFDCAF9DA72D2F9B8" box="[530,656,1549,1575]" pageId="23" pageNumber="474" value="2001-11-26">26/xi/2001</collectingDate>
</date>
; R/
<collectorName id="2696534ABA79DE2AFD6DF9DA7377F9B8" box="[693,821,1549,1575]" pageId="23" pageNumber="474">
V
<emphasis id="B917EA8EBA79DE2AFD17F9D97377F9B8" box="[719,821,1550,1575]" italics="true" pageId="23" pageNumber="474">Natureza</emphasis>
</collectorName>
;
<specimenCode id="DBC59EE7BA79DE2AFC9CF9DA739BF9B8" box="[836,985,1549,1575]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="23" pageNumber="474" type="Museum">MZSP 77990</specimenCode>
</materialsCitation>
<materialsCitation id="3B0B3CC1BA79DE2AFC38F9DA7480F9B8" box="[992,1218,1549,1576]" collectingDate="2001-11-26" collectionCode="LMUFRPE" collectorName="V Natureza" country="Brazil" elevation="500" isTailEnumeration="true" latitude="-6.24" location="Revizee Nordeste" longLatPrecision="21" longitude="-34.868332" pageId="23" pageNumber="474" specimenCode="LMUFRPE 022" specimenCount="1" stateProvince="Rio Grande do Norte" typeStatus="paratype">
[1]:
<specimenCode id="DBC59EE7BA79DE2AFBD7F9D97480F9B8" box="[1039,1218,1549,1576]" collectionCode="LMUFRPE" pageId="23" pageNumber="474">LMUFRPE 022</specimenCode>
</materialsCitation>
[1].
</paragraph>
<paragraph id="8BDC369CBA79DE2AFF1FF9E67540F9D3" blockId="23.[151,1437,1477,1972]" box="[199,1282,1585,1614]" pageId="23" pageNumber="474">
<materialsCitation id="3B0B3CC1BA79DE2AFF1FF9E67540F9D3" box="[199,1282,1585,1614]" country="Brazil" elevation="510" latitude="-6.234444" location="Revizee Nordeste" longLatPrecision="21" longitude="-34.87583" pageId="23" pageNumber="474" specimenCount="510" stateProvince="Rio Grande do Norte" typeStatus="holotype">
<emphasis id="B917EA8EBA79DE2AFF1FF9E67124F9D4" bold="true" box="[199,358,1585,1611]" pageId="23" pageNumber="474">
<typeStatus id="54D8883EBA79DE2AFF1FF9E67140F9D4" box="[199,258,1585,1611]" pageId="23" pageNumber="474">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F374760CBA79DE2AFEB4F9E671F7F9D4" box="[364,437,1585,1611]" name="Brazil" pageId="23" pageNumber="474">Brazil</collectingCountry>
:
<collectingRegion id="49A7F87EBA79DE2AFE18F9E672ECF9D3" box="[448,686,1585,1612]" country="Brazil" name="Rio Grande do Norte" pageId="23" pageNumber="474">Rio Grande do Norte</collectingRegion>
;
<location id="8EBC6047BA79DE2AFD62F9E673C6F9D4" LSID="urn:lsid:plazi:treatment:03CA878ABA79DE2AFF4FFAED754BF82B:8EBC6047BA79DE2AFD62F9E673C6F9D4" box="[698,900,1585,1611]" country="Brazil" latitude="-6.234444" longLatPrecision="21" longitude="-34.87583" name="Revizee Nordeste" pageId="23" pageNumber="474" stateProvince="Rio Grande do Norte">Revizee Nordeste</location>
;
<geoCoordinate id="EE57505BBA79DE2AFC57F9E47450F9D1" box="[911,1042,1587,1614]" degrees="06" direction="south" minutes="14" orientation="latitude" pageId="23" pageNumber="474" precision="15" seconds="04" value="-6.234444">06°14′04″S</geoCoordinate>
,
<geoCoordinate id="EE57505BBA79DE2AFBC6F9E474E5F9D1" box="[1054,1191,1587,1614]" degrees="34" direction="west" minutes="52" orientation="longitude" pageId="23" pageNumber="474" precision="15" seconds="33" value="-34.87583">34°52′33″W</geoCoordinate>
,
<specimenCount id="9D65FD15BA79DE2AFB6CF9E67540F9D3" box="[1204,1282,1585,1612]" count="510" pageId="23" pageNumber="474" type="generic">
<quantity id="4C9B9B79BA79DE2AFB6CF9E674BCF9D3" box="[1204,1278,1585,1612]" metricMagnitude="2" metricUnit="m" metricValue="5.1" pageId="23" pageNumber="474" unit="m" value="510.0">
<elevation id="004ED1AFBA79DE2AFB6CF9E674BCF9D3" box="[1204,1278,1585,1612]" metricMagnitude="2" metricUnit="m" metricValue="5.1" pageId="23" pageNumber="474" unit="m" value="510.0">510 m</elevation>
</quantity>
.
