treatments-xml/data/03/93/80/03938025FF93A909A7D8B3E6FE14FE50.xml
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<document id="931B2BD7D73C6D32258B35A9B0D35D30" ID-CLB-Dataset="8556" ID-DOI="10.11646/zootaxa.4843.1.1" ID-GBIF-Dataset="a2aef684-44f1-429d-9dd5-dd394539fa36" ID-ISSN="1175-5326" ID-Zenodo-Dep="4405729" ID-ZooBank="57044928-1950-4525-850E-E381A52599A1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1609419131752" checkinUser="plazi" docAuthor="Salata, Sebastian &amp; Fisher, Brian L." docDate="2020" docId="03938025FF93A909A7D8B3E6FE14FE50" docLanguage="en" docName="zootaxa.4843.1.1.pdf" docOrigin="Zootaxa 4843 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Pheidole flavodepressa Salata &amp; Fisher 2020, sp. nov." docType="treatment" docVersion="9" lastPageNumber="32" masterDocId="FFAAF85DFF8EA916A74FB608FFC6FFFD" masterDocTitle="The ant genus Pheidole Westwood, 1839 (Hymenoptera: Formicidae) in Madagascar-taxonomic revision of the bessonii species-group" masterLastPageNumber="64" masterPageNumber="1" pageNumber="30" updateTime="1698850390174" updateUser="ExternalLinkService">
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<mods:title id="5838682B6DAA51D3BD831CB349895AA5">The ant genus Pheidole Westwood, 1839 (Hymenoptera: Formicidae) in Madagascar-taxonomic revision of the bessonii species-group</mods:title>
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<mods:name id="FCD220A2A615A27E6237E53C9C52859C" type="personal">
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<mods:namePart id="3A28254962072855D545FC02E19B6069">Salata, Sebastian</mods:namePart>
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<mods:name id="0190D680A949A2CA21CB5215A73CBA93" type="personal">
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<mods:roleTerm id="0358F769378B9F20C6236F6FE2F3E8B1">Author</mods:roleTerm>
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<mods:namePart id="E1E2DEDD07018CDB4A6E0498EDA4B0C8">Fisher, Brian L.</mods:namePart>
<mods:nameIdentifier id="EC43ABBB64091C1C29A4EDFCC89EB919" type="ORCID">0000-0002-4653-3270</mods:nameIdentifier>
<mods:nameIdentifier id="D549F7A00C7C97D30BE89406A781527B" type="email">bfisher@calacademy.org</mods:nameIdentifier>
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<mods:title id="7296CF04C97C9208B4024E32D8E0233E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D293B75F8CB9B7658523404B254152AD">
<mods:date id="A60E4AE03B999625CA6F15BF3EF7E1B8">2020</mods:date>
<mods:detail id="FFAAA3DD592E4CE59B87AB01CCADC22B" type="pubDate">
<mods:number id="0EAC905AEC2B3A810E99038CF557ABB2">2020-08-31</mods:number>
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<mods:number id="8D9C5733FCC5D9B89735BC13F9216B88">4843</mods:number>
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<mods:classification id="F6B2661AC7B15E63D99FEAA69D2C6014">journal article</mods:classification>
<mods:identifier id="BB77735B12357E1DAA652EEB5C701768" type="CLB-Dataset">8556</mods:identifier>
<mods:identifier id="3DD3B57253DBD74F9E4E19E6ECF8F559" type="DOI">10.11646/zootaxa.4843.1.1</mods:identifier>
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<treatment id="03938025FF93A909A7D8B3E6FE14FE50" ID-DOI="http://doi.org/10.5281/zenodo.4488387" ID-GBIF-Taxon="177202126" ID-Zenodo-Dep="4488387" LSID="urn:lsid:plazi:treatment:03938025FF93A909A7D8B3E6FE14FE50" httpUri="http://treatment.plazi.org/id/03938025FF93A909A7D8B3E6FE14FE50" lastPageId="31" lastPageNumber="32" pageId="29" pageNumber="30">
<subSubSection id="C32062B8FF93A90BA7D8B3E6FDD4F9F4" box="[151,530,1518,1545]" pageId="29" pageNumber="30" type="nomenclature">
<paragraph id="8B853133FF93A90BA7D8B3E6FDD4F9F4" blockId="29.[151,530,1518,1581]" box="[151,530,1518,1545]" pageId="29" pageNumber="30">
<heading id="D0CD865FFF93A90BA7D8B3E6FDD4F9F4" bold="true" box="[151,530,1518,1545]" fontSize="11" level="2" pageId="29" pageNumber="30" reason="4">
<emphasis id="B94EED21FF93A90BA7D8B3E6FDD4F9F4" bold="true" box="[151,530,1518,1545]" pageId="29" pageNumber="30">
<taxonomicName id="4C3A4AB0FF93A90BA7D8B3E6FE6BF9F4" authority="Salata &amp; Fisher, 2020" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[151,429,1518,1545]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="flavodepressa" status="sp. nov.">
<emphasis id="B94EED21FF93A90BA7D8B3E6FE6BF9F4" bold="true" box="[151,429,1518,1545]" italics="true" pageId="29" pageNumber="30">Pheidole flavodepressa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D505AFF93A90BA6FBB3E7FDD4F9F4" box="[436,530,1519,1545]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FF93A90BA7D8B01BFE50F9D0" box="[151,406,1554,1581]" pageId="29" pageNumber="30" type="description">
<paragraph id="8B853133FF93A90BA7D8B01BFE50F9D0" blockId="29.[151,530,1518,1581]" box="[151,406,1554,1581]" pageId="29" pageNumber="30">
