610 lines
76 KiB
XML
610 lines
76 KiB
XML
<document ID-ISSN="2305-2562" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683576867786" checkinUser="felipe" docAuthor="Irwin, Michael E. & Wiegmann, Brian M." docDate="2001" docId="03A21212FFB1C43CFE9111E1FEB3FA32" docLanguage="en" docName="AfrInvertebr.42.1.225–253.pdf" docOrigin="African Invertebrates 42" docStyle="DocumentStyle:B78C63EC8CEBBB75364A82B35BB1F83D.3:AfrInvertebr.2008-2015.journal_article.0cover" docStyleId="B78C63EC8CEBBB75364A82B35BB1F83D" docStyleName="AfrInvertebr.2008-2015.journal_article.0cover" docStyleVersion="3" docTitle="Tongamya stuckenbergi Irwin & Wiegmann 2001, sp. n." docType="treatment" docVersion="2" lastPageNumber="247" masterDocId="FF9B6A6AFFA1C42BFFDA145BFFACFFCC" masterDocTitle="A review of the southern African genus Tongamya (Diptera: Asiloidea: Mydidae: Megascelinae), with a molecular assessment of the phylogenetic placement of Tongamya and the Megascelinae" masterLastPageNumber="253" masterPageNumber="225" pageNumber="241" updateTime="1683597725632" updateUser="felipe" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A review of the southern African genus Tongamya (Diptera: Asiloidea: Mydidae: Megascelinae), with a molecular assessment of the phylogenetic placement of Tongamya and the Megascelinae</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Irwin, Michael E.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wiegmann, Brian M.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>African Invertebrates</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2001</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2001-12-31</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>42</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>225</mods:start>
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<mods:end>253</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="ISSN">2305-2562</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:03A21212FFB1C43CFE9111E1FEB3FA32" httpUri="http://treatment.plazi.org/id/03A21212FFB1C43CFE9111E1FEB3FA32" lastPageId="23" lastPageNumber="247" pageId="16" pageNumber="241">
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<subSubSection pageId="16" pageNumber="241" type="nomenclature">
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<paragraph blockId="16.[144,1104,1466,1672]" box="[331,916,1466,1491]" pageId="16" pageNumber="241">
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<heading box="[331,916,1466,1491]" centered="true" fontSize="11" level="2" pageId="16" pageNumber="241" reason="2">
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<taxonomicName ID-CoL="7CH7S" ID-ENA="160320" authority="Irwin & Wiegmann" authorityName="Irwin & Wiegmann" authorityYear="2001" box="[331,845,1466,1491]" class="Insecta" family="Mydidae" genus="Tongamya" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="241" phylum="Arthropoda" rank="species" species="stuckenbergi" status="sp. n.">
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<emphasis bold="true" box="[331,617,1466,1491]" pageId="16" pageNumber="241">Tongamya stuckenbergi</emphasis>
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Irwin & Wiegmann
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</taxonomicName>
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,
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<taxonomicNameLabel box="[856,916,1466,1491]" pageId="16" pageNumber="241" rank="species">sp. n.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<paragraph blockId="16.[144,1104,1466,1672]" box="[308,939,1506,1531]" pageId="16" pageNumber="241">
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Habitus illustrations of male (
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<figureCitation box="[642,709,1506,1531]" captionStart="Fig" captionStartId="7.[182,215,1623,1643]" captionTargetBox="[232,1101,709,1599]" captionTargetId="figure-9@7.[205,1112,709,1626]" captionTargetPageId="7" captionText="Fig. 2. Habitus view of male Tongamya stuckenbergi sp. n. (MEI 109134). 2a. Magnified view of male terminalia, lateral view." pageId="16" pageNumber="241">Fig. 2</figureCitation>
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) and female (
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<figureCitation box="[863,930,1506,1531]" captionStart="Fig" captionStartId="9.[182,215,1307,1327]" captionTargetBox="[240,1094,203,1280]" captionTargetId="figure-125@9.[211,1115,203,1299]" captionTargetPageId="9" captionText="Fig. 3. Habitus view of female Tongamya stuckenbergi sp. n. (MEI 109135). 3a. Magnified view of female terminalia, lateral view." pageId="16" pageNumber="241">Fig. 3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="23" lastPageNumber="248" pageId="16" pageNumber="241" type="description">
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<paragraph blockId="16.[144,1104,1466,1672]" pageId="16" pageNumber="241">
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Etiology: This species is named for Brian R. Stuckenberg, mentor and life-long friend of
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<taxonomicName authorityName="Xu & Norell" authorityYear="2004" box="[178,232,1575,1600]" class="Reptilia" family="Troodontidae" genus="Mei" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="16" pageNumber="251" phylum="Chordata" rank="genus">MEI</taxonomicName>
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, in honour of a lifetime of meritorious contributions to African and World Dipterology.
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</paragraph>
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<paragraph blockId="16.[144,1104,1466,1672]" box="[144,1104,1647,1672]" pageId="16" pageNumber="241">
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Type Material: Male
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<typeStatus box="[382,477,1647,1672]" pageId="16" pageNumber="241">holotype</typeStatus>
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(
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<emphasis box="[496,636,1648,1672]" italics="true" pageId="16" pageNumber="241">
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<taxonomicName authorityName="Xu & Norell" authorityYear="2004" box="[496,545,1648,1672]" class="Reptilia" family="Troodontidae" genus="Mei" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="16" pageNumber="241" phylum="Chordata" rank="genus">MEI</taxonomicName>
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109134
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</emphasis>
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),
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<specimenCount box="[661,765,1647,1672]" pageId="16" pageNumber="241" type="male">two male</specimenCount>
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<typeStatus box="[775,879,1647,1672]" pageId="16" pageNumber="241">paratypes</typeStatus>
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(
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<emphasis box="[897,1038,1648,1672]" italics="true" pageId="16" pageNumber="241">
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<taxonomicName authorityName="Xu & Norell" authorityYear="2004" box="[897,946,1648,1672]" class="Reptilia" family="Troodontidae" genus="Mei" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="16" pageNumber="241" phylum="Chordata" rank="genus">MEI</taxonomicName>
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109136
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</emphasis>
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;
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<taxonomicName authorityName="Xu & Norell" authorityYear="2004" box="[1055,1104,1648,1672]" class="Reptilia" family="Troodontidae" genus="Mei" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="16" pageNumber="241" phylum="Chordata" rank="genus">
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<emphasis box="[1055,1104,1648,1672]" italics="true" pageId="16" pageNumber="241">MEI</emphasis>
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</taxonomicName>
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</paragraph>
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<caption pageId="17" pageNumber="242" startId="17.[182,215,872,892]" targetBox="[229,1091,203,850]" targetPageId="17" targetType="figure">
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<paragraph blockId="17.[182,1141,872,916]" pageId="17" pageNumber="242">
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Fig. 14. Male terminalia, lateral view, of
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<taxonomicName authority="Stuckenberg, 1966" authorityName="Stuckenberg" authorityYear="1966" box="[560,914,872,892]" class="Insecta" family="Mydidae" genus="Tongamya" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="242" phylum="Arthropoda" rank="species" species="miranda">
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<emphasis box="[560,737,872,892]" italics="true" pageId="17" pageNumber="242">Tongamya miranda</emphasis>
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<bibRefCitation author="STUCKENBERG, B. R." box="[744,914,872,892]" pageId="17" pageNumber="242" pagination="106 - 120" refId="ref12950" refString="STUCKENBERG, B. R. 1966. A remarkable new genus and species of Apioceridae in South Africa (Diptera). Revue du Zoologie par Botanie du Africa 73: 106 - 120." type="journal article" year="1966">Stuckenberg, 1966</bibRefCitation>
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</taxonomicName>
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(
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<emphasis box="[928,1043,872,892]" italics="true" pageId="17" pageNumber="242">
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<taxonomicName authorityName="Xu & Norell" authorityYear="2004" box="[928,968,872,892]" class="Reptilia" family="Troodontidae" genus="Mei" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="17" pageNumber="242" phylum="Chordata" rank="genus">MEI</taxonomicName>
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089484
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</emphasis>
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) depicting tergite 8, sternite 8, and major sclerites of the genitalia.
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</paragraph>
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</caption>
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<paragraph blockId="17.[181,1142,953,1674]" pageId="17" pageNumber="242">
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<emphasis box="[181,262,953,977]" italics="true" pageId="17" pageNumber="242">089484</emphasis>
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), and
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<specimenCount box="[335,456,953,978]" pageId="17" pageNumber="242" type="female">one female</specimenCount>
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<typeStatus box="[465,558,953,978]" pageId="17" pageNumber="242">paratype</typeStatus>
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(
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<emphasis box="[575,713,953,977]" italics="true" pageId="17" pageNumber="242">
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<taxonomicName authorityName="Xu & Norell" authorityYear="2004" box="[575,624,953,977]" class="Reptilia" family="Troodontidae" genus="Mei" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="17" pageNumber="242" phylum="Chordata" rank="genus">MEI</taxonomicName>
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109135
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</emphasis>
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) from the same locality and collection date (see material examined). The females from
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<collectingCountry box="[708,788,985,1010]" name="Angola" pageId="17" pageNumber="242">Angola</collectingCountry>
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(
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<emphasis box="[804,939,985,1009]" italics="true" pageId="17" pageNumber="242">
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<taxonomicName authorityName="Xu & Norell" authorityYear="2004" box="[804,853,985,1009]" class="Reptilia" family="Troodontidae" genus="Mei" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="17" pageNumber="242" phylum="Chordata" rank="genus">MEI</taxonomicName>
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116621
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</emphasis>
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) and Etosha (
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<emphasis italics="true" pageId="17" pageNumber="242">
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<taxonomicName authorityName="Xu & Norell" authorityYear="2004" box="[1093,1142,985,1009]" class="Reptilia" family="Troodontidae" genus="Mei" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="17" pageNumber="242" phylum="Chordata" rank="genus">MEI</taxonomicName>
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108188
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</emphasis>
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) are included in the material examined below, but are not included as type material; upon discovery of males of these populations, they may prove to be a distinct species.