</specimenCount>
</materialsCitation>
</paragraph>
<paragraph id="8BDC369CBA79DE2AFF1FF98272CBF9F0" blockId="23.[151,1437,1477,1972]" box="[199,649,1621,1647]" pageId="23" pageNumber="474">
<emphasis id="B917EA8EBA79DE2AFF1FF98271F3F9F0" bold="true" box="[199,433,1621,1647]" pageId="23" pageNumber="474">Material examined.</emphasis>
Only the
<typeStatus id="54D8883EBA79DE2AFDF8F98272C4F9F0" box="[544,646,1621,1647]" pageId="23" pageNumber="474" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C3796517BA79DE2AFF1FF9AE713CF943" pageId="23" pageNumber="474" type="description">
<paragraph id="8BDC369CBA79DE2AFF1FF9AE713CF943" blockId="23.[151,1437,1477,1972]" pageId="23" pageNumber="474">
<emphasis id="B917EA8EBA79DE2AFF1FF9AE713CF90C" bold="true" box="[199,382,1657,1683]" pageId="23" pageNumber="474">Measurements.</emphasis>
<typeStatus id="54D8883EBA79DE2AFE5CF9AE71ADF90C" box="[388,495,1657,1683]" pageId="23" pageNumber="474" type="holotype">Holotype</typeStatus>
, MZSP 77988: 3 whorls; SL=
<quantity id="4C9B9B79BA79DE2AFC9DF9AE73EEF90B" box="[837,940,1657,1684]" metricMagnitude="-3" metricUnit="m" metricValue="1.08" pageId="23" pageNumber="474" unit="mm" value="1.08">1.08 mm</quantity>
; BWL=
<quantity id="4C9B9B79BA79DE2AFBD4F9AE7431F90B" box="[1036,1139,1657,1684]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="23" pageNumber="474" unit="mm" value="0.8">0.80 mm</quantity>
; AL=
<quantity id="4C9B9B79BA79DE2AFB60F9AE755DF90B" box="[1208,1311,1657,1684]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" pageId="23" pageNumber="474" unit="mm" value="0.54">0.54 mm</quantity>
; SW=
<quantity id="4C9B9B79BA79DE2AFAB3F9AE7084F927" metricMagnitude="-4" metricUnit="m" metricValue="7.4" pageId="23" pageNumber="474" unit="mm" value="0.74">0.74 mm</quantity>
; AW=
<quantity id="4C9B9B79BA79DE2AFEC9F94A7135F927" box="[273,375,1693,1720]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="23" pageNumber="474" unit="mm" value="0.5">0.50 mm</quantity>
.
<typeStatus id="54D8883EBA79DE2AFE59F94971A1F927" box="[385,483,1694,1720]" pageId="23" pageNumber="474" type="paratype">Paratype</typeStatus>
LMUFRPE 022: 3 whorls; SL=
<quantity id="4C9B9B79BA79DE2AFC95F94A73F1F927" box="[845,947,1693,1720]" metricMagnitude="-3" metricUnit="m" metricValue="1.08" pageId="23" pageNumber="474" unit="mm" value="1.08">1.08 mm</quantity>
; BWL=
<quantity id="4C9B9B79BA79DE2AFBC9F94A7435F927" box="[1041,1143,1693,1720]" metricMagnitude="-4" metricUnit="m" metricValue="8.1" pageId="23" pageNumber="474" unit="mm" value="0.81">0.81 mm</quantity>
; AL=
<quantity id="4C9B9B79BA79DE2AFB63F94A7563F927" box="[1211,1313,1693,1720]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" pageId="23" pageNumber="474" unit="mm" value="0.54">0.54 mm</quantity>
; SW=
<quantity id="4C9B9B79BA79DE2AFAB3F94A7084F943" metricMagnitude="-4" metricUnit="m" metricValue="7.1" pageId="23" pageNumber="474" unit="mm" value="0.71">0.71 mm</quantity>
; AW=
<quantity id="4C9B9B79BA79DE2AFECBF9167138F943" box="[275,378,1729,1756]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="23" pageNumber="474" unit="mm" value="0.5">0.50 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3796517BA79DE2AFF1FF932754BF82B" pageId="23" pageNumber="474" type="discussion">
<paragraph id="8BDC369CBA79DE2AFF1FF932722CF810" blockId="23.[151,1437,1477,1972]" pageId="23" pageNumber="474">
<emphasis id="B917EA8EBA79DE2AFF1FF9327179F960" bold="true" box="[199,315,1765,1791]" pageId="23" pageNumber="474">Remarks.</emphasis>
<taxonomicName id="4C634D1FBA79DE2AFE9BF932724EF960" authorityName="Barros, Lima &amp; Francisco" authorityYear="2007" box="[323,524,1765,1791]" class="Gastropoda" family="Eulimidae" genus="Aclis" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="474" phylum="Mollusca" rank="species" species="macrostoma">
<emphasis id="B917EA8EBA79DE2AFE9BF932724EF960" box="[323,524,1765,1791]" italics="true" pageId="23" pageNumber="474">Aclis macrostoma</emphasis>
</taxonomicName>
is known only by the
<typeStatus id="54D8883EBA79DE2AFCD5F931737FF89F" box="[781,829,1766,1792]" pageId="23" pageNumber="474">type</typeStatus>
series and has conspicuous characteristics among the taxa of the present study, such as the broad elliptical aperture, having about half the total size of the shell, and the outer lip vastly expanded in comparison to the other species (
<bibRefCitation id="EFF24B6DBA79DE2AFCEEF8FA7442F8D8" author="Barros, J. C. N. &amp; Lima, S. F. B. &amp; Francisco, J. A." box="[822,1024,1837,1863]" pageId="23" pageNumber="474" pagination="61 - 68" refId="ref23513" refString="Barros, J. C. N., Lima, S. F. B. &amp; Francisco, J. A. (2007) Two new species of Aclis (Mollusca: Gastropoda: Aclididae) from the continental slope of northeast Brazil. Zootaxa, 1614 (1), 61 - 68. https: // doi. org / 10.11646 / zootaxa. 1614.1.6" type="journal article" year="2007">
Barros
<emphasis id="B917EA8EBA79DE2AFC50F8F973FDF8D8" box="[904,959,1837,1863]" italics="true" pageId="23" pageNumber="474">et al.</emphasis>
2007
</bibRefCitation>
:
<figureCitation id="13582A19BA79DE2AFBD1F8FA7416F8D8" box="[1033,1108,1837,1863]" captionStart="FIGURE 1" captionStartId="6.[151,250,1295,1319]" captionTargetBox="[151,1435,181,1271]" captionTargetId="figure-23@6.[151,1435,181,1271]" captionTargetPageId="6" captionText="FIGURE 1. Terminology of some of the shell features used in descriptions in the present study. A. Microscopic spiral striae of the teleoconch indicated by arrows. B. Spiral keels of the teleoconch indicated by arrows. C. Umbilical fissure indicated by arrow. D. Umbilicus indicated by arrow." figureDoi="http://doi.org/10.5281/zenodo.10954422" httpUri="https://zenodo.org/record/10954422/files/figure.png" pageId="23" pageNumber="474">Figs. 1</figureCitation>
,
<figureCitation id="13582A19BA79DE2AFBB8F8FA7432F8D7" box="[1120,1136,1837,1864]" captionStart="FIGURE 5" captionStartId="12.[151,250,1825,1849]" captionTargetBox="[151,1435,597,1801]" captionTargetId="figure-218@12.[151,1435,597,1801]" captionTargetPageId="12" captionText="FIGURE 5. Aclis kanelaAbsalão, 2009. A, D–G. MNRJ 28293. B–C. MNRJ 26713. A–C. Shell in frontal view; rectangle in A indicates detail in E. D. Detail of body whorl in frontal view. E, F. Detail of teleoconch sculpture; rectangle in E indicates detail in F. G. Detail of apical whorls. Scale bars: A–C. 1 mm; D. 500 μm; E–F. 50 μm; G. 200 μm." figureDoi="http://doi.org/10.5281/zenodo.10954434" httpUri="https://zenodo.org/record/10954434/files/figure.png" pageId="23" pageNumber="474">5</figureCitation>
). Although
<typeStatus id="54D8883EBA79DE2AFB2AF8F97560F8D7" box="[1266,1314,1838,1864]" pageId="23" pageNumber="474">type</typeStatus>
specimens have a tiny size, the expanded aperture, well-formed inner lip and the relatively sizeable umbilical fissure indicate that this may represent an adult specimen.
