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<document id="882B1AAEFBE2AB5D45B57DF59FE43471" ID-CLB-Dataset="31657" ID-DOI="10.5281/zenodo.322670" ID-GBIF-Dataset="7ddec036-fcb1-4703-ada3-670f5fc8f25d" ID-ISSN="0035-418" ID-Zenodo-Dep="322670" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1509607194732" checkinUser="plazi" docAuthor="Gabriel Biffi" docDate="2017" docId="03C1CF7FFFD6CB34FC33FBA80EDDF885" docLanguage="en" docName="11_Biffi.pdf" docOrigin="Revue suisse de Zoologie 124 (1)" docStyle="DocumentStyle:225056F0A23C51DDC06AC27EAEA15D5B.2:RevSuisseZool.2015-2019.journal_article" docStyleId="225056F0A23C51DDC06AC27EAEA15D5B" docStyleName="RevSuisseZool.2015-2019.journal_article" docStyleVersion="2" docTitle="Paramaronius unituberculatus Gabriel Biffi 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="118" masterDocId="FFF8B707FFD6CB37FFA5FFEC0C2DFFD8" masterDocTitle="A new Paramaronius species from Argentina (Coleoptera: Cantharidae)" masterLastPageNumber="118" masterPageNumber="115" pageNumber="115" updateTime="1698470657999" updateUser="ExternalLinkService">
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<mods:title id="8789D3D0EAF6346210F5DBBB6921A7B2">A new Paramaronius species from Argentina (Coleoptera: Cantharidae)</mods:title>
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<mods:name id="F84546C17D12A41A24948CFBF1A9D4E5" type="personal">
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<mods:namePart id="5C0B66A22621326324FEB0A420EC8580">Gabriel Biffi</mods:namePart>
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<mods:date id="515B79F7901E52D7E848BC1F3350270A">2017</mods:date>
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<mods:number id="A2D059CE25D06F6D2E3AC6DD92444640">2017-03-30</mods:number>
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<treatment id="03C1CF7FFFD6CB34FC33FBA80EDDF885" ID-DOI="http://doi.org/10.5281/zenodo.6030401" ID-GBIF-Taxon="137227715" ID-Zenodo-Dep="6030401" LSID="urn:lsid:plazi:treatment:03C1CF7FFFD6CB34FC33FBA80EDDF885" httpUri="http://treatment.plazi.org/id/03C1CF7FFFD6CB34FC33FBA80EDDF885" lastPageId="3" lastPageNumber="118" pageId="0" pageNumber="115">
<subSubSection id="C3722DE2FFD6CB37FC33FBA8096BFB84" box="[918,1350,1091,1116]" pageId="0" pageNumber="115" type="nomenclature">
<paragraph id="8BD77E69FFD6CB37FC33FBA8096BFB84" blockId="0.[918,1350,1091,1149]" box="[918,1350,1091,1116]" pageId="0" pageNumber="115">
<heading id="D09FC905FFD6CB37FC33FBA8096BFB84" bold="true" box="[918,1350,1091,1116]" centered="true" fontSize="10" level="0" pageId="0" pageNumber="115" reason="4">
<emphasis id="B91CA27BFFD6CB37FC33FBA8096BFB84" bold="true" box="[918,1350,1091,1116]" pageId="0" pageNumber="115">
<taxonomicName id="4C6805EAFFD6CB37FC33FBA808C7FB83" authority="Gabriel Biffi, 2017" authorityName="Gabriel Biffi" authorityYear="2017" box="[918,1258,1091,1116]" class="Insecta" family="Cantharidae" genus="Paramaronius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="unituberculatus" status="sp. nov.">
<emphasis id="B91CA27BFFD6CB37FC33FBA808C7FB83" bold="true" box="[918,1258,1091,1116]" italics="true" pageId="0" pageNumber="115">Paramaronius unituberculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22F1F00FFD6CB37FB54FBA8096BFB84" box="[1265,1350,1092,1116]" pageId="0" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3722DE2FFD6CB37FB9EFB88088DFBA5" box="[1083,1184,1124,1149]" pageId="0" pageNumber="115" type="description">
<paragraph id="8BD77E69FFD6CB37FB9EFB88088DFBA5" blockId="0.[918,1350,1091,1149]" box="[1083,1184,1124,1149]" pageId="0" pageNumber="115">
<figureCitation id="135362ECFFD6CB37FB9EFB88088DFBA5" box="[1083,1184,1124,1149]" captionStart-0="Figs 1 - 4" captionStart-1="Figs 5 - 8" captionStart-2="Figs 9 - 12" captionStartId-0="1.[151,192,1250,1271]" captionStartId-1="2.[151,192,1707,1728]" captionStartId-2="3.[151,192,847,868]" captionTargetBox-0="[187,1434,254,1201]" captionTargetBox-1="[149,1439,259,1662]" captionTargetBox-2="[184,1432,258,781]" captionTargetId-0="figure@1.[149,1439,254,1244]" captionTargetId-1="figure@2.[148,1439,254,1669]" captionTargetId-2="figure@3.[148,1448,250,821]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="Figs 1 - 4. Paramaronius unituberculatus sp. nov. (1) Dorsal habitus of male (holotype, NMB). (2) Dorsal habitus of female (paratype, MZSP). (3) Detail of the elytra of male. (4) Detail of the elytra of female." captionText-1="Figs 5 - 8. Aedeagus of Paramaronius unituberculatus sp. nov. (holotype, NMB). (5) Dorsal view. (6) Left view. (7) Ventral view. (8) Right view. Scale = 0.5 mm." captionText-2="Figs 9 - 12. Paramaronius unituberculatus sp. nov. (9) Seventh ventrite of the male (holotype, NMB). (10) Seventh ventrite of the female (paratype, MZSP). (11) Right paramere of aedeagus (holotype, NMB). (12) Right coxite of female (paratype, MZSP). Scales: 9 - 10 = 0.5 mm, 11 - 12 = 0.2 mm." httpUri-0="https://zenodo.org/record/1040489/files/figure.png" httpUri-1="https://zenodo.org/record/1040491/files/figure.png" httpUri-2="https://zenodo.org/record/1040493/files/figure.png" pageId="0" pageNumber="115">
Figs
<docIssue id="CF0B0186FFD6CB37FBD5FB890857FBA5" box="[1136,1146,1125,1149]" pageId="0" pageNumber="115">1</docIssue>
-12
</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3722DE2FFD6CB37FC9AFB7F095DFAE2" pageId="0" pageNumber="115" type="materials_examined">
<paragraph id="8BD77E69FFD6CB37FC9AFB7F095DFAE2" blockId="0.[831,1437,1171,1258]" lastBlockId="0.[831,1437,1282,1339]" pageId="0" pageNumber="115">
<materialsCitation id="3B007434FFD6CB37FC9AFB7F0FB0FB32" ID-GBIF-Occurrence="1677457971" collectingDate="1982-02-08" collectionCode="NMB" collectorName="H &amp; A Howden" country="Argentina" location="Cerro San Bernardo" pageId="0" pageNumber="115" specimenCount="1" specimenCount-male="1" stateProvince="Salta" typeStatus="holotype">
<emphasis id="B91CA27BFFD6CB37FC9AFB7F0F9EFB73" bold="true" box="[831,947,1171,1195]" pageId="0" pageNumber="115">
<typeStatus id="54D3C0CBFFD6CB37FC9AFB7F0F80FB73" box="[831,941,1171,1195]" pageId="0" pageNumber="115" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectionCode id="ED79E6ACFFD6CB37FC63FB79082BFB74" box="[966,1030,1173,1196]" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="0" pageNumber="115">NMB</collectionCode>
,
<specimenCount id="9D6EB5E0FFD6CB37FBBAFB78086DFB74" box="[1055,1088,1172,1196]" pageId="0" pageNumber="115" type="male">1♂</specimenCount>
;
<collectingCountry id="F37F3EF9FFD6CB37FBFFFB7808D6FB74" box="[1114,1275,1172,1196]" name="Argentina" pageId="0" pageNumber="115">ARGENTINA</collectingCountry>
,
<emphasis id="B91CA27BFFD6CB37FAB6FB7F0966FB73" box="[1299,1355,1171,1195]" italics="true" pageId="0" pageNumber="115">
<collectingRegion id="49ACB08BFFD6CB37FAB6FB7F0966FB73" box="[1299,1355,1171,1195]" country="Argentina" name="Salta" pageId="0" pageNumber="115">Salta</collectingRegion>
</emphasis>
,
<collectingRegion id="49ACB08BFFD6CB37FAC3FB7F09B1FB72" box="[1382,1436,1171,1194]" country="Argentina" name="Salta" pageId="0" pageNumber="115">Salta</collectingRegion>
(
<location id="8EB728B2FFD6CB37FCE2FB580817FB12" LSID="urn:lsid:plazi:treatment:03C1CF7FFFD6CB34FC33FBA80EDDF885:8EB728B2FFD6CB37FCE2FB580817FB12" box="[839,1082,1203,1228]" country="Argentina" name="Cerro San Bernardo" pageId="0" pageNumber="115" stateProvince="Salta">Cerro San Bernardo</location>
),
<collectingDate id="EF92A141FFD6CB37FBF3FB5F0894FB12" box="[1110,1209,1203,1226]" pageId="0" pageNumber="115" value="1982-02-08">
<date id="FFD658A9FFD6CB37FBF3FB5F0894FB12" box="[1110,1209,1203,1226]" pageId="0" pageNumber="115" value="1982-02-08">8.ii.1982</date>
</collectingDate>
, 1450 m,
<collectorName id="269D1BBFFFD6CB37FAE7FB590FB0FB32" pageId="0" pageNumber="115">H &amp; A Howden</collectorName>
</materialsCitation>
.
<materialsCitation id="3B007434FFD6CB37FC9AFAEE0832FAE2" ID-GBIF-Occurrence="1677457973" collectingDate="1949-01" collectionCode="MZSP" collectorName="Wigodzinsky" country="Argentina" location="Salta" pageId="0" pageNumber="115" specimenCode="MZSP 7062" specimenCount="1" specimenCount-female="1" stateProvince="Salta" typeStatus="paratype">
<typeStatus id="54D3C0CBFFD6CB37FC9AFAEE0F9EFAC2" box="[831,947,1282,1306]" pageId="0" pageNumber="115" type="paratype">
<emphasis id="B91CA27BFFD6CB37FC9AFAEE0F9EFAC2" bold="true" box="[831,947,1282,1306]" pageId="0" pageNumber="115">Paratypes</emphasis>
</typeStatus>
:
<specimenCode id="DBCED612FFD6CB37FC69FAEF0871FAC2" box="[972,1116,1283,1307]" collectionCode="MZSP" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="0" pageNumber="115">MZSP 7062</specimenCode>
;
<specimenCount id="9D6EB5E0FFD6CB37FBD5FAEF08BFFAC3" box="[1136,1170,1283,1307]" pageId="0" pageNumber="115" type="female">1♀</specimenCount>
;
<collectingCountry id="F37F3EF9FFD6CB37FB0CFAEF0964FAC3" box="[1193,1353,1283,1307]" name="Argentina" pageId="0" pageNumber="115">ARGENTINA</collectingCountry>
,
<collectingRegion id="49ACB08BFFD6CB37FAFBFAEE09BBFAC2" box="[1374,1430,1282,1306]" country="Argentina" name="Salta" pageId="0" pageNumber="115">
<emphasis id="B91CA27BFFD6CB37FAFBFAEE09BBFAC2" box="[1374,1430,1282,1306]" italics="true" pageId="0" pageNumber="115">Salta</emphasis>
</collectingRegion>
,
<collectingDate id="EF92A141FFD6CB37FC9AFACF0FABFAE2" box="[831,902,1315,1338]" pageId="0" pageNumber="115" value="1949-01">
<date id="FFD658A9FFD6CB37FC9AFACF0FABFAE2" box="[831,902,1315,1338]" pageId="0" pageNumber="115" value="1949-01">i.1949</date>
</collectingDate>
,
<collectorName id="269D1BBFFFD6CB37FC35FACF0832FAE2" box="[912,1055,1315,1338]" pageId="0" pageNumber="115">Wigodzinsky</collectorName>
</materialsCitation>
.
<materialsCitation id="3B007434FFD6CB37FB99FAC80941FAE2" ID-GBIF-Occurrence="1677457972" box="[1084,1388,1315,1339]" collectingDate="1945-02" collectionCode="NMB" country="Argentina" location="Amblayo" pageId="0" pageNumber="115" specimenCount="1" specimenCount-female="1" stateProvince="Salta" typeStatus="paratype">
<collectionCode id="ED79E6ACFFD6CB37FB99FAC80851FAE3" box="[1084,1148,1316,1339]" country="Switzerland" httpUri="http://grbio.org/cool/8wbz-jkc3" name="Naturhistorishes Museum" pageId="0" pageNumber="115">NMB</collectionCode>
;
<specimenCount id="9D6EB5E0FFD6CB37FB22FACF0884FAE3" box="[1159,1193,1315,1339]" pageId="0" pageNumber="115" type="female">1♀</specimenCount>
<location id="8EB728B2FFD6CB37FB0BFACF0939FAE2" LSID="urn:lsid:plazi:treatment:03C1CF7FFFD6CB34FC33FBA80EDDF885:8EB728B2FFD6CB37FB0BFACF0939FAE2" box="[1198,1300,1315,1338]" country="Argentina" name="Amblayo" pageId="0" pageNumber="115" stateProvince="Salta">Amblayo</location>
,
<collectingDate id="EF92A141FFD6CB37FABBFACF0941FAE2" box="[1310,1388,1315,1338]" pageId="0" pageNumber="115" value="1945-02">
<date id="FFD658A9FFD6CB37FABBFACF0941FAE2" box="[1310,1388,1315,1338]" pageId="0" pageNumber="115" value="1945-02">ii.1945</date>
</collectingDate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3722DE2FFD6CB37FC9AFABD08ACFA11" pageId="0" pageNumber="115" type="diagnosis">
<paragraph id="8BD77E69FFD6CB37FC9AFABD08ACFA11" blockId="0.[831,1437,1361,1481]" pageId="0" pageNumber="115">
<emphasis id="B91CA27BFFD6CB37FC9AFABD0F95FAB1" bold="true" box="[831,952,1361,1385]" pageId="0" pageNumber="115">Diagnosis:</emphasis>
<taxonomicName id="4C6805EAFFD6CB37FC60FABE0938FAB1" authority="Gabriel Biffi, 2017" authorityName="Gabriel Biffi" authorityYear="2017" box="[965,1301,1361,1385]" class="Insecta" family="Cantharidae" genus="Paramaronius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="115" phylum="Arthropoda" rank="species" species="unituberculatus" status="sp. nov.">
<emphasis id="B91CA27BFFD6CB37FC60FABE0938FAB1" box="[965,1301,1361,1385]" italics="true" pageId="0" pageNumber="115">Paramaronius unituberculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22F1F00FFD6CB37FA86FABF0950FAB2" box="[1315,1405,1363,1386]" pageId="0" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
is characterized by the single apical tubercle in the elytra of males and by the shapes of aedeagus and the seventh ventrite of males and females.
</paragraph>
</subSubSection>
<subSubSection id="C3722DE2FFD6CB37FC9AFA0C0FBEF980" pageId="0" pageNumber="115" type="etymology">
<paragraph id="8BD77E69FFD6CB37FC9AFA0C0FBEF980" blockId="0.[831,1437,1504,1624]" pageId="0" pageNumber="115">
<emphasis id="B91CA27BFFD6CB37FC9AFA0C0FE9FA20" bold="true" box="[831,964,1504,1528]" pageId="0" pageNumber="115">Etymology:</emphasis>
The specific epithet is a derivative of the Latin words
<emphasis id="B91CA27BFFD6CB37FC7DF9ED0FD6F9C0" box="[984,1019,1537,1560]" italics="true" pageId="0" pageNumber="115">uni</emphasis>
(one) +
<emphasis id="B91CA27BFFD6CB37FBCBF9EC08D0F9C0" box="[1134,1277,1536,1560]" italics="true" pageId="0" pageNumber="115">tuberculatum</emphasis>
(tuberculate), referring to the single tubercle on the apex of each elytron.
</paragraph>
</subSubSection>
<subSubSection id="C3722DE2FFD6CB35FC9AF9830E19F870" lastPageId="2" lastPageNumber="117" pageId="0" pageNumber="115" type="description">
<paragraph id="8BD77E69FFD6CB36FC9AF9830E1BFA00" blockId="0.[831,1437,1647,1927]" lastBlockId="1.[151,757,1377,2073]" lastPageId="1" lastPageNumber="116" pageId="0" pageNumber="115">
<emphasis id="B91CA27BFFD6CB37FC9AF9830FE3F95F" bold="true" box="[831,974,1647,1671]" pageId="0" pageNumber="115">Description:</emphasis>
Head occipital region, center of clypeus and apex of mandibles light to dark brown; vertex, frons, sides of clypeus, genae and base of mandibles pale yellow; antennae light to dark brown, lighter on ventral side of three proximal antennomeres. Labial and maxillary palpi light brown, darker on last palpomeres. Pronotum pale yellow with two large dark brown lateral patches.
<taxonomicName id="4C6805EAFFD6CB37FC3AF8BC0826F8BF" box="[927,1035,1872,1895]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Corynexochida" pageId="0" pageNumber="115" phylum="Arthropoda" rank="genus">Scutellum</taxonomicName>
pale yellow. Elytra dark brown with yellow apical patch. Hind wings dark brown. Thorax and legs light brown, darker on the dorsal surface of femora and pro- and mesothoracic tarsi. Abdomen dark brown, yellow on the margins of each ventrite and tergite; last ventrite completely brown.
</paragraph>
<footnote id="E8736267FFD6CB37FF32F7ED0DFBF7E0" pageId="0" pageNumber="115">
<paragraph id="8BD77E69FFD6CB37FF32F7ED0DFBF7E0" blockId="0.[151,499,2049,2106]" pageId="0" pageNumber="115">
Manuscript accepted 0 9.01.
<docYear id="930E9292FFD6CB37FE1EF7EE0DDEF7C2" box="[443,499,2050,2074]" pageId="0" pageNumber="115">2017</docYear>
DOI: 10.5281/zenodo.
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</paragraph>
</footnote>
<caption id="DF172EE1FFD7CB36FF32FB0E0F90FACC" httpUri="https://zenodo.org/record/1040489/files/figure.png" pageId="1" pageNumber="116" targetBox="[187,1434,254,1201]" targetPageId="1">
<paragraph id="8BD77E69FFD7CB36FF32FB0E0F90FACC" blockId="1.[151,1437,1249,1301]" pageId="1" pageNumber="116">
Figs 1-4.
<taxonomicName id="4C6805EAFFD7CB36FF5FFB0E0E32FB2E" authority="Gabriel Biffi, 2017" authorityName="Gabriel Biffi" authorityYear="2017" box="[250,543,1249,1271]" class="Insecta" family="Cantharidae" genus="Paramaronius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="116" phylum="Arthropoda" rank="species" species="unituberculatus" status="sp. nov.">
<emphasis id="B91CA27BFFD7CB36FF5FFB0E0E32FB2E" box="[250,543,1249,1271]" italics="true" pageId="1" pageNumber="116">Paramaronius unituberculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22F1F00FFD7CB36FD80FB0F0E5DFB20" box="[549,624,1251,1272]" pageId="1" pageNumber="116" rank="species">sp. nov.</taxonomicNameLabel>
(1) Dorsal habitus of male (holotype, NMB). (2) Dorsal habitus of female (paratype, MZSP). (3) Detail of the elytra of male. (4) Detail of the elytra of female.
</paragraph>
</caption>
<paragraph id="8BD77E69FFD7CB36FF32FA0C0FA8F840" blockId="1.[151,757,1377,2073]" lastBlockId="1.[831,1437,1377,2073]" pageId="1" pageNumber="116">
<emphasis id="B91CA27BFFD7CB36FF32FA0C0CE3FA20" box="[151,206,1504,1528]" italics="true" pageId="1" pageNumber="116">Male</emphasis>
(
<figureCitation id="135362ECFFD7CB36FF7CFA0D0D38FA21" box="[217,277,1505,1529]" captionStart="Figs 1 - 4" captionStartId="1.[151,192,1250,1271]" captionTargetBox="[187,1434,254,1201]" captionTargetId="figure@1.[149,1439,254,1244]" captionTargetPageId="1" captionText="Figs 1 - 4. Paramaronius unituberculatus sp. nov. (1) Dorsal habitus of male (holotype, NMB). (2) Dorsal habitus of female (paratype, MZSP). (3) Detail of the elytra of male. (4) Detail of the elytra of female." httpUri="https://zenodo.org/record/1040489/files/figure.png" pageId="1" pageNumber="116">Fig. 1</figureCitation>
): head densely pubescent; vertex and occipital region flat, lateral margins slightly arcuate behind the eyes.
<emphasis id="B91CA27BFFD7CB36FF70F9CC0D17F9EF" box="[213,314,1568,1591]" italics="true" pageId="1" pageNumber="116">Antennae</emphasis>
long, with two antennomeres surpassing the apex of elytra; antennomere I slightly swollen apically, antennomere II three times shorter than I, antennomere III twice longer than II, antennomeres IV-VIII subequal in length, slightly longer than I, antennomeres VIII-XI progressively decreasing in length towards the apex.
<emphasis id="B91CA27BFFD7CB36FF32F90D0D2CF920" box="[151,257,1761,1784]" italics="true" pageId="1" pageNumber="116">Pronotum</emphasis>
as long as wide; anterior and posterior margins rounded; lateral margins slightly sinuate; anterior and posterior angles directed upwards.
<taxonomicName id="4C6805EAFFD7CB36FDB0F8CC0E53F8E0" box="[533,638,1824,1848]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Corynexochida" pageId="1" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="B91CA27BFFD7CB36FDB0F8CC0E53F8E0" box="[533,638,1824,1848]" italics="true" pageId="1" pageNumber="116">Scutellum</emphasis>
</taxonomicName>
triangular, apex widely rounded.
<emphasis id="B91CA27BFFD7CB36FE22F8AC0DE7F880" box="[391,458,1856,1880]" italics="true" pageId="1" pageNumber="116">Elytra</emphasis>
(
<figureCitation id="135362ECFFD7CB36FE7CF8AD0E34F881" box="[473,537,1857,1881]" captionStart="Figs 1 - 4" captionStartId="1.[151,192,1250,1271]" captionTargetBox="[187,1434,254,1201]" captionTargetId="figure@1.[149,1439,254,1244]" captionTargetPageId="1" captionText="Figs 1 - 4. Paramaronius unituberculatus sp. nov. (1) Dorsal habitus of male (holotype, NMB). (2) Dorsal habitus of female (paratype, MZSP). (3) Detail of the elytra of male. (4) Detail of the elytra of female." httpUri="https://zenodo.org/record/1040489/files/figure.png" pageId="1" pageNumber="116">Fig. 3</figureCitation>
) densely pubescent, short, each elytron 2.6 times longer than wide; internal margins dehiscent from apical half, apex rounded; each elytron with a deep and broad hollow meeting medially; a dense fringe of long setae on the anterior wall and margin of hollows; dorsal surface with a large apical tubercle.
<emphasis id="B91CA27BFFD7CB36FF32F7ED0CE4F7C0" box="[151,201,2049,2072]" italics="true" pageId="1" pageNumber="116">Legs</emphasis>
slender; tibiae longer than femora; tarsomeres gradually increasing in size from pro- to metathoracic leg; first metatarsomere 1.85 times longer than second and 3.65 times longer than third.
<emphasis id="B91CA27BFFD7CB36FB13FA4C0936FA60" box="[1206,1307,1440,1464]" italics="true" pageId="1" pageNumber="116">Abdomen</emphasis>
: abdominal glandular pores slightly prominent. Seventh abdominal ventrite (
<figureCitation id="135362ECFFD7CB36FC02FA0D0FC0FA21" box="[935,1005,1505,1529]" captionStart="Figs 9 - 12" captionStartId="3.[151,192,847,868]" captionTargetBox="[184,1432,258,781]" captionTargetId="figure@3.[148,1448,250,821]" captionTargetPageId="3" captionText="Figs 9 - 12. Paramaronius unituberculatus sp. nov. (9) Seventh ventrite of the male (holotype, NMB). (10) Seventh ventrite of the female (paratype, MZSP). (11) Right paramere of aedeagus (holotype, NMB). (12) Right coxite of female (paratype, MZSP). Scales: 9 - 10 = 0.5 mm, 11 - 12 = 0.2 mm." httpUri="https://zenodo.org/record/1040493/files/figure.png" pageId="1" pageNumber="116">Fig. 9</figureCitation>
) slightly wider than long, pubescent, distal margin slightly emarginated laterally and strongly emarginated and rounded at middle.
<emphasis id="B91CA27BFFD7CB36FB57F9CC0977F9E0" box="[1266,1370,1568,1592]" italics="true" pageId="1" pageNumber="116">Aedeagus</emphasis>
(
<figureCitation id="135362ECFFD7CB36FAD4F9CD0F4EF980" captionStart="Figs 5 - 8" captionStartId="2.[151,192,1707,1728]" captionTargetBox="[149,1439,259,1662]" captionTargetId="figure@2.[148,1439,254,1669]" captionTargetPageId="2" captionText="Figs 5 - 8. Aedeagus of Paramaronius unituberculatus sp. nov. (holotype, NMB). (5) Dorsal view. (6) Left view. (7) Ventral view. (8) Right view. Scale = 0.5 mm." httpUri="https://zenodo.org/record/1040491/files/figure.png" pageId="1" pageNumber="116">Fig. 5-8</figureCitation>
) with a large right prolongation of tegmen, broadly sinuous on posterior margin, covered with long and thick setae; base and dorsal surface with projecting apophysis; left setiferous prolongation long and curved, slightly clubbed, with few setae at apex; right paramere (
<figureCitation id="135362ECFFD7CB36FAEDF92D09B9F901" box="[1352,1428,1729,1753]" captionStart="Figs 9 - 12" captionStartId="3.[151,192,847,868]" captionTargetBox="[184,1432,258,781]" captionTargetId="figure@3.[148,1448,250,821]" captionTargetPageId="3" captionText="Figs 9 - 12. Paramaronius unituberculatus sp. nov. (9) Seventh ventrite of the male (holotype, NMB). (10) Seventh ventrite of the female (paratype, MZSP). (11) Right paramere of aedeagus (holotype, NMB). (12) Right coxite of female (paratype, MZSP). Scales: 9 - 10 = 0.5 mm, 11 - 12 = 0.2 mm." httpUri="https://zenodo.org/record/1040493/files/figure.png" pageId="1" pageNumber="116">Fig. 11</figureCitation>
) long, curved, narrowing upwards and divided in two opposite processes reaching both apophysis of tegmen; left paramere flat, long, divided apically and covering the base of dorsal surface of median lobe; median lobe long and slightly curved to the right; left surface of tegmen rough.
</paragraph>
<paragraph id="8BD77E69FFD7CB35FC9AF84C0E3BF8A0" blockId="1.[831,1437,1377,2073]" lastBlockId="2.[151,756,1825,1912]" lastPageId="2" lastPageNumber="117" pageId="1" pageNumber="116">
<emphasis id="B91CA27BFFD7CB36FC9AF84C0FA2F860" box="[831,911,1952,1976]" italics="true" pageId="1" pageNumber="116">Female</emphasis>
(
<figureCitation id="135362ECFFD7CB36FC01F84D0FC1F861" box="[932,1004,1953,1977]" captionStart="Figs 1 - 4" captionStartId="1.[151,192,1250,1271]" captionTargetBox="[187,1434,254,1201]" captionTargetId="figure@1.[149,1439,254,1244]" captionTargetPageId="1" captionText="Figs 1 - 4. Paramaronius unituberculatus sp. nov. (1) Dorsal habitus of male (holotype, NMB). (2) Dorsal habitus of female (paratype, MZSP). (3) Detail of the elytra of male. (4) Detail of the elytra of female." httpUri="https://zenodo.org/record/1040489/files/figure.png" pageId="1" pageNumber="116">Fig. 2</figureCitation>
): slightly smaller than the male; eyes smaller and more rounded; dorsal surface of elytra (
<figureCitation id="135362ECFFD7CB36FCE2F80D0FBAF821" box="[839,919,2017,2041]" captionStart="Figs 1 - 4" captionStartId="1.[151,192,1250,1271]" captionTargetBox="[187,1434,254,1201]" captionTargetId="figure@1.[149,1439,254,1244]" captionTargetPageId="1" captionText="Figs 1 - 4. Paramaronius unituberculatus sp. nov. (1) Dorsal habitus of male (holotype, NMB). (2) Dorsal habitus of female (paratype, MZSP). (3) Detail of the elytra of male. (4) Detail of the elytra of female." httpUri="https://zenodo.org/record/1040489/files/figure.png" pageId="1" pageNumber="116">Fig. 4</figureCitation>
) without sculptural modifications; seventh abdominal ventrite (
<figureCitation id="135362ECFFD7CB36FB82F7ED0851F7C1" box="[1063,1148,2049,2073]" captionStart="Figs 9 - 12" captionStartId="3.[151,192,847,868]" captionTargetBox="[184,1432,258,781]" captionTargetId="figure@3.[148,1448,250,821]" captionTargetPageId="3" captionText="Figs 9 - 12. Paramaronius unituberculatus sp. nov. (9) Seventh ventrite of the male (holotype, NMB). (10) Seventh ventrite of the female (paratype, MZSP). (11) Right paramere of aedeagus (holotype, NMB). (12) Right coxite of female (paratype, MZSP). Scales: 9 - 10 = 0.5 mm, 11 - 12 = 0.2 mm." httpUri="https://zenodo.org/record/1040493/files/figure.png" pageId="1" pageNumber="116">Fig. 10</figureCitation>
) narrowed apicad, distal margin almost straight and with a small notch at middle; coxites (
<figureCitation id="135362ECFFD4CB35FF56F8AD0D6FF881" box="[243,322,1857,1881]" captionStart="Figs 9 - 12" captionStartId="3.[151,192,847,868]" captionTargetBox="[184,1432,258,781]" captionTargetId="figure@3.[148,1448,250,821]" captionTargetPageId="3" captionText="Figs 9 - 12. Paramaronius unituberculatus sp. nov. (9) Seventh ventrite of the male (holotype, NMB). (10) Seventh ventrite of the female (paratype, MZSP). (11) Right paramere of aedeagus (holotype, NMB). (12) Right coxite of female (paratype, MZSP). Scales: 9 - 10 = 0.5 mm, 11 - 12 = 0.2 mm." httpUri="https://zenodo.org/record/1040493/files/figure.png" pageId="2" pageNumber="117">Fig. 12</figureCitation>
) small and membranous, base long and slender; styles short, wider apically.
</paragraph>
<caption id="DF172EE1FFD4CB35FF32F9470E21F906" httpUri="https://zenodo.org/record/1040491/files/figure.png" pageId="2" pageNumber="117" targetBox="[149,1439,259,1662]" targetPageId="2">
<paragraph id="8BD77E69FFD4CB35FF32F9470E21F906" blockId="2.[151,1436,1707,1758]" pageId="2" pageNumber="117">
Figs 5-8. Aedeagus of
<taxonomicName id="4C6805EAFFD4CB35FEDFF9470E8DF918" authority="Gabriel Biffi, 2017" authorityName="Gabriel Biffi" authorityYear="2017" box="[378,672,1707,1728]" class="Insecta" family="Cantharidae" genus="Paramaronius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="species" species="unituberculatus" status="sp. nov.">
<emphasis id="B91CA27BFFD4CB35FEDFF9470E8DF918" box="[378,672,1707,1728]" italics="true" pageId="2" pageNumber="117">Paramaronius unituberculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22F1F00FFD4CB35FD03F9400EDFF919" box="[678,754,1708,1729]" pageId="2" pageNumber="117" rank="species">sp. nov.</taxonomicNameLabel>
(holotype, NMB). (5) Dorsal view. (6) Left view. (7) Ventral view. (8) Right view. Scale = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BD77E69FFD4CB35FF32F8630E19F870" blockId="2.[151,564,1935,1960]" box="[151,564,1935,1960]" pageId="2" pageNumber="117">
<emphasis id="B91CA27BFFD4CB35FF32F8630D09F87F" bold="true" box="[151,292,1935,1959]" pageId="2" pageNumber="117">Dimensions:</emphasis>
Total length 7.5-9.0 mm.
</paragraph>
</subSubSection>
<subSubSection id="C3722DE2FFD4CB34FF32F8520E59FBB5" lastPageId="3" lastPageNumber="118" pageId="2" pageNumber="117" type="discussion">
<paragraph id="8BD77E69FFD4CB34FF32F8520E59FBB5" blockId="2.[151,756,1982,2070]" lastBlockId="3.[151,756,1014,1133]" lastPageId="3" lastPageNumber="118" pageId="2" pageNumber="117">
<emphasis id="B91CA27BFFD4CB35FF32F8520D24F80E" bold="true" box="[151,265,1982,2006]" pageId="2" pageNumber="117">Remarks:</emphasis>
<taxonomicName id="4C6805EAFFD4CB35FEBDF8530E47F80E" authority="Gabriel Biffi, 2017" authorityName="Gabriel Biffi" authorityYear="2017" box="[280,618,1982,2006]" class="Insecta" family="Cantharidae" genus="Paramaronius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="species" species="unituberculatus" status="sp. nov.">
<emphasis id="B91CA27BFFD4CB35FEBDF8530E47F80E" box="[280,618,1982,2006]" italics="true" pageId="2" pageNumber="117">Paramaronius unituberculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22F1F00FFD4CB35FDDDF82C0EF9F80F" box="[632,724,1984,2007]" pageId="2" pageNumber="117" rank="species">sp. nov.</taxonomicNameLabel>
is similar to
<emphasis id="B91CA27BFFD4CB35FEA8F8330D5AF82E" box="[269,375,2014,2038]" italics="true" pageId="2" pageNumber="117">P. kraatzi</emphasis>
(Pic, 1938) and
<taxonomicName id="4C6805EAFFD4CB35FD99F8330CE1F7CE" authority="Wittmer, 1963" authorityName="Wittmer" authorityYear="1963" class="Insecta" family="Cantharidae" genus="Paramaronius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="species" species="freyi">
<emphasis id="B91CA27BFFD4CB35FD99F8330EA6F82E" box="[572,651,2014,2038]" italics="true" pageId="2" pageNumber="117">P. freyi</emphasis>
<bibRefCitation id="EFF90398FFD4CB35FD33F8330CE1F7CE" author="Wittmer W." journalOrPublisher="Frey. Entomologische Arbeiten aus dem Museum Georg Frey" pageId="2" pageNumber="117" pagination="128 - 135" part="14" refString="Wittmer W. 1963. Neue neotropische Malacodermata aus dem Museum G. Frey. Entomologische Arbeiten aus dem Museum Georg Frey 14: 128 - 135." title="Neue neotropische Malacodermata aus dem Museum G" type="journal article" year="1963">
Wittmer,
<quantity id="4C90D38CFFD4CB35FF32F8130CE1F7CE" box="[151,204,2047,2070]" metricMagnitude="1" metricUnit="m" metricValue="4.98602" pageId="2" pageNumber="117" unit="in" value="1963.0">1963</quantity>
</bibRefCitation>
</taxonomicName>
in the typical elytral sculpture and in the rugosity on the left surface of the tegmen. It differs from these species by the colouration of head, antennae, pronotum and legs, apex of elytra with just one tubercle (two tubercles in
<emphasis id="B91CA27BFFD4CB35FC69F86D081BF840" box="[972,1078,1920,1944]" italics="true" pageId="2" pageNumber="117">P. kraatzi</emphasis>
and three in
<taxonomicName id="4C6805EAFFD4CB35FB76F86D090FF840" authorityName="Wittmer" authorityYear="1963" box="[1235,1314,1920,1944]" class="Insecta" family="Cantharidae" genus="Paramaronius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="species" species="freyi">
<emphasis id="B91CA27BFFD4CB35FB76F86D090FF840" box="[1235,1314,1920,1944]" italics="true" pageId="2" pageNumber="117">P. freyi</emphasis>
</taxonomicName>
), shape of seventh abdominal ventrite of males and females, shape of aedeagus and parameres, and less roughness of tegmen. Males of
<emphasis id="B91CA27BFFD4CB35FBAAF80D08E3F820" box="[1039,1230,2017,2040]" italics="true" pageId="2" pageNumber="117">P. impressipennis</emphasis>
(Pic, 1906), from
<collectingCountry id="F37F3EF9FFD4CB35FC9AF7ED0F87F7C0" box="[831,938,2049,2072]" name="Argentina" pageId="2" pageNumber="117">Argentina</collectingCountry>
and
<collectingCountry id="F37F3EF9FFD4CB35FC48F7ED081FF7C0" box="[1005,1074,2049,2072]" name="Brazil" pageId="2" pageNumber="117">Brazil</collectingCountry>
, differ from
<taxonomicName id="4C6805EAFFD4CB35FB6AF7ED09B0F7C0" authority="Gabriel Biffi, 2017" authorityName="Gabriel Biffi" authorityYear="2017" box="[1231,1437,2048,2073]" class="Insecta" family="Cantharidae" genus="Paramaronius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="117" phylum="Arthropoda" rank="species" species="unituberculatus" status="sp. nov.">
<emphasis id="B91CA27BFFD4CB35FB6AF7ED08F2F7C0" box="[1231,1247,2049,2072]" italics="true" pageId="2" pageNumber="117">P</emphasis>
.
<emphasis id="B91CA27BFFD4CB35FB51F7EC09B0F7C0" box="[1268,1437,2048,2072]" italics="true" pageId="2" pageNumber="117">unituberculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22F1F00FFD5CB34FF32FC1B0CC1FBD6" box="[151,236,1015,1038]" pageId="3" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
by the presence of elytra with one longitudinal ridge on posterior half, without deep grooves and tubercles. Females differ especially by the shape of distal margin of the last abdominal ventrites.
</paragraph>
</subSubSection>
<caption id="DF172EE1FFD5CB34FF32FCA30EA5FC47" httpUri="https://zenodo.org/record/1040493/files/figure.png" pageId="3" pageNumber="118" targetBox="[184,1432,258,781]" targetPageId="3">
<paragraph id="8BD77E69FFD5CB34FF32FCA30EA5FC47" blockId="3.[151,1437,847,927]" pageId="3" pageNumber="118">
Figs 9-12.
<taxonomicName id="4C6805EAFFD5CB34FEA7FCBC0E05FCBC" authority="Gabriel Biffi, 2017" authorityName="Gabriel Biffi" authorityYear="2017" box="[258,552,847,869]" class="Insecta" family="Cantharidae" genus="Paramaronius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="118" phylum="Arthropoda" rank="species" species="unituberculatus" status="sp. nov.">
<emphasis id="B91CA27BFFD5CB34FEA7FCBC0E05FCBC" box="[258,552,847,869]" italics="true" pageId="3" pageNumber="118">Paramaronius unituberculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22F1F00FFD5CB34FD8AFCBC0E56FCBD" box="[559,635,848,869]" pageId="3" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
(9) Seventh ventrite of the male (holotype, NMB). (10) Seventh ventrite of the female (paratype, MZSP). (11) Right paramere of aedeagus (holotype, NMB). (12) Right coxite of female (paratype, MZSP). Scales: 9-10 = 0.5 mm, 11-12 = 0.2 mm.
</paragraph>
</caption>
<subSubSection id="C3722DE2FFD5CB34FF32FB690EDDF885" pageId="3" pageNumber="118" type="key">
<paragraph id="8BD77E69FFD5CB34FF32FB690E9DFB64" blockId="3.[151,757,1157,1885]" pageId="3" pageNumber="118">
The most recent identification key (
<bibRefCitation id="EFF90398FFD5CB34FD80FB690E8DFB44" author="Biffi G." box="[549,672,1157,1181]" journalOrPublisher="ZooKeys" pageId="3" pageNumber="118" pagination="49 - 69" part="516" refString="Biffi G. 2015. Three new species and distributional records for Paramaronius Wittmer (Coleoptera, Cantharidae, Chauliognathinae). ZooKeys 516: 49 - 69." title="Three new species and distributional records for Paramaronius Wittmer (Coleoptera, Cantharidae, Chauliognathinae)" type="journal article" year="2015">Biffi, 2015</bibRefCitation>
) has to be modified to include the new species as follows:
</paragraph>
<paragraph id="8BD77E69FFD5CB34FF32FB280EC2FAE4" blockId="3.[151,757,1157,1885]" pageId="3" pageNumber="118">
<emphasis id="B91CA27BFFD5CB34FF32FB280C89FB04" bold="true" box="[151,164,1220,1244]" pageId="3" pageNumber="118">4</emphasis>
Head, pronotum and legs completely testaceous; antennae testaceous, darker from antennomere 7 to apex; males: each elytron with one median and two apical tubercles ....................
<emphasis id="B91CA27BFFD5CB34FDE8FAC90EC2FAE4" bold="true" box="[589,751,1316,1340]" pageId="3" pageNumber="118">
<emphasis id="B91CA27BFFD5CB34FDE8FAC90E99FAE4" bold="true" box="[589,692,1316,1340]" italics="true" pageId="3" pageNumber="118">P. kraatzi</emphasis>
(Pic)
</emphasis>
</paragraph>
<paragraph id="8BD77E69FFD5CB34FF32FAAA0EC2FA64" blockId="3.[151,757,1157,1885]" pageId="3" pageNumber="118">
Head dark brown to black; pronotum completely yellow or with two brown patches on lateral margin; males: each elytron with one or three apical tubercles ................................................
<emphasis id="B91CA27BFFD5CB34FD70FA480EC2FA64" bold="true" box="[725,751,1444,1468]" pageId="3" pageNumber="118">4a</emphasis>
</paragraph>
<paragraph id="8BD77E69FFD5CB34FF32FA280EC2F944" blockId="3.[151,757,1157,1885]" pageId="3" pageNumber="118">
<emphasis id="B91CA27BFFD5CB34FF32FA280C9CFA04" bold="true" box="[151,177,1476,1500]" pageId="3" pageNumber="118">4a</emphasis>
Head black; pronotum completely yellow or with two brown spots on lateral margins; legs yellow with brown spots on apex of femora and tibiae, tarsi brown; antennae brown, slightly lighter on two first and three last antennomeres; males: each elytron with one median and three apical tubercles.....................................
<taxonomicName id="4C6805EAFFD5CB34FD99F9690EC2F944" authority="Wittmer" authorityName="Wittmer" authorityYear="1963" box="[572,751,1668,1692]" class="Insecta" family="Cantharidae" genus="Paramaronius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="118" phylum="Arthropoda" rank="species" species="freyi">
<emphasis id="B91CA27BFFD5CB34FD99F9690EC2F944" bold="true" box="[572,751,1668,1692]" pageId="3" pageNumber="118">
<emphasis id="B91CA27BFFD5CB34FD99F9690EA5F944" bold="true" box="[572,648,1668,1692]" italics="true" pageId="3" pageNumber="118">P. freyi</emphasis>
Wittmer
</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BD77E69FFD5CB34FF32F94A0EDDF885" blockId="3.[151,757,1157,1885]" pageId="3" pageNumber="118">
Head dark brown; pronotum with two brown spots on lateral margins; legs light brown, darker on dorsal surface of femora, tarsi brown; antenna completely dark brown; males: each elytron with low median and apical tubercle (
<figureCitation id="135362ECFFD5CB34FD9AF8C90E50F8E6" box="[575,637,1829,1854]" captionStart="Figs 1 - 4" captionStartId="1.[151,192,1250,1271]" captionTargetBox="[187,1434,254,1201]" captionTargetId="figure@1.[149,1439,254,1244]" captionTargetPageId="1" captionText="Figs 1 - 4. Paramaronius unituberculatus sp. nov. (1) Dorsal habitus of male (holotype, NMB). (2) Dorsal habitus of female (paratype, MZSP). (3) Detail of the elytra of male. (4) Detail of the elytra of female." httpUri="https://zenodo.org/record/1040489/files/figure.png" pageId="3" pageNumber="118">Fig. 3</figureCitation>
) ............... ..................................
<taxonomicName id="4C6805EAFFD5CB34FE68F8A90EBAF884" authority="Gabriel Biffi, 2017" authorityName="Gabriel Biffi" authorityYear="2017" box="[461,663,1860,1884]" class="Insecta" family="Cantharidae" genus="Paramaronius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="118" phylum="Arthropoda" rank="species" species="unituberculatus" status="sp. nov.">
<emphasis id="B91CA27BFFD5CB34FE68F8A90EBAF884" bold="true" box="[461,663,1860,1884]" italics="true" pageId="3" pageNumber="118">P. unituberculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22F1F00FFD5CB34FD38F8AA0EDDF885" box="[669,752,1862,1885]" pageId="3" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>