<figureCitation id="13012DB6FF93A90BA7D8B01BFF21F9D0" box="[151,231,1554,1581]" captionStart="FIGURE 15" captionStartId="31.[151,250,1975,1999]" captionTargetBox="[151,1436,453,1946]" captionTargetId="figure-133@31.[151,1436,453,1952]" captionTargetPageId="31" captionText="FIGURE 15. Pheidole flavodepressa, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0230800) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0243625)." figureDoi="http://doi.org/10.5281/zenodo.4405769" httpUri="https://zenodo.org/record/4405769/files/figure.png" pageId="29" pageNumber="30">Figs 15</figureCitation>
AF, 27G, 28G
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FF93A90BA7D8B053FB69F8B0" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="8B853133FF93A90BA7D8B053FCF2F940" blockId="29.[151,1437,1626,2013]" pageId="29" pageNumber="30">
<materialsCitation id="3B523B6EFF93A90BA7D8B053FB29F964" ID-GBIF-Occurrence="3027954591" accessionNumber="BLF27044" collectingDate="2011-05-04" collectionCode="CASC" collectorName="B. L. Fisher" country="Madagascar" elevation="225" latitude="-14.104" location="Makirovana forest" longLatPrecision="55" longitude="50.03574" pageId="29" pageNumber="30" specimenCount="1" stateProvince="Antsiranana" typeStatus="holotype">
<typeStatus id="54818F91FF93A90BA7D8B053FEE9F988" box="[151,303,1627,1653]" pageId="29" pageNumber="30" type="holotype">HOLOTYPE</typeStatus>
: 1s.,
<collectingCountry id="F32D71A3FF93A90BA628B053FE36F988" box="[359,496,1627,1653]" name="Madagascar" pageId="29" pageNumber="30">Madagascar</collectingCountry>
,
<collectingRegion id="49FEFFD1FF93A90BA6B6B053FD45F988" box="[505,643,1627,1653]" country="Madagascar" name="Antsiranana" pageId="29" pageNumber="30">Antsiranana</collectingRegion>
,
<location id="8EE567E8FF93A90BA5C1B053FC99F988" LSID="urn:lsid:plazi:treatment:03938025FF93A909A7D8B3E6FE14FE50:8EE567E8FF93A90BA5C1B053FC99F988" box="[654,863,1627,1653]" country="Madagascar" latitude="-14.104" longLatPrecision="55" longitude="50.03574" name="Makirovana forest" pageId="29" pageNumber="30" stateProvince="Antsiranana">Makirovana forest</location>
,
<geoCoordinate id="EE0E57F4FF93A90BA425B053FC06F988" box="[874,960,1627,1653]" degrees="14.104" direction="south" orientation="latitude" pageId="29" pageNumber="30" precision="55" value="-14.104">-14.104</geoCoordinate>
<geoCoordinate id="EE0E57F4FF93A90BA488B052FBF5F988" box="[967,1075,1626,1653]" degrees="50.03574" direction="east" orientation="longitude" pageId="29" pageNumber="30" precision="1" value="50.03574">50.03574</geoCoordinate>
,
<quantity id="4CC29CD6FF93A90BA372B052FB40F988" box="[1085,1158,1626,1653]" metricMagnitude="2" metricUnit="m" metricValue="2.25" pageId="29" pageNumber="30" unit="m" value="225.0">
<elevation id="0017D600FF93A90BA372B052FB40F988" box="[1085,1158,1626,1653]" metricMagnitude="2" metricUnit="m" metricValue="2.25" pageId="29" pageNumber="30" unit="m" value="225.0">225 m</elevation>
</quantity>
,
<date id="FF8417F3FF93A90BA3DEB053FAD9F988" box="[1169,1311,1627,1653]" pageId="29" pageNumber="30" value="2011-05-04">
<collectingDate id="EFC0EE1BFF93A90BA3DEB053FAD9F988" box="[1169,1311,1627,1653]" pageId="29" pageNumber="30" value="2011-05-04">4-May-2011</collectingDate>
</date>
, rainforest, sifted litter (leaf mold, rotten wood),
<collectorName id="26CF54E5FF93A90BA577B077FD7BF964" box="[568,701,1663,1689]" pageId="29" pageNumber="30">B. L. Fisher</collectorName>
<emphasis id="B94EED21FF93A90BA58BB077FD33F964" box="[708,757,1663,1689]" italics="true" pageId="29" pageNumber="30">et al</emphasis>
. leg. BLF27044, CASENT0243625 (
<collectionCode id="ED2BA9F6FF93A90BA3D7B077FB21F964" box="[1176,1255,1663,1689]" pageId="29" pageNumber="30">CASC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B523B6EFF93A90BA3B5B077FCE9F940" ID-GBIF-Occurrence="3027954431" accessionNumber="BLF27044" collectingDate="2011-05-04" collectionCode="CASC" collectorName="B. L. Fisher" country="Madagascar" elevation="225" latitude="-14.104" location="Makirovana forest" longLatPrecision="55" longitude="50.03574" pageId="29" pageNumber="30" specimenCount="1" stateProvince="Antsiranana" typeStatus="paratype">
<typeStatus id="54818F91FF93A90BA3B5B077FA5FF964" box="[1274,1433,1663,1689]" pageId="29" pageNumber="30" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="9D3CFABAFF93A90BA7D8B0ABFF06F940" box="[151,192,1699,1725]" pageId="29" pageNumber="30" type="generic">1w.</specimenCount>
, the same data as holotype, CASENT0230800 (
<collectionCode id="ED2BA9F6FF93A90BA597B0ABFCE1F940" box="[728,807,1699,1725]" pageId="29" pageNumber="30">CASC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B853133FF93A90BA788B0CEFE71F8D4" blockId="29.[151,1437,1626,2013]" pageId="29" pageNumber="30">
<emphasis id="B94EED21FF93A90BA788B0CEFE47F91D" bold="true" box="[199,385,1734,1760]" pageId="29" pageNumber="30">Other material.</emphasis>
<materialsCitation id="3B523B6EFF93A90BA6C7B0CFFEAAF8F8" ID-GBIF-Occurrence="3027954614" collectionCode="CASC" collectorName="B. L. Fisher" country="Madagascar" elevation="225" latitude="-14.104" location="Makirovana forest" longLatPrecision="55" longitude="50.03574" pageId="29" pageNumber="30" specimenCount="4" stateProvince="Antsiranana">
<collectingCountry id="F32D71A3FF93A90BA6C7B0CFFDD7F91C" box="[392,529,1735,1761]" name="Madagascar" pageId="29" pageNumber="30">Madagascar</collectingCountry>
.
<collectingRegion id="49FEFFD1FF93A90BA555B0CFFD60F91C" box="[538,678,1735,1761]" country="Madagascar" name="Antsiranana" pageId="29" pageNumber="30">Antsiranana</collectingRegion>
:
<specimenCount id="9D3CFABAFF93A90BA5FFB0CFFD1CF91C" box="[688,730,1735,1761]" pageId="29" pageNumber="30" type="generic">4w.</specimenCount>
, 2s.,
<location id="8EE567E8FF93A90BA45CB0CFFC22F91C" LSID="urn:lsid:plazi:treatment:03938025FF93A909A7D8B3E6FE14FE50:8EE567E8FF93A90BA45CB0CFFC22F91C" box="[787,996,1735,1761]" country="Madagascar" latitude="-14.104" longLatPrecision="55" longitude="50.03574" name="Makirovana forest" pageId="29" pageNumber="30" stateProvince="Antsiranana">Makirovana forest</location>
,
<geoCoordinate id="EE0E57F4FF93A90BA4A1B0CFFB83F91C" box="[1006,1093,1735,1761]" degrees="14.104" direction="south" orientation="latitude" pageId="29" pageNumber="30" precision="55" value="-14.104">-14.104</geoCoordinate>
<geoCoordinate id="EE0E57F4FF93A90BA303B0CEFB7EF91C" box="[1100,1208,1734,1761]" degrees="50.03574" direction="east" orientation="longitude" pageId="29" pageNumber="30" precision="1" value="50.03574">50.03574</geoCoordinate>
,
<quantity id="4CC29CD6FF93A90BA38DB0CEFACDF91C" box="[1218,1291,1734,1761]" metricMagnitude="2" metricUnit="m" metricValue="2.25" pageId="29" pageNumber="30" unit="m" value="225.0">
<elevation id="0017D600FF93A90BA38DB0CEFACDF91C" box="[1218,1291,1734,1761]" metricMagnitude="2" metricUnit="m" metricValue="2.25" pageId="29" pageNumber="30" unit="m" value="225.0">225 m</elevation>
</quantity>
,
<collectorName id="26CF54E5FF93A90BA258B0CFFA5AF91C" box="[1303,1436,1735,1761]" pageId="29" pageNumber="30">B. L. Fisher</collectorName>
<emphasis id="B94EED21FF93A90BA7D8B0E3FF0CF8F8" box="[151,202,1771,1797]" italics="true" pageId="29" pageNumber="30">et al</emphasis>
. leg. (
<collectionCode id="ED2BA9F6FF93A90BA65AB0E3FEA2F8F8" box="[277,356,1771,1797]" pageId="29" pageNumber="30">CASC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B523B6EFF93A90BA636B0E3FE75F8D4" ID-GBIF-Occurrence="3027954440" collectionCode="CASC" collectorName="R. S. Manongarivo &amp; B. L. Fisher" country="Madagascar" elevation="1175" latitude="-13.99833" location="Antanambao" longLatPrecision="1" longitude="48.42833" pageId="29" pageNumber="30" specimenCount="1" stateProvince="Antsiranana">
<specimenCount id="9D3CFABAFF93A90BA636B0E3FE64F8F8" box="[377,418,1771,1797]" pageId="29" pageNumber="30" type="generic">1w.</specimenCount>
,
<collectorName id="26CF54E5FF93A90BA6FEB0E3FD42F8F8" box="[433,644,1771,1797]" pageId="29" pageNumber="30">R.S. Manongarivo</collectorName>
,
<quantity id="4CC29CD6FF93A90BA5DEB0E2FD29F8F8" box="[657,751,1770,1797]" metricMagnitude="4" metricUnit="m" metricValue="1.45" pageId="29" pageNumber="30" unit="km" value="14.5">14.5 km</quantity>
220° SW
<location id="8EE567E8FF93A90BA427B0E3FC3CF8F8" LSID="urn:lsid:plazi:treatment:03938025FF93A909A7D8B3E6FE14FE50:8EE567E8FF93A90BA427B0E3FC3CF8F8" box="[872,1018,1771,1797]" country="Madagascar" latitude="-13.99833" longLatPrecision="1" longitude="48.42833" name="Antanambao" pageId="29" pageNumber="30" stateProvince="Antsiranana">Antanambao</location>
,
<geoCoordinate id="EE0E57F4FF93A90BA348B0E3FBBFF8F8" box="[1031,1145,1771,1797]" degrees="13.99833" direction="south" orientation="latitude" pageId="29" pageNumber="30" precision="1" value="-13.99833">-13.99833</geoCoordinate>
<geoCoordinate id="EE0E57F4FF93A90BA3CDB0E3FB28F8F8" box="[1154,1262,1771,1797]" degrees="48.42833" direction="east" orientation="longitude" pageId="29" pageNumber="30" precision="1" value="48.42833">48.42833</geoCoordinate>
,
<quantity id="4CC29CD6FF93A90BA3B4B0E2FA95F8F8" box="[1275,1363,1770,1797]" metricMagnitude="3" metricUnit="m" metricValue="1.175" pageId="29" pageNumber="30" unit="m" value="1175.0">
<elevation id="0017D600FF93A90BA3B4B0E2FA95F8F8" box="[1275,1363,1770,1797]" metricMagnitude="3" metricUnit="m" metricValue="1.175" pageId="29" pageNumber="30" unit="m" value="1175.0">1175 m</elevation>
</quantity>
,
<collectorName id="26CF54E5FF93A90BA22EB0E3FF1BF8D4" pageId="29" pageNumber="30">B. L. Fisher</collectorName>
<emphasis id="B94EED21FF93A90BA7ABB107FED3F8D4" box="[228,277,1807,1833]" italics="true" pageId="29" pageNumber="30">et al</emphasis>
. leg. (
<collectionCode id="ED2BA9F6FF93A90BA614B107FE6CF8D4" box="[347,426,1807,1833]" pageId="29" pageNumber="30">CASC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B853133FF93A90BA788B13AFB69F8B0" blockId="29.[151,1437,1626,2013]" box="[199,1199,1842,1869]" pageId="29" pageNumber="30">
<emphasis id="B94EED21FF93A90BA788B13AFE60F8B0" bold="true" box="[199,422,1842,1869]" pageId="29" pageNumber="30">Geographic range.</emphasis>
<collectingCountry id="F32D71A3FF93A90BA6E2B13BFDF0F8B0" box="[429,566,1843,1869]" name="Madagascar" pageId="29" pageNumber="30">Madagascar</collectingCountry>
,
<collectingRegion id="49FEFFD1FF93A90BA570B13BFD0FF8B0" box="[575,713,1843,1869]" country="Madagascar" name="Antsiranana" pageId="29" pageNumber="30">Antsiranana</collectingRegion>
, Makirovana forest and R.S. Manongarivo.
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FF93A908A788B15EFC08FE08" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="diagnosis">
<paragraph id="8B853133FF93A908A788B15EFC08FE08" blockId="29.[151,1437,1626,2013]" lastBlockId="30.[151,1437,151,2013]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
<emphasis id="B94EED21FF93A90BA788B15EFDC4F88C" bold="true" box="[199,514,1878,1905]" pageId="29" pageNumber="30">Diagnosis. Major workers.</emphasis>
Head in full-face view sub-rectangular, elongated, not widening posteriorly, lateral margins relatively straight with very deep posteromedian concavity; side of head with moderately dense, long, subdecumbent to suberect pilosity; medial frons with dense and thick costulae; interspaces between costulae distinctly rugulate; frons laterally with thick and dense rugae; interspaces between rugae distinctly rugulate; occipital lobes with thick and dense rugae; interspaces between rugae distinctly rugulate; sculpture not weakening posteriorly; scape, when laid back, exceeding the midlength of head by two-fifths of its length; inner hypostomal tooth indistinct and bulge-like; outer hypostomal tooth lobe-like, high, and wide, with top arching posterolaterally; median tooth absent; promesonotum short, angular, and moderately low; posterior mesonotum smoothly declining towards propodeum; mesonotal process indistinct and tubercle-like; mesosoma punctate; promesonotum with additional thin, dense, and transverse rugae; katepisternum and lateral sides of propodeum with additional thick and dense rugulae; gaster shagreened; body orange.
<emphasis id="B94EED21FF90A908A526B766FCE3FE74" bold="true" box="[617,805,366,393]" pageId="30" pageNumber="31">Minor workers.</emphasis>
Occiput elongated forming moderately long and narrow neck; nuchal collar distinct; head sculpture smooth; scape, when laid back, exceeding the posterior head margin by three-fifths of its length; promesonotum low, very long, and arched; promesonotal groove present; metanotal groove distinct; propodeal spines long and thin; mesosoma smooth; body yellow.
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FF90A908A788B7F6FF7FF8F8" pageId="30" pageNumber="31" type="description">
<paragraph id="8B853133FF90A908A788B7F6FBEAFD78" blockId="30.[151,1437,151,2013]" pageId="30" pageNumber="31">
<emphasis id="B94EED21FF90A908A788B7F6FDDEFDE4" bold="true" box="[199,536,510,537]" pageId="30" pageNumber="31">Description. Major workers.</emphasis>
Measurements (n=3): HL: 1.861.89 (1.88); HW: 1.561.66 (1.6); SL: 1.331.41 (1.37); EL: 0.20.23 (0.22); WL: 1.681.76 (1.71); PSL: 0.40.42 (0.41); MTL: 1.531.54 (1.54); PNW: 0.640.66 (0.65); PTW: 0.180.2 (0.19); PPW: 0.530.58 (0.56); CI: 113.7119.3 (117.3); SI: 83.690.2 (85.8); PSLI: 21.0 22.3 (21.9); PPI: 31.235.8 (33.8); PNI: 39.642.3 (40.7); MTI: 92.898.9 (95.9).
</paragraph>
<paragraph id="8B853133FF90A908A788B487FBB7FAA8" blockId="30.[151,1437,151,2013]" pageId="30" pageNumber="31">
<emphasis id="B94EED21FF90A908A788B487FEC8FD54" bold="true" box="[199,270,655,681]" pageId="30" pageNumber="31">Head.</emphasis>
In full-face view sub-rectangular, elongated, not widening posteriorly, lateral sides relatively straight with very deep posteromedian concavity (
<figureCitation id="13012DB6FF90A908A525B4BBFD16FD30" box="[618,720,690,717]" captionStart="FIGURE 15" captionStartId="31.[151,250,1975,1999]" captionTargetBox="[151,1436,453,1946]" captionTargetId="figure-133@31.[151,1436,453,1952]" captionTargetPageId="31" captionText="FIGURE 15. Pheidole flavodepressa, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0230800) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0243625)." figureDoi="http://doi.org/10.5281/zenodo.4405769" httpUri="https://zenodo.org/record/4405769/files/figure.png" pageId="30" pageNumber="31">Fig. 15B</figureCitation>
). In lateral view sub-oval, not depressed posteriorly; ventral and dorsal margins convex; inner hypostomal tooth not visible. Side of head with moderately dense, long, subdecumbent to suberect pilosity; head dorsum with relatively dense, long, suberect to erect pilosity. Medial frons with dense and thick costulae; interspaces between costulae distinctly rugulate. Frons laterally with thick and dense rugae; interspaces between rugae distinctly rugulate. Occipital lobes with thick and dense rugae; interspaces between rugae distinctly rugulate; sculpture not weakening posteriorly. Gena with dense and moderately thick costulae; interspaces between costulae mostly smooth. Sides posterolateral from eyes with dense but very thin network of rugoreticulae; sculpture not weakening posteriorly. Center of clypeus shiny with few rugae, lateral sides with distinct rugulae; median notch present, moderately wide and shallow; median longitudinal carina present; lateral longitudinal carinae present. Scape, when laid back, exceeding the midlength of head by two-fifths of its length; pilosity subdecumbent to erect (
<figureCitation id="13012DB6FF90A908A7B1B213FE4DFBC8" box="[254,395,1050,1077]" captionStart="FIGURE 15" captionStartId="31.[151,250,1975,1999]" captionTargetBox="[151,1436,453,1946]" captionTargetId="figure-133@31.[151,1436,453,1952]" captionTargetPageId="31" captionText="FIGURE 15. Pheidole flavodepressa, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0230800) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0243625)." figureDoi="http://doi.org/10.5281/zenodo.4405769" httpUri="https://zenodo.org/record/4405769/files/figure.png" pageId="30" pageNumber="31">Fig. 15B, D</figureCitation>
). Inner hypostomal tooth indistinct, bulge-like; outer hypostomal tooth lobe-like, high, and wide, with top arching posterolaterally; median tooth absent (
<figureCitation id="13012DB6FF90A908A40CB237FC60FBA4" box="[835,934,1087,1113]" captionStart="FIGURE 27" captionStartId="60.[151,250,1604,1628]" captionTargetBox="[151,1436,1137,1581]" captionTargetId="figure-432@60.[151,1436,1137,1581]" captionTargetPageId="60" captionText="FIGURE 27. Major worker, hypostomal tooth. Pheidole antsahabe (A). Pheidole atsirakambiaty (B). Pheidole bessonii (C). Pheidole clara (D). Pheidole decollata (E). Pheidole flammea (F). Pheidole flavodepressa (G). Pheidole grallatrix (H). Pheidole madecassa (I). Pheidole mantadioflava (J). Pheidole maro (K). Pheidole oswaldi (L). Pheidole ovalinoda (M). Pheidole similis (N). Pheidole tenebrovulgaris (O). Pheidole uranus (P). Pheidole voreios (Q). Pheidole zirafy (R)." figureDoi="http://doi.org/10.5281/zenodo.4405803" httpUri="https://zenodo.org/record/4405803/files/figure.png" pageId="30" pageNumber="31">Fig. 27G</figureCitation>
).
<emphasis id="B94EED21FF90A908A4F2B237FB84FBA4" bold="true" box="[957,1090,1087,1113]" pageId="30" pageNumber="31">Mesosoma.</emphasis>
In lateral view, promesonotum short, angular, and moderately low; posterior mesonotum smoothly declining towards propodeum; mesonotal process indistinct and tubercle-like; promesonotal groove absent; metanotal groove indistinct; propodeal spines long, with narrow base and acute top; humeral tubercle weakly produced (
<figureCitation id="13012DB6FF90A908A4E8B2A3FBD7FB38" box="[935,1041,1194,1221]" captionStart="FIGURE 15" captionStartId="31.[151,250,1975,1999]" captionTargetBox="[151,1436,453,1946]" captionTargetId="figure-133@31.[151,1436,453,1952]" captionTargetPageId="31" captionText="FIGURE 15. Pheidole flavodepressa, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0230800) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0243625)." figureDoi="http://doi.org/10.5281/zenodo.4405769" httpUri="https://zenodo.org/record/4405769/files/figure.png" pageId="30" pageNumber="31">Fig. 15D</figureCitation>
). Surface punctate; promesonotum with additional thin, dense, and transverse rugae; katepisternum and lateral sides of propodeum with additional thick and dense rugulae. Pilosity dense, long, and erect (
<figureCitation id="13012DB6FF90A908A59AB2FBFC9AFAF0" box="[725,860,1266,1293]" captionStart="FIGURE 15" captionStartId="31.[151,250,1975,1999]" captionTargetBox="[151,1436,453,1946]" captionTargetId="figure-133@31.[151,1436,453,1952]" captionTargetPageId="31" captionText="FIGURE 15. Pheidole flavodepressa, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0230800) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0243625)." figureDoi="http://doi.org/10.5281/zenodo.4405769" httpUri="https://zenodo.org/record/4405769/files/figure.png" pageId="30" pageNumber="31">Fig. 15D, F</figureCitation>
).
<emphasis id="B94EED21FF90A908A43FB2FAFC0EFAF1" bold="true" box="[880,968,1266,1292]" pageId="30" pageNumber="31">Petiole.</emphasis>
In rear view node dorsoventrally slightly convex (
<figureCitation id="13012DB6FF90A908A7B9B31FFEBCFACC" box="[246,378,1302,1329]" captionStart="FIGURE 15" captionStartId="31.[151,250,1975,1999]" captionTargetBox="[151,1436,453,1946]" captionTargetId="figure-133@31.[151,1436,453,1952]" captionTargetPageId="31" captionText="FIGURE 15. Pheidole flavodepressa, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0230800) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0243625)." figureDoi="http://doi.org/10.5281/zenodo.4405769" httpUri="https://zenodo.org/record/4405769/files/figure.png" pageId="30" pageNumber="31">Fig. 15D, F</figureCitation>
).
<emphasis id="B94EED21FF90A908A6C2B31EFE24FACD" bold="true" box="[397,482,1302,1328]" pageId="30" pageNumber="31">Gaster.</emphasis>
Dull and shagreened; pilosity moderately sparse, long, and erect (
<figureCitation id="13012DB6FF90A908A3F2B31FFA86FACC" box="[1213,1344,1302,1329]" captionStart="FIGURE 15" captionStartId="31.[151,250,1975,1999]" captionTargetBox="[151,1436,453,1946]" captionTargetId="figure-133@31.[151,1436,453,1952]" captionTargetPageId="31" captionText="FIGURE 15. Pheidole flavodepressa, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0230800) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0243625)." figureDoi="http://doi.org/10.5281/zenodo.4405769" httpUri="https://zenodo.org/record/4405769/files/figure.png" pageId="30" pageNumber="31">Fig. 15D, F</figureCitation>
).
<emphasis id="B94EED21FF90A908A21CB31EFA5AFACD" bold="true" box="[1363,1436,1302,1328]" pageId="30" pageNumber="31">Color.</emphasis>
Orange, antennae and legs yellow, gaster and mandibles brownish-orange (
<figureCitation id="13012DB6FF90A908A491B333FBA2FAA8" box="[990,1124,1338,1365]" captionStart="FIGURE 15" captionStartId="31.[151,250,1975,1999]" captionTargetBox="[151,1436,453,1946]" captionTargetId="figure-133@31.[151,1436,453,1952]" captionTargetPageId="31" captionText="FIGURE 15. Pheidole flavodepressa, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0230800) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0243625)." figureDoi="http://doi.org/10.5281/zenodo.4405769" httpUri="https://zenodo.org/record/4405769/files/figure.png" pageId="30" pageNumber="31">Fig. 15D, F</figureCitation>
).
</paragraph>
<paragraph id="8B853133FF90A908A788B356FB1BFA18" blockId="30.[151,1437,151,2013]" pageId="30" pageNumber="31">
<emphasis id="B94EED21FF90A908A788B356FDD8FA84" bold="true" box="[199,542,1374,1401]" pageId="30" pageNumber="31">Description. Minor workers.</emphasis>
Measurements (n=5): HL: 0.840.91 (0.87); HW: 0.480.51 (0.49); SL: 1.26 1.36 (1.31); EL: 0.140.16 (0.15); WL: 1.121.18 (1.14); PSL: 0.20.24 (0.22); MTL: 1.141.25 (1.19); PNW: 0.370.38 (0.38); PTW: 0.070.09 (0.08); PPW: 0.130.16 (0.15); CI: 173.5185.3 (177.1); SI: 257.9279.6 (266.9); PSLI: 22.827.6 (25.7); PPI: 49.062.0 (56.0); PNI: 74.479.6 (76.9); MTI: 236.1256.3 (242.2).
</paragraph>
<paragraph id="8B853133FF90A908A788B3E7FF7FF8F8" blockId="30.[151,1437,151,2013]" pageId="30" pageNumber="31">
<emphasis id="B94EED21FF90A908A788B3E7FEC8F9F4" bold="true" box="[199,270,1519,1545]" pageId="30" pageNumber="31">Head.</emphasis>
In full-face view oval, posterior region elongated forming moderately long and narrow neck; nuchal collar distinct (
<figureCitation id="13012DB6FF90A908A60DB01BFE6AF9D0" box="[322,428,1554,1581]" captionStart="FIGURE 15" captionStartId="31.[151,250,1975,1999]" captionTargetBox="[151,1436,453,1946]" captionTargetId="figure-133@31.[151,1436,453,1952]" captionTargetPageId="31" captionText="FIGURE 15. Pheidole flavodepressa, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0230800) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0243625)." figureDoi="http://doi.org/10.5281/zenodo.4405769" httpUri="https://zenodo.org/record/4405769/files/figure.png" pageId="30" pageNumber="31">Fig. 15A</figureCitation>
). Pilosity relatively sparse, moderately long, subdecumbent to erect. Sculpture shiny and smooth. Clypeus with median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, exceeding the posterior head margin by three-fifths of its length; pilosity dense, subdecumbent to erect (
<figureCitation id="13012DB6FF90A908A220B053FF29F964" captionStart="FIGURE 15" captionStartId="31.[151,250,1975,1999]" captionTargetBox="[151,1436,453,1946]" captionTargetId="figure-133@31.[151,1436,453,1952]" captionTargetPageId="31" captionText="FIGURE 15. Pheidole flavodepressa, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0230800) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0243625)." figureDoi="http://doi.org/10.5281/zenodo.4405769" httpUri="https://zenodo.org/record/4405769/files/figure.png" pageId="30" pageNumber="31">Fig. 15A, C</figureCitation>
).
<emphasis id="B94EED21FF90A908A649B077FE4DF964" bold="true" box="[262,395,1663,1689]" pageId="30" pageNumber="31">Mesosoma.</emphasis>
In lateral view, promesonotum low, very long, and arched; promesonotal groove present; metanotal groove distinct; propodeal spines long, thin, and triangular (
<figureCitation id="13012DB6FF90A908A4F6B0ABFBE7F940" box="[953,1057,1698,1725]" captionStart="FIGURE 15" captionStartId="31.[151,250,1975,1999]" captionTargetBox="[151,1436,453,1946]" captionTargetId="figure-133@31.[151,1436,453,1952]" captionTargetPageId="31" captionText="FIGURE 15. Pheidole flavodepressa, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0230800) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0243625)." figureDoi="http://doi.org/10.5281/zenodo.4405769" httpUri="https://zenodo.org/record/4405769/files/figure.png" pageId="30" pageNumber="31">Fig. 15C</figureCitation>
). Sculpture smooth. Pilosity very sparse, long, and erect (
<figureCitation id="13012DB6FF90A908A6D0B0CFFDE2F91C" box="[415,548,1734,1761]" captionStart="FIGURE 15" captionStartId="31.[151,250,1975,1999]" captionTargetBox="[151,1436,453,1946]" captionTargetId="figure-133@31.[151,1436,453,1952]" captionTargetPageId="31" captionText="FIGURE 15. Pheidole flavodepressa, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0230800) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0243625)." figureDoi="http://doi.org/10.5281/zenodo.4405769" httpUri="https://zenodo.org/record/4405769/files/figure.png" pageId="30" pageNumber="31">Fig. 15C, E</figureCitation>
).
<emphasis id="B94EED21FF90A908A578B0CEFD4AF91D" bold="true" box="[567,652,1734,1760]" pageId="30" pageNumber="31">Gaster.</emphasis>
With sparse and erect pilosity (
<figureCitation id="13012DB6FF90A908A4A3B0CFFBB7F91C" box="[1004,1137,1734,1761]" captionStart="FIGURE 15" captionStartId="31.[151,250,1975,1999]" captionTargetBox="[151,1436,453,1946]" captionTargetId="figure-133@31.[151,1436,453,1952]" captionTargetPageId="31" captionText="FIGURE 15. Pheidole flavodepressa, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0230800) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0243625)." figureDoi="http://doi.org/10.5281/zenodo.4405769" httpUri="https://zenodo.org/record/4405769/files/figure.png" pageId="30" pageNumber="31">Fig. 15C, E</figureCitation>
).
<emphasis id="B94EED21FF90A908A3CBB0CEFB0BF91D" bold="true" box="[1156,1229,1734,1760]" pageId="30" pageNumber="31">Color.</emphasis>
Yellow (
<figureCitation id="13012DB6FF90A908A27CB0CFFF6AF8F8" captionStart="FIGURE 15" captionStartId="31.[151,250,1975,1999]" captionTargetBox="[151,1436,453,1946]" captionTargetId="figure-133@31.[151,1436,453,1952]" captionTargetPageId="31" captionText="FIGURE 15. Pheidole flavodepressa, full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT0230800) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT0243625)." figureDoi="http://doi.org/10.5281/zenodo.4405769" httpUri="https://zenodo.org/record/4405769/files/figure.png" pageId="30" pageNumber="31">Fig. 15C, E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FF90A908A788B106FC73F8B0" pageId="30" pageNumber="31" type="biology_ecology">
<paragraph id="8B853133FF90A908A788B106FC73F8B0" blockId="30.[151,1437,151,2013]" pageId="30" pageNumber="31">
<emphasis id="B94EED21FF90A908A788B106FEE1F8D5" bold="true" box="[199,295,1806,1832]" pageId="30" pageNumber="31">Biology.</emphasis>
The species was collected between
<quantity id="4CC29CD6FF90A908A5F5B106FC80F8D4" box="[698,838,1806,1833]" metricMagnitude="2" metricUnit="m" metricValue="7.0" metricValueMax="11.75" metricValueMin="2.25" pageId="30" pageNumber="31" unit="m" value="700.0" valueMax="1175.0" valueMin="225.0">2251175 m</quantity>
elevation, in rainforest and montane rainforest. Nesting preferences are unknown. Workers were collected from sifted litter.
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FF90A909A788B15EFE6EFE98" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31" type="discussion">
<paragraph id="8B853133FF90A909A788B15EFE6EFE98" blockId="30.[151,1437,151,2013]" lastBlockId="31.[151,1437,151,429]" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
<emphasis id="B94EED21FF90A908A788B15EFE89F88D" bold="true" box="[199,335,1878,1904]" pageId="30" pageNumber="31">Comments.</emphasis>
<taxonomicName id="4C3A4AB0FF90A908A617B15FFD98F88C" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[344,606,1879,1905]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="flavodepressa">
<emphasis id="B94EED21FF90A908A617B15FFD98F88C" box="[344,606,1879,1905]" italics="true" pageId="30" pageNumber="31">Pheidole flavodepressa</emphasis>
</taxonomicName>
is known from two localities in
<collectingRegion id="49FEFFD1FF90A908A49CB15FFB9CF88C" box="[979,1114,1879,1905]" country="Madagascar" name="Antsiranana" pageId="30" pageNumber="31">Antsiranana</collectingRegion>
and is most similar to parapatric
<emphasis id="B94EED21FF90A908A7AFB173FE82F868" box="[224,324,1915,1941]" italics="true" pageId="30" pageNumber="31">
<taxonomicName id="4C3A4AB0FF90A908A7AFB173FE86F868" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[224,320,1915,1941]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="zirafy">P. zirafy</taxonomicName>
.
</emphasis>
Majors of
<taxonomicName id="4C3A4AB0FF90A908A689B173FDB8F868" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[454,638,1915,1941]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="flavodepressa">
<emphasis id="B94EED21FF90A908A689B173FDB8F868" box="[454,638,1915,1941]" italics="true" pageId="30" pageNumber="31">P. flavodepressa</emphasis>
</taxonomicName>
can be separated based on orange body coloration, side of head with moderately dense, long and subdecumbent to suberect pilosity, and occipital lobes with thick and dense rugae with distinctly rugulate interspaces between rugae. Majors of
<taxonomicName id="4C3A4AB0FF90A908A46CB1CBFC47F820" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[803,897,1987,2013]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="zirafy">
<emphasis id="B94EED21FF90A908A46CB1CBFC47F820" box="[803,897,1987,2013]" italics="true" pageId="30" pageNumber="31">P. zirafy</emphasis>
</taxonomicName>
have blackish brown body, moderately sparse, short and appressed to decumbent pilosity on side of head, and occipital lobes with rugae that are thin, weakening posteriorly, and archposterolaterally, with mostly smooth interspaces between rugae. Minors of
<taxonomicName id="4C3A4AB0FF91A909A3ABB6B3FA5AFF28" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[1252,1436,187,213]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="flavodepressa">
<emphasis id="B94EED21FF91A909A3ABB6B3FA5AFF28" box="[1252,1436,187,213]" italics="true" pageId="31" pageNumber="32">P. flavodepressa</emphasis>
</taxonomicName>
have yellow body coloration, shorter and thicker neck, and smooth mesosoma. In contrast,
<taxonomicName id="4C3A4AB0FF91A909A3C0B6D7FB2CFF04" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[1167,1258,223,249]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="zirafy">
<emphasis id="B94EED21FF91A909A3C0B6D7FB2CFF04" box="[1167,1258,223,249]" italics="true" pageId="31" pageNumber="32">P. zirafy</emphasis>
</taxonomicName>
has dark brown minors with longer and thinner neck and mostly sculptured mesosoma. Minor workers of
<taxonomicName id="4C3A4AB0FF91A909A332B70BFAF4FEE0" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[1149,1330,259,285]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="flavodepressa">
<emphasis id="B94EED21FF91A909A332B70BFAF4FEE0" box="[1149,1330,259,285]" italics="true" pageId="31" pageNumber="32">P. flavodepressa</emphasis>
</taxonomicName>
may also be confused with those of
<emphasis id="B94EED21FF91A909A6F1B72FFD79FEBC" box="[446,703,295,321]" italics="true" pageId="31" pageNumber="32">
<taxonomicName id="4C3A4AB0FF91A909A6F1B72FFD89FEBC" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[446,591,295,321]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="antsahabe">P. antsahabe</taxonomicName>
,
<taxonomicName id="4C3A4AB0FF91A909A515B72FFD79FEBC" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[602,703,295,321]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="similis">P. similis</taxonomicName>
</emphasis>
,
<taxonomicName id="4C3A4AB0FF91A909A582B72FFC4CFEBC" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[717,906,295,321]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="mantadioflava">
<emphasis id="B94EED21FF91A909A582B72FFC4CFEBC" box="[717,906,295,321]" italics="true" pageId="31" pageNumber="32">P. mantadioflava</emphasis>
</taxonomicName>
, and
<emphasis id="B94EED21FF91A909A488B72FFBE2FEBC" box="[967,1060,295,321]" italics="true" pageId="31" pageNumber="32">
<taxonomicName id="4C3A4AB0FF91A909A488B72FFBE7FEBC" authorityName="Salata &amp; Fisher" authorityYear="2020" box="[967,1057,295,321]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="clara">P. clara</taxonomicName>
,
</emphasis>
but differ from them in distinctly longer and thinner neck.
</paragraph>
</subSubSection>
<subSubSection id="C32062B8FF91A909A788B767FE14FE50" pageId="31" pageNumber="32" type="etymology">
<paragraph id="8B853133FF91A909A788B767FE14FE50" blockId="31.[151,1437,151,429]" pageId="31" pageNumber="32">
<emphasis id="B94EED21FF91A909A788B767FE88FE74" bold="true" box="[199,334,367,393]" pageId="31" pageNumber="32">Etymology.</emphasis>
Latin for yellow and deep, in reference to the yellow body coloration and deep posteromedian concavity in major workers.
</paragraph>
</subSubSection>
</treatment>
</document>