</paragraph>
<paragraph id="8BDC369CBA79DE2AFF1FF84E754BF82B" blockId="23.[151,1437,1477,1972]" box="[199,1289,1945,1972]" pageId="23" pageNumber="474">
<emphasis id="B917EA8EBA79DE2AFF1FF84E71B6F82C" bold="true" box="[199,500,1945,1971]" pageId="23" pageNumber="474">Geographic Distribution.</emphasis>
<materialsCitation id="3B0B3CC1BA79DE2AFE22F84E754BF82B" box="[506,1289,1945,1972]" country="Brazil" elevation="443" location="From" pageId="23" pageNumber="474" specimenCount="1" stateProvince="Rio Grande do Norte">
<collectingCountry id="F374760CBA79DE2AFE22F84E7201F82C" box="[506,579,1945,1971]" name="Brazil" pageId="23" pageNumber="474">Brazil</collectingCountry>
:
<collectingRegion id="49A7F87EBA79DE2AFD96F84E737BF82B" box="[590,825,1945,1972]" country="Brazil" name="Rio Grande do Norte" pageId="23" pageNumber="474">Rio Grande do Norte</collectingRegion>
(
<bibRefCitation id="EFF24B6DBA79DE2AFC90F84E7455F82C" author="Barros, J. C. N. &amp; Lima, S. F. B. &amp; Francisco, J. A." box="[840,1047,1945,1971]" pageId="23" pageNumber="474" pagination="61 - 68" refId="ref23513" refString="Barros, J. C. N., Lima, S. F. B. &amp; Francisco, J. A. (2007) Two new species of Aclis (Mollusca: Gastropoda: Aclididae) from the continental slope of northeast Brazil. Zootaxa, 1614 (1), 61 - 68. https: // doi. org / 10.11646 / zootaxa. 1614.1.6" type="journal article" year="2007">
Barros
<emphasis id="B917EA8EBA79DE2AFC43F84D7391F82C" box="[923,979,1945,1971]" italics="true" pageId="23" pageNumber="474">et al.</emphasis>
2007
</bibRefCitation>
).
<location id="8EBC6047BA79DE2AFBF1F84D7424F82B" LSID="urn:lsid:plazi:treatment:03CA878ABA79DE2AFF4FFAED754BF82B:8EBC6047BA79DE2AFBF1F84D7424F82B" box="[1065,1126,1946,1972]" country="Brazil" name="From" pageId="23" pageNumber="474" stateProvince="Rio Grande do Norte">From</location>
<quantity id="4C9B9B79BA79DE2AFBB5F84E7547F82B" box="[1133,1285,1945,1972]" metricMagnitude="2" metricUnit="m" metricValue="4.425" metricValueMax="5.1" metricValueMin="3.75" pageId="23" pageNumber="474" unit="m" value="442.5" valueMax="510.0" valueMin="375.0">
<elevation id="004ED1AFBA79DE2AFBB5F84E7547F82B" box="[1133,1285,1945,1972]" metricMagnitude="2" metricUnit="m" metricValue="4.425" metricValueMax="5.1" metricValueMin="3.75" pageId="23" pageNumber="474" unit="m" value="442.5" valueMax="510.0" valueMin="375.0">375 to 510 m</elevation>
</quantity>
.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>
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