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<document id="35A9A5C5992AB26E0AB6DC8731FAD290" ID="10.11646/zootaxa.4175.6.5" ID-DOI="10.11646/zootaxa.4175.6.5" ID-GBIF-Dataset="e79fd18a-fa17-4b4b-8490-1e50489bb71b" ID-ISSN="1175-5326" ID-Zenodo-Dep="262702" ID-ZooBank="DD402425-F055-4CC9-B7BF-4710F4CA96CF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1476866464250" checkinUser="plazi" docAuthor="Iwan, Dariusz" docDate="2016" docId="03A72351FFBBBB5EB5E353EF088A9476" docLanguage="en" docName="zootaxa.4175.6.5.pdf" docOrigin="Zootaxa 4175 (6)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Atrocrates" docType="key" docVersion="7" lastPageNumber="584" masterDocId="FF9E5B29FFB6BB53B57456670D169309" masterDocTitle="Species groups in the genus Atrocrates Koch, 1956 (Coleoptera: Tenebrionidae: Pedinini)" masterLastPageNumber="587" masterPageNumber="571" pageNumber="584" updateTime="1698390191471" updateUser="plazi">
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<mods:title id="F4E696E582502B3FFB020E39E95D62EE">Species groups in the genus Atrocrates Koch, 1956 (Coleoptera: Tenebrionidae: Pedinini)</mods:title>
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<mods:date id="DDA070EE9626E8AF2A01768E253D97C3">2016</mods:date>
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<treatment id="03A72351FFBBBB5EB5E353EF088A9476" ID-DOI="http://doi.org/10.5281/zenodo.6087560" ID-GBIF-Taxon="124996583" ID-Zenodo-Dep="6087560" LSID="urn:lsid:plazi:treatment:03A72351FFBBBB5EB5E353EF088A9476" httpUri="http://treatment.plazi.org/id/03A72351FFBBBB5EB5E353EF088A9476" lastPageNumber="584" pageId="13" pageNumber="584">
<subSubSection id="C314C1CCFFBBBB5EB5E353EF0F4396A8" box="[151,597,1415,1442]" pageId="13" pageNumber="584" type="nomenclature">
<paragraph id="8BB19247FFBBBB5EB5E353EF0F4396A8" blockId="13.[151,597,1415,1442]" box="[151,597,1415,1442]" pageId="13" pageNumber="584">
<heading id="D0F9252BFFBBBB5EB5E353EF0F4396A8" bold="true" box="[151,597,1415,1442]" fontSize="11" level="1" pageId="13" pageNumber="584" reason="1">
<emphasis id="B97A4E55FFBBBB5EB5E353EF0F4396A8" bold="true" box="[151,597,1415,1442]" pageId="13" pageNumber="584">
Key for species groups of
<taxonomicName id="4C0EE9C4FFBBBB5EB4AC53E00F4396A8" box="[472,597,1415,1441]" class="Insecta" family="Tenebrionidae" genus="Atrocrates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="584" phylum="Arthropoda" rank="genus">
<emphasis id="B97A4E55FFBBBB5EB4AC53E00F4396A8" bold="true" box="[472,597,1415,1441]" italics="true" pageId="13" pageNumber="584">Atrocrates</emphasis>
</taxonomicName>
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<paragraph id="8BB19247FFBBBB5EB5E353A9088A96ED" blockId="13.[151,1436,1486,1919]" box="[151,1436,1486,1508]" pageId="13" pageNumber="584">
<keyLead id="30FF3172FFBBBB5EB5E353A9088A96ED" box="[151,1436,1486,1508]" pageId="13" pageNumber="584">
1. Mid part of mentum widened anteriad (
<figureCitation id="13358EC2FFBBBB5EB72E53A90FB696ED" box="[602,672,1486,1508]" captionStart="FIGURES 23 – 30" captionStartId="6.[151,264,1964,1987]" captionTargetBox="[202,1356,193,1919]" captionTargetId="figure@6.[202,1357,193,1921]" captionTargetPageId="6" captionText="FIGURES 23 – 30. Atrocrates striatus (Quensel, 1806), male (23 – 26). A. occultator Iwan, 1998, male (27 – 30). Mentum (23, 27). Pronotum (24, 28). Mid tibia, ventral (25, 30). Mid tibia, inside (26). Pronotal posteriori angle and elytral humerus (29)." httpUri="https://zenodo.org/record/262706/files/figure.png" pageId="13" pageNumber="584">Fig. 27</figureCitation>
)...............................................
<emphasis id="B97A4E55FFBBBB5EB18053A9088A96ED" bold="true" box="[1268,1436,1486,1508]" pageId="13" pageNumber="584">
<taxonomicName id="4C0EE9C4FFBBBB5EB18053A9084196ED" box="[1268,1367,1486,1508]" class="Insecta" family="Tenebrionidae" genus="Atrocrates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="584" phylum="Arthropoda" rank="species" species="occultator">
<emphasis id="B97A4E55FFBBBB5EB18053A9084196ED" bold="true" box="[1268,1367,1486,1508]" italics="true" pageId="13" pageNumber="584">occultator</emphasis>
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-group
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</paragraph>
<paragraph id="8BB19247FFBBBB5EB5E3538B088A9508" blockId="13.[151,1436,1486,1919]" box="[151,1436,1515,1537]" pageId="13" pageNumber="584">
<keyLead id="30FF3172FFBBBB5EB5E3538B088A9508" box="[151,1436,1515,1537]" pageId="13" pageNumber="584">
-. Mid part of mentum narrowed anteriad (
<figureCitation id="13358EC2FFBBBB5EB717538C0FB69508" box="[611,672,1515,1537]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1966,1989]" captionTargetBox="[198,1349,193,1914]" captionTargetId="figure@2.[198,1349,193,1915]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Atrocrates garbarczyki Iwan, 1998, male (1 – 4), A. pliskoae Iwan, 1998 (5 – 8). Head, ventral (1). Pronotum (2, 7). Fore leg (3). Mid leg (4). Mentum (5). Mid tibia (6). Pronotal posterior angle (8)." httpUri="https://zenodo.org/record/262703/files/figure.png" pageId="13" pageNumber="584">Figs 5</figureCitation>
,
<figureCitation id="13358EC2FFBBBB5EB7DF538C0FAF9508" box="[683,697,1515,1537]" captionStart="FIGURES 9 – 14" captionStartId="4.[151,264,1736,1759]" captionTargetBox="[151,1436,267,1712]" captionTargetId="figure@4.[151,1436,267,1715]" captionTargetPageId="4" captionText="FIGURES 9 – 14. Atrocrates bredasdorpensis Iwan, 1998, male. Mentum (9). Pronotum (10). Pronotal posterior angle (11). Pronotal posterior angle and elytral humerus (12). Fore leg (13). Fore tibia (14)." httpUri="https://zenodo.org/record/262704/files/figure.png" pageId="13" pageNumber="584">9</figureCitation>
,
<figureCitation id="13358EC2FFBBBB5EB7B7538C0E179508" box="[707,769,1515,1537]" captionStart="FIGURES 15 – 22" captionStartId="5.[151,264,1956,1979]" captionTargetBox="[207,1360,193,1919]" captionTargetId="figure@5.[207,1361,193,1921]" captionTargetPageId="5" captionText="FIGURES 15 – 22. Atrocrates splendidus Iwan, 1998, male (15 – 17). A. robustus Iwan, 1998 (18, 19); A. libidinosus Iwan, 1998 (20 – 22). Mentum (15, 20). Pronotum (16, 21). Elytra (17). Fore tibia (18). Apex of fore tibia (19). Mid tibia (22)." httpUri="https://zenodo.org/record/262705/files/figure.png" pageId="13" pageNumber="584">15, 20</figureCitation>
,
<figureCitation id="13358EC2FFBBBB5EB67F538C0E339508" box="[779,805,1515,1537]" captionStart="FIGURES 23 – 30" captionStartId="6.[151,264,1964,1987]" captionTargetBox="[202,1356,193,1919]" captionTargetId="figure@6.[202,1357,193,1921]" captionTargetPageId="6" captionText="FIGURES 23 – 30. Atrocrates striatus (Quensel, 1806), male (23 – 26). A. occultator Iwan, 1998, male (27 – 30). Mentum (23, 27). Pronotum (24, 28). Mid tibia, ventral (25, 30). Mid tibia, inside (26). Pronotal posteriori angle and elytral humerus (29)." httpUri="https://zenodo.org/record/262706/files/figure.png" pageId="13" pageNumber="584">23</figureCitation>
,
<figureCitation id="13358EC2FFBBBB5EB65B538C0E5F9508" box="[815,841,1515,1537]" captionStart="FIGURES 31 – 38" captionStartId="7.[151,264,1962,1985]" captionTargetBox="[209,1377,193,1940]" captionTargetId="figure@7.[209,1378,193,1941]" captionTargetPageId="7" captionText="FIGURES 31 – 38. Atrocrates galbasi sp. nov., male. Head, dorsal (31). Eye, dorsal (32). Antenna (33). Mentum (34). Pronotum (35). Elytral humerus (36). Prothorax, ventral (37). Meso- and metathorax ventral (38)." httpUri="https://zenodo.org/record/262707/files/figure.png" pageId="13" pageNumber="584">34</figureCitation>
,
<figureCitation id="13358EC2FFBBBB5EB627538C0E7A9508" box="[851,876,1515,1537]" captionStart="FIGURES 47 – 54" captionStartId="10.[151,264,1962,1985]" captionTargetBox="[209,1377,193,1940]" captionTargetId="figure@10.[209,1378,193,1941]" captionTargetPageId="10" captionText="FIGURES 47 – 54. Atrocrates matthewsi sp. nov., male. Head, dorsal (47). Eye, dorsal (48). Head, ventral (49). Mentum (50). Antenna (51). Pronotum (52). Basal part of elytra (53). Posterior part of elytra (54)." httpUri="https://zenodo.org/record/262709/files/figure.png" pageId="13" pageNumber="584">50</figureCitation>
)...........................................
<emphasis id="B97A4E55FFBBBB5EB0E4538C088A9508" bold="true" box="[1424,1436,1515,1537]" pageId="13" pageNumber="584">2</emphasis>
</keyLead>
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<keyStep id="30FA8AE2FFBBBB5EB5E3506F088A9550" pageId="13" pageNumber="584">
<paragraph id="8BB19247FFBBBB5EB5E3506F088A9535" blockId="13.[151,1436,1486,1919]" pageId="13" pageNumber="584">
<keyLead id="30FF3172FFBBBB5EB5E3506F088A9535" pageId="13" pageNumber="584">
2. Pronotal sides rounded or parallelsided basally, widest at 1/3-1/2 from base; posterior pronotal angles not produced posteriad (
<figureCitation id="13358EC2FFBBBB5EB59350410C309535" box="[231,294,1574,1596]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1966,1989]" captionTargetBox="[198,1349,193,1914]" captionTargetId="figure@2.[198,1349,193,1915]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Atrocrates garbarczyki Iwan, 1998, male (1 – 4), A. pliskoae Iwan, 1998 (5 – 8). Head, ventral (1). Pronotum (2, 7). Fore leg (3). Mid leg (4). Mentum (5). Mid tibia (6). Pronotal posterior angle (8)." httpUri="https://zenodo.org/record/262703/files/figure.png" pageId="13" pageNumber="584">Figs 2</figureCitation>
,
<figureCitation id="13358EC2FFBBBB5EB44450410C5C9535" box="[304,330,1574,1596]" captionStart="FIGURES 9 – 14" captionStartId="4.[151,264,1736,1759]" captionTargetBox="[151,1436,267,1712]" captionTargetId="figure@4.[151,1436,267,1715]" captionTargetPageId="4" captionText="FIGURES 9 – 14. Atrocrates bredasdorpensis Iwan, 1998, male. Mentum (9). Pronotum (10). Pronotal posterior angle (11). Pronotal posterior angle and elytral humerus (12). Fore leg (13). Fore tibia (14)." httpUri="https://zenodo.org/record/262704/files/figure.png" pageId="13" pageNumber="584">10</figureCitation>
,
<figureCitation id="13358EC2FFBBBB5EB42050410C789535" box="[340,366,1574,1596]" captionStart="FIGURES 15 – 22" captionStartId="5.[151,264,1956,1979]" captionTargetBox="[207,1360,193,1919]" captionTargetId="figure@5.[207,1361,193,1921]" captionTargetPageId="5" captionText="FIGURES 15 – 22. Atrocrates splendidus Iwan, 1998, male (15 – 17). A. robustus Iwan, 1998 (18, 19); A. libidinosus Iwan, 1998 (20 – 22). Mentum (15, 20). Pronotum (16, 21). Elytra (17). Fore tibia (18). Apex of fore tibia (19). Mid tibia (22)." httpUri="https://zenodo.org/record/262705/files/figure.png" pageId="13" pageNumber="584">16</figureCitation>
,
<figureCitation id="13358EC2FFBBBB5EB40C50410C849535" box="[376,402,1574,1596]" captionStart="FIGURES 31 – 38" captionStartId="7.[151,264,1962,1985]" captionTargetBox="[209,1377,193,1940]" captionTargetId="figure@7.[209,1378,193,1941]" captionTargetPageId="7" captionText="FIGURES 31 – 38. Atrocrates galbasi sp. nov., male. Head, dorsal (31). Eye, dorsal (32). Antenna (33). Mentum (34). Pronotum (35). Elytral humerus (36). Prothorax, ventral (37). Meso- and metathorax ventral (38)." httpUri="https://zenodo.org/record/262707/files/figure.png" pageId="13" pageNumber="584">35</figureCitation>
,
<figureCitation id="13358EC2FFBBBB5EB4E850410CA39535" box="[412,437,1574,1596]" captionStart="FIGURES 47 – 54" captionStartId="10.[151,264,1962,1985]" captionTargetBox="[209,1377,193,1940]" captionTargetId="figure@10.[209,1378,193,1941]" captionTargetPageId="10" captionText="FIGURES 47 – 54. Atrocrates matthewsi sp. nov., male. Head, dorsal (47). Eye, dorsal (48). Head, ventral (49). Mentum (50). Antenna (51). Pronotum (52). Basal part of elytra (53). Posterior part of elytra (54)." httpUri="https://zenodo.org/record/262709/files/figure.png" pageId="13" pageNumber="584">52</figureCitation>
)................................................................................
<emphasis id="B97A4E55FFBBBB5EB0E45041088A9535" bold="true" box="[1424,1436,1574,1596]" pageId="13" pageNumber="584">3</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BB19247FFBBBB5EB5E35023088A9550" blockId="13.[151,1436,1486,1919]" box="[151,1436,1603,1625]" pageId="13" pageNumber="584">
<keyLead id="30FF3172FFBBBB5EB5E35023088A9550" box="[151,1436,1603,1625]" pageId="13" pageNumber="584">
-. Pronotal sides narrowed anteriad, widest at base; posterior pronotal angles produced posteriad (
<figureCitation id="13358EC2FFBBBB5EB106502409A79550" box="[1138,1201,1603,1625]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1966,1989]" captionTargetBox="[198,1349,193,1914]" captionTargetId="figure@2.[198,1349,193,1915]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Atrocrates garbarczyki Iwan, 1998, male (1 – 4), A. pliskoae Iwan, 1998 (5 – 8). Head, ventral (1). Pronotum (2, 7). Fore leg (3). Mid leg (4). Mentum (5). Mid tibia (6). Pronotal posterior angle (8)." httpUri="https://zenodo.org/record/262703/files/figure.png" pageId="13" pageNumber="584">Figs 7</figureCitation>
,
<figureCitation id="13358EC2FFBBBB5EB1CF502409C39550" box="[1211,1237,1603,1625]" captionStart="FIGURES 15 – 22" captionStartId="5.[151,264,1956,1979]" captionTargetBox="[207,1360,193,1919]" captionTargetId="figure@5.[207,1361,193,1921]" captionTargetPageId="5" captionText="FIGURES 15 – 22. Atrocrates splendidus Iwan, 1998, male (15 – 17). A. robustus Iwan, 1998 (18, 19); A. libidinosus Iwan, 1998 (20 – 22). Mentum (15, 20). Pronotum (16, 21). Elytra (17). Fore tibia (18). Apex of fore tibia (19). Mid tibia (22)." httpUri="https://zenodo.org/record/262705/files/figure.png" pageId="13" pageNumber="584">21</figureCitation>
,
<figureCitation id="13358EC2FFBBBB5EB1AB502409EE9550" box="[1247,1272,1603,1625]" captionStart="FIGURES 23 – 30" captionStartId="6.[151,264,1964,1987]" captionTargetBox="[202,1356,193,1919]" captionTargetId="figure@6.[202,1357,193,1921]" captionTargetPageId="6" captionText="FIGURES 23 – 30. Atrocrates striatus (Quensel, 1806), male (23 – 26). A. occultator Iwan, 1998, male (27 – 30). Mentum (23, 27). Pronotum (24, 28). Mid tibia, ventral (25, 30). Mid tibia, inside (26). Pronotal posteriori angle and elytral humerus (29)." httpUri="https://zenodo.org/record/262706/files/figure.png" pageId="13" pageNumber="584">24</figureCitation>
)..........
<emphasis id="B97A4E55FFBBBB5EB0E45024088A9550" bold="true" box="[1424,1436,1603,1625]" pageId="13" pageNumber="584">6</emphasis>
</keyLead>
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<keyStep id="30FA8AE2FFBBBB5EB5E35007088A959D" pageId="13" pageNumber="584">
<paragraph id="8BB19247FFBBBB5EB5E35007088A957E" blockId="13.[151,1436,1486,1919]" box="[151,1436,1632,1655]" pageId="13" pageNumber="584">
<keyLead id="30FF3172FFBBBB5EB5E35007088A957E" box="[151,1436,1632,1655]" pageId="13" pageNumber="584">
3. Male mid tibia simple (
<figureCitation id="13358EC2FFBBBB5EB4B550070CEB957F" box="[449,509,1632,1654]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1966,1989]" captionTargetBox="[198,1349,193,1914]" captionTargetId="figure@2.[198,1349,193,1915]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Atrocrates garbarczyki Iwan, 1998, male (1 – 4), A. pliskoae Iwan, 1998 (5 – 8). Head, ventral (1). Pronotum (2, 7). Fore leg (3). Mid leg (4). Mentum (5). Mid tibia (6). Pronotal posterior angle (8)." httpUri="https://zenodo.org/record/262703/files/figure.png" pageId="13" pageNumber="584">Fig. 4</figureCitation>
)................................................................
<emphasis id="B97A4E55FFBBBB5EB0775006088A957E" bold="true" box="[1283,1436,1633,1655]" pageId="13" pageNumber="584">
<taxonomicName id="4C0EE9C4FFBBBB5EB07750060841957E" box="[1283,1367,1633,1655]" class="Insecta" family="Tenebrionidae" genus="Atrocrates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="584" phylum="Arthropoda" rank="species" species="bellamyi">
<emphasis id="B97A4E55FFBBBB5EB07750060841957E" bold="true" box="[1283,1367,1633,1655]" italics="true" pageId="13" pageNumber="584">bellamyi</emphasis>
</taxonomicName>
-group
</emphasis>
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</paragraph>
<paragraph id="8BB19247FFBBBB5EB5E35018088A959D" blockId="13.[151,1436,1486,1919]" box="[151,1436,1662,1684]" pageId="13" pageNumber="584">
<keyLead id="30FF3172FFBBBB5EB5E35018088A959D" box="[151,1436,1662,1684]" pageId="13" pageNumber="584">
-. Male mid tibia with denticle (
<figureCitation id="13358EC2FFBBBB5EB48950190F5E959D" box="[509,584,1662,1684]" captionStart="FIGURES 9 – 14" captionStartId="4.[151,264,1736,1759]" captionTargetBox="[151,1436,267,1712]" captionTargetId="figure@4.[151,1436,267,1715]" captionTargetPageId="4" captionText="FIGURES 9 – 14. Atrocrates bredasdorpensis Iwan, 1998, male. Mentum (9). Pronotum (10). Pronotal posterior angle (11). Pronotal posterior angle and elytral humerus (12). Fore leg (13). Fore tibia (14)." httpUri="https://zenodo.org/record/262704/files/figure.png" pageId="13" pageNumber="584">Figs 14</figureCitation>
,
<figureCitation id="13358EC2FFBBBB5EB72550190F99959D" box="[593,655,1662,1684]" captionStart="FIGURES 15 – 22" captionStartId="5.[151,264,1956,1979]" captionTargetBox="[207,1360,193,1919]" captionTargetId="figure@5.[207,1361,193,1921]" captionTargetPageId="5" captionText="FIGURES 15 – 22. Atrocrates splendidus Iwan, 1998, male (15 – 17). A. robustus Iwan, 1998 (18, 19); A. libidinosus Iwan, 1998 (20 – 22). Mentum (15, 20). Pronotum (16, 21). Elytra (17). Fore tibia (18). Apex of fore tibia (19). Mid tibia (22)." httpUri="https://zenodo.org/record/262705/files/figure.png" pageId="13" pageNumber="584">18, 19</figureCitation>
,
<figureCitation id="13358EC2FFBBBB5EB7ED50190FC1959D" box="[665,727,1662,1684]" captionStart="FIGURES 39 – 46" captionStartId="8.[151,264,1964,1987]" captionTargetBox="[209,1377,193,1940]" captionTargetId="figure@8.[209,1378,193,1941]" captionTargetPageId="8" captionText="FIGURES 39 – 46. Atrocrates galbasi sp. nov., male. 4 th and 5 th abdominal ventrites (39). Fore leg (40). Fore tibia, dorsal (41). Fore tibia, ventral (42). Mid tibia, dorsal (43). Mid tibia, inside (44). Mid tibia, ventral (45). Hind tibia, ventral (46)." httpUri="https://zenodo.org/record/262708/files/figure.png" pageId="13" pageNumber="584">44, 45</figureCitation>
,
<figureCitation id="13358EC2FFBBBB5EB79550190FEC959D" box="[737,762,1662,1684]" captionStart="FIGURES 55 – 62" captionStartId="11.[151,264,1934,1957]" captionTargetBox="[217,1354,193,1894]" captionTargetId="figure@11.[217,1355,193,1895]" captionTargetPageId="11" captionText="FIGURES 55 – 62. Atrocrates matthewsi sp. nov., female. Prothorax, ventral (55). Meso- and metathorax ventral (56). Meso- and metathorax ventral and abdomen (57). 3 rd, 4 th and 5 th abdominal ventrites (58). Fore tibia, ventral (59). Mid tibia, ventral (60). Hind tibia, ventral (61). Apex of fore tibia, ventral (62)." httpUri="https://zenodo.org/record/262710/files/figure.png" pageId="13" pageNumber="584">60</figureCitation>
).....................................................
<emphasis id="B97A4E55FFBBBB5EB0E45019088A959D" bold="true" box="[1424,1436,1662,1684]" pageId="13" pageNumber="584">4</emphasis>
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<keyStep id="30FA8AE2FFBBBB5EB5E350FC088A95C6" pageId="13" pageNumber="584">
<paragraph id="8BB19247FFBBBB5EB5E350FC088A95B8" blockId="13.[151,1436,1486,1919]" box="[151,1436,1691,1713]" pageId="13" pageNumber="584">
<keyLead id="30FF3172FFBBBB5EB5E350FC088A95B8" box="[151,1436,1691,1713]" pageId="13" pageNumber="584">
4. Elytral humeri obtuse (
<figureCitation id="13358EC2FFBBBB5EB4C950FC0F1495B8" box="[445,514,1691,1713]" captionStart="FIGURES 15 – 22" captionStartId="5.[151,264,1956,1979]" captionTargetBox="[207,1360,193,1919]" captionTargetId="figure@5.[207,1361,193,1921]" captionTargetPageId="5" captionText="FIGURES 15 – 22. Atrocrates splendidus Iwan, 1998, male (15 – 17). A. robustus Iwan, 1998 (18, 19); A. libidinosus Iwan, 1998 (20 – 22). Mentum (15, 20). Pronotum (16, 21). Elytra (17). Fore tibia (18). Apex of fore tibia (19). Mid tibia (22)." httpUri="https://zenodo.org/record/262705/files/figure.png" pageId="13" pageNumber="584">Fig. 17</figureCitation>
)............................................................
<emphasis id="B97A4E55FFBBBB5EB18750FC088A95B8" bold="true" box="[1267,1436,1691,1713]" pageId="13" pageNumber="584">
<taxonomicName id="4C0EE9C4FFBBBB5EB18750FC084195B8" box="[1267,1367,1691,1713]" class="Insecta" family="Tenebrionidae" genus="Atrocrates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="584" phylum="Arthropoda" rank="species" species="coconatae">
<emphasis id="B97A4E55FFBBBB5EB18750FC084195B8" bold="true" box="[1267,1367,1691,1713]" italics="true" pageId="13" pageNumber="584">coconatae</emphasis>
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-group
</emphasis>
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</paragraph>
<paragraph id="8BB19247FFBBBB5EB5E350DE088A95C6" blockId="13.[151,1436,1486,1919]" box="[151,1436,1720,1743]" pageId="13" pageNumber="584">
<keyLead id="30FF3172FFBBBB5EB5E350DE088A95C6" box="[151,1436,1720,1743]" pageId="13" pageNumber="584">
<emphasis id="B97A4E55FFBBBB5EB5E350DE0D8995C6" bold="true" box="[151,159,1721,1743]" pageId="13" pageNumber="584">-</emphasis>
. Elytral humeri rectangular (
<figureCitation id="13358EC2FFBBBB5EB49E50DF0F2395C7" box="[490,565,1720,1742]" captionStart="FIGURES 9 – 14" captionStartId="4.[151,264,1736,1759]" captionTargetBox="[151,1436,267,1712]" captionTargetId="figure@4.[151,1436,267,1715]" captionTargetPageId="4" captionText="FIGURES 9 – 14. Atrocrates bredasdorpensis Iwan, 1998, male. Mentum (9). Pronotum (10). Pronotal posterior angle (11). Pronotal posterior angle and elytral humerus (12). Fore leg (13). Fore tibia (14)." httpUri="https://zenodo.org/record/262704/files/figure.png" pageId="13" pageNumber="584">Figs 12</figureCitation>
,
<figureCitation id="13358EC2FFBBBB5EB74B50DF0F4F95C7" box="[575,601,1720,1742]" captionStart="FIGURES 31 – 38" captionStartId="7.[151,264,1962,1985]" captionTargetBox="[209,1377,193,1940]" captionTargetId="figure@7.[209,1378,193,1941]" captionTargetPageId="7" captionText="FIGURES 31 – 38. Atrocrates galbasi sp. nov., male. Head, dorsal (31). Eye, dorsal (32). Antenna (33). Mentum (34). Pronotum (35). Elytral humerus (36). Prothorax, ventral (37). Meso- and metathorax ventral (38)." httpUri="https://zenodo.org/record/262707/files/figure.png" pageId="13" pageNumber="584">36</figureCitation>
,
<figureCitation id="13358EC2FFBBBB5EB71750DF0F6A95C7" box="[611,636,1720,1742]" captionStart="FIGURES 47 – 54" captionStartId="10.[151,264,1962,1985]" captionTargetBox="[209,1377,193,1940]" captionTargetId="figure@10.[209,1378,193,1941]" captionTargetPageId="10" captionText="FIGURES 47 – 54. Atrocrates matthewsi sp. nov., male. Head, dorsal (47). Eye, dorsal (48). Head, ventral (49). Mentum (50). Antenna (51). Pronotum (52). Basal part of elytra (53). Posterior part of elytra (54)." httpUri="https://zenodo.org/record/262709/files/figure.png" pageId="13" pageNumber="584">53</figureCitation>
)...............................................................
<emphasis id="B97A4E55FFBBBB5EB0E450DE088A95C6" bold="true" box="[1424,1436,1721,1743]" pageId="13" pageNumber="584">5</emphasis>
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</paragraph>
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<keyStep id="30FA8AE2FFBBBB5EB5E350B1088A9400" pageId="13" pageNumber="584">
<paragraph id="8BB19247FFBBBB5EB5E350B1088A95E5" blockId="13.[151,1436,1486,1919]" box="[151,1436,1750,1772]" pageId="13" pageNumber="584">
<keyLead id="30FF3172FFBBBB5EB5E350B1088A95E5" box="[151,1436,1750,1772]" pageId="13" pageNumber="584">
5. Elytral humeri protruding outwards (
<figureCitation id="13358EC2FFBBBB5EB73650B10F9195E5" box="[578,647,1750,1772]" captionStart="FIGURES 9 – 14" captionStartId="4.[151,264,1736,1759]" captionTargetBox="[151,1436,267,1712]" captionTargetId="figure@4.[151,1436,267,1715]" captionTargetPageId="4" captionText="FIGURES 9 – 14. Atrocrates bredasdorpensis Iwan, 1998, male. Mentum (9). Pronotum (10). Pronotal posterior angle (11). Pronotal posterior angle and elytral humerus (12). Fore leg (13). Fore tibia (14)." httpUri="https://zenodo.org/record/262704/files/figure.png" pageId="13" pageNumber="584">Fig. 12</figureCitation>
).................................................
<emphasis id="B97A4E55FFBBBB5EB18450B1088A95E5" bold="true" box="[1264,1436,1750,1772]" pageId="13" pageNumber="584">
<taxonomicName id="4C0EE9C4FFBBBB5EB18450B1084095E5" box="[1264,1366,1750,1772]" class="Insecta" family="Tenebrionidae" genus="Atrocrates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="584" phylum="Arthropoda" rank="species" species="bisinuatus">
<emphasis id="B97A4E55FFBBBB5EB18450B1084095E5" bold="true" box="[1264,1366,1750,1772]" italics="true" pageId="13" pageNumber="584">bisinuatus</emphasis>
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-group
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BB19247FFBBBB5EB5E35094088A9400" blockId="13.[151,1436,1486,1919]" box="[151,1436,1779,1801]" pageId="13" pageNumber="584">
<keyLead id="30FF3172FFBBBB5EB5E35094088A9400" box="[151,1436,1779,1801]" pageId="13" pageNumber="584">
<emphasis id="B97A4E55FFBBBB5EB5E350940D899400" bold="true" box="[151,159,1779,1801]" pageId="13" pageNumber="584">-</emphasis>
. Elytral humeri not protruding outwards (
<figureCitation id="13358EC2FFBBBB5EB71250940FA79400" box="[614,689,1779,1801]" captionStart="FIGURES 31 – 38" captionStartId="7.[151,264,1962,1985]" captionTargetBox="[209,1377,193,1940]" captionTargetId="figure@7.[209,1378,193,1941]" captionTargetPageId="7" captionText="FIGURES 31 – 38. Atrocrates galbasi sp. nov., male. Head, dorsal (31). Eye, dorsal (32). Antenna (33). Mentum (34). Pronotum (35). Elytral humerus (36). Prothorax, ventral (37). Meso- and metathorax ventral (38)." httpUri="https://zenodo.org/record/262707/files/figure.png" pageId="13" pageNumber="584">Figs 36</figureCitation>
,
<figureCitation id="13358EC2FFBBBB5EB7CF50940FC29400" box="[699,724,1779,1801]" captionStart="FIGURES 47 – 54" captionStartId="10.[151,264,1962,1985]" captionTargetBox="[209,1377,193,1940]" captionTargetId="figure@10.[209,1378,193,1941]" captionTargetPageId="10" captionText="FIGURES 47 – 54. Atrocrates matthewsi sp. nov., male. Head, dorsal (47). Eye, dorsal (48). Head, ventral (49). Mentum (50). Antenna (51). Pronotum (52). Basal part of elytra (53). Posterior part of elytra (54)." httpUri="https://zenodo.org/record/262709/files/figure.png" pageId="13" pageNumber="584">53</figureCitation>
)..............................................
<emphasis id="B97A4E55FFBBBB5EB0645094088A9400" bold="true" box="[1296,1436,1779,1801]" pageId="13" pageNumber="584">
<taxonomicName id="4C0EE9C4FFBBBB5EB064509408409400" box="[1296,1366,1779,1801]" class="Insecta" family="Tenebrionidae" genus="Atrocrates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="584" phylum="Arthropoda" rank="species" species="galbasi">
<emphasis id="B97A4E55FFBBBB5EB064509408409400" bold="true" box="[1296,1366,1779,1801]" italics="true" pageId="13" pageNumber="584">galbasi</emphasis>
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-group
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</keyStep>
<keyStep id="30FA8AE2FFBBBB5EB5E35177088A944D" pageId="13" pageNumber="584">
<paragraph id="8BB19247FFBBBB5EB5E35177088A942E" blockId="13.[151,1436,1486,1919]" box="[151,1436,1808,1831]" pageId="13" pageNumber="584">
<keyLead id="30FF3172FFBBBB5EB5E35177088A942E" box="[151,1436,1808,1831]" pageId="13" pageNumber="584">
6. Male mid tibiae S-like, with large, preapical denticle (
<figureCitation id="13358EC2FFBBBB5EB79351770E3A942F" box="[743,812,1808,1830]" captionStart="FIGURES 15 – 22" captionStartId="5.[151,264,1956,1979]" captionTargetBox="[207,1360,193,1919]" captionTargetId="figure@5.[207,1361,193,1921]" captionTargetPageId="5" captionText="FIGURES 15 – 22. Atrocrates splendidus Iwan, 1998, male (15 – 17). A. robustus Iwan, 1998 (18, 19); A. libidinosus Iwan, 1998 (20 – 22). Mentum (15, 20). Pronotum (16, 21). Elytra (17). Fore tibia (18). Apex of fore tibia (19). Mid tibia (22)." httpUri="https://zenodo.org/record/262705/files/figure.png" pageId="13" pageNumber="584">Fig. 22</figureCitation>
).....................................
<emphasis id="B97A4E55FFBBBB5EB18F5176088A942E" bold="true" box="[1275,1436,1809,1831]" pageId="13" pageNumber="584">
<taxonomicName id="4C0EE9C4FFBBBB5EB18F51760841942E" box="[1275,1367,1809,1831]" class="Insecta" family="Tenebrionidae" genus="Atrocrates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="584" phylum="Arthropoda" rank="species" species="formosus">
<emphasis id="B97A4E55FFBBBB5EB18F51760841942E" bold="true" box="[1275,1367,1809,1831]" italics="true" pageId="13" pageNumber="584">formosus</emphasis>
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-group
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BB19247FFBBBB5EB5E35148088A944D" blockId="13.[151,1436,1486,1919]" box="[151,1436,1838,1860]" pageId="13" pageNumber="584">
<keyLead id="30FF3172FFBBBB5EB5E35148088A944D" box="[151,1436,1838,1860]" pageId="13" pageNumber="584">
-. Male mid tibiae slightly widened, with small denticle (
<figureCitation id="13358EC2FFBBBB5EB79951490E3D944D" box="[749,811,1838,1860]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1966,1989]" captionTargetBox="[198,1349,193,1914]" captionTargetId="figure@2.[198,1349,193,1915]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Atrocrates garbarczyki Iwan, 1998, male (1 – 4), A. pliskoae Iwan, 1998 (5 – 8). Head, ventral (1). Pronotum (2, 7). Fore leg (3). Mid leg (4). Mentum (5). Mid tibia (6). Pronotal posterior angle (8)." httpUri="https://zenodo.org/record/262703/files/figure.png" pageId="13" pageNumber="584">Figs 6</figureCitation>
,
<figureCitation id="13358EC2FFBBBB5EB64151490E58944D" box="[821,846,1838,1860]" captionStart="FIGURES 23 – 30" captionStartId="6.[151,264,1964,1987]" captionTargetBox="[202,1356,193,1919]" captionTargetId="figure@6.[202,1357,193,1921]" captionTargetPageId="6" captionText="FIGURES 23 – 30. Atrocrates striatus (Quensel, 1806), male (23 – 26). A. occultator Iwan, 1998, male (27 – 30). Mentum (23, 27). Pronotum (24, 28). Mid tibia, ventral (25, 30). Mid tibia, inside (26). Pronotal posteriori angle and elytral humerus (29)." httpUri="https://zenodo.org/record/262706/files/figure.png" pageId="13" pageNumber="584">25</figureCitation>
)..............................................
<emphasis id="B97A4E55FFBBBB5EB0E45149088A944D" bold="true" box="[1424,1436,1838,1860]" pageId="13" pageNumber="584">7</emphasis>
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<keyStep id="30FA8AE2FFBBBB5EB5E3512B088A9476" pageId="13" pageNumber="584">
<paragraph id="8BB19247FFBBBB5EB5E3512B088A9468" blockId="13.[151,1436,1486,1919]" box="[151,1436,1867,1889]" pageId="13" pageNumber="584">
<keyLead id="30FF3172FFBBBB5EB5E3512B088A9468" box="[151,1436,1867,1889]" pageId="13" pageNumber="584">
7. Male mid tibiae with preapical denticle (
<figureCitation id="13358EC2FFBBBB5EB712512C0FD99468" box="[614,719,1867,1889]" captionStart="FIGURES 23 – 30" captionStartId="6.[151,264,1964,1987]" captionTargetBox="[202,1356,193,1919]" captionTargetId="figure@6.[202,1357,193,1921]" captionTargetPageId="6" captionText="FIGURES 23 – 30. Atrocrates striatus (Quensel, 1806), male (23 – 26). A. occultator Iwan, 1998, male (27 – 30). Mentum (23, 27). Pronotum (24, 28). Mid tibia, ventral (25, 30). Mid tibia, inside (26). Pronotal posteriori angle and elytral humerus (29)." httpUri="https://zenodo.org/record/262706/files/figure.png" pageId="13" pageNumber="584">Fig. 25, 26</figureCitation>
)...........................................
<emphasis id="B97A4E55FFBBBB5EB199512C088A9468" bold="true" box="[1261,1436,1867,1889]" pageId="13" pageNumber="584">
<taxonomicName id="4C0EE9C4FFBBBB5EB199512C08419468" box="[1261,1367,1867,1889]" class="Insecta" family="Tenebrionidae" genus="Atrocrates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="584" phylum="Arthropoda" rank="species" species="evestigator">
<emphasis id="B97A4E55FFBBBB5EB199512C08419468" bold="true" box="[1261,1367,1867,1889]" italics="true" pageId="13" pageNumber="584">evestigator</emphasis>
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-group
</emphasis>
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</paragraph>
<paragraph id="8BB19247FFBBBB5EB5E3510E088A9476" blockId="13.[151,1436,1486,1919]" box="[151,1436,1896,1919]" pageId="13" pageNumber="584">
<keyLead id="30FF3172FFBBBB5EB5E3510E088A9476" box="[151,1436,1896,1919]" pageId="13" pageNumber="584">
-. Male mid tibiae with apical denticle (
<figureCitation id="13358EC2FFBBBB5EB733510F0F979477" box="[583,641,1896,1918]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1966,1989]" captionTargetBox="[198,1349,193,1914]" captionTargetId="figure@2.[198,1349,193,1915]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Atrocrates garbarczyki Iwan, 1998, male (1 – 4), A. pliskoae Iwan, 1998 (5 – 8). Head, ventral (1). Pronotum (2, 7). Fore leg (3). Mid leg (4). Mentum (5). Mid tibia (6). Pronotal posterior angle (8)." httpUri="https://zenodo.org/record/262703/files/figure.png" pageId="13" pageNumber="584">Fig. 6</figureCitation>
)....................................................
<emphasis id="B97A4E55FFBBBB5EB077510E088A9476" bold="true" box="[1283,1436,1897,1919]" pageId="13" pageNumber="584">
<taxonomicName id="4C0EE9C4FFBBBB5EB077510E08419476" box="[1283,1367,1897,1919]" class="Insecta" family="Tenebrionidae" genus="Eurynotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="584" phylum="Arthropoda" rank="species" species="capensis">
<emphasis id="B97A4E55FFBBBB5EB077510E08419476" bold="true" box="[1283,1367,1897,1919]" italics="true" pageId="13" pageNumber="584">capensis</emphasis>
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-group
</emphasis>
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</paragraph>
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</subSubSection>
</treatment>
</document>
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