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</paragraph>
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<paragraph blockId="17.[181,1142,953,1674]" pageId="17" pageNumber="242">
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Distribution: The
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<typeStatus box="[380,428,1089,1114]" pageId="17" pageNumber="242">type</typeStatus>
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material is from the westernmost edge of the Kalahari Desert in
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<collectingCountry box="[181,277,1121,1146]" name="Namibia" pageId="17" pageNumber="242">Namibia</collectingCountry>
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(
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<figureCitation box="[293,363,1121,1146]" captionStart="Figs 4–6. 4" captionStartId="12.[144,183,1371,1391]" captionTargetBox="[179,1075,206,1337]" captionTargetId="figure-81@12.[179,1075,203,1356]" captionTargetPageId="12" captionText="Figs 4–6. 4. Photograph of habitat where type material of Tongamya stuckenbergi sp. n. was collected (2 km S Otjisume Farm, Otjinene District, Namibia); note that the vegetation is not fully leafed. 5–6. Heads of male Tongamya, frontal views. 5. T. miranda Stuckenberg, 1966 (MEI 116615). 6. T. stuckenbergi sp. n. (MEI 109134)." pageId="17" pageNumber="242">Fig. 4</figureCitation>
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||
). Two females that we currently place in this taxon are from northern
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<collectingCountry box="[181,278,1153,1178]" name="Namibia" pageId="17" pageNumber="242">Namibia</collectingCountry>
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(Etosha) and southwestern
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<collectingCountry box="[604,687,1153,1178]" name="Angola" pageId="17" pageNumber="242">Angola</collectingCountry>
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(
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<figureCitation box="[706,793,1153,1178]" captionStart="Fig" captionStartId="16.[144,177,1043,1063]" captionTargetBox="[170,1081,203,1026]" captionTargetId="figure-210@16.[170,1081,203,1026]" captionTargetPageId="16" captionText="Fig. 13. Map of southern Africa depicting the known distribution of Tongamya miranda Stuckenberg, 1966 (squares) and Tongamya stuckenbergi sp. n. (circles)." pageId="17" pageNumber="242">Fig. 13</figureCitation>
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). (Note: Slight morphological differences separate the
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<specimenCount box="[447,570,1185,1210]" pageId="17" pageNumber="242" type="female">two female</specimenCount>
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specimens (
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<collectingCountry box="[705,789,1185,1210]" name="Angola" pageId="17" pageNumber="242">Angola</collectingCountry>
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and
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<collectingCountry box="[840,938,1185,1210]" name="Namibia" pageId="17" pageNumber="242">Namibia</collectingCountry>
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) from the material at the
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<typeStatus box="[249,297,1217,1242]" pageId="17" pageNumber="242">type</typeStatus>
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locality. These
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<specimenCount box="[473,609,1217,1242]" pageId="17" pageNumber="242" type="female">two females</specimenCount>
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extend the range delimited by the
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<typeStatus box="[995,1043,1217,1242]" pageId="17" pageNumber="242">type</typeStatus>
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material considerably northward; the differences between the
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<typeStatus box="[788,836,1249,1274]" pageId="17" pageNumber="242">type</typeStatus>
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population and that of the northern specimens are discussed above (see ‘Diagnoses of the species within
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<taxonomicName box="[181,297,1313,1337]" class="Insecta" family="Mydidae" genus="Tongamya" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="242" phylum="Arthropoda" rank="genus">
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<emphasis box="[181,297,1313,1337]" italics="true" pageId="17" pageNumber="242">Tongamya</emphasis>
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</taxonomicName>
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’). Males of these northern populations are unknown.)
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</paragraph>
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<paragraph blockId="17.[181,1142,953,1674]" pageId="17" pageNumber="242">
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Material Examined:
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<collectingCountry box="[412,533,1353,1378]" name="Angola" pageId="17" pageNumber="242">ANGOLA</collectingCountry>
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,
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<emphasis box="[546,772,1353,1377]" italics="true" pageId="17" pageNumber="242">Ambundos Province</emphasis>
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,
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<collectingRegion box="[787,988,1353,1378]" country="Angola" name="Benguela" pageId="17" pageNumber="242">Benguela District</collectingRegion>
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,
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<specimenCount box="[1000,1101,1353,1378]" pageId="17" pageNumber="242" type="female">1 female</specimenCount>
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,
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<quantity metricMagnitude="4" metricUnit="m" metricValue="4.8" pageId="17" pageNumber="242" unit="km" value="48.0">48 km</quantity>
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S.
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<collectingRegion box="[262,369,1385,1410]" country="Angola" name="Benguela" pageId="17" pageNumber="242">Benguela</collectingRegion>
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[-13.0113º, +13.4039º],
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<date box="[661,795,1385,1410]" pageId="17" pageNumber="242" value="1971-04-21">21-IV-1971</date>
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, H. D. Brown, (
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<emphasis box="[991,1135,1385,1409]" italics="true" pageId="17" pageNumber="242">
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<taxonomicName authorityName="Xu & Norell" authorityYear="2004" box="[991,1041,1385,1409]" class="Reptilia" family="Troodontidae" genus="Mei" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="17" pageNumber="242" phylum="Chordata" rank="genus">MEI</taxonomicName>
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||
116621
|
||
</emphasis>
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||
, NMSA).
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||
<collectingCountry box="[284,410,1417,1442]" name="Namibia" pageId="17" pageNumber="242">NAMIBIA</collectingCountry>
|
||
,
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<specimenCount box="[420,518,1417,1442]" pageId="17" pageNumber="242" type="female">1 female</specimenCount>
|
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, Etosha National Park, Mushara [-18.6167º, +16.8833º],
|
||
<date box="[181,302,1449,1474]" pageId="17" pageNumber="242" value="1986-10-14">14-X-1986</date>
|
||
to
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||
<date box="[333,465,1449,1474]" pageId="17" pageNumber="242" value="1986-11-15">15-XI-1986</date>
|
||
, E. Griffin, (
|
||
<emphasis box="[599,736,1449,1473]" italics="true" pageId="17" pageNumber="242">
|
||
<taxonomicName authorityName="Xu & Norell" authorityYear="2004" box="[599,648,1449,1473]" class="Reptilia" family="Troodontidae" genus="Mei" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="17" pageNumber="242" phylum="Chordata" rank="genus">MEI</taxonomicName>
|
||
108188
|
||
</emphasis>
|
||
, SMWN);
|
||
<specimenCount box="[855,937,1449,1474]" pageId="17" pageNumber="242" type="male">3 males</specimenCount>
|
||
<specimenCount box="[942,1039,1449,1474]" pageId="17" pageNumber="242" type="female">1 female</specimenCount>
|
||
,
|
||
<collectingRegion inferredBySuffix="true" pageId="17" pageNumber="242">Otjinene District</collectingRegion>
|
||
,
|
||
<quantity box="[278,334,1481,1506]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="242" unit="km" value="2.0">2 km</quantity>
|
||
S. Otjisume Farm [-21.3117, +19.3222],
|
||
<date box="[793,928,1481,1506]" pageId="17" pageNumber="242" value="1996-11-27">27-XI-1996</date>
|
||
, hand netted, M. E. Irwin,
|
||
<quantity box="[259,349,1513,1538]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="17" pageNumber="242" unit="m" value="1400.0">1400 m</quantity>
|
||
, (
|
||
<emphasis box="[374,518,1513,1537]" italics="true" pageId="17" pageNumber="242">
|
||
<taxonomicName authorityName="Xu & Norell" authorityYear="2004" box="[374,424,1513,1537]" class="Reptilia" family="Troodontidae" genus="Mei" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="17" pageNumber="242" phylum="Chordata" rank="genus">MEI</taxonomicName>
|
||
109134
|
||
</emphasis>
|
||
[
|
||
<typeStatus box="[534,640,1513,1538]" pageId="17" pageNumber="242">holotype</typeStatus>
|
||
], SMWN;
|
||
<emphasis box="[767,911,1513,1537]" italics="true" pageId="17" pageNumber="242">
|
||
<taxonomicName authorityName="Xu & Norell" authorityYear="2004" box="[767,817,1513,1537]" class="Reptilia" family="Troodontidae" genus="Mei" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="17" pageNumber="242" phylum="Chordata" rank="genus">MEI</taxonomicName>
|
||
109136
|
||
</emphasis>
|
||
[
|
||
<typeStatus box="[927,1032,1513,1538]" pageId="17" pageNumber="242">paratype</typeStatus>
|
||
], NMSA;
|
||
<emphasis box="[181,321,1545,1569]" italics="true" pageId="17" pageNumber="242">
|
||
<taxonomicName authorityName="Xu & Norell" authorityYear="2004" box="[181,231,1545,1569]" class="Reptilia" family="Troodontidae" genus="Mei" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="17" pageNumber="242" phylum="Chordata" rank="genus">MEI</taxonomicName>
|
||
089484
|
||
</emphasis>
|
||
[
|
||
<typeStatus box="[333,435,1545,1570]" pageId="17" pageNumber="242">paratype</typeStatus>
|
||
] [thorax macerated for DNA], NCSU, [remainder of specimen], MEIC;
|
||
<emphasis box="[266,407,1577,1601]" italics="true" pageId="17" pageNumber="242">
|
||
<taxonomicName authorityName="Xu & Norell" authorityYear="2004" box="[266,316,1577,1601]" class="Reptilia" family="Troodontidae" genus="Mei" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="17" pageNumber="242" phylum="Chordata" rank="genus">MEI</taxonomicName>
|
||
109135
|
||
</emphasis>
|
||
[
|
||
<typeStatus box="[421,526,1577,1602]" pageId="17" pageNumber="242">paratype</typeStatus>
|
||
], NMSA).
|
||
</paragraph>
|
||
<paragraph blockId="17.[181,1142,953,1674]" lastBlockId="18.[144,1105,206,1674]" lastPageId="18" lastPageNumber="243" pageId="17" pageNumber="242">
|
||
Description: The following description is, for the most part, based on the
|
||
<typeStatus box="[984,1082,1617,1642]" pageId="17" pageNumber="242">holotype</typeStatus>
|
||
male (
|
||
<emphasis box="[191,328,1649,1673]" italics="true" pageId="17" pageNumber="242">
|
||
<taxonomicName authorityName="Xu & Norell" authorityYear="2004" box="[191,240,1649,1673]" class="Reptilia" family="Troodontidae" genus="Mei" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="17" pageNumber="242" phylum="Chordata" rank="genus">MEI</taxonomicName>
|
||
109134
|
||
</emphasis>
|
||
).The internal male terminalia are described from a dissected male
|
||
<typeStatus box="[1047,1142,1649,1674]" pageId="17" pageNumber="242">paratype</typeStatus>
|
||
(
|
||
<emphasis box="[153,290,206,230]" italics="true" pageId="18" pageNumber="243">
|
||
<taxonomicName authorityName="Xu & Norell" authorityYear="2004" box="[153,202,206,230]" class="Reptilia" family="Troodontidae" genus="Mei" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="18" pageNumber="243" phylum="Chordata" rank="genus">MEI</taxonomicName>
|
||
089484
|
||
</emphasis>
|
||
). Where appropriate within the description, morphological variation based on the other male
|
||
<typeStatus box="[351,447,238,263]" pageId="18" pageNumber="243">paratype</typeStatus>
|
||
(
|
||
<emphasis box="[465,607,238,262]" italics="true" pageId="18" pageNumber="243">
|
||
<taxonomicName authorityName="Xu & Norell" authorityYear="2004" box="[465,515,238,262]" class="Reptilia" family="Troodontidae" genus="Mei" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="18" pageNumber="243" phylum="Chordata" rank="genus">MEI</taxonomicName>
|
||
109136
|
||
</emphasis>
|
||
) is provided. The single topotypical female specimen (
|
||
<emphasis box="[263,401,270,294]" italics="true" pageId="18" pageNumber="243">
|
||
<taxonomicName authorityName="Xu & Norell" authorityYear="2004" box="[263,313,270,294]" class="Reptilia" family="Troodontidae" genus="Mei" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="18" pageNumber="243" phylum="Chordata" rank="genus">MEI</taxonomicName>
|
||
109135
|
||
</emphasis>
|
||
) is described below the male description; comparisons are made with the two more northern females (
|
||
<emphasis box="[554,694,302,326]" italics="true" pageId="18" pageNumber="243">
|
||
<taxonomicName authorityName="Xu & Norell" authorityYear="2004" box="[554,604,302,326]" class="Reptilia" family="Troodontidae" genus="Mei" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="18" pageNumber="243" phylum="Chordata" rank="genus">MEI</taxonomicName>
|
||
108188
|
||
</emphasis>
|
||
and
|
||
<emphasis box="[746,884,302,326]" italics="true" pageId="18" pageNumber="243">
|
||
<taxonomicName authorityName="Xu & Norell" authorityYear="2004" box="[746,796,302,326]" class="Reptilia" family="Troodontidae" genus="Mei" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="18" pageNumber="243" phylum="Chordata" rank="genus">MEI</taxonomicName>
|
||
116621
|
||
</emphasis>
|
||
), tentatively placed within this taxon.
|
||
</paragraph>
|
||
<paragraph blockId="18.[144,1105,206,1674]" pageId="18" pageNumber="243">
|
||
<emphasis box="[144,202,373,397]" italics="true" pageId="18" pageNumber="243">Male</emphasis>
|
||
(
|
||
<figureCitation box="[220,292,373,398]" captionStart="Fig" captionStartId="7.[182,215,1623,1643]" captionTargetBox="[232,1101,709,1599]" captionTargetId="figure-9@7.[205,1112,709,1626]" captionTargetPageId="7" captionText="Fig. 2. Habitus view of male Tongamya stuckenbergi sp. n. (MEI 109134). 2a. Magnified view of male terminalia, lateral view." pageId="18" pageNumber="243">Fig. 2</figureCitation>
|
||
): Body length, excluding antennae, 11.0 mm (
|
||
<emphasis box="[831,975,373,397]" italics="true" pageId="18" pageNumber="243">
|
||
<taxonomicName authorityName="Xu & Norell" authorityYear="2004" box="[831,881,373,397]" class="Reptilia" family="Troodontidae" genus="Mei" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="18" pageNumber="243" phylum="Chordata" rank="genus">MEI</taxonomicName>
|
||
109134
|
||
</emphasis>
|
||
);
|
||
<quantity box="[1002,1104,373,398]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="18" pageNumber="243" unit="mm" value="11.4">11.4 mm</quantity>
|
||
(
|
||
<emphasis box="[153,294,405,429]" italics="true" pageId="18" pageNumber="243">
|
||
<taxonomicName authorityName="Xu & Norell" authorityYear="2004" box="[153,203,405,429]" class="Reptilia" family="Troodontidae" genus="Mei" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="18" pageNumber="243" phylum="Chordata" rank="genus">MEI</taxonomicName>
|
||
109136
|
||
</emphasis>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="18.[144,1105,206,1674]" pageId="18" pageNumber="243">
|
||
<emphasis box="[144,203,444,468]" italics="true" pageId="18" pageNumber="243">Head</emphasis>
|
||
: Head large (
|
||
<figureCitation box="[353,425,444,469]" captionStart="Fig" captionStartId="7.[182,215,1623,1643]" captionTargetBox="[232,1101,709,1599]" captionTargetId="figure-9@7.[205,1112,709,1626]" captionTargetPageId="7" captionText="Fig. 2. Habitus view of male Tongamya stuckenbergi sp. n. (MEI 109134). 2a. Magnified view of male terminalia, lateral view." pageId="18" pageNumber="243">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation box="[437,452,444,469]" captionStart="Figs 4–6. 4" captionStartId="12.[144,183,1371,1391]" captionTargetBox="[179,1075,206,1337]" captionTargetId="figure-81@12.[179,1075,203,1356]" captionTargetPageId="12" captionText="Figs 4–6. 4. Photograph of habitat where type material of Tongamya stuckenbergi sp. n. was collected (2 km S Otjisume Farm, Otjinene District, Namibia); note that the vegetation is not fully leafed. 5–6. Heads of male Tongamya, frontal views. 5. T. miranda Stuckenberg, 1966 (MEI 116615). 6. T. stuckenbergi sp. n. (MEI 109134)." pageId="18" pageNumber="243">6</figureCitation>
|
||
), wider above, narrowing below (
|
||
<figureCitation box="[826,908,444,469]" captionStart="Figs 11–12" captionStartId="15.[182,220,729,749]" captionTargetBox="[241,1054,205,703]" captionTargetId="figure-286@15.[241,1054,205,703]" captionTargetPageId="15" captionText="Figs 11–12. Head profiles (lateral views) of male Tongamya. 11. T.miranda Stuckenberg, 1966 (MEI 116615). 12. T. stuckenbergi sp. n. (MEI 109134)." pageId="18" pageNumber="243">Fig. 12</figureCitation>
|
||
). Ocellar tubercle visible in frontal view(
|
||
<figureCitation box="[379,441,476,501]" captionStart="Figs 4–6. 4" captionStartId="12.[144,183,1371,1391]" captionTargetBox="[179,1075,206,1337]" captionTargetId="figure-81@12.[179,1075,203,1356]" captionTargetPageId="12" captionText="Figs 4–6. 4. Photograph of habitat where type material of Tongamya stuckenbergi sp. n. was collected (2 km S Otjisume Farm, Otjinene District, Namibia); note that the vegetation is not fully leafed. 5–6. Heads of male Tongamya, frontal views. 5. T. miranda Stuckenberg, 1966 (MEI 116615). 6. T. stuckenbergi sp. n. (MEI 109134)." pageId="18" pageNumber="243">Fig.6</figureCitation>
|
||
),formed by a transversely raised, rounded ridge.Frons narrowly triangular (
|
||
<figureCitation box="[263,329,508,533]" captionStart="Figs 4–6. 4" captionStartId="12.[144,183,1371,1391]" captionTargetBox="[179,1075,206,1337]" captionTargetId="figure-81@12.[179,1075,203,1356]" captionTargetPageId="12" captionText="Figs 4–6. 4. Photograph of habitat where type material of Tongamya stuckenbergi sp. n. was collected (2 km S Otjisume Farm, Otjinene District, Namibia); note that the vegetation is not fully leafed. 5–6. Heads of male Tongamya, frontal views. 5. T. miranda Stuckenberg, 1966 (MEI 116615). 6. T. stuckenbergi sp. n. (MEI 109134)." pageId="18" pageNumber="243">Fig. 6</figureCitation>
|
||
). Pruinescence light yellowish-grey overall. Antennae short, attached at lower third of head (
|
||
<figureCitation box="[371,453,540,565]" captionStart="Figs 7–10" captionStartId="13.[182,221,1360,1380]" captionTargetBox="[229,1072,205,1327]" captionTargetId="figure-88@13.[200,1107,205,1338]" captionTargetPageId="13" captionText="Figs 7–10. Heads of female Tongamya. 7–8. Frontal views, showing vertex, frons with anterior ocellus, antennae, face, and mouthparts. 9–10. Lateral views, showing vertex and ocellar tubercle, compound eye, frons, antennae, parafacials, genae, and occiput. 7, 9. T. miranda Stuckenberg, 1966 (MEI 113930). 8, 10. T. stuckenbergi (MEI 109135)." pageId="18" pageNumber="243">Fig. 10</figureCitation>
|
||
); ratio of scape:pedicel:flagellum, 4:2:9. Apex of flagellum subpyriform with off-centered fovea; stylus one-segmented. Compound eyes occupying most of head (
|
||
<figureCitation box="[299,379,604,629]" captionStart="Figs 11–12" captionStartId="15.[182,220,729,749]" captionTargetBox="[241,1054,205,703]" captionTargetId="figure-286@15.[241,1054,205,703]" captionTargetPageId="15" captionText="Figs 11–12. Head profiles (lateral views) of male Tongamya. 11. T.miranda Stuckenberg, 1966 (MEI 116615). 12. T. stuckenbergi sp. n. (MEI 109134)." pageId="18" pageNumber="243">Fig. 12</figureCitation>
|
||
); eye facets divided at level of antennae; lower facets smaller (
|
||
<figureCitation captionStart="Figs 4–6. 4" captionStartId="12.[144,183,1371,1391]" captionTargetBox="[179,1075,206,1337]" captionTargetId="figure-81@12.[179,1075,203,1356]" captionTargetPageId="12" captionText="Figs 4–6. 4. Photograph of habitat where type material of Tongamya stuckenbergi sp. n. was collected (2 km S Otjisume Farm, Otjinene District, Namibia); note that the vegetation is not fully leafed. 5–6. Heads of male Tongamya, frontal views. 5. T. miranda Stuckenberg, 1966 (MEI 116615). 6. T. stuckenbergi sp. n. (MEI 109134)." pageId="18" pageNumber="243">Fig. 6</figureCitation>
|
||
). Anterior ocellus enlarged; posterior ocelli one-third as broad and far removed from anterior ocellus (
|
||
<figureCitation box="[320,384,668,693]" captionStart="Figs 4–6. 4" captionStartId="12.[144,183,1371,1391]" captionTargetBox="[179,1075,206,1337]" captionTargetId="figure-81@12.[179,1075,203,1356]" captionTargetPageId="12" captionText="Figs 4–6. 4. Photograph of habitat where type material of Tongamya stuckenbergi sp. n. was collected (2 km S Otjisume Farm, Otjinene District, Namibia); note that the vegetation is not fully leafed. 5–6. Heads of male Tongamya, frontal views. 5. T. miranda Stuckenberg, 1966 (MEI 116615). 6. T. stuckenbergi sp. n. (MEI 109134)." pageId="18" pageNumber="243">Fig. 6</figureCitation>
|
||
). Occiput with short, scattered, outwardly radiating pale blond setae. Mouthparts greatly reduced; medially placed labium directed upward, cleft apically, forming a pair of elongate labial palps. Elongate, thinly pointed, spine-like structure, formed by union of two robust, convergent setae, directed outward and upward.Small, one-segmented maxillary palp flanking lateral edges of prementum.
|
||
</paragraph>
|
||
<paragraph blockId="18.[144,1105,206,1674]" pageId="18" pageNumber="243">
|
||
<emphasis box="[144,225,835,859]" italics="true" pageId="18" pageNumber="243">Thorax</emphasis>
|
||
: Tanish yellow, including legs with well defined dark brown patches on upper portion of antepronotum, outer lateral edges of basisternum, flanking anteriomesal edge of postalar callus, lateral edges of scutellum, anterolateral margins of postnotum, anteroventral portion of anepisternum, outer portion of anterior basalare, ventral most axillary sclerite, anterior and posterior edges of sclerite supporting anterior spiracle, anterior half of subalar sclerite, anterior edge of sclerite supporting posterior spiracle, lower half of both katepisternum and meron, anterobasal and apical margins of fore- and middle coxae, anterolateral face of hind coxa, dorso-apical tips of hind femora, and apical tarsomeres of all legs. Fine, sparse, tan pile confined to pronotum, outer lateral half of basisternum, and anterior face of forecoxa. Pile in a broad lateral sweep from postpronotal lobe to attachment of wing; parallel, longitudinal vitae of slightly shorter pile along flanks of mesonotum. Postpronotal lobes enlarged, tanish-yellow, with thin dusting of tan pruinescence and thin, dense, pale tan hairs. Presternum bare, with deep, ventro-medial groove. Scutellum deep, strongly transverse, yellow; dorsally flattened, mottled with tan pruinescence. Mesopleuron bare except pruinescence on ventral halves of katepisternum and meron, posterodorsal thirds of katepimeron, katatergite, outer lateral half of basisternum, and anterior face of middle coxa. Halter
|
||
<quantity box="[905,991,1346,1371]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="18" pageNumber="243" unit="mm" value="1.4">1.4 mm</quantity>
|
||
in length; stem tan, knob white.
|
||
</paragraph>
|
||
<paragraph blockId="18.[144,1105,206,1674]" pageId="18" pageNumber="243">
|
||
<emphasis box="[144,212,1417,1441]" italics="true" pageId="18" pageNumber="243">Wings</emphasis>
|
||
(
|
||
<figureCitation box="[226,297,1417,1442]" captionStart="Fig" captionStartId="7.[182,215,1623,1643]" captionTargetBox="[232,1101,709,1599]" captionTargetId="figure-9@7.[205,1112,709,1626]" captionTargetPageId="7" captionText="Fig. 2. Habitus view of male Tongamya stuckenbergi sp. n. (MEI 109134). 2a. Magnified view of male terminalia, lateral view." pageId="18" pageNumber="243">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation box="[308,324,1417,1442]" captionStart="Fig" captionStartId="9.[182,215,1307,1327]" captionTargetBox="[240,1094,203,1280]" captionTargetId="figure-125@9.[211,1115,203,1299]" captionTargetPageId="9" captionText="Fig. 3. Habitus view of female Tongamya stuckenbergi sp. n. (MEI 109135). 3a. Magnified view of female terminalia, lateral view." pageId="18" pageNumber="243">3</figureCitation>
|
||
): Length
|
||
<quantity box="[431,518,1417,1442]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="18" pageNumber="243" unit="mm" value="7.2">7.2 mm</quantity>
|
||
, of
|
||
<typeStatus box="[557,653,1417,1442]" pageId="18" pageNumber="243">paratype</typeStatus>
|
||
8.0 mm, 3 times longer than wide. Wing membrane transversely wrinkled, cloudy white; alula narrow. R
|
||
<subScript attach="left" box="[880,905,1466,1480]" fontSize="6" pageId="18" pageNumber="243">2+3</subScript>
|
||
, R
|
||
<subScript attach="left" box="[941,949,1466,1480]" fontSize="6" pageId="18" pageNumber="243">4</subScript>
|
||
and R
|
||
<subScript attach="left" box="[1029,1037,1466,1480]" fontSize="6" pageId="18" pageNumber="243">5</subScript>
|
||
+M
|
||
<subScript attach="left" box="[1078,1103,1466,1480]" fontSize="6" pageId="18" pageNumber="243">1+2</subScript>
|
||
ending in R
|
||
<subScript attach="left" box="[273,281,1498,1512]" fontSize="6" pageId="18" pageNumber="243">1</subScript>
|
||
, joining R
|
||
<subScript attach="left" box="[400,408,1498,1512]" fontSize="6" pageId="18" pageNumber="243">1</subScript>
|
||
almost at costa; CuA
|
||
<subScript attach="left" box="[647,655,1498,1512]" fontSize="6" pageId="18" pageNumber="243">2</subScript>
|
||
+M
|
||
<subScript attach="left" box="[696,704,1498,1512]" fontSize="6" pageId="18" pageNumber="243">3</subScript>
|
||
curving upward to end in R
|
||
<subScript attach="left" box="[1015,1023,1498,1512]" fontSize="6" pageId="18" pageNumber="243">5</subScript>
|
||
+M
|
||
<subScript attach="left" box="[1064,1089,1498,1512]" fontSize="6" pageId="18" pageNumber="243">1+2</subScript>
|
||
.. Costa weakening gradually from base of junction with Sc to wing tip. Costa, subcosta, R
|
||
<subScript attach="left" box="[162,170,1562,1576]" fontSize="6" pageId="18" pageNumber="243">1</subScript>
|
||
, CuA
|
||
<subScript attach="left" box="[241,249,1562,1576]" fontSize="6" pageId="18" pageNumber="243">2</subScript>
|
||
and, to a lesser extent, R
|
||
<subScript attach="left" box="[554,579,1562,1576]" fontSize="6" pageId="18" pageNumber="243">2+3</subScript>
|
||
and M
|
||
<subScript attach="left" box="[667,675,1562,1576]" fontSize="6" pageId="18" pageNumber="243">1</subScript>
|
||
pale brown; remaining veins almost colourless. Vein CuA
|
||
<subScript attach="left" box="[382,390,1594,1608]" fontSize="6" pageId="18" pageNumber="243">1</subScript>
|
||
curves upward to join M
|
||
<subScript attach="left" box="[672,680,1594,1608]" fontSize="6" pageId="18" pageNumber="243">3</subScript>
|
||
; m
|
||
<subScript attach="left" box="[717,725,1594,1608]" fontSize="6" pageId="18" pageNumber="243">3</subScript>
|
||
cell closed.
|
||
</paragraph>
|
||
<paragraph blockId="18.[144,1105,206,1674]" lastBlockId="19.[181,1142,206,391]" lastPageId="19" lastPageNumber="244" pageId="18" pageNumber="243">
|
||
<emphasis box="[144,197,1617,1641]" italics="true" pageId="18" pageNumber="243">Legs</emphasis>
|
||
(
|
||
<figureCitation box="[211,280,1617,1642]" captionStart="Fig" captionStartId="7.[182,215,1623,1643]" captionTargetBox="[232,1101,709,1599]" captionTargetId="figure-9@7.[205,1112,709,1626]" captionTargetPageId="7" captionText="Fig. 2. Habitus view of male Tongamya stuckenbergi sp. n. (MEI 109134). 2a. Magnified view of male terminalia, lateral view." pageId="18" pageNumber="243">Fig. 2</figureCitation>
|
||
): Surfaces of legs covered with dense, short, recumbent, yellow-tan hairs; hind tibial anteroventral setae short. Fore- and middle coxae with erect, sparse, yellow pile on anterior surface; hind coxa with similar pile on outer lateral surface. Hind legs conspicuously longer and thicker. Fore- and middle tibiae swollen apically; hind tibia greatly thickened apically, broadening to 2 times as wide at apex. Fore- and middle basitarsus thin, elongate, about 6 times longer than wide; hind basitarsus stout, about 2.5 times longer than wide, almost uniformly thick throughout, slightly narrower apically and almost as thick as apex of hind tibia.
|
||
</paragraph>
|
||
<paragraph blockId="19.[181,1142,407,752]" pageId="19" pageNumber="244">
|
||
<emphasis box="[181,287,408,432]" italics="true" pageId="19" pageNumber="244">Abdomen</emphasis>
|
||
(
|
||
<figureCitation box="[302,371,407,432]" captionStart="Fig" captionStartId="7.[182,215,1623,1643]" captionTargetBox="[232,1101,709,1599]" captionTargetId="figure-9@7.[205,1112,709,1626]" captionTargetPageId="7" captionText="Fig. 2. Habitus view of male Tongamya stuckenbergi sp. n. (MEI 109134). 2a. Magnified view of male terminalia, lateral view." pageId="19" pageNumber="244">Fig. 2</figureCitation>
|
||
): Elongate, tapering, 2.5 times longer than length of thorax. Tergite 1 swollen laterally, forming single broad, raised, tan callus across posterior half. Prominent circular spiracle imbedded within each posterodorsal corner of sternite 1. Spiracles of segments 3–7 set laterally in pleural membranes; sclerotised brown rings surround spiracular openings. Abdomen greasy in appearance, lacking pruinescence, with short, appressed, sparsely spaced pale yellow hairs, except for callus and terminalia, which have denser, longer hairs. T2–T7 dark, shiny brown dorsally, pale tan at lateral margins and as narrow wedges along outer posterior margins of T3–T7. Sternites and pleural areas pale tan; anteromedial portion of S2–S3 each with a small, dark brown, inverted ‘V’ marking, its arms coalesce into irregularly shaped blotches in similar positions on S4–S7.
|
||
</paragraph>
|
||
<paragraph blockId="19.[181,1142,769,1498]" pageId="19" pageNumber="244">
|
||
<emphasis box="[181,367,769,793]" italics="true" pageId="19" pageNumber="244">Male terminalia</emphasis>
|
||
: Erect, yellow on preserved specimen. Tergite 8 (
|
||
<figureCitation box="[954,1039,769,794]" captionStart="Fig" captionStartId="17.[182,215,872,892]" captionTargetBox="[229,1091,203,850]" captionTargetId="figure-288@17.[229,1091,203,850]" captionTargetPageId="17" captionText="Fig. 14. Male terminalia, lateral view, of Tongamya miranda Stuckenberg, 1966 (MEI 089484) depicting tergite 8, sternite 8, and major sclerites of the genitalia." pageId="19" pageNumber="244">Fig. 14</figureCitation>
|
||
) lacking vestiture, arched, medially narrow, anteromesally heavily brown pigmented, posteromesally weakly pigmented. Sternite 8 broad, deep, shovel-shaped, weakly pigmented, without vestiture. Epandrium (
|
||
<figureCitation box="[652,732,865,890]" captionStart="Figs 15–22" captionStartId="20.[144,183,1360,1380]" captionTargetBox="[221,1010,203,1340]" captionTargetId="figure-74@20.[221,1020,203,1354]" captionTargetPageId="20" captionText="Figs 15–22. Sclerites of the male genitalia. 15, 17, 19, 21. Epandria, cerci, and hypoprocts, dorsal view. 16, 18, 20, 22. Hypandria and ventral plates of gonopods, ventral view. 15, 16. Megascelus albovillosus Artigas, 1973 (MEI 108216). 17, 18. Neorhaphiomidas hardyi Norris, 1936 (MEI 118404). 19, 20. Tongamya miranda Stuckenberg, 1966 (MEI 116615). 21, 22. Tongamya stuckenbergi sp. n. (MEI 089484)." pageId="19" pageNumber="244">Fig. 21</figureCitation>
|
||
) divided by broad median biconcave cleft into 2 subtriangular sclerites anteriorly connected by sclerotised strap; brown with thin, sparse, pale tan hairs over posteriodorsal four-fifths. Elongate, flattened, semiforked plate (herein termed ‘ventral plate of gonopod’) lying beneath enlarged gonopod (
|
||
<figureCitation box="[190,282,993,1018]" captionStart="Fig" captionStartId="17.[182,215,872,892]" captionTargetBox="[229,1091,203,850]" captionTargetId="figure-288@17.[229,1091,203,850]" captionTargetPageId="17" captionText="Fig. 14. Male terminalia, lateral view, of Tongamya miranda Stuckenberg, 1966 (MEI 089484) depicting tergite 8, sternite 8, and major sclerites of the genitalia." pageId="19" pageNumber="244">Figs. 14</figureCitation>
|
||
,
|
||
<figureCitation box="[294,325,993,1018]" captionStart="Figs 15–22" captionStartId="20.[144,183,1360,1380]" captionTargetBox="[221,1010,203,1340]" captionTargetId="figure-74@20.[221,1020,203,1354]" captionTargetPageId="20" captionText="Figs 15–22. Sclerites of the male genitalia. 15, 17, 19, 21. Epandria, cerci, and hypoprocts, dorsal view. 16, 18, 20, 22. Hypandria and ventral plates of gonopods, ventral view. 15, 16. Megascelus albovillosus Artigas, 1973 (MEI 108216). 17, 18. Neorhaphiomidas hardyi Norris, 1936 (MEI 118404). 19, 20. Tongamya miranda Stuckenberg, 1966 (MEI 116615). 21, 22. Tongamya stuckenbergi sp. n. (MEI 089484)." pageId="19" pageNumber="244">22</figureCitation>
|
||
), heretofore assumed to be hypandrium in
|
||
<taxonomicName box="[800,916,993,1017]" class="Insecta" family="Mydidae" genus="Tongamya" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="244" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[800,916,993,1017]" italics="true" pageId="19" pageNumber="244">Tongamya</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation author="STUCKENBERG, B. R." box="[930,1137,993,1018]" pageId="19" pageNumber="244" pagination="106 - 120" refId="ref12950" refString="STUCKENBERG, B. R. 1966. A remarkable new genus and species of Apioceridae in South Africa (Diptera). Revue du Zoologie par Botanie du Africa 73: 106 - 120." type="journal article" year="1966">Stuckenberg 1966</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="YEATES, D. K. & IRWIN, M. E." box="[181,417,1025,1050]" pageId="19" pageNumber="244" pagination="247 - 301" refId="ref13418" refString="YEATES, D. K. & IRWIN, M. E. 1996. Apioceridae (Insecta: Diptera): Cladistic reappraisal and biogeography. Zoological Journal of the Linnaean Society 116: 247 - 301." type="journal article" year="1996">Yeates & Irwin 1996</bibRefCitation>
|
||
) and
|
||
<taxonomicName authorityName="Philippi" authorityYear="1865" box="[481,613,1025,1049]" class="Insecta" family="Mydidae" genus="Megascelus" kingdom="Animalia" order="Diptera" pageId="19" pageNumber="244" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[481,613,1025,1049]" italics="true" pageId="19" pageNumber="244">Megascelus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation author="ARTIGAS, J. N." box="[628,776,1025,1050]" pageId="19" pageNumber="244" pagination="97 - 122" refId="ref11437" refString="ARTIGAS, J. N. 1970. Los apioceratidos de Chile con la description de cinco nuevas especias (Diptera-Apioceratidae). Boletin da Sociedad Biologia de Concepcion 42: 97 - 122." type="journal article" year="1970">Artigas 1970</bibRefCitation>
|
||
1973). Our interpretation is that hypandrium may be a small, scoop-shaped, well sclerotised plate lying anterior to ventral plate of gonopod. If true, hypandrium bare, broadly scoop-shaped, overlapping ventral plate of gonopod (
|
||
<figureCitation box="[395,481,1121,1146]" captionStart="Figs 15–22" captionStartId="20.[144,183,1360,1380]" captionTargetBox="[221,1010,203,1340]" captionTargetId="figure-74@20.[221,1020,203,1354]" captionTargetPageId="20" captionText="Figs 15–22. Sclerites of the male genitalia. 15, 17, 19, 21. Epandria, cerci, and hypoprocts, dorsal view. 16, 18, 20, 22. Hypandria and ventral plates of gonopods, ventral view. 15, 16. Megascelus albovillosus Artigas, 1973 (MEI 108216). 17, 18. Neorhaphiomidas hardyi Norris, 1936 (MEI 118404). 19, 20. Tongamya miranda Stuckenberg, 1966 (MEI 116615). 21, 22. Tongamya stuckenbergi sp. n. (MEI 089484)." pageId="19" pageNumber="244">Fig. 22</figureCitation>
|
||
), supporting a pair of thin, strongly sclerotised arms that extend laterally and dorsally, and attach to lateral fold where gonopods and epandrium hinge (
|
||
<figureCitation box="[259,342,1185,1210]" captionStart="Fig" captionStartId="17.[182,215,872,892]" captionTargetBox="[229,1091,203,850]" captionTargetId="figure-288@17.[229,1091,203,850]" captionTargetPageId="17" captionText="Fig. 14. Male terminalia, lateral view, of Tongamya miranda Stuckenberg, 1966 (MEI 089484) depicting tergite 8, sternite 8, and major sclerites of the genitalia." pageId="19" pageNumber="244">Fig. 14</figureCitation>
|
||
). Gonopod consisting of large, paired, robust gonocoxites (
|
||
<figureCitation box="[1005,1086,1185,1210]" captionStart="Fig" captionStartId="17.[182,215,872,892]" captionTargetBox="[229,1091,203,850]" captionTargetId="figure-288@17.[229,1091,203,850]" captionTargetPageId="17" captionText="Fig. 14. Male terminalia, lateral view, of Tongamya miranda Stuckenberg, 1966 (MEI 089484) depicting tergite 8, sternite 8, and major sclerites of the genitalia." pageId="19" pageNumber="244">Fig. 14</figureCitation>
|
||
) and ventral plate of gonopod; gonostyli lost. Brown stripe along dorsolateral edge of each gonocoxite. Aedeagus consists of a slightly upcurved tube, widely open anteriorly and posteriorly, with a strong, dorsocentral, vertically oriented, rigid extension to base of hypoproct (
|
||
<figureCitation box="[307,391,1313,1338]" captionStart="Fig" captionStartId="17.[182,215,872,892]" captionTargetBox="[229,1091,203,850]" captionTargetId="figure-288@17.[229,1091,203,850]" captionTargetPageId="17" captionText="Fig. 14. Male terminalia, lateral view, of Tongamya miranda Stuckenberg, 1966 (MEI 089484) depicting tergite 8, sternite 8, and major sclerites of the genitalia." pageId="19" pageNumber="244">Figs 14</figureCitation>
|
||
,
|
||
<figureCitation box="[401,433,1313,1338]" captionStart="Figs 23–30" captionStartId="21.[182,221,1359,1379]" captionTargetBox="[240,1052,203,1337]" captionTargetId="figure-84@21.[240,1080,185,1337]" captionTargetPageId="21" captionText="Figs 23–30. Sclerites of the male genitalia. 23, 25, 27, 29. Aedeagus, lateral view. 24, 26, 28, 30. Aedeagus, dorsal view. 23, 24. Megascelus albovillosus Artigas, 1973 (MEI 108216). 25, 26. Neorhaphiomidas hardyi Norris, 1936 (MEI 118403). 27, 28. Tongamya miranda Stuckenberg, 1966 (MEI 116615). 29, 30. Tongamya stuckenbergi sp. n. (MEI 089484)." pageId="19" pageNumber="244">29</figureCitation>
|
||
). Distiphallus narrowing and flattening along apical third, flaring somewhat at apex (
|
||
<figureCitation box="[387,517,1345,1370]" captionStart="Figs 23–30" captionStartId="21.[182,221,1359,1379]" captionTargetBox="[240,1052,203,1337]" captionTargetId="figure-84@21.[240,1080,185,1337]" captionTargetPageId="21" captionText="Figs 23–30. Sclerites of the male genitalia. 23, 25, 27, 29. Aedeagus, lateral view. 24, 26, 28, 30. Aedeagus, dorsal view. 23, 24. Megascelus albovillosus Artigas, 1973 (MEI 108216). 25, 26. Neorhaphiomidas hardyi Norris, 1936 (MEI 118403). 27, 28. Tongamya miranda Stuckenberg, 1966 (MEI 116615). 29, 30. Tongamya stuckenbergi sp. n. (MEI 089484)." pageId="19" pageNumber="244">Figs. 29, 30</figureCitation>
|
||
). Ejaculatory apodeme small (
|
||
<figureCitation box="[841,921,1345,1370]" captionStart="Figs 23–30" captionStartId="21.[182,221,1359,1379]" captionTargetBox="[240,1052,203,1337]" captionTargetId="figure-84@21.[240,1080,185,1337]" captionTargetPageId="21" captionText="Figs 23–30. Sclerites of the male genitalia. 23, 25, 27, 29. Aedeagus, lateral view. 24, 26, 28, 30. Aedeagus, dorsal view. 23, 24. Megascelus albovillosus Artigas, 1973 (MEI 108216). 25, 26. Neorhaphiomidas hardyi Norris, 1936 (MEI 118403). 27, 28. Tongamya miranda Stuckenberg, 1966 (MEI 116615). 29, 30. Tongamya stuckenbergi sp. n. (MEI 089484)." pageId="19" pageNumber="244">Fig. 29</figureCitation>
|
||
). Lateral ejaculatory process (
|
||
<bibRefCitation author="SINCLAIR, B. J. & CUMMING, J. M. & WOOD, D. M." box="[283,508,1377,1402]" pageId="19" pageNumber="244" pagination="407 - 432" refId="ref12815" refString="SINCLAIR, B. J., CUMMING, J. M. & WOOD, D. M. 1993. Homology and phylogenetic implications of male genitalia in Diptera-Lower Brachycera. Entomologica Scandinavica 24: 407 - 432." type="journal article" year="1993">
|
||
Sinclair
|
||
<emphasis box="[382,441,1377,1401]" italics="true" pageId="19" pageNumber="244">et al.</emphasis>
|
||
1993
|
||
</bibRefCitation>
|
||
) prominent, horseshoe shaped. Cerci form a shallowly roofed, bilobed sclerite (
|
||
<figureCitation box="[457,541,1409,1434]" captionStart="Figs 15–22" captionStartId="20.[144,183,1360,1380]" captionTargetBox="[221,1010,203,1340]" captionTargetId="figure-74@20.[221,1020,203,1354]" captionTargetPageId="20" captionText="Figs 15–22. Sclerites of the male genitalia. 15, 17, 19, 21. Epandria, cerci, and hypoprocts, dorsal view. 16, 18, 20, 22. Hypandria and ventral plates of gonopods, ventral view. 15, 16. Megascelus albovillosus Artigas, 1973 (MEI 108216). 17, 18. Neorhaphiomidas hardyi Norris, 1936 (MEI 118404). 19, 20. Tongamya miranda Stuckenberg, 1966 (MEI 116615). 21, 22. Tongamya stuckenbergi sp. n. (MEI 089484)." pageId="19" pageNumber="244">Fig. 21</figureCitation>
|
||
). Hypoproct horizontally flattened, thin, extending to apex of cerci (
|
||
<figureCitation box="[360,451,1441,1466]" captionStart="Fig" captionStartId="17.[182,215,872,892]" captionTargetBox="[229,1091,203,850]" captionTargetId="figure-288@17.[229,1091,203,850]" captionTargetPageId="17" captionText="Fig. 14. Male terminalia, lateral view, of Tongamya miranda Stuckenberg, 1966 (MEI 089484) depicting tergite 8, sternite 8, and major sclerites of the genitalia." pageId="19" pageNumber="244">Fig. 14</figureCitation>
|
||
). Basally, hypoproct curves ventrad, uniting solidly with dorsocentral portion of aedeagus.
|
||
</paragraph>
|
||
<paragraph blockId="19.[181,1142,1514,1667]" lastBlockId="21.[181,1142,1485,1670]" lastPageId="21" lastPageNumber="246" pageId="19" pageNumber="244">
|
||
<emphasis box="[181,263,1515,1539]" italics="true" pageId="19" pageNumber="244">Female</emphasis>
|
||
(
|
||
<emphasis box="[278,414,1515,1539]" italics="true" pageId="19" pageNumber="244">
|
||
<taxonomicName authorityName="Xu & Norell" authorityYear="2004" box="[278,327,1515,1539]" class="Reptilia" family="Troodontidae" genus="Mei" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="19" pageNumber="244" phylum="Chordata" rank="genus">MEI</taxonomicName>
|
||
109135
|
||
</emphasis>
|
||
): Length, excluding antennae,
|
||
<quantity box="[750,850,1514,1539]" metricMagnitude="-2" metricUnit="m" metricValue="1.58" pageId="19" pageNumber="244" unit="mm" value="15.8">15.8 mm</quantity>
|
||
. Similar to male but larger. Head proportionally smaller, 1.5 times wider than high. In lateral view, head narrow above, widest at level of antennae, narrowing below (
|
||
<figureCitation box="[683,764,1578,1603]" captionStart="Figs 7–10" captionStartId="13.[182,221,1360,1380]" captionTargetBox="[229,1072,205,1327]" captionTargetId="figure-88@13.[200,1107,205,1338]" captionTargetPageId="13" captionText="Figs 7–10. Heads of female Tongamya. 7–8. Frontal views, showing vertex, frons with anterior ocellus, antennae, face, and mouthparts. 9–10. Lateral views, showing vertex and ocellar tubercle, compound eye, frons, antennae, parafacials, genae, and occiput. 7, 9. T. miranda Stuckenberg, 1966 (MEI 113930). 8, 10. T. stuckenbergi (MEI 109135)." pageId="19" pageNumber="244">Fig. 10</figureCitation>
|
||
). Frons wider than high, shallowly concave, with patchy, sparse, greyish-white pruinescence around anterior ocellus; with numerous, vertically oriented, subparallel, irregular grooves over frons and vertex (
|
||
<figureCitation box="[1063,1127,1642,1667]" captionStart="Figs 7–10" captionStartId="13.[182,221,1360,1380]" captionTargetBox="[229,1072,205,1327]" captionTargetId="figure-88@13.[200,1107,205,1338]" captionTargetPageId="13" captionText="Figs 7–10. Heads of female Tongamya. 7–8. Frontal views, showing vertex, frons with anterior ocellus, antennae, face, and mouthparts. 9–10. Lateral views, showing vertex and ocellar tubercle, compound eye, frons, antennae, parafacials, genae, and occiput. 7, 9. T. miranda Stuckenberg, 1966 (MEI 113930). 8, 10. T. stuckenbergi (MEI 109135)." pageId="19" pageNumber="244">Fig. 8</figureCitation>
|
||
). Vertex yellow-tan with vertical swelling laterally (
|
||
<figureCitation box="[693,759,1515,1540]" captionStart="Figs 7–10" captionStartId="13.[182,221,1360,1380]" captionTargetBox="[229,1072,205,1327]" captionTargetId="figure-88@13.[200,1107,205,1338]" captionTargetPageId="13" captionText="Figs 7–10. Heads of female Tongamya. 7–8. Frontal views, showing vertex, frons with anterior ocellus, antennae, face, and mouthparts. 9–10. Lateral views, showing vertex and ocellar tubercle, compound eye, frons, antennae, parafacials, genae, and occiput. 7, 9. T. miranda Stuckenberg, 1966 (MEI 113930). 8, 10. T. stuckenbergi (MEI 109135)." pageId="20" pageNumber="245">Fig. 8</figureCitation>
|
||
) (area between ocellar tubercle and compound eye of Etosha and
|
||
<collectingCountry box="[501,581,1547,1572]" name="Angola" pageId="20" pageNumber="245">Angola</collectingCountry>
|
||
specimens not swollen); vertex elevated on sides above eye margins, forming a central, raised, tan ocellar tubercle (more prominent in Etosha andAngola specimens).Genae slightly swollen, cream-white, with thin white pruinescence. Compound eyes dichoptic, bare, small, undivided; facets uniform in size (
|
||
<figureCitation box="[923,997,1643,1668]" captionStart="Figs 7–10" captionStartId="13.[182,221,1360,1380]" captionTargetBox="[229,1072,205,1327]" captionTargetId="figure-88@13.[200,1107,205,1338]" captionTargetPageId="13" captionText="Figs 7–10. Heads of female Tongamya. 7–8. Frontal views, showing vertex, frons with anterior ocellus, antennae, face, and mouthparts. 9–10. Lateral views, showing vertex and ocellar tubercle, compound eye, frons, antennae, parafacials, genae, and occiput. 7, 9. T. miranda Stuckenberg, 1966 (MEI 113930). 8, 10. T. stuckenbergi (MEI 109135)." pageId="20" pageNumber="245">Fig.10</figureCitation>
|
||
).Anterior ocellus enlarged. Posterior ocelli smaller, situated laterally on ocellar tubercle, visible in frontal view (
|
||
<figureCitation box="[330,397,1517,1542]" captionStart="Figs 7–10" captionStartId="13.[182,221,1360,1380]" captionTargetBox="[229,1072,205,1327]" captionTargetId="figure-88@13.[200,1107,205,1338]" captionTargetPageId="13" captionText="Figs 7–10. Heads of female Tongamya. 7–8. Frontal views, showing vertex, frons with anterior ocellus, antennae, face, and mouthparts. 9–10. Lateral views, showing vertex and ocellar tubercle, compound eye, frons, antennae, parafacials, genae, and occiput. 7, 9. T. miranda Stuckenberg, 1966 (MEI 113930). 8, 10. T. stuckenbergi (MEI 109135)." pageId="21" pageNumber="246">Fig. 8</figureCitation>
|
||
). Wing length
|
||
<quantity box="[562,650,1517,1542]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="21" pageNumber="246" unit="mm" value="7.8">7.8 mm</quantity>
|
||
, width
|
||
<quantity box="[732,820,1517,1542]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="21" pageNumber="246" unit="mm" value="2.6">2.6 mm</quantity>
|
||
. Hind tibiae and basitarsi not swollen (
|
||
<figureCitation box="[281,347,1549,1574]" captionStart="Fig" captionStartId="9.[182,215,1307,1327]" captionTargetBox="[240,1094,203,1280]" captionTargetId="figure-125@9.[211,1115,203,1299]" captionTargetPageId="9" captionText="Fig. 3. Habitus view of female Tongamya stuckenbergi sp. n. (MEI 109135). 3a. Magnified view of female terminalia, lateral view." pageId="21" pageNumber="246">Fig. 3</figureCitation>
|
||
). Abdomen greasy in appearance, dark reddish-brown; swollen over first five segments (widest at segment 3), tapering rapidly at segment 6, with segments 7, 8 and genitalia cylindrical and half as thick as segment 3 (
|
||
<figureCitation box="[722,785,1613,1638]" captionStart="Fig" captionStartId="9.[182,215,1307,1327]" captionTargetBox="[240,1094,203,1280]" captionTargetId="figure-125@9.[211,1115,203,1299]" captionTargetPageId="9" captionText="Fig. 3. Habitus view of female Tongamya stuckenbergi sp. n. (MEI 109135). 3a. Magnified view of female terminalia, lateral view." pageId="21" pageNumber="246">Fig. 3</figureCitation>
|
||
); white marking on posterolateral corners of T2–T4 (
|
||
<figureCitation box="[384,449,1645,1670]" captionStart="Fig" captionStartId="9.[182,215,1307,1327]" captionTargetBox="[240,1094,203,1280]" captionTargetId="figure-125@9.[211,1115,203,1299]" captionTargetPageId="9" captionText="Fig. 3. Habitus view of female Tongamya stuckenbergi sp. n. (MEI 109135). 3a. Magnified view of female terminalia, lateral view." pageId="21" pageNumber="246">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption pageId="20" pageNumber="245" startId="20.[144,183,1360,1380]" targetBox="[221,1010,203,1340]" targetPageId="20" targetType="figure">
|
||
<paragraph blockId="20.[144,1104,1360,1476]" pageId="20" pageNumber="245">
|
||
Figs 15–22. Sclerites of the male genitalia. 15, 17, 19, 21. Epandria, cerci, and hypoprocts, dorsal view. 16, 18, 20, 22. Hypandria and ventral plates of gonopods, ventral view. 15, 16.
|
||
<taxonomicName authority="Artigas, 1973" authorityName="Artigas" authorityYear="1973" class="Insecta" family="Mydidae" genus="Megascelus" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="245" phylum="Arthropoda" rank="species" species="albovillosus">
|
||
<emphasis box="[888,1104,1384,1404]" italics="true" pageId="20" pageNumber="245">Megascelus albovillosus</emphasis>
|
||
Artigas, 1973
|
||
</taxonomicName>
|
||
(
|
||
<emphasis box="[378,492,1408,1428]" italics="true" pageId="20" pageNumber="245">
|
||
<taxonomicName authorityName="Xu & Norell" authorityYear="2004" box="[378,418,1408,1428]" class="Reptilia" family="Troodontidae" genus="Mei" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="20" pageNumber="245" phylum="Chordata" rank="genus">MEI</taxonomicName>
|
||
108216
|
||
</emphasis>
|
||
). 17, 18.
|
||
<taxonomicName authority="Norris, 1936" authorityName="Norris" authorityYear="1936" box="[581,929,1408,1428]" class="Insecta" family="Mydidae" genus="Neorhaphiomidas" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="245" phylum="Arthropoda" rank="species" species="hardyi">
|
||
<emphasis box="[581,807,1408,1428]" italics="true" pageId="20" pageNumber="245">Neorhaphiomidas hardyi</emphasis>
|
||
<bibRefCitation author="NORRIS, K. R." box="[814,929,1408,1428]" pageId="20" pageNumber="245" pagination="49 - 70" refId="ref12597" refString="NORRIS, K. R. 1936. New species of Apioceridae (Diptera) from Western Australia. Journal of the Royal Society of Western Australia 22: 49 - 70." type="journal article" year="1936">Norris, 1936</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis box="[943,1056,1408,1428]" italics="true" pageId="20" pageNumber="245">
|
||
<taxonomicName authorityName="Xu & Norell" authorityYear="2004" box="[943,983,1408,1428]" class="Reptilia" family="Troodontidae" genus="Mei" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="20" pageNumber="245" phylum="Chordata" rank="genus">MEI</taxonomicName>
|
||
118404
|
||
</emphasis>
|
||
). 19, 20.
|
||
<taxonomicName authority="Stuckenberg, 1966" authorityName="Stuckenberg" authorityYear="1966" box="[274,624,1432,1452]" class="Insecta" family="Mydidae" genus="Tongamya" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="245" phylum="Arthropoda" rank="species" species="miranda">
|
||
<emphasis box="[274,450,1432,1452]" italics="true" pageId="20" pageNumber="245">Tongamya miranda</emphasis>
|
||
<bibRefCitation author="STUCKENBERG, B. R." box="[455,624,1432,1452]" pageId="20" pageNumber="245" pagination="106 - 120" refId="ref12950" refString="STUCKENBERG, B. R. 1966. A remarkable new genus and species of Apioceridae in South Africa (Diptera). Revue du Zoologie par Botanie du Africa 73: 106 - 120." type="journal article" year="1966">Stuckenberg, 1966</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis box="[637,749,1432,1452]" italics="true" pageId="20" pageNumber="245">
|
||
<taxonomicName authorityName="Xu & Norell" authorityYear="2004" box="[637,677,1432,1452]" class="Reptilia" family="Troodontidae" genus="Mei" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="20" pageNumber="245" phylum="Chordata" rank="genus">MEI</taxonomicName>
|
||
116615
|
||
</emphasis>
|
||
). 21, 22.
|
||
<taxonomicName authority="Irwin & Wiegmann, 2001" authorityName="Irwin & Wiegmann" authorityYear="2001" box="[836,1050,1432,1452]" class="Insecta" family="Mydidae" genus="Tongamya" kingdom="Animalia" order="Diptera" pageId="20" pageNumber="245" phylum="Arthropoda" rank="species" species="stuckenbergi" status="sp. n.">
|
||
<emphasis box="[836,1050,1432,1452]" italics="true" pageId="20" pageNumber="245">Tongamya stuckenbergi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1055,1104,1432,1452]" pageId="20" pageNumber="245" rank="species">sp. n.</taxonomicNameLabel>
|
||
(
|
||
<emphasis box="[247,361,1456,1476]" italics="true" pageId="20" pageNumber="245">
|
||
<taxonomicName authorityName="Xu & Norell" authorityYear="2004" box="[247,287,1456,1476]" class="Reptilia" family="Troodontidae" genus="Mei" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="20" pageNumber="245" phylum="Chordata" rank="genus">MEI</taxonomicName>
|
||
089484
|
||
</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="21" pageNumber="246" startId="21.[182,221,1359,1379]" targetBox="[240,1052,203,1337]" targetPageId="21" targetType="figure">
|
||
<paragraph blockId="21.[182,1142,1359,1451]" pageId="21" pageNumber="246">
|
||
Figs 23–30. Sclerites of the male genitalia. 23, 25, 27, 29. Aedeagus, lateral view. 24, 26, 28, 30. Aedeagus, dorsal view. 23, 24.
|
||
<taxonomicName authority="Artigas, 1973" authorityName="Artigas" authorityYear="1973" box="[496,897,1383,1403]" class="Insecta" family="Mydidae" genus="Megascelus" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="246" phylum="Arthropoda" rank="species" species="albovillosus">
|
||
<emphasis box="[496,746,1383,1403]" italics="true" pageId="21" pageNumber="246">Megascelus albovillosus</emphasis>
|
||
Artigas, 1973
|
||
</taxonomicName>
|
||
(
|
||
<emphasis box="[916,1044,1383,1403]" italics="true" pageId="21" pageNumber="246">
|
||
<taxonomicName authorityName="Xu & Norell" authorityYear="2004" box="[916,959,1383,1403]" class="Reptilia" family="Troodontidae" genus="Mei" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="21" pageNumber="246" phylum="Chordata" rank="genus">MEI</taxonomicName>
|
||
108216
|
||
</emphasis>
|
||
). 25, 26.
|
||
<taxonomicName authority="Norris, 1936" authorityName="Norris" authorityYear="1936" box="[278,625,1407,1427]" class="Insecta" family="Mydidae" genus="Neorhaphiomidas" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="246" phylum="Arthropoda" rank="species" species="hardyi">
|
||
<emphasis box="[278,504,1407,1427]" italics="true" pageId="21" pageNumber="246">Neorhaphiomidas hardyi</emphasis>
|
||
<bibRefCitation author="NORRIS, K. R." box="[510,625,1407,1427]" pageId="21" pageNumber="246" pagination="49 - 70" refId="ref12597" refString="NORRIS, K. R. 1936. New species of Apioceridae (Diptera) from Western Australia. Journal of the Royal Society of Western Australia 22: 49 - 70." type="journal article" year="1936">Norris, 1936</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis box="[639,752,1407,1427]" italics="true" pageId="21" pageNumber="246">
|
||
<taxonomicName authorityName="Xu & Norell" authorityYear="2004" box="[639,679,1407,1427]" class="Reptilia" family="Troodontidae" genus="Mei" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="21" pageNumber="246" phylum="Chordata" rank="genus">MEI</taxonomicName>
|
||
118403
|
||
</emphasis>
|
||
). 27, 28.
|
||
<taxonomicName authority="Stuckenberg, 1966" authorityName="Stuckenberg" authorityYear="1966" class="Insecta" family="Mydidae" genus="Tongamya" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="246" phylum="Arthropoda" rank="species" species="miranda">
|
||
<emphasis box="[840,1017,1407,1427]" italics="true" pageId="21" pageNumber="246">Tongamya miranda</emphasis>
|
||
<bibRefCitation author="STUCKENBERG, B. R." pageId="21" pageNumber="246" pagination="106 - 120" refId="ref12950" refString="STUCKENBERG, B. R. 1966. A remarkable new genus and species of Apioceridae in South Africa (Diptera). Revue du Zoologie par Botanie du Africa 73: 106 - 120." type="journal article" year="1966">Stuckenberg, 1966</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis box="[336,448,1431,1451]" italics="true" pageId="21" pageNumber="246">
|
||
<taxonomicName authorityName="Xu & Norell" authorityYear="2004" box="[336,376,1431,1451]" class="Reptilia" family="Troodontidae" genus="Mei" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="21" pageNumber="246" phylum="Chordata" rank="genus">MEI</taxonomicName>
|
||
116615
|
||
</emphasis>
|
||
). 29, 30.
|
||
<taxonomicName authority="Irwin & Wiegmann, 2001" authorityName="Irwin & Wiegmann" authorityYear="2001" box="[535,750,1431,1451]" class="Insecta" family="Mydidae" genus="Tongamya" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="246" phylum="Arthropoda" rank="species" species="stuckenbergi" status="sp. n.">
|
||
<emphasis box="[535,750,1431,1451]" italics="true" pageId="21" pageNumber="246">Tongamya stuckenbergi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[755,804,1431,1451]" pageId="21" pageNumber="246" rank="species">sp. n.</taxonomicNameLabel>
|
||
(
|
||
<emphasis box="[817,931,1431,1451]" italics="true" pageId="21" pageNumber="246">
|
||
<taxonomicName authorityName="Xu & Norell" authorityYear="2004" box="[817,857,1431,1451]" class="Reptilia" family="Troodontidae" genus="Mei" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="21" pageNumber="246" phylum="Chordata" rank="genus">MEI</taxonomicName>
|
||
089484
|
||
</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="22" pageNumber="247" startId="22.[144,177,888,908]" targetBox="[181,1062,203,867]" targetPageId="22" targetType="figure">
|
||
<paragraph blockId="22.[144,1104,888,956]" pageId="22" pageNumber="247">
|
||
Fig. 31.Auxiliary spiracle of female tergite 8 in
|
||
<taxonomicName authority="Irwin & Wiegmann, 2001" authorityName="Irwin & Wiegmann" authorityYear="2001" box="[570,783,888,908]" class="Insecta" family="Mydidae" genus="Tongamya" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="247" phylum="Arthropoda" rank="species" species="stuckenbergi" status="sp. n.">
|
||
<emphasis box="[570,783,888,908]" italics="true" pageId="22" pageNumber="247">Tongamya stuckenbergi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[787,835,888,908]" pageId="22" pageNumber="247" rank="species">sp. n.</taxonomicNameLabel>
|
||
(
|
||
<emphasis box="[846,959,888,908]" italics="true" pageId="22" pageNumber="247">
|
||
<taxonomicName authorityName="Xu & Norell" authorityYear="2004" box="[846,886,888,908]" class="Reptilia" family="Troodontidae" genus="Mei" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="22" pageNumber="247" phylum="Chordata" rank="genus">MEI</taxonomicName>
|
||
108188
|
||
</emphasis>
|
||
), showing cross section of tergite 8, the invaginated spiracular opening, and some larger internal tracheae. Orientation: the anterior end of the insect is to the left.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="22.[144,1104,1005,1670]" lastBlockId="23.[181,1142,1413,1534]" lastPageId="23" lastPageNumber="248" pageId="22" pageNumber="247">
|
||
<emphasis box="[144,353,1005,1029]" italics="true" pageId="22" pageNumber="247">Female terminalia</emphasis>
|
||
: Segment 8 telescoped within segment 7. Spiracles inset at weakly pigmented posterior area of segment 8; spiracular opening smaller than spiracles of other segments; characterised by thin trachea extending inward for a considerable distance, then expanding greatly within segment (
|
||
<figureCitation box="[700,781,1101,1126]" captionStart="Fig" captionStartId="22.[144,177,888,908]" captionTargetBox="[181,1062,203,867]" captionTargetId="figure-265@22.[181,1068,203,867]" captionTargetPageId="22" captionText="Fig. 31.Auxiliary spiracle of female tergite 8 in Tongamya stuckenbergi sp. n. (MEI 108188), showing cross section of tergite 8, the invaginated spiracular opening, and some larger internal tracheae. Orientation: the anterior end of the insect is to the left." pageId="22" pageNumber="247">Fig. 31</figureCitation>
|
||
) (termed ‘auxiliary spiracle’ by
|
||
<bibRefCitation author="MCALPINE, J. F." box="[177,417,1133,1158]" pageId="22" pageNumber="247" pagination="9 - 63" refId="ref12538" refString="MCALPINE, J. F. 1981. Morphology and terminology - Adults. In: McAlpine, J. F., et al., eds., Manual of Nearctic Diptera. Volume 1. Ottawa: Agriculture Canada, Research Branch. (Monograph; no. 27) pp. 9 - 63." type="journal article" year="1981">
|
||
McAlpine
|
||
<emphasis box="[297,352,1133,1157]" italics="true" pageId="22" pageNumber="247">et al.</emphasis>
|
||
1981
|
||
</bibRefCitation>
|
||
). Tergite 8 (T8) elongate, ‘U’ shaped in cross section, tapering to shovel-shaped towards apex. Anterodorsal third relatively smooth and bare; rest of dorsum with short, sparse hairs. Sternite 8 (S8) similar in shape to T8 but shorter, narrower, ventrally smooth in texture with scattered, short, erect hairs. Posteriorly truncate, slightly infolded, forming shallow invagination. Tergite 9+10 (T9+10) heavily sclerotised, comprised of upper, anterior component (T9) and two ventrolateral, posterior sclerites (acanthophorites, T10) (
|
||
<figureCitation box="[513,601,1325,1350]" captionStart="Fig" captionStartId="23.[182,215,1317,1337]" captionTargetBox="[193,1121,203,1288]" captionTargetId="figure-66@23.[181,1121,203,1288]" captionTargetPageId="23" captionText="Fig. 32. Female genitalia of Tongamya stuckenbergi sp. n. (MEI 108188) depicting internal weakly sclerotised structures, dorsal view. T9 = tergite 9." pageId="22" pageNumber="247">Figs 32</figureCitation>
|
||
,
|
||
<figureCitation box="[612,644,1325,1350]" captionStart="Fig" captionStartId="26.[144,177,839,859]" captionTargetBox="[280,965,203,820]" captionTargetId="figure-436@26.[280,965,203,820]" captionTargetPageId="26" captionText="Fig. 35. Female genitalia of Tongamya stuckenbergi sp. n. (MEI 109135), lateral view, depicting position of tergite 9 (T9), acanthophorite with digging spines, cercus, and anterior attachment to furca." pageId="22" pageNumber="247">35</figureCitation>
|
||
). T9 dome-shaped (
|
||
<figureCitation box="[865,949,1325,1350]" captionStart="Fig" captionStartId="26.[144,177,839,859]" captionTargetBox="[280,965,203,820]" captionTargetId="figure-436@26.[280,965,203,820]" captionTargetPageId="26" captionText="Fig. 35. Female genitalia of Tongamya stuckenbergi sp. n. (MEI 109135), lateral view, depicting position of tergite 9 (T9), acanthophorite with digging spines, cercus, and anterior attachment to furca." pageId="22" pageNumber="247">Fig. 35</figureCitation>
|
||
), divided into 2 narrowly fused dorsolateral sclerites (
|
||
<figureCitation box="[582,666,1357,1382]" captionStart="Fig" captionStartId="23.[182,215,1317,1337]" captionTargetBox="[193,1121,203,1288]" captionTargetId="figure-66@23.[181,1121,203,1288]" captionTargetPageId="23" captionText="Fig. 32. Female genitalia of Tongamya stuckenbergi sp. n. (MEI 108188) depicting internal weakly sclerotised structures, dorsal view. T9 = tergite 9." pageId="22" pageNumber="247">Fig. 32</figureCitation>
|
||
). Sternite 9 (furca, S9) (
|
||
<figureCitation box="[930,1010,1357,1382]" captionStart="Fig" captionStartId="23.[182,215,1317,1337]" captionTargetBox="[193,1121,203,1288]" captionTargetId="figure-66@23.[181,1121,203,1288]" captionTargetPageId="23" captionText="Fig. 32. Female genitalia of Tongamya stuckenbergi sp. n. (MEI 108188) depicting internal weakly sclerotised structures, dorsal view. T9 = tergite 9." pageId="22" pageNumber="247">Fig. 32</figureCitation>
|
||
) framed by longitudinal, opposing, membrane-spanded struts. Spermathecal elements (
|
||
<figureCitation box="[1015,1095,1389,1414]" captionStart="Fig" captionStartId="23.[182,215,1317,1337]" captionTargetBox="[193,1121,203,1288]" captionTargetId="figure-66@23.[181,1121,203,1288]" captionTargetPageId="23" captionText="Fig. 32. Female genitalia of Tongamya stuckenbergi sp. n. (MEI 108188) depicting internal weakly sclerotised structures, dorsal view. T9 = tergite 9." pageId="22" pageNumber="247">Fig. 32</figureCitation>
|
||
) arise from gonopore as a common spermathecal duct. Two large, irregularly ovalshaped, weakly sclerotised spermathecae attach by thin spermathecal ducts to midpoint of common spermathecal duct, which extends as a twisted duct terminating in a spermathecal sac (
|
||
<figureCitation box="[358,443,1517,1542]" captionStart="Fig" captionStartId="23.[182,215,1317,1337]" captionTargetBox="[193,1121,203,1288]" captionTargetId="figure-66@23.[181,1121,203,1288]" captionTargetPageId="23" captionText="Fig. 32. Female genitalia of Tongamya stuckenbergi sp. n. (MEI 108188) depicting internal weakly sclerotised structures, dorsal view. T9 = tergite 9." pageId="22" pageNumber="247">Fig. 32</figureCitation>
|
||
) (
|
||
<bibRefCitation author="WINTERTON, S. L. & MERRITT, D. J. & O'TOOLE, A. & YEATES, D. K. & IRWIN, M. E." box="[470,727,1517,1542]" pageId="22" pageNumber="247" pagination="273 - 279" refId="ref13255" refString="WINTERTON, S. L., MERRITT, D. J., O'TOOLE, A., YEATES, D. K. & IRWIN, M. E. 1999. Morphology and histology of the spermathecal sac, a novel structure in the female reproductive system of Therevidae (Diptera: Asiloidea). International Journal of Insect Morphology and Embryology 28: 273 - 279." type="journal article" year="1999">
|
||
Winterton
|
||
<emphasis box="[596,656,1517,1541]" italics="true" pageId="22" pageNumber="247">et al.</emphasis>
|
||
1999
|
||
</bibRefCitation>
|
||
). Two elongate, slightly bulging accessory glands attach by thin ducts originating from 2 minute, parallel apertures at posterior end of furcal membrane (
|
||
<figureCitation box="[581,668,1581,1606]" captionStart="Fig" captionStartId="23.[182,215,1317,1337]" captionTargetBox="[193,1121,203,1288]" captionTargetId="figure-66@23.[181,1121,203,1288]" captionTargetPageId="23" captionText="Fig. 32. Female genitalia of Tongamya stuckenbergi sp. n. (MEI 108188) depicting internal weakly sclerotised structures, dorsal view. T9 = tergite 9." pageId="22" pageNumber="247">Fig. 32</figureCitation>
|
||
). T10 (acanthophorites) (
|
||
<figureCitation box="[961,1052,1581,1606]" captionStart="Fig" captionStartId="23.[182,215,1317,1337]" captionTargetBox="[193,1121,203,1288]" captionTargetId="figure-66@23.[181,1121,203,1288]" captionTargetPageId="23" captionText="Fig. 32. Female genitalia of Tongamya stuckenbergi sp. n. (MEI 108188) depicting internal weakly sclerotised structures, dorsal view. T9 = tergite 9." pageId="22" pageNumber="247">Figs 32</figureCitation>
|
||
,
|
||
<figureCitation box="[1066,1095,1581,1606]" captionStart="Fig" captionStartId="26.[144,177,839,859]" captionTargetBox="[280,965,203,820]" captionTargetId="figure-436@26.[280,965,203,820]" captionTargetPageId="26" captionText="Fig. 35. Female genitalia of Tongamya stuckenbergi sp. n. (MEI 109135), lateral view, depicting position of tergite 9 (T9), acanthophorite with digging spines, cercus, and anterior attachment to furca." pageId="22" pageNumber="247">35</figureCitation>
|
||
) paired sclerites, curving and widening ventrad, each baring circlet of 6–7 strong, flat, blunt spines (
|
||
<figureCitation box="[293,376,1645,1670]" captionStart="Fig" captionStartId="26.[144,177,839,859]" captionTargetBox="[280,965,203,820]" captionTargetId="figure-436@26.[280,965,203,820]" captionTargetPageId="26" captionText="Fig. 35. Female genitalia of Tongamya stuckenbergi sp. n. (MEI 109135), lateral view, depicting position of tergite 9 (T9), acanthophorite with digging spines, cercus, and anterior attachment to furca." pageId="22" pageNumber="247">Fig. 35</figureCitation>
|
||
). Hypoproct at posterior margin of furca, extends posteromesally as a small, weakly sclerotised, long-haired structure, terminating between paired lateral cerci (
|
||
<figureCitation box="[340,429,1445,1470]" captionStart="Fig" captionStartId="23.[182,215,1317,1337]" captionTargetBox="[193,1121,203,1288]" captionTargetId="figure-66@23.[181,1121,203,1288]" captionTargetPageId="23" captionText="Fig. 32. Female genitalia of Tongamya stuckenbergi sp. n. (MEI 108188) depicting internal weakly sclerotised structures, dorsal view. T9 = tergite 9." pageId="23" pageNumber="248">Fig. 32</figureCitation>
|
||
). Cerci sausage-shaped, thinly sclerotised, longhaired, lying laterally within and conforming to inner basal curve of paired acanthophorites (
|
||
<figureCitation box="[190,275,1509,1534]" captionStart="Fig" captionStartId="26.[144,177,839,859]" captionTargetBox="[280,965,203,820]" captionTargetId="figure-436@26.[280,965,203,820]" captionTargetPageId="26" captionText="Fig. 35. Female genitalia of Tongamya stuckenbergi sp. n. (MEI 109135), lateral view, depicting position of tergite 9 (T9), acanthophorite with digging spines, cercus, and anterior attachment to furca." pageId="23" pageNumber="248">Fig. 35</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |