405 lines
55 KiB
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405 lines
55 KiB
XML
<document id="6928B4820A9D6E2729AC6BBEC62EC69E" ID-CLB-Dataset="20988" ID-DOI="10.5852/ejt.2020.713" ID-GBIF-Dataset="1d5fdead-d612-4c66-b482-4f051f990351" ID-Zenodo-Dep="4020752" ID-ZooBank="A6CB58F5-1ECC-47F0-AA07-798844AF80A7" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1599662866733" checkinUser="valdenar" docAuthor="Antić, Dragan Ž. & Reip, Hans S." docDate="2020" docId="03A963622845FFB62DFC19AD0EFCFAFE" docLanguage="en" docName="ejt-713_antic_reip.pdf" docOrigin="European Journal of Taxonomy 713" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Leucogeorgia satunini Antić & Reip 2020, comb. nov." docType="treatment" docVersion="9" lastPageNumber="28" masterDocId="FF901B1A285DFFAD2C24183E0F08FFEE" masterDocTitle="The millipede genus Leucogeorgia Verhoeff, 1930 in the Caucasus, with descriptions of eleven new species, erection of a new monotypic genus and notes on the tribe Leucogeorgiini (Diplopoda: Julida: Julidae)" masterLastPageNumber="106" masterPageNumber="1" pageNumber="25" updateTime="1698845858502" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="E9D3474270604DCE956223C4B9422F62">The millipede genus Leucogeorgia Verhoeff, 1930 in the Caucasus, with descriptions of eleven new species, erection of a new monotypic genus and notes on the tribe Leucogeorgiini (Diplopoda: Julida: Julidae)</mods:title>
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<mods:roleTerm id="0B4B0B8E276699E194CE1CF8BA6D1210">Author</mods:roleTerm>
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<mods:namePart id="B47ACE151EFCF7F7C8BC34B7111BC12C">Antić, Dragan Ž.</mods:namePart>
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<mods:affiliation id="3E63CF0A088E9535F528F5431C8025AB">University of Belgrade - Faculty of Biology, Institute of Zoology, Studentski Trg 16, 11000 Belgrade, Serbia. Serbian Biospeleological Society, Trg Dositeja Obradovića 2, 21000 Novi Sad, Serbia. Department of Soil Zoology, Senckenberg Museum of Natural History Görlitz, Am Museum 1, 02826 Görlitz, Germany. Thüringer Höhlenverein e. V., Johann-Sebastian-Bach-Str. 16 a, 98529 Suhl, Germany.</mods:affiliation>
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<mods:nameIdentifier id="EB345C943906CAD346AD0394D6176524" type="email">dragan.antic@bio.bg.ac.rs</mods:nameIdentifier>
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<mods:namePart id="DEBA8825EE754F368375544C74E22450">Reip, Hans S.</mods:namePart>
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<mods:nameIdentifier id="AADC479B9081F053ABE16FF78526D694" type="email">reip@myriapoden-info.de</mods:nameIdentifier>
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<mods:title id="B38FFDA199DCCFFA12FA3B291F5BFDF5">European Journal of Taxonomy</mods:title>
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<mods:part id="F380DF0916CD1352C1D88224D5246000">
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<mods:date id="47840664F9352B4E0EEDA35F7A8E4628">2020</mods:date>
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<mods:number id="17B9044B387D9B4319979B53FB4C6206">2020-09-07</mods:number>
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<mods:number id="9796E8B51509F26016F5851143ACCD41">713</mods:number>
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<treatment id="03A963622845FFB62DFC19AD0EFCFAFE" ID-DOI="http://doi.org/10.5281/zenodo.4335844" ID-GBIF-Taxon="167523048" ID-Zenodo-Dep="4335844" LSID="urn:lsid:plazi:treatment:03A963622845FFB62DFC19AD0EFCFAFE" httpUri="http://treatment.plazi.org/id/03A963622845FFB62DFC19AD0EFCFAFE" lastPageId="27" lastPageNumber="28" pageId="24" pageNumber="25">
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<subSubSection id="C31A81FF2845FFB52DFC19AD0B54FE40" box="[472,1116,403,430]" pageId="24" pageNumber="25" type="nomenclature">
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<paragraph id="8BBFD2742845FFB52DFC19AD0B54FE40" blockId="24.[472,1116,403,465]" box="[472,1116,403,430]" pageId="24" pageNumber="25">
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<heading id="D0F765182845FFB52DFC19AD0B54FE40" box="[472,1116,403,430]" centered="true" fontSize="11" level="2" pageId="24" pageNumber="25" reason="2">
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<taxonomicName id="4C00A9F72845FFB52DFC19AD0CDBFE40" ID-CoL="849PY" authority="(Lohmander, 1936)" authorityName="Antić & Reip" authorityYear="2020" baseAuthorityName="Lohmander" baseAuthorityYear="1936" box="[472,979,403,430]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="satunini" status="comb. nov.">
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<emphasis id="B9740E662845FFB52DFC19AD0DE0FE43" bold="true" box="[472,744,403,429]" italics="true" pageId="24" pageNumber="25">Leucogeorgia satunini</emphasis>
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(
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<bibRefCitation id="EF91AF852845FFB52EDC19AD0CC3FE40" author="Lohmander H." box="[760,971,403,430]" pageId="24" pageNumber="25" pagination="1 - 196" refId="ref41985" refString="Lohmander H. 1936. Uber die Diplopoden des Kaukasusgebietes. Goteborgs Kungliga Vetenskaps- och Vitterhets-Samhalles Handlingar, Series 5 B 5 (1): 1 - 196." type="journal article" year="1936">Lohmander, 1936</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel id="A247B31D2845FFB52FFE19AD0B54FE40" box="[986,1116,403,430]" pageId="24" pageNumber="25" rank="species">comb. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C31A81FF2845FFB52EB719880CA9FE3F" box="[659,929,438,465]" pageId="24" pageNumber="25" type="description">
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<paragraph id="8BBFD2742845FFB52EB719880CA9FE3F" blockId="24.[472,1116,403,465]" box="[659,929,438,465]" pageId="24" pageNumber="25">
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<figureCitation id="133BCEF12845FFB52EB719880DFBFE3E" box="[659,755,438,464]" captionStart="Fig" captionStartId="5.[189,232,1788,1814]" captionTargetBox="[245,1342,265,1760]" captionTargetId="figure@5.[245,1342,265,1760]" captionTargetPageId="5" captionText="Fig. 2. Gonopods, mesal view. A. Leucogeorgia lobata sp. nov., right gonopods, paratype ♂ from Kalitshona Cave (ZMUM). B. L. longipes Verhoeff, 1930, right gonopods, ♂ from Dolabistavi Cave (SMNG). C. L. prometheus sp. nov., left gonopods, non-type ♂ from Sataplia II Cave (ZMUM). D. L. satunini (Lohmander, 1936) comb. nov., right gonopods, ♂ from Kelasurskaya Cave (ZMUM). Abbreviations: am = accessory membrane; f = flagellum; l = lobe; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum." figureDoi="http://doi.org/10.5281/zenodo.4020756" httpUri="https://zenodo.org/record/4020756/files/figure.png" pageId="24" pageNumber="25">Figs 2D</figureCitation>
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,
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<figureCitation id="133BCEF12845FFB52EDB19880C43FE3F" box="[767,843,438,465]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="25.[189,232,1895,1921]" captionStartId-1="26.[189,232,1858,1884]" captionTargetBox-0="[277,1311,265,1869]" captionTargetBox-1="[255,1333,265,1830]" captionTargetId-0="figure@25.[277,1311,265,1869]" captionTargetId-1="figure@26.[255,1333,265,1830]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionText-0="Fig. 14. Leucogeorgia satunini (Lohmander, 1936) comb. nov., ♀ and ♂ from Kelasurskaya Cave (ZMUM). A. ♀, habitus, lateral view. B. ♀, anterior part of body, lateral view. C. ♂, midbody rings, lateral view. D. ♂, body ring 7, lateral view. E. ♂, body ring 7, ventral view. F. ♂, posterior part of body, lateral view. Scale bars: 1 mm." captionText-1="Fig. 15. Leucogeorgia satunini (Lohmander, 1936) comb. nov., ♂ from Kelasurskaya Cave (ZMUM). A. Head, ventral view. B. Right gonopods, mesal view. C. Left gonopods, mesal view. D. Left gonopods, lateral view. Abbreviations: am = accessory membrane; f = flagellum; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bars: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4020782" figureDoi-1="http://doi.org/10.5281/zenodo.4020784" httpUri-0="https://zenodo.org/record/4020782/files/figure.png" httpUri-1="https://zenodo.org/record/4020784/files/figure.png" pageId="24" pageNumber="25">14–15</figureCitation>
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,
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<figureCitation id="133BCEF12845FFB52F7319880CA9FE3F" box="[855,929,438,465]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="96.[189,232,1921,1947]" captionStartId-1="97.[189,232,1991,2017]" captionTargetBox-0="[189,1399,979,1893]" captionTargetBox-1="[189,1399,890,1963]" captionTargetId-0="figure@96.[189,1399,979,1893]" captionTargetId-1="figure@97.[189,1399,890,1963]" captionTargetPageId-0="96" captionTargetPageId-1="97" captionText-0="Fig. 56. General distribution of the Caucasian leucogeorgiinines. The white dashed line hypothetically indicates the boundary between the northwestern and southeastern groups of species of the genus Leucogeorgia Verhoeff, 1930." captionText-1="Fig. 57. Distribution of the northwestern group of the genus Leucogeorgia Verhoeff, 1930." figureDoi-0="http://doi.org/10.5281/zenodo.4020873" figureDoi-1="http://doi.org/10.5281/zenodo.4020875" httpUri-0="https://zenodo.org/record/4020873/files/figure.png" httpUri-1="https://zenodo.org/record/4020875/files/figure.png" pageId="24" pageNumber="25">56–57</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31A81FF2845FFB52C9919C60C0FFDFD" box="[189,775,504,531]" pageId="24" pageNumber="25" type="reference_group">
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||
<paragraph id="8BBFD2742845FFB52C9919C60C0FFDFD" blockId="24.[189,775,504,531]" box="[189,775,504,531]" pageId="24" pageNumber="25">
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||
<treatmentCitationGroup id="AB10F55A2845FFB52C9919C60C0FFDFD" box="[189,775,504,531]" pageId="24" pageNumber="25">
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<treatmentCitation id="0AA1F4652845FFB52C9919C60C0CFDFD" author="Lohmander H." box="[189,772,504,531]" page="95" pageId="24" pageNumber="25" year="1936">
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<taxonomicName id="4C00A9F72845FFB52C9919C60C0CFDFD" ID-CoL="7RRDC" authority="Lohmander, 1936: 95" authorityName="Lohmander" authorityPageNumber="95" authorityYear="1936" box="[189,772,504,531]" class="Diplopoda" family="Julidae" genus="Archileucogeorgia" kingdom="Animalia" order="Julida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="satunini">
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<emphasis id="B9740E662845FFB52C9919C60EF5FDFC" box="[189,509,504,530]" italics="true" pageId="24" pageNumber="25">Archileucogeorgia satunini</emphasis>
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<bibRefCitation id="EF91AF852845FFB52E2019C60C0CFDFD" author="Lohmander H." box="[516,772,504,531]" pageId="24" pageNumber="25" pagination="1 - 196" refId="ref41985" refString="Lohmander H. 1936. Uber die Diplopoden des Kaukasusgebietes. Goteborgs Kungliga Vetenskaps- och Vitterhets-Samhalles Handlingar, Series 5 B 5 (1): 1 - 196." type="journal article" year="1936">Lohmander, 1936: 95</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31A81FF2845FFB52C991A070BC3FB58" pageId="24" pageNumber="25" type="diagnosis">
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<paragraph id="8BBFD2742845FFB52C991A070E3EFDBD" blockId="24.[189,310,569,595]" box="[189,310,569,595]" pageId="24" pageNumber="25">
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<heading id="D0F765182845FFB52C991A070E3EFDBD" bold="true" box="[189,310,569,595]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
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<emphasis id="B9740E662845FFB52C991A070E3EFDBD" bold="true" box="[189,310,569,595]" pageId="24" pageNumber="25">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BBFD2742845FFB52C991A560BC3FB58" blockId="24.[189,1399,616,1206]" pageId="24" pageNumber="25">
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This species belongs to the group of
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<taxonomicName id="4C00A9F72845FFB52E491A570C07FD6C" authorityName="Verhoeff" authorityYear="1930" box="[621,783,617,642]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
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<emphasis id="B9740E662845FFB52E491A570C07FD6C" box="[621,783,617,642]" italics="true" pageId="24" pageNumber="25">Leucogeorgia</emphasis>
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</taxonomicName>
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spp. without modified mouthparts and with teeth on the mesomeral claw (vs their absence in
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<taxonomicName id="4C00A9F72845FFB52E8D1AB20C4FFD4B" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[681,839,651,677]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="golovatchi" status="sp. nov.">
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<emphasis id="B9740E662845FFB52E8D1AB20C4FFD4B" box="[681,839,651,677]" italics="true" pageId="24" pageNumber="25">L. golovatchi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A247B31D2845FFB52F741AB20CA7FD48" box="[848,943,652,678]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
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,
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<taxonomicName id="4C00A9F72845FFB52F9A1AB20B22FD4B" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[958,1066,651,677]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="lobata" status="sp. nov.">
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<emphasis id="B9740E662845FFB52F9A1AB20B22FD4B" box="[958,1066,651,677]" italics="true" pageId="24" pageNumber="25">L. lobata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A247B31D2845FFB528171AB20B98FD48" box="[1075,1168,652,678]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
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and
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<taxonomicName id="4C00A9F72845FFB528E91AB20A7FFD4B" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[1229,1399,651,677]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="prometheus" status="sp. nov.">
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<emphasis id="B9740E662845FFB528E91AB20A7FFD4B" box="[1229,1399,651,677]" italics="true" pageId="24" pageNumber="25">L. prometheus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A247B31D2845FFB52C991A910E14FD27" box="[189,284,687,713]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
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).
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<taxonomicName id="4C00A9F72845FFB52D151A910D32FD26" authorityName="Antić & Reip" authorityYear="2020" baseAuthorityName="Lohmander" baseAuthorityYear="1936" box="[305,570,687,712]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="satunini">
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<emphasis id="B9740E662845FFB52D151A910D32FD26" box="[305,570,687,712]" italics="true" pageId="24" pageNumber="25">Leucogeorgia satunini</emphasis>
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</taxonomicName>
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differs from
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<taxonomicName id="4C00A9F72845FFB52EF21A910C65FD26" authorityName="Antić & Reip" authorityYear="2020" baseAuthorityName="Lohmander" baseAuthorityYear="1936" box="[726,877,686,712]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="abchasica">
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<emphasis id="B9740E662845FFB52EF21A910C65FD26" box="[726,877,686,712]" italics="true" pageId="24" pageNumber="25">L. abchasica</emphasis>
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</taxonomicName>
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by having a mesomeral lamella of the same height all along and the presence of a posterior rise (vs height of the mesomeral lamella gradually decreasing caudad, also lacking a posterior rise in
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<taxonomicName id="4C00A9F72845FFB52F0B1ACB0CC1FCE1" authorityName="Antić & Reip" authorityYear="2020" baseAuthorityName="Lohmander" baseAuthorityYear="1936" box="[815,969,757,783]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="abchasica">
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<emphasis id="B9740E662845FFB52F0B1ACB0CC1FCE1" box="[815,969,757,783]" italics="true" pageId="24" pageNumber="25">L. abchasica</emphasis>
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</taxonomicName>
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), and by having a rounded ventral margin of male body ring 7 without strong teeth in ventral view (vs subtriangular ventral margin of male body ring 7 with strong teeth in ventral view in
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<taxonomicName id="4C00A9F72845FFB52ED41B020C8EFCBB" authorityName="Antić & Reip" authorityYear="2020" baseAuthorityName="Lohmander" baseAuthorityYear="1936" box="[752,902,827,853]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="abchasica">
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<emphasis id="B9740E662845FFB52ED41B020C8EFCBB" box="[752,902,827,853]" italics="true" pageId="24" pageNumber="25">L. abchasica</emphasis>
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</taxonomicName>
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).
|
||
<taxonomicName id="4C00A9F72845FFB52FBB1B020BAFFCBB" authorityName="Antić & Reip" authorityYear="2020" baseAuthorityName="Lohmander" baseAuthorityYear="1936" box="[927,1191,828,853]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="satunini">
|
||
<emphasis id="B9740E662845FFB52FBB1B020BAFFCBB" box="[927,1191,828,853]" italics="true" pageId="24" pageNumber="25">Leucogeorgia satunini</emphasis>
|
||
</taxonomicName>
|
||
differs from both
|
||
<taxonomicName id="4C00A9F72845FFB52C991B610E34FC96" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[189,316,862,888]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="borealis" status="sp. nov.">
|
||
<emphasis id="B9740E662845FFB52C991B610E34FC96" box="[189,316,862,888]" italics="true" pageId="24" pageNumber="25">L. borealis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D2845FFB52D611B610EA9FC97" box="[325,417,863,889]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C00A9F72845FFB52DF91B610D5DFC96" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[477,597,862,888]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="oculata" status="sp. nov.">
|
||
<emphasis id="B9740E662845FFB52DF91B610D5DFC96" box="[477,597,862,888]" italics="true" pageId="24" pageNumber="25">L. oculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D2845FFB52E7A1B610DB2FC97" box="[606,698,863,889]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by having an elongate and erect mesomeral claw (vs stout in
|
||
<taxonomicName id="4C00A9F72845FFB52CF91BBC0E54FC75" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[221,348,897,923]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="borealis" status="sp. nov.">
|
||
<emphasis id="B9740E662845FFB52CF91BBC0E54FC75" box="[221,348,897,923]" italics="true" pageId="24" pageNumber="25">L. borealis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D2845FFB52D411BBC0EC9FC72" box="[357,449,898,924]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C00A9F72845FFB52DD91BBC0D7EFC75" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[509,630,897,923]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="oculata" status="sp. nov.">
|
||
<emphasis id="B9740E662845FFB52DD91BBC0D7EFC75" box="[509,630,897,923]" italics="true" pageId="24" pageNumber="25">L. oculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D2845FFB52E5B1BBC0DD7FC72" box="[639,735,898,924]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
|
||
).
|
||
<taxonomicName id="4C00A9F72845FFB52ED01BBC0CF6FC75" authorityName="Antić & Reip" authorityYear="2020" baseAuthorityName="Lohmander" baseAuthorityYear="1936" box="[756,1022,898,923]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="satunini">
|
||
<emphasis id="B9740E662845FFB52ED01BBC0CF6FC75" box="[756,1022,898,923]" italics="true" pageId="24" pageNumber="25">Leucogeorgia satunini</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C00A9F72845FFB528B91BBC0BE4FC75" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[1181,1260,898,923]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="gioi" status="sp. nov.">
|
||
<emphasis id="B9740E662845FFB528B91BBC0BE4FC75" box="[1181,1260,898,923]" italics="true" pageId="24" pageNumber="25">L. gioi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D2845FFB528D01BBC0A59FC72" box="[1268,1361,898,924]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by having a more robust and strongly denticulate mesomeral claw (vs mesomeral claw being more slender and with just a few teeth in
|
||
<taxonomicName id="4C00A9F72845FFB52E3A1BF70D67FC0C" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[542,623,969,994]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="gioi" status="sp. nov.">
|
||
<emphasis id="B9740E662845FFB52E3A1BF70D67FC0C" box="[542,623,969,994]" italics="true" pageId="24" pageNumber="25">L. gioi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D2845FFB52E5E1BF60DD2FC0C" box="[634,730,968,994]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
|
||
) and a not too strongly denticulate mesomeral lamella (vs mesomeral lamella strongly denticulate in
|
||
<taxonomicName id="4C00A9F72845FFB52EFB1BD20C25FBEB" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[735,813,1004,1029]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="gioi" status="sp. nov.">
|
||
<emphasis id="B9740E662845FFB52EFB1BD20C25FBEB" box="[735,813,1004,1029]" italics="true" pageId="24" pageNumber="25">L. gioi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D2845FFB52F101BD20C9CFBE8" box="[820,916,1004,1030]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
|
||
). From
|
||
<taxonomicName id="4C00A9F72845FFB52FD41BD20B6FFBEB" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[1008,1127,1003,1029]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="oculata" status="sp. nov.">
|
||
<emphasis id="B9740E662845FFB52FD41BD20B6FFBEB" box="[1008,1127,1003,1029]" italics="true" pageId="24" pageNumber="25">L. oculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D2845FFB5284B1BD20BC5FBE8" box="[1135,1229,1004,1030]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C00A9F72845FFB528FE1BD20A50FBEB" authorityName="Antić & Reip" authorityYear="2020" baseAuthorityName="Lohmander" baseAuthorityYear="1936" box="[1242,1368,1004,1029]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="satunini">
|
||
<emphasis id="B9740E662845FFB528FE1BD20A50FBEB" box="[1242,1368,1004,1029]" italics="true" pageId="24" pageNumber="25">L. satunini</emphasis>
|
||
</taxonomicName>
|
||
in addition differs by the absence of both pigmenation and ommatidia (vs a pigmented body and ommatidia present in
|
||
<taxonomicName id="4C00A9F72845FFB52D1C1C0C0EA7FBA5" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[312,431,1073,1099]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="oculata" status="sp. nov.">
|
||
<emphasis id="B9740E662845FFB52D1C1C0C0EA7FBA5" box="[312,431,1073,1099]" italics="true" pageId="24" pageNumber="25">L. oculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D2845FFB52D931C0C0D1EFBA2" box="[439,534,1074,1100]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
|
||
). Besides all this,
|
||
<taxonomicName id="4C00A9F72845FFB52EC81C0C0C62FBA5" authorityName="Antić & Reip" authorityYear="2020" baseAuthorityName="Lohmander" baseAuthorityYear="1936" box="[748,874,1074,1099]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="satunini">
|
||
<emphasis id="B9740E662845FFB52EC81C0C0C62FBA5" box="[748,874,1074,1099]" italics="true" pageId="24" pageNumber="25">L. satunini</emphasis>
|
||
</taxonomicName>
|
||
differs from all of the previously mentioned species by the presence of longer midbody metazonal setae, these being ca 10% of vertical diameter of rings (vs either midbody metazonal setae 5–6 % of vertical diameter of rings in
|
||
<taxonomicName id="4C00A9F72845FFB528731C470BE4FB7C" authorityName="Antić & Reip" authorityYear="2020" baseAuthorityName="Lohmander" baseAuthorityYear="1936" box="[1111,1260,1144,1170]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="abchasica">
|
||
<emphasis id="B9740E662845FFB528731C470BE4FB7C" box="[1111,1260,1144,1170]" italics="true" pageId="24" pageNumber="25">L. abchasica</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C00A9F72845FFB528DD1C470A7EFB7C" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[1273,1398,1144,1170]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="borealis" status="sp. nov.">
|
||
<emphasis id="B9740E662845FFB528DD1C470A7EFB7C" box="[1273,1398,1144,1170]" italics="true" pageId="24" pageNumber="25">L. borealis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D2845FFB52C991CA20E10FB58" box="[189,280,1180,1206]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C00A9F72845FFB52D741CA20E95FB5B" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[336,413,1180,1205]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="gioi" status="sp. nov.">
|
||
<emphasis id="B9740E662845FFB52D741CA20E95FB5B" box="[336,413,1180,1205]" italics="true" pageId="24" pageNumber="25">L. gioi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D2845FFB52D801CA20EF7FB58" box="[420,511,1180,1206]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
|
||
or metazonal setae completely absent in
|
||
<taxonomicName id="4C00A9F72845FFB52FC51CA20B50FB5B" authority="Antić & Reip, 2020" authorityName="Antić & Reip" authorityYear="2020" box="[993,1112,1179,1205]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="oculata" status="sp. nov.">
|
||
<emphasis id="B9740E662845FFB52FC51CA20B50FB5B" box="[993,1112,1179,1205]" italics="true" pageId="24" pageNumber="25">L. oculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D2845FFB528441CA20BB6FB58" box="[1120,1214,1180,1206]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31A81FF2845FFB52C991CE20A7FFAA7" pageId="24" pageNumber="25" type="materials_examined">
|
||
<paragraph id="8BBFD2742845FFB52C991CE20EA2FB18" blockId="24.[189,426,1244,1270]" box="[189,426,1244,1270]" pageId="24" pageNumber="25">
|
||
<heading id="D0F765182845FFB52C991CE20EA2FB18" bold="true" box="[189,426,1244,1270]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
|
||
<emphasis id="B9740E662845FFB52C991CE20EA2FB18" bold="true" box="[189,426,1244,1270]" pageId="24" pageNumber="25">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742845FFB52C991D350A7FFAA7" blockId="24.[189,1399,1289,1353]" pageId="24" pageNumber="25">
|
||
<materialsCitation id="3B68D8292845FFB52C991D350D4CFACB" ID-GBIF-Occurrence="2858644344" box="[189,580,1291,1317]" location="Gulripsh District" pageId="24" pageNumber="25" specimenCount="1" stateProvince="ABKHAZIA">
|
||
<collectingRegion id="49C41C962845FFB52C991D350E5DFACB" box="[189,341,1291,1317]" country="Georgia" name="Abkhazia" pageId="24" pageNumber="25">ABKHAZIA</collectingRegion>
|
||
–
|
||
<location id="8EDF84AF2845FFB52D4B1D350D4CFACB" LSID="urn:lsid:plazi:treatment:03A963622845FFB62DFC19AD0EFCFAFE:8EDF84AF2845FFB52D4B1D350D4CFACB" box="[367,580,1291,1317]" name="Gulripsh District" pageId="24" pageNumber="25" stateProvince="ABKHAZIA">
|
||
<emphasis id="B9740E662845FFB52D4B1D350D4CFACB" bold="true" box="[367,580,1291,1317]" pageId="24" pageNumber="25">Gulripsh District</emphasis>
|
||
</location>
|
||
</materialsCitation>
|
||
<emphasis id="B9740E662845FFB52E6E1D350D5CFACB" bold="true" box="[586,596,1291,1317]" pageId="24" pageNumber="25">•</emphasis>
|
||
<materialsCitation id="3B68D8292845FFB52E7E1D350A7BFAA7" ID-GBIF-Occurrence="2858644398" collectedFrom="Tsebeldinsky karst Massif" collectingDate="2014-09-03" collectionCode="ZMUM" collectorName="I. S. Turbanov" county="Tsebeldinsky" elevation="95" latitude="43.02" location="Kelasurskaya Cave" longLatPrecision="688" longitude="41.14" municipality="Massif" pageId="24" pageNumber="25" specimenCount="12" specimenCount-female="3" specimenCount-male="1" stateProvince="Abkhazia">
|
||
<specimenCount id="9D0619FD2845FFB52E7E1D350D8DFACB" box="[602,645,1289,1317]" pageId="24" pageNumber="25" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D0619FD2845FFB52EB61D350DDCFACB" box="[658,724,1291,1317]" pageId="24" pageNumber="25" type="female">3 ♀♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D0619FD2845FFB52EC41D350C22FACB" box="[736,810,1291,1317]" pageId="24" pageNumber="25">8 juvs</specimenCount>
|
||
;
|
||
<collectedFrom id="381C91892845FFB52F171D350B6DFACB" box="[819,1125,1291,1317]" pageId="24" pageNumber="25">Tsebeldinsky karst Massif</collectedFrom>
|
||
,
|
||
<locationDeviation id="9A0DCB562845FFB528541D350E18FAA6" pageId="24" pageNumber="25">near Nizhniy Kyalasur village</locationDeviation>
|
||
,
|
||
<location id="8EDF84AF2845FFB52D3F1D100EF5FAA6" LSID="urn:lsid:plazi:treatment:03A963622845FFB62DFC19AD0EFCFAFE:8EDF84AF2845FFB52D3F1D100EF5FAA6" box="[283,509,1326,1352]" county="Tsebeldinsky" latitude="43.02" longLatPrecision="688" longitude="41.14" municipality="Massif" name="Kelasurskaya Cave" pageId="24" pageNumber="25" stateProvince="Abkhazia">Kelasurskaya Cave</location>
|
||
, depth -
|
||
<quantity id="4CF87F912845FFB52E781D100D94FAA7" box="[604,668,1326,1353]" metricMagnitude="1" metricUnit="m" metricValue="9.5" pageId="24" pageNumber="25" unit="m" value="95.0">
|
||
<elevation id="002D35472845FFB52E781D100D94FAA7" box="[604,668,1326,1353]" metricMagnitude="1" metricUnit="m" metricValue="9.5" pageId="24" pageNumber="25" unit="m" value="95.0">95 m</elevation>
|
||
</quantity>
|
||
;
|
||
<geoCoordinate id="EE34B4B32845FFB52E8C1D100C1DFAA7" box="[680,789,1326,1353]" degrees="43.02" direction="north" orientation="latitude" pageId="24" pageNumber="25" precision="555" value="43.02">43.02° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE34B4B32845FFB52F051D100C83FAA7" box="[801,907,1326,1353]" degrees="41.14" direction="east" orientation="longitude" pageId="24" pageNumber="25" precision="555" value="41.14">41.14° E</geoCoordinate>
|
||
;
|
||
<date id="FFBEF4B42845FFB52FB31D100B2EFAA6" box="[919,1062,1326,1352]" pageId="24" pageNumber="25" value="2014-09-03">
|
||
<collectingDate id="EFFA0D5C2845FFB52FB31D100B2EFAA6" box="[919,1062,1326,1352]" pageId="24" pageNumber="25" value="2014-09-03">3 Sep. 2014</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="26F5B7A22845FFB528161D100BDAFAA6" box="[1074,1234,1326,1352]" pageId="24" pageNumber="25">I.S. Turbanov</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED114AB12845FFB529301D110A7BFAA7" box="[1300,1395,1327,1353]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="24" pageNumber="25">ZMUM</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31A81FF2845FFB62C991D510CC0FC68" lastPageId="27" lastPageNumber="28" pageId="24" pageNumber="25" type="description">
|
||
<paragraph id="8BBFD2742845FFB52C991D510E63FA67" blockId="24.[189,363,1391,1417]" box="[189,363,1391,1417]" pageId="24" pageNumber="25">
|
||
<heading id="D0F765182845FFB52C991D510E63FA67" bold="true" box="[189,363,1391,1417]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
|
||
<emphasis id="B9740E662845FFB52C991D510E63FA67" bold="true" box="[189,363,1391,1417]" pageId="24" pageNumber="25">Redescription</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742845FFB52C991DA00BA0FA10" blockId="24.[189,1399,1438,1535]" pageId="24" pageNumber="25">
|
||
SIZE AND NUMBER OF BODY RINGS. Male
|
||
<quantity id="4CF87F912845FFB52EA01DA00DD0FA57" box="[644,728,1438,1465]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="24" pageNumber="25" unit="mm" value="17.0">17 mm</quantity>
|
||
long, vertical diameter of largest body ring
|
||
<quantity id="4CF87F912845FFB528C31DA00A27FA57" box="[1255,1327,1438,1465]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="24" pageNumber="25" unit="mm" value="1.0">1 mm</quantity>
|
||
, body with 43 podous rings + 1 apodous ring + telson. Females
|
||
<quantity id="4CF87F912845FFB52F7A1DFF0CFAFA32" box="[862,1010,1473,1500]" metricMagnitude="-2" metricUnit="m" metricValue="1.525" metricValueMax="1.65" metricValueMin="1.4" pageId="24" pageNumber="25" unit="mm" value="15.25" valueMax="16.5" valueMin="14.0">14–16.5 mm</quantity>
|
||
long, vertical diameter of largest body ring
|
||
<quantity id="4CF87F912845FFB52D121DDA0EDDFA11" box="[310,469,1508,1535]" metricMagnitude="-3" metricUnit="m" metricValue="1.075" metricValueMax="1.2" metricValueMin="0.95" pageId="24" pageNumber="25" unit="mm" value="1.075" valueMax="1.2" valueMin="0.95">0.95–1.2 mm</quantity>
|
||
, body with 38–40 podous rings + 1–2 apodous rings + telson.
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742845FFB52C991E180DDDF9AE" blockId="24.[189,725,1574,1600]" box="[189,725,1574,1600]" pageId="24" pageNumber="25">
|
||
COLOUR (
|
||
<figureCitation id="133BCEF12845FFB52D0C1E180E77F9AE" box="[296,383,1574,1600]" captionStart="Fig" captionStartId="25.[189,232,1895,1921]" captionTargetBox="[277,1311,265,1869]" captionTargetId="figure@25.[277,1311,265,1869]" captionTargetPageId="25" captionText="Fig. 14. Leucogeorgia satunini (Lohmander, 1936) comb. nov., ♀ and ♂ from Kelasurskaya Cave (ZMUM). A. ♀, habitus, lateral view. B. ♀, anterior part of body, lateral view. C. ♂, midbody rings, lateral view. D. ♂, body ring 7, lateral view. E. ♂, body ring 7, ventral view. F. ♂, posterior part of body, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4020782" httpUri="https://zenodo.org/record/4020782/files/figure.png" pageId="24" pageNumber="25">Fig. 14</figureCitation>
|
||
). Yellowish white in alcohol.
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742845FFB52C991E560C60F896" blockId="24.[189,1399,1639,1912]" pageId="24" pageNumber="25">
|
||
HEAD (
|
||
<figureCitation id="133BCEF12845FFB52D2B1E590E74F96C" box="[271,380,1639,1666]" captionStart="Fig" captionStartId="25.[189,232,1895,1921]" captionTargetBox="[277,1311,265,1869]" captionTargetId="figure@25.[277,1311,265,1869]" captionTargetPageId="25" captionText="Fig. 14. Leucogeorgia satunini (Lohmander, 1936) comb. nov., ♀ and ♂ from Kelasurskaya Cave (ZMUM). A. ♀, habitus, lateral view. B. ♀, anterior part of body, lateral view. C. ♂, midbody rings, lateral view. D. ♂, body ring 7, lateral view. E. ♂, body ring 7, ventral view. F. ♂, posterior part of body, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4020782" httpUri="https://zenodo.org/record/4020782/files/figure.png" pageId="24" pageNumber="25">Figs 14B</figureCitation>
|
||
,
|
||
<figureCitation id="133BCEF12845FFB52DA81E590EB6F96C" box="[396,446,1639,1666]" captionStart="Fig" captionStartId="26.[189,232,1858,1884]" captionTargetBox="[255,1333,265,1830]" captionTargetId="figure@26.[255,1333,265,1830]" captionTargetPageId="26" captionText="Fig. 15. Leucogeorgia satunini (Lohmander, 1936) comb. nov., ♂ from Kelasurskaya Cave (ZMUM). A. Head, ventral view. B. Right gonopods, mesal view. C. Left gonopods, mesal view. D. Left gonopods, lateral view. Abbreviations: am = accessory membrane; f = flagellum; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4020784" httpUri="https://zenodo.org/record/4020784/files/figure.png" pageId="24" pageNumber="25">15A</figureCitation>
|
||
). Without ommatidia. Frontal setae absent. Labrum with three labral teeth, four supralabral setae and 14 labral setae. Gnathochilarium with rhomboid promentum. Lamellae linguales with 4+4 setae, stipites with 3+3 distolateral and 5+4 medial setae. Antennae
|
||
<quantity id="4CF87F912845FFB5284B1E900BC4F926" box="[1135,1228,1710,1737]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="24" pageNumber="25" unit="mm" value="1.5">1.5 mm</quantity>
|
||
long in male, their length 150% of vertical diameter of largest body ring. Lengths of antennomeres I–VIII (in mm): 0.08 (I), 0.34 (II), 0.27 (III), 0.24 (IV), 0.28 (V), 0.17 (VI), 0.1 (VII) and 0.02 (VIII). Length/width ratio of antennomeres I–VII: 0.8 (I), 3.4 (II), 2.5 (III), 2 (IV), 2.1 (V), 1.1 (VI) and 1 (VII). Antennomeres V and VI each with a terminal corolla of large sensilla basiconica bacilliformia; antennomere VII with a terminal corolla of small sensilla basiconica bacilliformia.
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742845FFB52C991F9E0CD8F832" blockId="24.[189,1399,1951,2013]" pageId="24" pageNumber="25">
|
||
BODY RINGS (
|
||
<figureCitation id="133BCEF12845FFB52D7C1FA10ECDF857" box="[344,453,1951,1978]" captionStart="Fig" captionStartId="25.[189,232,1895,1921]" captionTargetBox="[277,1311,265,1869]" captionTargetId="figure@25.[277,1311,265,1869]" captionTargetPageId="25" captionText="Fig. 14. Leucogeorgia satunini (Lohmander, 1936) comb. nov., ♀ and ♂ from Kelasurskaya Cave (ZMUM). A. ♀, habitus, lateral view. B. ♀, anterior part of body, lateral view. C. ♂, midbody rings, lateral view. D. ♂, body ring 7, lateral view. E. ♂, body ring 7, ventral view. F. ♂, posterior part of body, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4020782" httpUri="https://zenodo.org/record/4020782/files/figure.png" pageId="24" pageNumber="25">Fig. 14C</figureCitation>
|
||
). Entire metazonal area with longitudinal striations. Metazonal setae relatively long, length of midbody setae ca 10% of vertical diameter of rings.
|
||
</paragraph>
|
||
<caption id="DF7F82FC2844FFB42C991F590D2EF805" ID-DOI="http://doi.org/10.5281/zenodo.4020782" ID-Zenodo-Dep="4020782" httpUri="https://zenodo.org/record/4020782/files/figure.png" pageId="25" pageNumber="26" startId="25.[189,232,1895,1921]" targetBox="[277,1311,265,1869]" targetPageId="25">
|
||
<paragraph id="8BBFD2742844FFB42C991F590D2EF805" blockId="25.[189,1399,1893,2027]" pageId="25" pageNumber="26">
|
||
<emphasis id="B9740E662844FFB42C991F590E16F86F" bold="true" box="[189,286,1895,1921]" pageId="25" pageNumber="26">Fig. 14.</emphasis>
|
||
<taxonomicName id="4C00A9F72844FFB42D0E1F560C22F86C" authority="(Lohmander, 1936)" authorityName="Antić & Reip" authorityYear="2020" baseAuthorityName="Lohmander" baseAuthorityYear="1936" box="[298,810,1895,1922]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="satunini" status="comb. nov.">
|
||
<emphasis id="B9740E662844FFB42D0E1F560D3FF86F" box="[298,567,1896,1921]" italics="true" pageId="25" pageNumber="26">Leucogeorgia satunini</emphasis>
|
||
(
|
||
<bibRefCitation id="EF91AF852844FFB42E6E1F590C2AF86C" author="Lohmander H." box="[586,802,1895,1922]" pageId="25" pageNumber="26" pagination="1 - 196" refId="ref41985" refString="Lohmander H. 1936. Uber die Diplopoden des Kaukasusgebietes. Goteborgs Kungliga Vetenskaps- och Vitterhets-Samhalles Handlingar, Series 5 B 5 (1): 1 - 196." type="journal article" year="1936">Lohmander, 1936</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D2844FFB42F131F590CB7F86C" box="[823,959,1895,1922]" pageId="25" pageNumber="26" rank="species">comb. nov.</taxonomicNameLabel>
|
||
, ♀ and ♂ from Kelasurskaya Cave (ZMUM).
|
||
<emphasis id="B9740E662844FFB42D191FB40E5AF84A" bold="true" box="[317,338,1930,1956]" pageId="25" pageNumber="26">A</emphasis>
|
||
. ♀, habitus, lateral view.
|
||
<emphasis id="B9740E662844FFB42EAD1FB40D95F84A" bold="true" box="[649,669,1930,1956]" pageId="25" pageNumber="26">B</emphasis>
|
||
. ♀, anterior part of body, lateral view.
|
||
<emphasis id="B9740E662844FFB428521FB40B83F84A" bold="true" box="[1142,1163,1930,1956]" pageId="25" pageNumber="26">C</emphasis>
|
||
. ♂, midbody rings, lateral view.
|
||
<emphasis id="B9740E662844FFB42D751F900E6EF826" bold="true" box="[337,358,1966,1992]" pageId="25" pageNumber="26">D</emphasis>
|
||
. ♂, body ring 7, lateral view.
|
||
<emphasis id="B9740E662844FFB42E9A1F900DDAF826" bold="true" box="[702,722,1966,1992]" pageId="25" pageNumber="26">E</emphasis>
|
||
. ♂, body ring 7, ventral view.
|
||
<emphasis id="B9740E662844FFB428161F900B4CF826" bold="true" box="[1074,1092,1966,1992]" pageId="25" pageNumber="26">F</emphasis>
|
||
. ♂, posterior part of body, lateral view. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7F82FC2847FFB72C991F7C0D23F804" ID-DOI="http://doi.org/10.5281/zenodo.4020784" ID-Zenodo-Dep="4020784" httpUri="https://zenodo.org/record/4020784/files/figure.png" pageId="26" pageNumber="27" startId="26.[189,232,1858,1884]" targetBox="[255,1333,265,1830]" targetPageId="26">
|
||
<paragraph id="8BBFD2742847FFB72C991F7C0D23F804" blockId="26.[189,1399,1857,2026]" pageId="26" pageNumber="27">
|
||
<emphasis id="B9740E662847FFB72C991F7C0E13F8B2" bold="true" box="[189,283,1858,1884]" pageId="26" pageNumber="27">Fig. 15.</emphasis>
|
||
<taxonomicName id="4C00A9F72847FFB72D071F7D0C11F8B3" authority="(Lohmander, 1936)" authorityName="Antić & Reip" authorityYear="2020" baseAuthorityName="Lohmander" baseAuthorityYear="1936" box="[291,793,1858,1885]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="satunini" status="comb. nov.">
|
||
<emphasis id="B9740E662847FFB72D071F7D0D25F8B2" box="[291,557,1859,1884]" italics="true" pageId="26" pageNumber="27">Leucogeorgia satunini</emphasis>
|
||
(
|
||
<bibRefCitation id="EF91AF852847FFB72E191F7C0C19F8B3" author="Lohmander H." box="[573,785,1858,1885]" pageId="26" pageNumber="27" pagination="1 - 196" refId="ref41985" refString="Lohmander H. 1936. Uber die Diplopoden des Kaukasusgebietes. Goteborgs Kungliga Vetenskaps- och Vitterhets-Samhalles Handlingar, Series 5 B 5 (1): 1 - 196." type="journal article" year="1936">Lohmander, 1936</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A247B31D2847FFB72F061F7C0CAFF8B3" box="[802,935,1858,1885]" pageId="26" pageNumber="27" rank="species">comb. nov.</taxonomicNameLabel>
|
||
, ♂ from Kelasurskaya Cave (ZMUM).
|
||
<emphasis id="B9740E662847FFB72C991F580FDAF86E" bold="true" box="[189,210,1894,1920]" pageId="26" pageNumber="27">A</emphasis>
|
||
. Head, ventral view.
|
||
<emphasis id="B9740E662847FFB72DE21F580ED2F86E" bold="true" box="[454,474,1894,1920]" pageId="26" pageNumber="27">B</emphasis>
|
||
. Right gonopods, mesal view.
|
||
<emphasis id="B9740E662847FFB72F181F580C59F86E" bold="true" box="[828,849,1894,1920]" pageId="26" pageNumber="27">C</emphasis>
|
||
. Left gonopods, mesal view.
|
||
<emphasis id="B9740E662847FFB728801F580BB1F86E" bold="true" box="[1188,1209,1894,1920]" pageId="26" pageNumber="27">D</emphasis>
|
||
. Left gonopods, lateral view. Abbreviations: am = accessory membrane; f = flagellum; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bars: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBFD2742846FFB62C9919320BBFFEA6" blockId="27.[189,1398,267,329]" pageId="27" pageNumber="28">
|
||
TELSON (
|
||
<figureCitation id="133BCEF12846FFB62D0619350E81FECB" box="[290,393,267,294]" captionStart="Fig" captionStartId="25.[189,232,1895,1921]" captionTargetBox="[277,1311,265,1869]" captionTargetId="figure@25.[277,1311,265,1869]" captionTargetPageId="25" captionText="Fig. 14. Leucogeorgia satunini (Lohmander, 1936) comb. nov., ♀ and ♂ from Kelasurskaya Cave (ZMUM). A. ♀, habitus, lateral view. B. ♀, anterior part of body, lateral view. C. ♂, midbody rings, lateral view. D. ♂, body ring 7, lateral view. E. ♂, body ring 7, ventral view. F. ♂, posterior part of body, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4020782" httpUri="https://zenodo.org/record/4020782/files/figure.png" pageId="27" pageNumber="28">Fig. 14F</figureCitation>
|
||
). Epiproct with a very short and blunt preanal process, covered with dorsal and lateral setae. Paraprocts rounded, with numerous setae. Hypoproct without any modifications.
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742846FFB62C9919460B52FE5B" blockId="27.[189,1399,375,437]" pageId="27" pageNumber="28">LEGS IN MALES. First pair of legs lost during SEM manipulations. Postfemoral and tibial ventral pads well developed on anterior legs, then gradually disappearing towards posterior legs.</paragraph>
|
||
<paragraph id="8BBFD2742846FFB62C9919DA0D0CFDCF" blockId="27.[189,1399,484,545]" pageId="27" pageNumber="28">
|
||
VENTRAL MARGIN OF MALE BODY RING 7 (
|
||
<figureCitation id="133BCEF12846FFB62EAA19DA0DE9FE10" box="[654,737,484,510]" captionStart="Fig" captionStartId="25.[189,232,1895,1921]" captionTargetBox="[277,1311,265,1869]" captionTargetId="figure@25.[277,1311,265,1869]" captionTargetPageId="25" captionText="Fig. 14. Leucogeorgia satunini (Lohmander, 1936) comb. nov., ♀ and ♂ from Kelasurskaya Cave (ZMUM). A. ♀, habitus, lateral view. B. ♀, anterior part of body, lateral view. C. ♂, midbody rings, lateral view. D. ♂, body ring 7, lateral view. E. ♂, body ring 7, ventral view. F. ♂, posterior part of body, lateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4020782" httpUri="https://zenodo.org/record/4020782/files/figure.png" pageId="27" pageNumber="28">Fig. 14</figureCitation>
|
||
D–E). Well-developed, rounded in lateral view, without strong teeth in ventral view.
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742846FFB62C991A6E0EB5FD84" blockId="27.[189,445,592,618]" box="[189,445,592,618]" pageId="27" pageNumber="28">PENES. Not examined.</paragraph>
|
||
<paragraph id="8BBFD2742846FFB62C991AA70CC0FC68" blockId="27.[189,1399,665,903]" pageId="27" pageNumber="28">
|
||
GONOPODS (
|
||
<figureCitation id="133BCEF12846FFB62D661AA70E95FD5D" box="[322,413,665,691]" captionStart="Fig" captionStartId="5.[189,232,1788,1814]" captionTargetBox="[245,1342,265,1760]" captionTargetId="figure@5.[245,1342,265,1760]" captionTargetPageId="5" captionText="Fig. 2. Gonopods, mesal view. A. Leucogeorgia lobata sp. nov., right gonopods, paratype ♂ from Kalitshona Cave (ZMUM). B. L. longipes Verhoeff, 1930, right gonopods, ♂ from Dolabistavi Cave (SMNG). C. L. prometheus sp. nov., left gonopods, non-type ♂ from Sataplia II Cave (ZMUM). D. L. satunini (Lohmander, 1936) comb. nov., right gonopods, ♂ from Kelasurskaya Cave (ZMUM). Abbreviations: am = accessory membrane; f = flagellum; l = lobe; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum." figureDoi="http://doi.org/10.5281/zenodo.4020756" httpUri="https://zenodo.org/record/4020756/files/figure.png" pageId="27" pageNumber="28">Figs 2D</figureCitation>
|
||
,
|
||
<figureCitation id="133BCEF12846FFB62D8E1AA70ECFFD5A" box="[426,455,665,692]" captionStart="Fig" captionStartId="26.[189,232,1858,1884]" captionTargetBox="[255,1333,265,1830]" captionTargetId="figure@26.[255,1333,265,1830]" captionTargetPageId="26" captionText="Fig. 15. Leucogeorgia satunini (Lohmander, 1936) comb. nov., ♂ from Kelasurskaya Cave (ZMUM). A. Head, ventral view. B. Right gonopods, mesal view. C. Left gonopods, mesal view. D. Left gonopods, lateral view. Abbreviations: am = accessory membrane; f = flagellum; m = mesomere; mc = mesomeral claw; ml = mesomeral lamella; o = opisthomere; p = promere; pl = protective lamella; s = solenomere; v = velum. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4020784" httpUri="https://zenodo.org/record/4020784/files/figure.png" pageId="27" pageNumber="28">15</figureCitation>
|
||
B–D). Promere (p) long and slender, slightly wider in central part, with a flagellum (f); apical part spatulate, with denticulate margins; basal half with two developed ridges. Mesomere (m) with a well-developed and denticulate mesomeral claw (mc); mesomeral lamella of same height as entire length, with a posterior rise, distal margin smooth, posterior part finely fimbriate. Opisthomere (o) bipartite. Anterior branch of o with a solenomere (s) with a medium-sized tip, and a well-developed and fimbriate velum (v). Posterior branch of o in form of a shield-like protective lamella (pl). Mesomere and opisthomere connected basally with an accessory membrane (am).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31A81FF2846FFB62C991B8B0B07FC10" pageId="27" pageNumber="28" type="materials_examined">
|
||
<paragraph id="8BBFD2742846FFB62C991B8B0E55FC21" blockId="27.[189,349,949,975]" box="[189,349,949,975]" pageId="27" pageNumber="28">
|
||
<heading id="D0F765182846FFB62C991B8B0E55FC21" bold="true" box="[189,349,949,975]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
|
||
<emphasis id="B9740E662846FFB62C991B8B0E55FC21" bold="true" box="[189,349,949,975]" pageId="27" pageNumber="28">
|
||
<typeStatus id="54BB6CD62846FFB62C991B8B0FF2FC21" box="[189,250,949,975]" pageId="27" pageNumber="28">Type</typeStatus>
|
||
locality
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742846FFB62C991BDA0B07FC10" blockId="27.[189,1039,996,1022]" box="[189,1039,996,1022]" pageId="27" pageNumber="28">
|
||
Canyon of the Kelasuri River, Sukhum and Gulripsh districts,
|
||
<collectingRegion id="49C41C962846FFB62FB31BDA0B03FC10" box="[919,1035,996,1022]" country="Georgia" name="Abkhazia" pageId="27" pageNumber="28">Abkhazia</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31A81FF2846FFB62C991C120BFBFB98" box="[189,1267,1068,1142]" pageId="27" pageNumber="28" type="distribution">
|
||
<paragraph id="8BBFD2742846FFB62C991C120E5EFBA8" blockId="27.[189,342,1068,1094]" box="[189,342,1068,1094]" pageId="27" pageNumber="28">
|
||
<heading id="D0F765182846FFB62C991C120E5EFBA8" bold="true" box="[189,342,1068,1094]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
|
||
<emphasis id="B9740E662846FFB62C991C120E5EFBA8" bold="true" box="[189,342,1068,1094]" pageId="27" pageNumber="28">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742846FFB62C991C620BFBFB98" blockId="27.[189,1267,1115,1142]" box="[189,1267,1115,1142]" pageId="27" pageNumber="28">
|
||
Known only from three localities in Sukhum and Gulripsh districts (
|
||
<figureCitation id="133BCEF12846FFB62FFA1C650B3DFB98" box="[990,1077,1115,1142]" captionStart="Fig" captionStartId="97.[189,232,1991,2017]" captionTargetBox="[189,1399,890,1963]" captionTargetId="figure@97.[189,1399,890,1963]" captionTargetPageId="97" captionText="Fig. 57. Distribution of the northwestern group of the genus Leucogeorgia Verhoeff, 1930." figureDoi="http://doi.org/10.5281/zenodo.4020875" httpUri="https://zenodo.org/record/4020875/files/figure.png" pageId="27" pageNumber="28">Fig. 57</figureCitation>
|
||
, black triangle).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31A81FF2846FFB62C991C9A0EFCFAFE" pageId="27" pageNumber="28" type="discussion">
|
||
<paragraph id="8BBFD2742846FFB62C991C9A0E27FB50" blockId="27.[189,303,1188,1214]" box="[189,303,1188,1214]" pageId="27" pageNumber="28">
|
||
<heading id="D0F765182846FFB62C991C9A0E27FB50" bold="true" box="[189,303,1188,1214]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
|
||
<emphasis id="B9740E662846FFB62C991C9A0E27FB50" bold="true" box="[189,303,1188,1214]" pageId="27" pageNumber="28">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BBFD2742846FFB62C991CED0EFCFAFE" blockId="27.[189,1399,1235,1296]" pageId="27" pageNumber="28">
|
||
In the Kelasurskaya Cave, this species lives together with
|
||
<taxonomicName id="4C00A9F72846FFB62F7C1CEA0CE4FB03" authorityName="Antić & Reip" authorityYear="2020" baseAuthorityName="Lohmander" baseAuthorityYear="1936" box="[856,1004,1235,1261]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="abchasica">
|
||
<emphasis id="B9740E662846FFB62F7C1CEA0CE4FB03" box="[856,1004,1235,1261]" italics="true" pageId="27" pageNumber="28">L. abchasica</emphasis>
|
||
</taxonomicName>
|
||
and an unidentified
|
||
<taxonomicName id="4C00A9F72846FFB628F11CEA0A7FFB03" authorityName="Verhoeff" authorityYear="1930" box="[1237,1399,1236,1261]" class="Diplopoda" family="Julidae" genus="Leucogeorgia" kingdom="Animalia" order="Julida" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9740E662846FFB628F11CEA0A7FFB03" box="[1237,1399,1236,1261]" italics="true" pageId="27" pageNumber="28">Leucogeorgia</emphasis>
|
||
</taxonomicName>
|
||
with modified mouthparts.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |