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<document id="208A974AD9B24148BBD7E9126A114B31" ID-CLB-Dataset="3967" ID-DOI="10.11646/zootaxa.5054.1.1" ID-GBIF-Dataset="c7ac2c61-e8c5-483c-83ae-8af28635f4ac" ID-ISSN="1175-5326" ID-Zenodo-Dep="5577421" ID-ZooBank="75417CFA-BF40-4B6F-A317-DEBFB447893D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1634627545673" checkinUser="plazi" docAuthor="Kluge, Nikita J." docDate="2021" docId="03936656F610E0D37AB0FCFFFEAAFD9A" docLanguage="en" docName="zootaxa.5054.1.1.pdf" docOrigin="Zootaxa 5054 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Centroptella (Chopralla) papuanica Kluge 2021, sp. n." docType="treatment" docVersion="13" lastPageNumber="130" masterDocId="FFAA1E2EF66BE0527A27FF80FFFBFFCD" masterDocTitle="Review of Centroptella Braasch &amp; Soldán 1980 (Ephemeroptera, Baetidae)" masterLastPageNumber="144" masterPageNumber="1" pageNumber="124" updateTime="1699460234914" updateUser="ExternalLinkService">
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<mods:title id="BC5313BBA7C45DD23099B32032F3C696">Review of Centroptella Braasch &amp; Soldán 1980 (Ephemeroptera, Baetidae)</mods:title>
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<mods:namePart id="B73A56ABA691D29D4D11226999702C7E">Kluge, Nikita J.</mods:namePart>
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<mods:date id="9B5FC24C2DD6378DB20CADA2A5610A4D">2021</mods:date>
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<mods:number id="FE439D00A94ACA363EF36E76D7479C6C">2021-10-19</mods:number>
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<mods:number id="6206115F2F6733ED71889819A873B0B3">5054</mods:number>
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<mods:classification id="B3885C7594EFC4EE3DD723D47B7ED176">journal article</mods:classification>
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<mods:identifier id="7ACD128FB06DA2205DD8460087F7EDFA" type="DOI">10.11646/zootaxa.5054.1.1</mods:identifier>
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<treatment id="03936656F610E0D37AB0FCFFFEAAFD9A" ID-DOI="http://doi.org/10.5281/zenodo.5781535" ID-GBIF-Taxon="189136132" ID-Zenodo-Dep="5781535" LSID="urn:lsid:plazi:treatment:03936656F610E0D37AB0FCFFFEAAFD9A" httpUri="http://treatment.plazi.org/id/03936656F610E0D37AB0FCFFFEAAFD9A" lastPageId="129" lastPageNumber="130" pageId="123" pageNumber="124">
<subSubSection id="C32084CBF610E0297AB0FCFFFD3DFC57" box="[151,710,895,922]" pageId="123" pageNumber="124" type="nomenclature">
<paragraph id="8B85D740F610E0297AB0FCFFFD3DFC57" blockId="123.[151,710,895,958]" box="[151,710,895,922]" pageId="123" pageNumber="124">
<emphasis id="B94E0B52F610E0297AB0FCFFFD3DFC57" bold="true" box="[151,710,895,922]" pageId="123" pageNumber="124">
3.4.
<taxonomicName id="4C3AACC3F610E0297AECFCFFFD87FC54" authority="Kluge, 2021" authorityName="Kluge" authorityYear="2021" box="[203,636,895,922]" class="Insecta" family="Baetidae" genus="Centroptella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="papuanica" status="sp. nov." subGenus="Chopralla">
<emphasis id="B94E0B52F610E0297AECFCFFFE9AFC57" bold="true" box="[203,353,895,922]" italics="true" pageId="123" pageNumber="124">Centroptella</emphasis>
(
<emphasis id="B94E0B52F610E0297B55FCFFFE17FC57" bold="true" box="[370,492,895,922]" italics="true" pageId="123" pageNumber="124">Chopralla</emphasis>
)
<emphasis id="B94E0B52F610E0297BDAFCFFFD87FC54" bold="true" box="[509,636,895,921]" italics="true" pageId="123" pageNumber="124">papuanica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27DB629F610E02978A4FC00FD3DFC57" box="[643,710,896,922]" pageId="123" pageNumber="124" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C32084CBF610E0297AB0FC23FEB8FC73" box="[151,323,931,958]" pageId="123" pageNumber="124" type="description">
<paragraph id="8B85D740F610E0297AB0FC23FEB8FC73" blockId="123.[151,710,895,958]" box="[151,323,931,958]" pageId="123" pageNumber="124">
(
<figureCitation id="1301CBC5F610E0297AB8FC23FEC0FC73" box="[159,315,931,958]" captionStart-0="FIGURES 584–597" captionStart-1="FIGURES 598–606" captionStart-2="FIGURES 607–610" captionStart-3="FIGURES 611–617" captionStart-4="FIGURES 618–623" captionStart-5="FIGURES 624–625" captionStartId-0="122.[151,264,1851,1875]" captionStartId-1="124.[151,264,1899,1923]" captionStartId-2="125.[151,264,1430,1454]" captionStartId-3="126.[151,264,1830,1854]" captionStartId-4="127.[151,264,1691,1715]" captionStartId-5="128.[151,264,1809,1833]" captionTargetBox-0="[152,1425,193,1810]" captionTargetBox-1="[164,1423,273,1876]" captionTargetBox-2="[177,1421,191,1392]" captionTargetBox-3="[151,1436,181,1806]" captionTargetBox-4="[151,1436,181,1663]" captionTargetBox-5="[151,1435,1101,1786]" captionTargetId-0="figure-27@122.[151,1436,181,1827]" captionTargetId-1="figure-72@124.[164,1423,273,1876]" captionTargetId-2="figure-20@125.[164,1423,181,1406]" captionTargetId-3="figure-27@126.[151,1436,181,1806]" captionTargetId-4="figure-20@127.[151,1436,181,1667]" captionTargetId-5="figure-433@128.[151,1436,1101,1786]" captionTargetPageId-0="122" captionTargetPageId-1="124" captionTargetPageId-2="125" captionTargetPageId-3="126" captionTargetPageId-4="127" captionTargetPageId-5="128" captionText-0="FIGURES 584–597. Tergalii I–VII. 584–590, Centroptella (Chopralla) rufostriata sp. n.; 591–597, Centroptella (Chopralla) papuanica sp. n." captionText-1="FIGURES 598–606. Centroptella (Chopralla) papuanica sp. n., larval exuviae (holotype). 598, abdomen; 599, pronotum and mesonotum; 600, area of pronotum close to median suture with bifurcate seta corresponding to a on Fig. 14; 601, scale on metanotum with same magnification; 602, abdominal sternum VI and enlarged scales; 603, abdominal sternum IX and enlarged scales; 604, labrum; 605, labium; 606, abdominal terga VIII–IX and enlarged scales." captionText-2="FIGURES 607–610. Centroptella (Chopralla) papuanica sp. n. 607–608, left and right mandibles; 609, subimaginal gonostyli crumpled under larval cuticle; 610, larval abdominal segment X without caudalii (holotype). Abbreviations: gs1–gs3, segments of gonostyli; m.gs, gonostylar muscle; m.sg, styligeral muscle; pgs, larval protogonostylus; usg, unistyliger." captionText-3="FIGURES 611–617. Centroptella (Chopralla) papuanica sp. n., larval exuviae. 611–613, fore, middle and hind legs, with focus on anterior side; 614–616, their tibiae, with focus on posterior side (arrows show patella-tibial suture); 617, cercus." captionText-4="FIGURES 618–623. Centroptella (Chopralla) papuanica sp. n. 618, fore leg of female imago; 619, abdomen of female imago; 620, female imago; 621, subimaginal exuviae of mesopleuron; 622, subimaginal exuviae of mesonotum; 623, abdomen of male subimago extracted from larva (618, 619, 621, holotype)." captionText-5="FIGURES 624–625. Centroptella (Chopralla) papuanica sp. n., egg (extracted from holotype)." figureDoi-0="http://doi.org/10.5281/zenodo.5577620" figureDoi-1="http://doi.org/10.5281/zenodo.5577622" figureDoi-2="http://doi.org/10.5281/zenodo.5577624" figureDoi-3="http://doi.org/10.5281/zenodo.5577626" figureDoi-4="http://doi.org/10.5281/zenodo.5577628" figureDoi-5="http://doi.org/10.5281/zenodo.5577630" httpUri-0="https://zenodo.org/record/5577620/files/figure.png" httpUri-1="https://zenodo.org/record/5577622/files/figure.png" httpUri-2="https://zenodo.org/record/5577624/files/figure.png" httpUri-3="https://zenodo.org/record/5577626/files/figure.png" httpUri-4="https://zenodo.org/record/5577628/files/figure.png" httpUri-5="https://zenodo.org/record/5577630/files/figure.png" pageId="123" pageNumber="124">Figs 591–625</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32084CBF610E0297AB0FC6BFE9FFBBC" pageId="123" pageNumber="124" type="materials_examined">
<paragraph id="8B85D740F610E0297AB0FC6BFEB7FB80" blockId="123.[151,1437,1003,2038]" pageId="123" pageNumber="124">
<emphasis id="B94E0B52F610E0297AB0FC6BFE85FBC8" bold="true" box="[151,382,1003,1029]" pageId="123" pageNumber="124">Material examined.</emphasis>
<materialsCitation id="3B52DD1DF610E0297BA5FC6CFE2EFBE4" ID-GBIF-Occurrence="3390856387" collectingDate="2012-08-13" collectorName="N. Kluge &amp; L. Sheyko." country="Indonesia" location="Waena" municipality="Jayapura" pageId="123" pageNumber="124" specimenCount="1" specimenCount-female="1" stateProvince="Papua Barat" typeStatus="holotype">
<emphasis id="B94E0B52F610E0297BA5FC6CFE03FBCB" bold="true" box="[386,504,1004,1030]" pageId="123" pageNumber="124">
<typeStatus id="548169E2F610E0297BA5FC6CFE08FBCB" box="[386,499,1004,1030]" pageId="123" pageNumber="124" type="holotype">Holotype</typeStatus>
:
</emphasis>
L-S-I
<specimenCount id="9D3C1CC9F610E0297867FC6CFDB5FBCB" box="[576,590,1004,1030]" pageId="123" pageNumber="124" type="female">♀</specimenCount>
{specimen [IV](4)B}:
<collectingCountry id="F32D97D0F610E029796CFC6CFC1DFBCB" box="[843,998,1004,1030]" name="Indonesia" pageId="123" pageNumber="124">INDONESIA</collectingCountry>
,
<collectingRegion id="49FE19A2F610E02979C9FC6CFBCEFBCB" box="[1006,1077,1004,1030]" country="Indonesia" name="Papua Barat" pageId="123" pageNumber="124">Papua</collectingRegion>
,
<collectingMunicipality id="6BE14D3AF610E0297E19FC6CFB5FFBCB" box="[1086,1188,1004,1030]" pageId="123" pageNumber="124">Jayapura</collectingMunicipality>
,
<location id="8EE5819BF610E0297E8BFC6CFB01FBCB" LSID="urn:lsid:plazi:treatment:03936656F610E0D37AB0FCFFFEAAFD9A:8EE5819BF610E0297E8BFC6CFB01FBCB" box="[1196,1274,1004,1030]" country="Indonesia" municipality="Jayapura" name="Waena" pageId="123" pageNumber="124" stateProvince="Papua Barat">Waena</location>
,
<date id="FF84F180F610E0297F24FC6CFA63FBCB" box="[1283,1432,1004,1030]" pageId="123" pageNumber="124" value="2012-08-13">
<collectingDate id="EFC00868F610E0297F24FC6CFA63FBCB" box="[1283,1432,1004,1030]" pageId="123" pageNumber="124" value="2012-08-13">13.VIII.2012</collectingDate>
</date>
, coll.
<collectorName id="26CFB296F610E0297AE8FB90FECDFBE4" box="[207,310,1039,1066]" pageId="123" pageNumber="124">N. Kluge</collectorName>
&amp;
<collectorName id="26CFB296F610E0297B7DFB90FE2EFBE4" box="[346,469,1039,1066]" pageId="123" pageNumber="124">L. Sheyko.</collectorName>
</materialsCitation>
<materialsCitation id="3B52DD1DF610E0297BFCFB90FA6CFBE7" ID-GBIF-Occurrence="3390856314" box="[475,1431,1039,1066]" collectingDate="2012-08-08" collectingDateMax="2012-08-13" collectingDateMin="2012-08-08" collectorName="N. Kluge &amp; L. Sheyko" country="Indonesia" location="Waena" municipality="Jayapura" pageId="123" pageNumber="124" specimenCount="1" specimenCount-female="1" stateProvince="Papua Barat" typeStatus="paratype">
<emphasis id="B94E0B52F610E0297BFCFB90FDA7FBE7" bold="true" box="[475,604,1040,1066]" pageId="123" pageNumber="124">
<typeStatus id="548169E2F610E0297BFCFB90FDACFBE7" box="[475,599,1040,1066]" pageId="123" pageNumber="124" type="paratype">Paratypes</typeStatus>
:
</emphasis>
the same locality,
<date id="FF84F180F610E029790AFB90FC25FBE7" box="[813,990,1040,1066]" pageId="123" pageNumber="124" value="2012-08-08" valueMax="2012-08-13" valueMin="2012-08-08">
<collectingDate id="EFC00868F610E029790AFB90FC25FBE7" box="[813,990,1040,1066]" pageId="123" pageNumber="124" value="2012-08-08" valueMax="2012-08-13" valueMin="2012-08-08">8–13.VIII.2012</collectingDate>
</date>
, coll.
<collectorName id="26CFB296F610E0297E05FB90FB73FBE4" box="[1058,1160,1039,1066]" pageId="123" pageNumber="124">N. Kluge</collectorName>
&amp;
<collectorName id="26CFB296F610E0297E8BFB90FADFFBE4" box="[1196,1316,1039,1066]" pageId="123" pageNumber="124">L. Sheyko</collectorName>
: 1 L-S-I
<specimenCount id="9D3C1CC9F610E0297FAEFB90FA6CFBE7" box="[1417,1431,1040,1066]" pageId="123" pageNumber="124" type="female">♀</specimenCount>
</materialsCitation>
, 2 L/S
<specimenCount id="9D3C1CC9F610E0297AFFFBB4FF17FB83" box="[216,236,1076,1102]" pageId="123" pageNumber="124" type="male">♂</specimenCount>
,
<specimenCount id="9D3C1CC9F610E0297ADFFBB4FEB2FB80" box="[248,329,1075,1102]" pageId="123" pageNumber="124" type="larva">1 larva</specimenCount>
.
</paragraph>
<paragraph id="8B85D740F610E0297AE0FBD7FE9FFBBC" blockId="123.[151,1437,1003,2038]" box="[199,356,1111,1137]" pageId="123" pageNumber="124">
<emphasis id="B94E0B52F610E0297AE0FBD7FE9FFBBC" bold="true" box="[199,356,1111,1137]" pageId="123" pageNumber="124">Descriptions.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C32084CBF610E0D27AE0FBFCFC7DFED3" lastPageId="128" lastPageNumber="129" pageId="123" pageNumber="124" type="description">
<paragraph id="8B85D740F610E0297AE0FBFCFDA4F98B" blockId="123.[151,1437,1003,2038]" pageId="123" pageNumber="124">
<emphasis id="B94E0B52F610E0297AE0FBFCFEE8FB5B" bold="true" box="[199,275,1148,1174]" pageId="123" pageNumber="124">
<emphasis id="B94E0B52F610E0297AE0FBFCFEF0FB5B" bold="true" box="[199,267,1148,1174]" italics="true" pageId="123" pageNumber="124">Larva</emphasis>
.
</emphasis>
CUTICULAR COLORATION: Head with anterior side mostly colorless; genae brown; dorsal side of head brown. Pronotum and mesonotum with diffusive darker and lighter brown areas (
<figureCitation id="1301CBC5F610E0297E55FB1FFB2DFB77" box="[1138,1238,1183,1210]" captionStart="FIGURES 598–606" captionStartId="124.[151,264,1899,1923]" captionTargetBox="[164,1423,273,1876]" captionTargetId="figure-72@124.[164,1423,273,1876]" captionTargetPageId="124" captionText="FIGURES 598–606. Centroptella (Chopralla) papuanica sp. n., larval exuviae (holotype). 598, abdomen; 599, pronotum and mesonotum; 600, area of pronotum close to median suture with bifurcate seta corresponding to a on Fig. 14; 601, scale on metanotum with same magnification; 602, abdominal sternum VI and enlarged scales; 603, abdominal sternum IX and enlarged scales; 604, labrum; 605, labium; 606, abdominal terga VIII–IX and enlarged scales." figureDoi="http://doi.org/10.5281/zenodo.5577622" httpUri="https://zenodo.org/record/5577622/files/figure.png" pageId="123" pageNumber="124">Fig. 599</figureCitation>
). Thoracic pleura and metanotum brown with lighter areas; sterna colorless. Legs with coxae partly colorless, partly brown; trochanteres colorless. All femora on outer side mostly colorless, with diffusive longitudinal darkening at midlength nearer outer margin; fore and middle femora with outer side brown and with large brown macula on posterior side; hind femur with outer side colorless, with brown longitudinal macula on posterior side near inner margin; tibia of each leg diffusively darkened with brown along rows of long, fine setae; tarsus of each leg brown proximally and along rows of long, fine setae; claws brown (
<figureCitation id="1301CBC5F610E029787CFAF7FD07FA5F" box="[603,764,1399,1426]" captionStart="FIGURES 611–617" captionStartId="126.[151,264,1830,1854]" captionTargetBox="[151,1436,181,1806]" captionTargetId="figure-27@126.[151,1436,181,1806]" captionTargetPageId="126" captionText="FIGURES 611–617. Centroptella (Chopralla) papuanica sp. n., larval exuviae. 611–613, fore, middle and hind legs, with focus on anterior side; 614–616, their tibiae, with focus on posterior side (arrows show patella-tibial suture); 617, cercus." figureDoi="http://doi.org/10.5281/zenodo.5577626" httpUri="https://zenodo.org/record/5577626/files/figure.png" pageId="123" pageNumber="124">Figs 611–613</figureCitation>
). Abdominal terga with diffusive darker and lighter brown areas; each tergum II–III and V–VI with pair of transverse blanks located posteriad-laterad of medioposterior sigilla and with smaller, more anterior pair of transverse blanks located laterad of medioanterior sigilla; on terga II and IV blanks occupy most part of tergum; terga VII–VIII mostly light, with posterior and/or anterior margins bordered by darker brown; terga IX–X mostly brown; medio-anterior and medioposterior sigilla lighter than background (
<figureCitation id="1301CBC5F610E0297F48F987FF3EF98B" captionStart="FIGURES 598–606" captionStartId="124.[151,264,1899,1923]" captionTargetBox="[164,1423,273,1876]" captionTargetId="figure-72@124.[164,1423,273,1876]" captionTargetPageId="124" captionText="FIGURES 598–606. Centroptella (Chopralla) papuanica sp. n., larval exuviae (holotype). 598, abdomen; 599, pronotum and mesonotum; 600, area of pronotum close to median suture with bifurcate seta corresponding to a on Fig. 14; 601, scale on metanotum with same magnification; 602, abdominal sternum VI and enlarged scales; 603, abdominal sternum IX and enlarged scales; 604, labrum; 605, labium; 606, abdominal terga VIII–IX and enlarged scales." figureDoi="http://doi.org/10.5281/zenodo.5577622" httpUri="https://zenodo.org/record/5577622/files/figure.png" pageId="123" pageNumber="124">Fig. 598</figureCitation>
). Caudalii light brownish (
<figureCitation id="1301CBC5F610E0297BD7F9ABFDA8F98B" box="[496,595,1579,1606]" captionStart="FIGURES 611–617" captionStartId="126.[151,264,1830,1854]" captionTargetBox="[151,1436,181,1806]" captionTargetId="figure-27@126.[151,1436,181,1806]" captionTargetPageId="126" captionText="FIGURES 611–617. Centroptella (Chopralla) papuanica sp. n., larval exuviae. 611–613, fore, middle and hind legs, with focus on anterior side; 614–616, their tibiae, with focus on posterior side (arrows show patella-tibial suture); 617, cercus." figureDoi="http://doi.org/10.5281/zenodo.5577626" httpUri="https://zenodo.org/record/5577626/files/figure.png" pageId="123" pageNumber="124">Fig. 617</figureCitation>
).
</paragraph>
<paragraph id="8B85D740F610E0297AE0F9D0FD78F943" blockId="123.[151,1437,1003,2038]" pageId="123" pageNumber="124">
HYPODERMAL COLORATION: Reddish-brown stripe on posterior margin of metanotum and each abdominal tergum (as in winged stages—
<figureCitation id="1301CBC5F610E0297830F9F3FD8DF943" box="[535,630,1651,1678]" captionStart="FIGURES 618–623" captionStartId="127.[151,264,1691,1715]" captionTargetBox="[151,1436,181,1663]" captionTargetId="figure-20@127.[151,1436,181,1667]" captionTargetPageId="127" captionText="FIGURES 618–623. Centroptella (Chopralla) papuanica sp. n. 618, fore leg of female imago; 619, abdomen of female imago; 620, female imago; 621, subimaginal exuviae of mesopleuron; 622, subimaginal exuviae of mesonotum; 623, abdomen of male subimago extracted from larva (618, 619, 621, holotype)." figureDoi="http://doi.org/10.5281/zenodo.5577628" httpUri="https://zenodo.org/record/5577628/files/figure.png" pageId="123" pageNumber="124">Fig. 620</figureCitation>
).
</paragraph>
<paragraph id="8B85D740F610E0297AE0F918FBEAF847" blockId="123.[151,1437,1003,2038]" pageId="123" pageNumber="124">
SHAPE: Labrum either equally wide at base and middle, or slightly wider at base (
<figureCitation id="1301CBC5F610E0297E7EF917FB41F97F" box="[1113,1210,1687,1714]" captionStart="FIGURES 598–606" captionStartId="124.[151,264,1899,1923]" captionTargetBox="[164,1423,273,1876]" captionTargetId="figure-72@124.[164,1423,273,1876]" captionTargetPageId="124" captionText="FIGURES 598–606. Centroptella (Chopralla) papuanica sp. n., larval exuviae (holotype). 598, abdomen; 599, pronotum and mesonotum; 600, area of pronotum close to median suture with bifurcate seta corresponding to a on Fig. 14; 601, scale on metanotum with same magnification; 602, abdominal sternum VI and enlarged scales; 603, abdominal sternum IX and enlarged scales; 604, labrum; 605, labium; 606, abdominal terga VIII–IX and enlarged scales." figureDoi="http://doi.org/10.5281/zenodo.5577622" httpUri="https://zenodo.org/record/5577622/files/figure.png" pageId="123" pageNumber="124">Fig. 604</figureCitation>
). Mandibles: incisor of left mandible terminated with 4 denticles decreasing from most distal to most proximal; incisor of right mandible terminated with 3 denticles of subequal width, without 4th denticle; left and right incisors with outer-ventral denticle; prostheca of left mandible with 3 blunt processes and 3 pointed processes; prostheca of right mandible directed medially-distally, pressed to kinetodontium, curved and dentate apically, without long branch; median margin of right mandible proximad of prostheca with or without short seta-like processes (
<figureCitation id="1301CBC5F610E0297E1CF8CBFB25F8AB" box="[1083,1246,1867,1894]" captionStart="FIGURES 607–610" captionStartId="125.[151,264,1430,1454]" captionTargetBox="[177,1421,191,1392]" captionTargetId="figure-20@125.[164,1423,181,1406]" captionTargetPageId="125" captionText="FIGURES 607–610. Centroptella (Chopralla) papuanica sp. n. 607–608, left and right mandibles; 609, subimaginal gonostyli crumpled under larval cuticle; 610, larval abdominal segment X without caudalii (holotype). Abbreviations: gs1–gs3, segments of gonostyli; m.gs, gonostylar muscle; m.sg, styligeral muscle; pgs, larval protogonostylus; usg, unistyliger." figureDoi="http://doi.org/10.5281/zenodo.5577624" httpUri="https://zenodo.org/record/5577624/files/figure.png" pageId="123" pageNumber="124">Figs 607–608</figureCitation>
). Maxillary palp nearly as long as lacinia, 2-segmented, with 2nd much longer than 1st segment.
</paragraph>
<paragraph id="8B85D740F610E02E7AE0F814FCD5FF34" blockId="123.[151,1437,1003,2038]" lastBlockId="124.[151,1436,151,249]" lastPageId="124" lastPageNumber="125" pageId="123" pageNumber="124">
Pronotum without protuberances (in contrast to
<taxonomicName id="4C3AACC3F610E02978CFF814FC92F860" authorityName="Kluge, Godunko &amp; Svitok" authorityYear="2020" box="[744,873,1939,1966]" class="Insecta" family="Baetidae" genus="Centroptella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="ingridae">
<emphasis id="B94E0B52F610E02978CFF814FC92F860" box="[744,873,1939,1966]" italics="true" pageId="123" pageNumber="124">C. ingridae</emphasis>
</taxonomicName>
). Metanotum with vestiges of hind protoptera (in all 6 individuals examined). Forelegs longest, hind legs shortest; on fore leg tarsus (measured on outer side) as long as tibia; on middle and hind legs tarsus slightly shorter than tibia (
<figureCitation id="1301CBC5F610E02979A0F85BFBDDF83B" box="[903,1062,2011,2038]" captionStart="FIGURES 611–617" captionStartId="126.[151,264,1830,1854]" captionTargetBox="[151,1436,181,1806]" captionTargetId="figure-27@126.[151,1436,181,1806]" captionTargetPageId="126" captionText="FIGURES 611–617. Centroptella (Chopralla) papuanica sp. n., larval exuviae. 611–613, fore, middle and hind legs, with focus on anterior side; 614–616, their tibiae, with focus on posterior side (arrows show patella-tibial suture); 617, cercus." figureDoi="http://doi.org/10.5281/zenodo.5577626" httpUri="https://zenodo.org/record/5577626/files/figure.png" pageId="123" pageNumber="124">Figs 611–613</figureCitation>
). All femora parallel-sided, outer margin straight or slightly concave, inner margin slightly convex. Foreleg without patella-tibial suture, middle and hind legs with patella-tibial suture greatly shifted to apex of tibia (
<figureCitation id="1301CBC5F617E02E79A4FF3BFBDBFF1B" box="[899,1056,187,214]" captionStart="FIGURES 611–617" captionStartId="126.[151,264,1830,1854]" captionTargetBox="[151,1436,181,1806]" captionTargetId="figure-27@126.[151,1436,181,1806]" captionTargetPageId="126" captionText="FIGURES 611–617. Centroptella (Chopralla) papuanica sp. n., larval exuviae. 611–613, fore, middle and hind legs, with focus on anterior side; 614–616, their tibiae, with focus on posterior side (arrows show patella-tibial suture); 617, cercus." figureDoi="http://doi.org/10.5281/zenodo.5577626" httpUri="https://zenodo.org/record/5577626/files/figure.png" pageId="124" pageNumber="125">Figs 614–616</figureCitation>
) [see
<taxonomicName id="4C3AACC3F617E02E7E43FF3BFB0AFF18" authorityName=": Kluge, Godunko &amp; Svitok" authorityYear="2020" box="[1124,1265,187,213]" class="Insecta" family="Baetidae" genus="Centroptella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="B94E0B52F617E02E7E43FF3BFB0AFF18" box="[1124,1265,187,213]" italics="true" pageId="124" pageNumber="125">Centroptella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27DB629F617E02E7EDEFF3BFADAFF18" box="[1273,1313,187,213]" pageId="124" pageNumber="125" sensu="lato">s. l.</taxonomicNameLabel>
(2)]. Claw with two rows of 1–4 denticles in each [see
<taxonomicName id="4C3AACC3F617E02E78A4FF5FFD0DFF34" authorityName="Waltz &amp; McCafferty" authorityYear="1987" box="[643,758,223,249]" class="Insecta" family="Baetidae" genus="Chopralla" kingdom="Animalia" order="Ephemeroptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="genus">
<emphasis id="B94E0B52F617E02E78A4FF5FFD0DFF34" box="[643,758,223,249]" italics="true" pageId="124" pageNumber="125">Chopralla</emphasis>
</taxonomicName>
(2)].
</paragraph>
<caption id="DF4587C8F617E02E7AB0F8EBFC0BF822" ID-DOI="http://doi.org/10.5281/zenodo.5577622" ID-Zenodo-Dep="5577622" httpUri="https://zenodo.org/record/5577622/files/figure.png" pageId="124" pageNumber="125" startId="124.[151,264,1899,1923]" targetBox="[164,1423,273,1876]" targetPageId="124">
<paragraph id="8B85D740F617E02E7AB0F8EBFC0BF822" blockId="124.[151,1437,1899,2032]" pageId="124" pageNumber="125">
<emphasis id="B94E0B52F617E02E7AB0F8EBFE8BF84E" bold="true" box="[151,368,1899,1923]" pageId="124" pageNumber="125">FIGURES 598–606.</emphasis>
<taxonomicName id="4C3AACC3F617E02E7B5CF8EBFD09F84E" authority="Kluge, 2021" authorityName="Kluge" authorityYear="2021" box="[379,754,1899,1924]" class="Insecta" family="Baetidae" genus="Centroptella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="124" pageNumber="125" phylum="Arthropoda" rank="species" species="papuanica" status="sp. nov." subGenus="Chopralla">
<emphasis id="B94E0B52F617E02E7B5CF8EBFE01F84E" box="[379,506,1899,1923]" italics="true" pageId="124" pageNumber="125">Centroptella</emphasis>
(
<emphasis id="B94E0B52F617E02E782BF8EBFD8FF84E" box="[524,628,1899,1923]" italics="true" pageId="124" pageNumber="125">Chopralla</emphasis>
)
<emphasis id="B94E0B52F617E02E78A0F8ECFD09F84E" box="[647,754,1900,1923]" italics="true" pageId="124" pageNumber="125">papuanica</emphasis>
</taxonomicName>
<emphasis id="B94E0B52F617E02E78DBF8ECFCC2F849" bold="true" box="[764,825,1900,1924]" pageId="124" pageNumber="125">
<taxonomicNameLabel id="A27DB629F617E02E78DBF8ECFCC2F849" box="[764,825,1900,1924]" pageId="124" pageNumber="125" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, larval exuviae (holotype). 598, abdomen; 599, pronotum and mesonotum; 600, area of pronotum close to median suture with bifurcate seta corresponding to
<emphasis id="B94E0B52F617E02E7EA9F810FB60F865" bold="true" box="[1166,1179,1936,1960]" pageId="124" pageNumber="125">a</emphasis>
on Fig. 14; 601, scale on metanotum with same magnification; 602, abdominal sternum VI and enlarged scales; 603, abdominal sternum IX and enlarged scales; 604, labrum; 605, labium; 606, abdominal terga VIII–IX and enlarged scales.
</paragraph>
</caption>
<caption id="DF4587C8F616E02F7AB0FA16FB27FA3B" ID-DOI="http://doi.org/10.5281/zenodo.5577624" ID-Zenodo-Dep="5577624" httpUri="https://zenodo.org/record/5577624/files/figure.png" pageId="125" pageNumber="126" startId="125.[151,264,1430,1454]" targetBox="[177,1421,191,1392]" targetPageId="125">
<paragraph id="8B85D740F616E02F7AB0FA16FB27FA3B" blockId="125.[151,1436,1430,1527]" pageId="125" pageNumber="126">
<emphasis id="B94E0B52F616E02F7AB0FA16FE97FA63" bold="true" box="[151,364,1430,1454]" pageId="125" pageNumber="126">FIGURES 607–610.</emphasis>
<taxonomicName id="4C3AACC3F616E02F7B54FA16FD19FA63" authority="Kluge, 2021" authorityName="Kluge" authorityYear="2021" box="[371,738,1430,1454]" class="Insecta" family="Baetidae" genus="Centroptella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="papuanica" status="sp. nov." subGenus="Chopralla">
<emphasis id="B94E0B52F616E02F7B54FA16FE09FA63" box="[371,498,1430,1454]" italics="true" pageId="125" pageNumber="126">Centroptella</emphasis>
(
<emphasis id="B94E0B52F616E02F7826FA16FD92FA63" box="[513,617,1430,1454]" italics="true" pageId="125" pageNumber="126">Chopralla</emphasis>
)
<emphasis id="B94E0B52F616E02F7850FA17FD19FA63" box="[631,738,1431,1454]" italics="true" pageId="125" pageNumber="126">papuanica</emphasis>
</taxonomicName>
<emphasis id="B94E0B52F616E02F78CFFA16FCDAFA63" bold="true" box="[744,801,1430,1454]" pageId="125" pageNumber="126">
<taxonomicNameLabel id="A27DB629F616E02F78CFFA16FCDAFA63" box="[744,801,1430,1454]" pageId="125" pageNumber="126" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
607–608, left and right mandibles; 609, subimaginal gonostyli crumpled under larval cuticle; 610, larval abdominal segment X without caudalii (holotype). Abbreviations: gs1–gs3, segments of gonostyli; m.gs, gonostylar muscle; m.sg, styligeral muscle; pgs, larval protogonostylus; usg, unistyliger.
</paragraph>
</caption>
<paragraph id="8B85D740F616E02F7AE0F9A4FB10F8DA" blockId="125.[151,1437,1572,2031]" pageId="125" pageNumber="126">
Denticles on posterior margins of abdominal terga and sterna flat (not conic). Terga I–IV without denticles; terga V–IX with long triangular denticles; on tergum IX row of denticles interrupted medially, behind pair of submedian setae (
<figureCitation id="1301CBC5F616E02F7B32F9ECFE83F94A" box="[277,376,1644,1671]" captionStart="FIGURES 598–606" captionStartId="124.[151,264,1899,1923]" captionTargetBox="[164,1423,273,1876]" captionTargetId="figure-72@124.[164,1423,273,1876]" captionTargetPageId="124" captionText="FIGURES 598–606. Centroptella (Chopralla) papuanica sp. n., larval exuviae (holotype). 598, abdomen; 599, pronotum and mesonotum; 600, area of pronotum close to median suture with bifurcate seta corresponding to a on Fig. 14; 601, scale on metanotum with same magnification; 602, abdominal sternum VI and enlarged scales; 603, abdominal sternum IX and enlarged scales; 604, labrum; 605, labium; 606, abdominal terga VIII–IX and enlarged scales." figureDoi="http://doi.org/10.5281/zenodo.5577622" httpUri="https://zenodo.org/record/5577622/files/figure.png" pageId="125" pageNumber="126">Fig. 606</figureCitation>
). Posterior projection of tergum X with small, irregular denticles on median part, laterally with paired groups of 2–5 long denticles (
<figureCitation id="1301CBC5F616E02F7811F910FD62F966" box="[566,665,1680,1707]" captionStart="FIGURES 607–610" captionStartId="125.[151,264,1430,1454]" captionTargetBox="[177,1421,191,1392]" captionTargetId="figure-20@125.[164,1423,181,1406]" captionTargetPageId="125" captionText="FIGURES 607–610. Centroptella (Chopralla) papuanica sp. n. 607–608, left and right mandibles; 609, subimaginal gonostyli crumpled under larval cuticle; 610, larval abdominal segment X without caudalii (holotype). Abbreviations: gs1–gs3, segments of gonostyli; m.gs, gonostylar muscle; m.sg, styligeral muscle; pgs, larval protogonostylus; usg, unistyliger." figureDoi="http://doi.org/10.5281/zenodo.5577624" httpUri="https://zenodo.org/record/5577624/files/figure.png" pageId="125" pageNumber="126">Fig. 610</figureCitation>
). Sterna I–IV without denticles; sterna V–VIII with regular, pointed, triangular denticles (
<figureCitation id="1301CBC5F616E02F7BA7F934FE1FF902" box="[384,484,1716,1743]" captionStart="FIGURES 598–606" captionStartId="124.[151,264,1899,1923]" captionTargetBox="[164,1423,273,1876]" captionTargetId="figure-72@124.[164,1423,273,1876]" captionTargetPageId="124" captionText="FIGURES 598–606. Centroptella (Chopralla) papuanica sp. n., larval exuviae (holotype). 598, abdomen; 599, pronotum and mesonotum; 600, area of pronotum close to median suture with bifurcate seta corresponding to a on Fig. 14; 601, scale on metanotum with same magnification; 602, abdominal sternum VI and enlarged scales; 603, abdominal sternum IX and enlarged scales; 604, labrum; 605, labium; 606, abdominal terga VIII–IX and enlarged scales." figureDoi="http://doi.org/10.5281/zenodo.5577622" httpUri="https://zenodo.org/record/5577622/files/figure.png" pageId="125" pageNumber="126">Fig. 602</figureCitation>
). Posterior margin of sternum IX in female convex, with row of triangular denticles diminished medially (
<figureCitation id="1301CBC5F616E02F7BA9F958FE0BF93E" box="[398,496,1752,1779]" captionStart="FIGURES 598–606" captionStartId="124.[151,264,1899,1923]" captionTargetBox="[164,1423,273,1876]" captionTargetId="figure-72@124.[164,1423,273,1876]" captionTargetPageId="124" captionText="FIGURES 598–606. Centroptella (Chopralla) papuanica sp. n., larval exuviae (holotype). 598, abdomen; 599, pronotum and mesonotum; 600, area of pronotum close to median suture with bifurcate seta corresponding to a on Fig. 14; 601, scale on metanotum with same magnification; 602, abdominal sternum VI and enlarged scales; 603, abdominal sternum IX and enlarged scales; 604, labrum; 605, labium; 606, abdominal terga VIII–IX and enlarged scales." figureDoi="http://doi.org/10.5281/zenodo.5577622" httpUri="https://zenodo.org/record/5577622/files/figure.png" pageId="125" pageNumber="126">Fig. 603</figureCitation>
), in male without denticles between protogonostyli (
<figureCitation id="1301CBC5F616E02F7E65F958FB5EF93E" box="[1090,1189,1752,1779]" captionStart="FIGURES 607–610" captionStartId="125.[151,264,1430,1454]" captionTargetBox="[177,1421,191,1392]" captionTargetId="figure-20@125.[164,1423,181,1406]" captionTargetPageId="125" captionText="FIGURES 607–610. Centroptella (Chopralla) papuanica sp. n. 607–608, left and right mandibles; 609, subimaginal gonostyli crumpled under larval cuticle; 610, larval abdominal segment X without caudalii (holotype). Abbreviations: gs1–gs3, segments of gonostyli; m.gs, gonostylar muscle; m.sg, styligeral muscle; pgs, larval protogonostylus; usg, unistyliger." figureDoi="http://doi.org/10.5281/zenodo.5577624" httpUri="https://zenodo.org/record/5577624/files/figure.png" pageId="125" pageNumber="126">Fig. 609</figureCitation>
). Paraprocts with thin and colorless antero- median apodeme, with large pointed denticles on posterior margin (
<figureCitation id="1301CBC5F616E02F7E5BF97CFB24F8DA" box="[1148,1247,1788,1815]" captionStart="FIGURES 607–610" captionStartId="125.[151,264,1430,1454]" captionTargetBox="[177,1421,191,1392]" captionTargetId="figure-20@125.[164,1423,181,1406]" captionTargetPageId="125" captionText="FIGURES 607–610. Centroptella (Chopralla) papuanica sp. n. 607–608, left and right mandibles; 609, subimaginal gonostyli crumpled under larval cuticle; 610, larval abdominal segment X without caudalii (holotype). Abbreviations: gs1–gs3, segments of gonostyli; m.gs, gonostylar muscle; m.sg, styligeral muscle; pgs, larval protogonostylus; usg, unistyliger." figureDoi="http://doi.org/10.5281/zenodo.5577624" httpUri="https://zenodo.org/record/5577624/files/figure.png" pageId="125" pageNumber="126">Fig. 610</figureCitation>
).
</paragraph>
<paragraph id="8B85D740F616E02F7AE0F8A0FBE3F84E" blockId="125.[151,1437,1572,2031]" pageId="125" pageNumber="126">
Tergalius I narrow, elongate-ellipsoid; other tergalii wider, gradually changing in shape from tergalius II to tergalius VII (
<figureCitation id="1301CBC5F616E02F7B30F8C4FE4DF892" box="[279,438,1860,1887]" captionStart="FIGURES 584–597" captionStartId="122.[151,264,1851,1875]" captionTargetBox="[152,1425,193,1810]" captionTargetId="figure-27@122.[151,1436,181,1827]" captionTargetPageId="122" captionText="FIGURES 584–597. Tergalii I–VII. 584–590, Centroptella (Chopralla) rufostriata sp. n.; 591–597, Centroptella (Chopralla) papuanica sp. n." figureDoi="http://doi.org/10.5281/zenodo.5577620" httpUri="https://zenodo.org/record/5577620/files/figure.png" pageId="125" pageNumber="126">Figs 591–597</figureCitation>
). Each tergalius II–VII, besides costal and anal ribs, with straight and narrow middle rib, located on dorsal surface on background of main trachea (see
<taxonomicName id="4C3AACC3F616E02F796AF8E8FC21F84F" authorityName=": Kluge, Godunko &amp; Svitok" authorityYear="2020" box="[845,986,1896,1922]" class="Insecta" family="Baetidae" genus="Centroptella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="125" pageNumber="126" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="B94E0B52F616E02F796AF8E8FC21F84F" box="[845,986,1896,1922]" italics="true" pageId="125" pageNumber="126">Centroptella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27DB629F616E02F79C7F8E9FBF3F84E" box="[992,1032,1896,1923]" pageId="125" pageNumber="126" sensu="lato">s. l.</taxonomicNameLabel>
).
</paragraph>
<paragraph id="8B85D740F616E02F7AE0F80CFE95F822" blockId="125.[151,1437,1572,2031]" pageId="125" pageNumber="126">
Enlarged denticles on outer side of cercus not large (reaching or not reaching midlength of next segment), located on every 2nd segment and arranged by 1–3 on each side of secondary swimming setal row [see
<taxonomicName id="4C3AACC3F616E02F7F37F830FA66F807" authorityName=": Kluge, Godunko &amp; Svitok" authorityYear="2020" box="[1296,1437,1968,1994]" class="Insecta" family="Baetidae" genus="Centroptella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="125" pageNumber="126" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="B94E0B52F616E02F7F37F830FA66F807" box="[1296,1437,1968,1994]" italics="true" pageId="125" pageNumber="126">Centroptella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27DB629F616E02F7AB0F855FF44F823" box="[151,191,2004,2031]" pageId="125" pageNumber="126" sensu="lato">s. l.</taxonomicNameLabel>
(3)] (
<figureCitation id="1301CBC5F616E02F7AD8F854FE99F822" box="[255,354,2004,2031]" captionStart="FIGURES 611–617" captionStartId="126.[151,264,1830,1854]" captionTargetBox="[151,1436,181,1806]" captionTargetId="figure-27@126.[151,1436,181,1806]" captionTargetPageId="126" captionText="FIGURES 611–617. Centroptella (Chopralla) papuanica sp. n., larval exuviae. 611–613, fore, middle and hind legs, with focus on anterior side; 614–616, their tibiae, with focus on posterior side (arrows show patella-tibial suture); 617, cercus." figureDoi="http://doi.org/10.5281/zenodo.5577626" httpUri="https://zenodo.org/record/5577626/files/figure.png" pageId="125" pageNumber="126">Fig. 617</figureCitation>
).
</paragraph>
<caption id="DF4587C8F615E02C7AB0F8A6FADEF8AF" ID-DOI="http://doi.org/10.5281/zenodo.5577626" ID-Zenodo-Dep="5577626" httpUri="https://zenodo.org/record/5577626/files/figure.png" pageId="126" pageNumber="127" startId="126.[151,264,1830,1854]" targetBox="[151,1436,181,1806]" targetPageId="126">
<paragraph id="8B85D740F615E02C7AB0F8A6FADEF8AF" blockId="126.[151,1436,1830,1890]" pageId="126" pageNumber="127">
<emphasis id="B94E0B52F615E02C7AB0F8A6FE91F8F3" bold="true" box="[151,362,1830,1854]" pageId="126" pageNumber="127">FIGURES 611–617.</emphasis>
<taxonomicName id="4C3AACC3F615E02C7B57F8A6FD26F8F3" authority="Kluge, 2021" authorityName="Kluge" authorityYear="2021" box="[368,733,1830,1854]" class="Insecta" family="Baetidae" genus="Centroptella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="papuanica" status="sp. nov." subGenus="Chopralla">
<emphasis id="B94E0B52F615E02C7B57F8A6FE14F8F3" box="[368,495,1830,1854]" italics="true" pageId="126" pageNumber="127">Centroptella</emphasis>
(
<emphasis id="B94E0B52F615E02C7BDAF8A6FD9EF8F3" box="[509,613,1830,1854]" italics="true" pageId="126" pageNumber="127">Chopralla</emphasis>
)
<emphasis id="B94E0B52F615E02C7855F8A7FD26F8F3" box="[626,733,1831,1854]" italics="true" pageId="126" pageNumber="127">papuanica</emphasis>
</taxonomicName>
<emphasis id="B94E0B52F615E02C78C4F8A6FCE0F8F3" bold="true" box="[739,795,1830,1854]" pageId="126" pageNumber="127">
<taxonomicNameLabel id="A27DB629F615E02C78C4F8A6FCE0F8F3" box="[739,795,1830,1854]" pageId="126" pageNumber="127" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, larval exuviae. 611–613, fore, middle and hind legs, with focus on anterior side; 614–616, their tibiae, with focus on posterior side (arrows show patella-tibial suture); 617, cercus.
</paragraph>
</caption>
<caption id="DF4587C8F614E02D7AB0F91BFD26F936" ID-DOI="http://doi.org/10.5281/zenodo.5577628" ID-Zenodo-Dep="5577628" httpUri="https://zenodo.org/record/5577628/files/figure.png" pageId="127" pageNumber="128" startId="127.[151,264,1691,1715]" targetBox="[151,1436,181,1663]" targetPageId="127">
<paragraph id="8B85D740F614E02D7AB0F91BFD26F936" blockId="127.[151,1436,1691,1787]" pageId="127" pageNumber="128">
<emphasis id="B94E0B52F614E02D7AB0F91BFE96F97E" bold="true" box="[151,365,1691,1715]" pageId="127" pageNumber="128">FIGURES 618–623.</emphasis>
<taxonomicName id="4C3AACC3F614E02D7B55F91BFD1AF97E" authority="Kluge, 2021" authorityName="Kluge" authorityYear="2021" box="[370,737,1691,1715]" class="Insecta" family="Baetidae" genus="Centroptella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="127" pageNumber="128" phylum="Arthropoda" rank="species" species="papuanica" status="sp. nov." subGenus="Chopralla">
<emphasis id="B94E0B52F614E02D7B55F91BFE0AF97E" box="[370,497,1691,1715]" italics="true" pageId="127" pageNumber="128">Centroptella</emphasis>
(
<emphasis id="B94E0B52F614E02D7827F91BFD93F97E" box="[512,616,1691,1715]" italics="true" pageId="127" pageNumber="128">Chopralla</emphasis>
)
<emphasis id="B94E0B52F614E02D7851F91CFD1AF97E" box="[630,737,1692,1715]" italics="true" pageId="127" pageNumber="128">papuanica</emphasis>
</taxonomicName>
<emphasis id="B94E0B52F614E02D78C0F91BFCE4F97E" bold="true" box="[743,799,1691,1715]" pageId="127" pageNumber="128">
<taxonomicNameLabel id="A27DB629F614E02D78C0F91BFCE4F97E" box="[743,799,1691,1715]" pageId="127" pageNumber="128" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
618, fore leg of female imago; 619, abdomen of female imago; 620, female imago; 621, subimaginal exuviae of mesopleuron; 622, subimaginal exuviae of mesonotum; 623, abdomen of male subimago extracted from larva (618, 619, 621, holotype).
</paragraph>
</caption>
<paragraph id="8B85D740F614E02D7AE0F8A9FC18F835" blockId="127.[151,1437,1833,2040]" pageId="127" pageNumber="128">
SCALES: Scales on occipit, median parts of pronotum and mesonotum very small (with sockets 3–4 mkm and length 4–5 mkm) (
<figureCitation id="1301CBC5F614E02D7B44F8CDFE3EF8A5" box="[355,453,1869,1896]" captionStart="FIGURES 598–606" captionStartId="124.[151,264,1899,1923]" captionTargetBox="[164,1423,273,1876]" captionTargetId="figure-72@124.[164,1423,273,1876]" captionTargetPageId="124" captionText="FIGURES 598–606. Centroptella (Chopralla) papuanica sp. n., larval exuviae (holotype). 598, abdomen; 599, pronotum and mesonotum; 600, area of pronotum close to median suture with bifurcate seta corresponding to a on Fig. 14; 601, scale on metanotum with same magnification; 602, abdominal sternum VI and enlarged scales; 603, abdominal sternum IX and enlarged scales; 604, labrum; 605, labium; 606, abdominal terga VIII–IX and enlarged scales." figureDoi="http://doi.org/10.5281/zenodo.5577622" httpUri="https://zenodo.org/record/5577622/files/figure.png" pageId="127" pageNumber="128">Fig. 600</figureCitation>
). Lateral parts of pronotum, lateral parts of mesonotum and protoptera nearly lack scales. Thoracic pleura and median part of metanotum with scales in opercula-bearing sockets, with longitudinal striation (
<figureCitation id="1301CBC5F614E02D7A87F815FEF8F87D" box="[160,259,1941,1968]" captionStart="FIGURES 598–606" captionStartId="124.[151,264,1899,1923]" captionTargetBox="[164,1423,273,1876]" captionTargetId="figure-72@124.[164,1423,273,1876]" captionTargetPageId="124" captionText="FIGURES 598–606. Centroptella (Chopralla) papuanica sp. n., larval exuviae (holotype). 598, abdomen; 599, pronotum and mesonotum; 600, area of pronotum close to median suture with bifurcate seta corresponding to a on Fig. 14; 601, scale on metanotum with same magnification; 602, abdominal sternum VI and enlarged scales; 603, abdominal sternum IX and enlarged scales; 604, labrum; 605, labium; 606, abdominal terga VIII–IX and enlarged scales." figureDoi="http://doi.org/10.5281/zenodo.5577622" httpUri="https://zenodo.org/record/5577622/files/figure.png" pageId="127" pageNumber="128">Fig. 601</figureCitation>
). Femora, tibiae and tarsi densely covered with colorless, short, oval scales in operculae-bearing sockets (sockets 5–6 mkm). Scales on abdominal terga elongate and pointed, with longitudinal striation (
<figureCitation id="1301CBC5F614E02D7EFAF839FABAF819" box="[1245,1345,1977,2004]" captionStart="FIGURES 598–606" captionStartId="124.[151,264,1899,1923]" captionTargetBox="[164,1423,273,1876]" captionTargetId="figure-72@124.[164,1423,273,1876]" captionTargetPageId="124" captionText="FIGURES 598–606. Centroptella (Chopralla) papuanica sp. n., larval exuviae (holotype). 598, abdomen; 599, pronotum and mesonotum; 600, area of pronotum close to median suture with bifurcate seta corresponding to a on Fig. 14; 601, scale on metanotum with same magnification; 602, abdominal sternum VI and enlarged scales; 603, abdominal sternum IX and enlarged scales; 604, labrum; 605, labium; 606, abdominal terga VIII–IX and enlarged scales." figureDoi="http://doi.org/10.5281/zenodo.5577622" httpUri="https://zenodo.org/record/5577622/files/figure.png" pageId="127" pageNumber="128">Fig. 606</figureCitation>
). Scales on abdominal sterna either similar (
<figureCitation id="1301CBC5F614E02D7802F85DFD7DF835" box="[549,646,2013,2040]" captionStart="FIGURES 598–606" captionStartId="124.[151,264,1899,1923]" captionTargetBox="[164,1423,273,1876]" captionTargetId="figure-72@124.[164,1423,273,1876]" captionTargetPageId="124" captionText="FIGURES 598–606. Centroptella (Chopralla) papuanica sp. n., larval exuviae (holotype). 598, abdomen; 599, pronotum and mesonotum; 600, area of pronotum close to median suture with bifurcate seta corresponding to a on Fig. 14; 601, scale on metanotum with same magnification; 602, abdominal sternum VI and enlarged scales; 603, abdominal sternum IX and enlarged scales; 604, labrum; 605, labium; 606, abdominal terga VIII–IX and enlarged scales." figureDoi="http://doi.org/10.5281/zenodo.5577622" httpUri="https://zenodo.org/record/5577622/files/figure.png" pageId="127" pageNumber="128">Fig. 603</figureCitation>
), or partly colorless (
<figureCitation id="1301CBC5F614E02D7953F85DFC2CF835" box="[884,983,2013,2040]" captionStart="FIGURES 598–606" captionStartId="124.[151,264,1899,1923]" captionTargetBox="[164,1423,273,1876]" captionTargetId="figure-72@124.[164,1423,273,1876]" captionTargetPageId="124" captionText="FIGURES 598–606. Centroptella (Chopralla) papuanica sp. n., larval exuviae (holotype). 598, abdomen; 599, pronotum and mesonotum; 600, area of pronotum close to median suture with bifurcate seta corresponding to a on Fig. 14; 601, scale on metanotum with same magnification; 602, abdominal sternum VI and enlarged scales; 603, abdominal sternum IX and enlarged scales; 604, labrum; 605, labium; 606, abdominal terga VIII–IX and enlarged scales." figureDoi="http://doi.org/10.5281/zenodo.5577622" httpUri="https://zenodo.org/record/5577622/files/figure.png" pageId="127" pageNumber="128">Fig. 602</figureCitation>
).
</paragraph>
<paragraph id="8B85D740F6EBE0D27AE0FF17FC7DFED3" blockId="128.[151,1437,150,1078]" pageId="128" pageNumber="129">
STOUT SETAE: Outer side of femur with regular row of 5–8 long, clavate setae and 2 subapical setae of same form (
<figureCitation id="1301CBC5F6EBE0D27AF9FF3BFE84FF1B" box="[222,383,187,214]" captionStart="FIGURES 611–617" captionStartId="126.[151,264,1830,1854]" captionTargetBox="[151,1436,181,1806]" captionTargetId="figure-27@126.[151,1436,181,1806]" captionTargetPageId="126" captionText="FIGURES 611–617. Centroptella (Chopralla) papuanica sp. n., larval exuviae. 611–613, fore, middle and hind legs, with focus on anterior side; 614–616, their tibiae, with focus on posterior side (arrows show patella-tibial suture); 617, cercus." figureDoi="http://doi.org/10.5281/zenodo.5577626" httpUri="https://zenodo.org/record/5577626/files/figure.png" pageId="128" pageNumber="129">Figs 611–613</figureCitation>
). Inner-anterior side of each femur with small (about 10 mkm length), stout, blunt setae; no longer setae on inner-anterior side of fore femur. Tibia and tarsus with stout setae on inner side (
<figureCitation id="1301CBC5F6EBE0D27EC8FF5FFA6AFF37" box="[1263,1425,223,250]" captionStart="FIGURES 611–617" captionStartId="126.[151,264,1830,1854]" captionTargetBox="[151,1436,181,1806]" captionTargetId="figure-27@126.[151,1436,181,1806]" captionTargetPageId="126" captionText="FIGURES 611–617. Centroptella (Chopralla) papuanica sp. n., larval exuviae. 611–613, fore, middle and hind legs, with focus on anterior side; 614–616, their tibiae, with focus on posterior side (arrows show patella-tibial suture); 617, cercus." figureDoi="http://doi.org/10.5281/zenodo.5577626" httpUri="https://zenodo.org/record/5577626/files/figure.png" pageId="128" pageNumber="129">Figs 614–616</figureCitation>
). Outer-apical seta of tibia small, blunt and elongate (as in
<figureCitation id="1301CBC5F6EBE0D2793FFE83FC81FED3" box="[792,890,259,286]" captionStart="FIGURES 364–368" captionStartId="80.[151,264,1703,1727]" captionTargetBox="[151,1436,181,1679]" captionTargetId="figure-27@80.[151,1436,181,1679]" captionTargetPageId="80" captionText="FIGURES 364–368. Centroptella (Crassolus) ludmilae sp. n., larval exuviae. 364, apex of fore tibia (arrow shows outer-apical seta); 365, abdominal tergum V; 366, abdominal tergum III of holotype (MA, medioanterior sigilla; MP, medioposterior sigilla); 367, abdominal tergum X; 368, female abdominal sternum IX and paraprocts." figureDoi="http://doi.org/10.5281/zenodo.5577542" httpUri="https://zenodo.org/record/5577542/files/figure.png" pageId="128" pageNumber="129">Fig. 364</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32084CBF6EBE0D27AE0FEA7FB54FD64" pageId="128" pageNumber="129" type="materials_examined">
<paragraph id="8B85D740F6EBE0D27AE0FEA7FCECFDD4" blockId="128.[151,1437,150,1078]" pageId="128" pageNumber="129">
<collectionCode id="ED2B4F85F6EBE0D27AE0FEA7FEE3FE8C" box="[199,280,295,321]" pageId="128" pageNumber="129">LONG</collectionCode>
,
<collectionCode id="ED2B4F85F6EBE0D27B02FEA7FE98FE8C" box="[293,355,295,321]" pageId="128" pageNumber="129">FINE</collectionCode>
SETAE: Bifurcate setae of metasternum (i) (see
<taxonomicName id="4C3AACC3F6EBE0D279B0FEA7FBDFFE8C" authorityName=": Kluge, Godunko &amp; Svitok" authorityYear="2020" box="[919,1060,295,321]" class="Insecta" family="Baetidae" genus="Centroptella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="128" pageNumber="129" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="B94E0B52F6EBE0D279B0FEA7FBDFFE8C" box="[919,1060,295,321]" italics="true" pageId="128" pageNumber="129">Centroptella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27DB629F6EBE0D27E0AFEA7FBAEFE8F" box="[1069,1109,295,322]" pageId="128" pageNumber="129" sensu="lato">s. l.</taxonomicNameLabel>
) form pair of dense, regular rows partly with contiguous sockets (as in
<figureCitation id="1301CBC5F6EBE0D27855FECBFD28FEA8" box="[626,723,330,357]" captionStart="FIGURES 512–517" captionStartId="107.[151,264,1802,1826]" captionTargetBox="[151,1436,181,1778]" captionTargetId="figure-20@107.[151,1436,181,1778]" captionTargetPageId="107" captionText="FIGURES 512–517. Centroptella (Chopralla) ghatensis sp. n., exuviae of larval thorax. 512–513, scales and bifurcate setae on anterior part of mesonotum (corresponding to e on Fig. 14); 514–517, bifurcate setae on thoracic sternum (corresponding to h and i on Fig. 15)." figureDoi="http://doi.org/10.5281/zenodo.5577594" httpUri="https://zenodo.org/record/5577594/files/figure.png" pageId="128" pageNumber="129">Fig. 517</figureCitation>
). Posterior arm of U-shaped row of long setae on fore- and middle leg situated across tibia (
<figureCitation id="1301CBC5F6EBE0D27BF0FEEFFD8CFE44" box="[471,631,366,393]" captionStart="FIGURES 611–617" captionStartId="126.[151,264,1830,1854]" captionTargetBox="[151,1436,181,1806]" captionTargetId="figure-27@126.[151,1436,181,1806]" captionTargetPageId="126" captionText="FIGURES 611–617. Centroptella (Chopralla) papuanica sp. n., larval exuviae. 611–613, fore, middle and hind legs, with focus on anterior side; 614–616, their tibiae, with focus on posterior side (arrows show patella-tibial suture); 617, cercus." figureDoi="http://doi.org/10.5281/zenodo.5577626" httpUri="https://zenodo.org/record/5577626/files/figure.png" pageId="128" pageNumber="129">Figs 614–615</figureCitation>
); on hind leg longitudinal (
<figureCitation id="1301CBC5F6EBE0D2798FFEEFFBF3FE47" box="[936,1032,367,394]" captionStart="FIGURES 611–617" captionStartId="126.[151,264,1830,1854]" captionTargetBox="[151,1436,181,1806]" captionTargetId="figure-27@126.[151,1436,181,1806]" captionTargetPageId="126" captionText="FIGURES 611–617. Centroptella (Chopralla) papuanica sp. n., larval exuviae. 611–613, fore, middle and hind legs, with focus on anterior side; 614–616, their tibiae, with focus on posterior side (arrows show patella-tibial suture); 617, cercus." figureDoi="http://doi.org/10.5281/zenodo.5577626" httpUri="https://zenodo.org/record/5577626/files/figure.png" pageId="128" pageNumber="129">Fig. 616</figureCitation>
) [see
<taxonomicName id="4C3AACC3F6EBE0D27E6BFEEFFB44FE44" authorityName="Waltz &amp; McCafferty" authorityYear="1987" box="[1100,1215,367,393]" class="Insecta" family="Baetidae" genus="Chopralla" kingdom="Animalia" order="Ephemeroptera" pageId="128" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis id="B94E0B52F6EBE0D27E6BFEEFFB44FE44" box="[1100,1215,367,393]" italics="true" pageId="128" pageNumber="129">Chopralla</emphasis>
</taxonomicName>
(1)]. Dorsal side of each tarsus with long, fine setae situated irregularly and partly forming two longitudinal rows (as in
<figureCitation id="1301CBC5F6EBE0D27EDCFE13FA63FE60" box="[1275,1432,402,429]" captionStart="FIGURES 560–565" captionStartId="117.[151,264,1920,1944]" captionTargetBox="[164,1423,434,1897]" captionTargetId="figure-128@117.[164,1423,417,1897]" captionTargetPageId="117" captionText="FIGURES 560–565. Centroptella (Chopralla) rufostriata sp. n., larva. 560–561, fore and middle tibiae, anterior view (setae and sutures on opposite side shown by interrupted lines); 562–563, hind tibia and tarsus, anterior and outer view (bases of long, fine setae shown as rings). 564, abdominal tergum IX; 565, abdominal sternum VII." figureDoi="http://doi.org/10.5281/zenodo.5577610" httpUri="https://zenodo.org/record/5577610/files/figure.png" pageId="128" pageNumber="129">Figs 562–563</figureCitation>
; see
<taxonomicName id="4C3AACC3F6EBE0D27AE5FE37FEB4FE1C" authorityName=": Kluge, Godunko &amp; Svitok" authorityYear="2020" box="[194,335,439,465]" class="Insecta" family="Baetidae" genus="Centroptella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="128" pageNumber="129" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="B94E0B52F6EBE0D27AE5FE37FEB4FE1C" box="[194,335,439,465]" italics="true" pageId="128" pageNumber="129">Centroptella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27DB629F6EBE0D27B71FE37FE84FE1F" box="[342,383,439,466]" pageId="128" pageNumber="129" sensu="lato">s. l.</taxonomicNameLabel>
). Tergum VII with pair of fine, V-shaped setae sublaterally (g) (as in
<figureCitation id="1301CBC5F6EBE0D27EAAFE37FB0BFE1F" box="[1165,1264,439,466]" captionStart="FIGURES 404–406" captionStartId="88.[151,264,1792,1816]" captionTargetBox="[170,1417,181,1768]" captionTargetId="figure-27@88.[170,1417,181,1768]" captionTargetPageId="88" captionText="FIGURES 404–406. Centroptella (Crassolus) ingridae, larval cuticle of abdomen. 404, tergum III of holotype (MA, medioanterior sigillum; MP, medioposterior sigillum); 405, terga and sterna; 406, tergum VII (letter g corresponds to Fig. 14)." figureDoi="http://doi.org/10.5281/zenodo.5577558" httpUri="https://zenodo.org/record/5577558/files/figure.png" pageId="128" pageNumber="129">Fig. 406</figureCitation>
). Each sternum IV–V with pair of regular, transverse rows of long, fine, bifurcate setae with spaced sockets (j); other sterna either without such setae, or with few setae irregularly situated.
</paragraph>
<paragraph id="8B85D740F6EBE0D27AE0FDA3FCBFFD48" blockId="128.[151,1437,150,1078]" pageId="128" pageNumber="129">
<collectionCode id="ED2B4F85F6EBE0D27AE0FDA3FEEDFDF0" box="[199,278,547,573]" pageId="128" pageNumber="129">MALE</collectionCode>
GENITALIA IN LARVA: In last larval instar, developing subimaginal gonostyli folded under larval cuticle in «
<taxonomicName id="4C3AACC3F6EBE0D27ADDFDC7FE86FDAC" box="[250,381,583,609]" class="Insecta" family="Baetidae" genus="Nigrobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="128" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis id="B94E0B52F6EBE0D27ADDFDC7FE86FDAC" box="[250,381,583,609]" italics="true" pageId="128" pageNumber="129">Nigrobaetis</emphasis>
</taxonomicName>
-
<typeStatus id="548169E2F6EBE0D27BA2FDC7FE43FDAC" box="[389,440,583,609]" pageId="128" pageNumber="129">type</typeStatus>
» pose, with 2nd segment bent by convexity medially, and 3rd segment directed caudally and inserted into 2nd segment (
<figureCitation id="1301CBC5F6EBE0D27BD1FDEBFDACFD4B" box="[502,599,619,646]" captionStart="FIGURES 607–610" captionStartId="125.[151,264,1430,1454]" captionTargetBox="[177,1421,191,1392]" captionTargetId="figure-20@125.[164,1423,181,1406]" captionTargetPageId="125" captionText="FIGURES 607–610. Centroptella (Chopralla) papuanica sp. n. 607–608, left and right mandibles; 609, subimaginal gonostyli crumpled under larval cuticle; 610, larval abdominal segment X without caudalii (holotype). Abbreviations: gs1–gs3, segments of gonostyli; m.gs, gonostylar muscle; m.sg, styligeral muscle; pgs, larval protogonostylus; usg, unistyliger." figureDoi="http://doi.org/10.5281/zenodo.5577624" httpUri="https://zenodo.org/record/5577624/files/figure.png" pageId="128" pageNumber="129">Fig. 609</figureCitation>
) [see
<taxonomicName id="4C3AACC3F6EBE0D278BEFDEBFCF7FD48" authorityName="Waltz &amp; McCafferty" authorityYear="1987" box="[665,780,619,645]" class="Insecta" family="Baetidae" genus="Chopralla" kingdom="Animalia" order="Ephemeroptera" pageId="128" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis id="B94E0B52F6EBE0D278BEFDEBFCF7FD48" box="[665,780,619,645]" italics="true" pageId="128" pageNumber="129">Chopralla</emphasis>
</taxonomicName>
(5)].
</paragraph>
<paragraph id="8B85D740F6EBE0D27AE0FD0FFB54FD64" blockId="128.[151,1437,150,1078]" box="[199,1199,655,681]" pageId="128" pageNumber="129">
<collectionCode id="ED2B4F85F6EBE0D27AE0FD0FFE7BFD64" box="[199,384,655,681]" pageId="128" pageNumber="129">RESPIRATORY</collectionCode>
MOVEMENTS: Tergalii able to make rhythmic respiratory movements.
</paragraph>
</subSubSection>
<subSubSection id="C32084CBF6EBE0D37AE0FD32FD11FE65" lastPageId="129" lastPageNumber="130" pageId="128" pageNumber="129" type="description">
<paragraph id="8B85D740F6EBE0D27AE0FD32FA8BFCF4" blockId="128.[151,1437,150,1078]" pageId="128" pageNumber="129">
<emphasis id="B94E0B52F6EBE0D27AE0FD32FEB8FD00" bold="true" box="[199,323,690,717]" italics="true" pageId="128" pageNumber="129">Subimago.</emphasis>
CUTICULAR COLORATION: Head and antennae colorless, with microtrichia dark brown. Pronotum light brownish. Mesonotum light brown with certain sutures darker and medioparapsidal suture lighter (
<figureCitation id="1301CBC5F6EBE0D27F48FD57FF3EFCDB" captionStart="FIGURES 618–623" captionStartId="127.[151,264,1691,1715]" captionTargetBox="[151,1436,181,1663]" captionTargetId="figure-20@127.[151,1436,181,1667]" captionTargetPageId="127" captionText="FIGURES 618–623. Centroptella (Chopralla) papuanica sp. n. 618, fore leg of female imago; 619, abdomen of female imago; 620, female imago; 621, subimaginal exuviae of mesopleuron; 622, subimaginal exuviae of mesonotum; 623, abdomen of male subimago extracted from larva (618, 619, 621, holotype)." figureDoi="http://doi.org/10.5281/zenodo.5577628" httpUri="https://zenodo.org/record/5577628/files/figure.png" pageId="128" pageNumber="129">Fig. 622</figureCitation>
). Thoracic sterna and pleura with brown and colorless areas (
<figureCitation id="1301CBC5F6EBE0D2795BFD7BFC24FCDB" box="[892,991,763,790]" captionStart="FIGURES 618–623" captionStartId="127.[151,264,1691,1715]" captionTargetBox="[151,1436,181,1663]" captionTargetId="figure-20@127.[151,1436,181,1667]" captionTargetPageId="127" captionText="FIGURES 618–623. Centroptella (Chopralla) papuanica sp. n. 618, fore leg of female imago; 619, abdomen of female imago; 620, female imago; 621, subimaginal exuviae of mesopleuron; 622, subimaginal exuviae of mesonotum; 623, abdomen of male subimago extracted from larva (618, 619, 621, holotype)." figureDoi="http://doi.org/10.5281/zenodo.5577628" httpUri="https://zenodo.org/record/5577628/files/figure.png" pageId="128" pageNumber="129">Fig. 621</figureCitation>
). Legs light brownish, nearly colorless. Wings light, without brown circles at bases of microtrichia. Abdominal terga and sterna brown, cerci colorless.
</paragraph>
<paragraph id="8B85D740F6EBE0D27AE0FCC3FC92FC93" blockId="128.[151,1437,150,1078]" box="[199,873,835,862]" pageId="128" pageNumber="129">HYPODERMAL COLORATION: As in imago (see below).</paragraph>
<paragraph id="8B85D740F6EBE0D27AE0FCE7FF29FC68" blockId="128.[151,1437,150,1078]" pageId="128" pageNumber="129">
TEXTURE: On all legs of both sexes all tarsal segments entirely covered with pointed microlepides (as in
<figureCitation id="1301CBC5F6EBE0D27F57FCE7FF3EFC68" captionStart="FIGURES 387–392" captionStartId="84.[151,264,1786,1810]" captionTargetBox="[151,1436,181,1762]" captionTargetId="figure-27@84.[151,1436,181,1762]" captionTargetPageId="84" captionText="FIGURES 387–392. Centroptella (Crassolus) ludmilae sp. n. 387–388, female imago; 389, exuviae of subimaginal pleuron; 390, exuviae of subimaginal mesonotum; 391, exuviae of female subimaginal fore and hind legs. Abbreviations: T1–T5, tarsomeres." figureDoi="http://doi.org/10.5281/zenodo.5577550" httpUri="https://zenodo.org/record/5577550/files/figure.png" pageId="128" pageNumber="129">Fig. 391</figureCitation>
).
</paragraph>
<paragraph id="8B85D740F6EBE0D27AE0FC2FFC3AFBF8" blockId="128.[151,1437,150,1078]" pageId="128" pageNumber="129">
<emphasis id="B94E0B52F6EBE0D27AE0FC2FFEA4FC04" bold="true" box="[199,351,942,969]" italics="true" pageId="128" pageNumber="129">Imago, male.</emphasis>
Unknown. Judging by subimago developing under larval cuticle, tarsus of middle and hind legs with 1 apical spine on initial 3rd tarsomere (next after 1st+2nd tarsomere); abdomen light with posterior margins of some terga narrowly bordered with reddish-brown (
<figureCitation id="1301CBC5F6EBE0D27921FC77FC91FBDF" box="[774,874,1015,1042]" captionStart="FIGURES 618–623" captionStartId="127.[151,264,1691,1715]" captionTargetBox="[151,1436,181,1663]" captionTargetId="figure-20@127.[151,1436,181,1667]" captionTargetPageId="127" captionText="FIGURES 618–623. Centroptella (Chopralla) papuanica sp. n. 618, fore leg of female imago; 619, abdomen of female imago; 620, female imago; 621, subimaginal exuviae of mesopleuron; 622, subimaginal exuviae of mesonotum; 623, abdomen of male subimago extracted from larva (618, 619, 621, holotype)." figureDoi="http://doi.org/10.5281/zenodo.5577628" httpUri="https://zenodo.org/record/5577628/files/figure.png" pageId="128" pageNumber="129">Fig. 623</figureCitation>
); median projection of penial bridge shallow and truncate; sterno-styligeral muscle present (
<figureCitation id="1301CBC5F6EBE0D27853FB9BFD2FFBFB" box="[628,724,1051,1078]" captionStart="FIGURES 607–610" captionStartId="125.[151,264,1430,1454]" captionTargetBox="[177,1421,191,1392]" captionTargetId="figure-20@125.[164,1423,181,1406]" captionTargetPageId="125" captionText="FIGURES 607–610. Centroptella (Chopralla) papuanica sp. n. 607–608, left and right mandibles; 609, subimaginal gonostyli crumpled under larval cuticle; 610, larval abdominal segment X without caudalii (holotype). Abbreviations: gs1–gs3, segments of gonostyli; m.gs, gonostylar muscle; m.sg, styligeral muscle; pgs, larval protogonostylus; usg, unistyliger." figureDoi="http://doi.org/10.5281/zenodo.5577624" httpUri="https://zenodo.org/record/5577624/files/figure.png" pageId="128" pageNumber="129">Fig. 609</figureCitation>
) [see
<taxonomicName id="4C3AACC3F6EBE0D27931FB9BFC72FBF8" authorityName="Waltz &amp; McCafferty" authorityYear="1987" box="[790,905,1051,1077]" class="Insecta" family="Baetidae" genus="Chopralla" kingdom="Animalia" order="Ephemeroptera" pageId="128" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis id="B94E0B52F6EBE0D27931FB9BFC72FBF8" box="[790,905,1051,1077]" italics="true" pageId="128" pageNumber="129">Chopralla</emphasis>
</taxonomicName>
(7)].
</paragraph>
<caption id="DF4587C8F6EBE0D27AB0F891FB9DF8E4" ID-DOI="http://doi.org/10.5281/zenodo.5577630" ID-Zenodo-Dep="5577630" box="[151,1126,1809,1834]" httpUri="https://zenodo.org/record/5577630/files/figure.png" pageId="128" pageNumber="129" startId="128.[151,264,1809,1833]" targetBox="[151,1435,1101,1786]" targetPageId="128">
<paragraph id="8B85D740F6EBE0D27AB0F891FB9DF8E4" blockId="128.[151,1126,1809,1834]" box="[151,1126,1809,1834]" pageId="128" pageNumber="129">
<emphasis id="B94E0B52F6EBE0D27AB0F891FE95F8E4" bold="true" box="[151,366,1809,1833]" pageId="128" pageNumber="129">FIGURES 624–625.</emphasis>
<taxonomicName id="4C3AACC3F6EBE0D27B54F891FD18F8E4" authority="Kluge, 2021" authorityName="Kluge" authorityYear="2021" box="[371,739,1809,1833]" class="Insecta" family="Baetidae" genus="Centroptella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="128" pageNumber="129" phylum="Arthropoda" rank="species" species="papuanica" status="sp. nov." subGenus="Chopralla">
<emphasis id="B94E0B52F6EBE0D27B54F891FE09F8E4" box="[371,498,1809,1833]" italics="true" pageId="128" pageNumber="129">Centroptella</emphasis>
(
<emphasis id="B94E0B52F6EBE0D27826F891FD92F8E4" box="[513,617,1809,1833]" italics="true" pageId="128" pageNumber="129">Chopralla</emphasis>
)
<emphasis id="B94E0B52F6EBE0D2785FF892FD18F8E4" box="[632,739,1810,1833]" italics="true" pageId="128" pageNumber="129">papuanica</emphasis>
</taxonomicName>
<emphasis id="B94E0B52F6EBE0D278CDF891FCD8F8E4" bold="true" box="[746,803,1809,1833]" pageId="128" pageNumber="129">
<taxonomicNameLabel id="A27DB629F6EBE0D278CDF891FCD8F8E4" box="[746,803,1809,1833]" pageId="128" pageNumber="129" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, egg (extracted from holotype).
</paragraph>
</caption>
<paragraph id="8B85D740F6EBE0D37AE0F8D8FCEAFEF0" blockId="128.[151,1436,1879,2014]" lastBlockId="129.[151,1436,151,599]" lastPageId="129" lastPageNumber="130" pageId="128" pageNumber="129">
<emphasis id="B94E0B52F6EBE0D27AE0F8D8FE8BF8BF" bold="true" box="[199,368,1879,1906]" italics="true" pageId="128" pageNumber="129">Imago, female</emphasis>
(
<figureCitation id="1301CBC5F6EBE0D27BA6F8D7FDD8F8BF" box="[385,547,1879,1906]" captionStart="FIGURES 618–623" captionStartId="127.[151,264,1691,1715]" captionTargetBox="[151,1436,181,1663]" captionTargetId="figure-20@127.[151,1436,181,1667]" captionTargetPageId="127" captionText="FIGURES 618–623. Centroptella (Chopralla) papuanica sp. n. 618, fore leg of female imago; 619, abdomen of female imago; 620, female imago; 621, subimaginal exuviae of mesopleuron; 622, subimaginal exuviae of mesonotum; 623, abdomen of male subimago extracted from larva (618, 619, 621, holotype)." figureDoi="http://doi.org/10.5281/zenodo.5577628" httpUri="https://zenodo.org/record/5577628/files/figure.png" pageId="128" pageNumber="129">Figs 618–620</figureCitation>
). Head ochre. Thorax and abdomen contrastingly darker (ochre with reddishbrown) in dorsal half and lighter (pale ochre) in ventral half; pronotum and first 4 intersegmental membranes of abdominal terga with pair of transverse bright red stripes; posterior margins of terga narrowly bordered with reddish-brown. Wing with membrane colorless, veins pale ochre or colorless, extreme base of costal vein proximad of costal brace brownish. Femora of all legs ochre, femur of fore leg slightly darker apically; tibiae of all legs ochre, slightly darkened in proximal part (
<figureCitation id="1301CBC5F6EAE0D37802FF3AFD73FF18" box="[549,648,186,213]" captionStart="FIGURES 618–623" captionStartId="127.[151,264,1691,1715]" captionTargetBox="[151,1436,181,1663]" captionTargetId="figure-20@127.[151,1436,181,1667]" captionTargetPageId="127" captionText="FIGURES 618–623. Centroptella (Chopralla) papuanica sp. n. 618, fore leg of female imago; 619, abdomen of female imago; 620, female imago; 621, subimaginal exuviae of mesopleuron; 622, subimaginal exuviae of mesonotum; 623, abdomen of male subimago extracted from larva (618, 619, 621, holotype)." figureDoi="http://doi.org/10.5281/zenodo.5577628" httpUri="https://zenodo.org/record/5577628/files/figure.png" pageId="129" pageNumber="130">Fig. 618</figureCitation>
). Patella-tibial suture present on middle and hind legs, absent on forelegs (
<figureCitation id="1301CBC5F6EAE0D37AF4FF5DFEC9FF35" box="[211,306,221,248]" captionStart="FIGURES 618–623" captionStartId="127.[151,264,1691,1715]" captionTargetBox="[151,1436,181,1663]" captionTargetId="figure-20@127.[151,1436,181,1667]" captionTargetPageId="127" captionText="FIGURES 618–623. Centroptella (Chopralla) papuanica sp. n. 618, fore leg of female imago; 619, abdomen of female imago; 620, female imago; 621, subimaginal exuviae of mesopleuron; 622, subimaginal exuviae of mesonotum; 623, abdomen of male subimago extracted from larva (618, 619, 621, holotype)." figureDoi="http://doi.org/10.5281/zenodo.5577628" httpUri="https://zenodo.org/record/5577628/files/figure.png" pageId="129" pageNumber="130">Fig. 618</figureCitation>
) [as in male; see
<taxonomicName id="4C3AACC3F6EAE0D37BD2FF5DFD79FF3A" authorityName=": Kluge, Godunko &amp; Svitok" authorityYear="2020" box="[501,642,221,247]" class="Insecta" family="Baetidae" genus="Centroptella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="129" pageNumber="130" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="B94E0B52F6EAE0D37BD2FF5DFD79FF3A" box="[501,642,221,247]" italics="true" pageId="129" pageNumber="130">Centroptella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27DB629F6EAE0D378AFFF5EFD4BFF3A" box="[648,688,221,248]" pageId="129" pageNumber="130" sensu="lato">s. l.</taxonomicNameLabel>
(2)]. Tarsus of each leg with 1 apical spine on initial 3rd tarsomere (on foreleg—on tarsomere next after 2nd tarsomere, on middle and hind leg—on tarsomere next after 1st+2nd tarsomere) (as in Figs 691–692). Cerci unicolor pale ochre.
</paragraph>
<paragraph id="8B85D740F6EAE0D37AE0FEC7FF26FE49" blockId="129.[151,1436,151,599]" pageId="129" pageNumber="130">
<emphasis id="B94E0B52F6EAE0D37AE0FEC7FF0DFEAC" bold="true" box="[199,246,327,353]" italics="true" pageId="129" pageNumber="130">Egg</emphasis>
(
<figureCitation id="1301CBC5F6EAE0D37B2DFEC7FE54FEAC" box="[266,431,326,353]" captionStart="FIGURES 624–625" captionStartId="128.[151,264,1809,1833]" captionTargetBox="[151,1435,1101,1786]" captionTargetId="figure-433@128.[151,1436,1101,1786]" captionTargetPageId="128" captionText="FIGURES 624–625. Centroptella (Chopralla) papuanica sp. n., egg (extracted from holotype)." figureDoi="http://doi.org/10.5281/zenodo.5577630" httpUri="https://zenodo.org/record/5577630/files/figure.png" pageId="129" pageNumber="130">Figs 624–625</figureCitation>
). Oval; chorion with net-like relief stretched longitudinally. Micropile without sperm guide.
</paragraph>
<paragraph id="8B85D740F6EAE0D37AE0FE0DFD11FE65" blockId="129.[151,1436,151,599]" box="[199,746,397,424]" pageId="129" pageNumber="130">
<emphasis id="B94E0B52F6EAE0D37AE0FE0DFEB5FE6A" bold="true" box="[199,334,397,423]" pageId="129" pageNumber="130">Dimension.</emphasis>
Fore wing length of female
<quantity id="4CC27AA5F6EAE0D378AAFE0DFD1DFE65" box="[653,742,397,424]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="129" pageNumber="130" unit="mm" value="4.3">4.3 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C32084CBF6EAE0D37AE0FE30FEAAFD9A" pageId="129" pageNumber="130" type="distribution">
<paragraph id="8B85D740F6EAE0D37AE0FE30FE07FE07" blockId="129.[151,1436,151,599]" box="[199,508,432,459]" pageId="129" pageNumber="130">
<emphasis id="B94E0B52F6EAE0D37AE0FE30FE9AFE07" bold="true" box="[199,353,432,458]" pageId="129" pageNumber="130">Distribution.</emphasis>
New
<collectingCountry id="F32D97D0F6EAE0D37B84FE30FE03FE07" box="[419,504,432,458]" name="Guinea-Bissau" pageId="129" pageNumber="130">Guinea</collectingCountry>
.
</paragraph>
<paragraph id="8B85D740F6EAE0D37AE0FE53FEAAFD9A" blockId="129.[151,1436,151,599]" pageId="129" pageNumber="130">
<emphasis id="B94E0B52F6EAE0D37AE0FE53FE99FE20" bold="true" box="[199,354,467,493]" pageId="129" pageNumber="130">Comparison.</emphasis>
<taxonomicName id="4C3AACC3F6EAE0D37B4FFE53FD06FE20" authority="Kluge, 2021" authorityName="Kluge" authorityYear="2021" box="[360,765,467,494]" class="Insecta" family="Baetidae" genus="Centroptella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="129" pageNumber="130" phylum="Arthropoda" rank="species" species="papuanica" status="sp. nov." subGenus="Chopralla">
<emphasis id="B94E0B52F6EAE0D37B4FFE53FE0EFE20" box="[360,501,467,493]" italics="true" pageId="129" pageNumber="130">Centroptella</emphasis>
(
<emphasis id="B94E0B52F6EAE0D37823FE53FD8CFE20" box="[516,631,467,493]" italics="true" pageId="129" pageNumber="130">Chopralla</emphasis>
)
<emphasis id="B94E0B52F6EAE0D378A0FE54FD06FE20" box="[647,765,468,493]" italics="true" pageId="129" pageNumber="130">papuanica</emphasis>
</taxonomicName>
<emphasis id="B94E0B52F6EAE0D37924FE54FCB9FE23" bold="true" box="[771,834,468,494]" pageId="129" pageNumber="130">
<taxonomicNameLabel id="A27DB629F6EAE0D37924FE54FCB9FE23" box="[771,834,468,494]" pageId="129" pageNumber="130" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
differs from other species of
<taxonomicName id="4C3AACC3F6EAE0D37EA0FE53FB01FE20" authorityName="Waltz &amp; McCafferty" authorityYear="1987" box="[1159,1274,467,493]" class="Insecta" family="Baetidae" genus="Chopralla" kingdom="Animalia" order="Ephemeroptera" pageId="129" pageNumber="130" phylum="Arthropoda" rank="genus">
<emphasis id="B94E0B52F6EAE0D37EA0FE53FB01FE20" box="[1159,1274,467,493]" italics="true" pageId="129" pageNumber="130">Chopralla</emphasis>
</taxonomicName>
in structure of right prostheca, which is pressed to kinetodontium and lacks long branches (
<figureCitation id="1301CBC5F6EAE0D379D2FE77FBA3FDDF" box="[1013,1112,503,530]" captionStart="FIGURES 607–610" captionStartId="125.[151,264,1430,1454]" captionTargetBox="[177,1421,191,1392]" captionTargetId="figure-20@125.[164,1423,181,1406]" captionTargetPageId="125" captionText="FIGURES 607–610. Centroptella (Chopralla) papuanica sp. n. 607–608, left and right mandibles; 609, subimaginal gonostyli crumpled under larval cuticle; 610, larval abdominal segment X without caudalii (holotype). Abbreviations: gs1–gs3, segments of gonostyli; m.gs, gonostylar muscle; m.sg, styligeral muscle; pgs, larval protogonostylus; usg, unistyliger." figureDoi="http://doi.org/10.5281/zenodo.5577624" httpUri="https://zenodo.org/record/5577624/files/figure.png" pageId="129" pageNumber="130">Fig. 608</figureCitation>
). The same structure of right prostheca occurs in all
<taxonomicName id="4C3AACC3F6EAE0D37BBBFD9AFDF7FDF9" box="[412,524,538,564]" class="Insecta" family="Baetidae" genus="Crassolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="129" pageNumber="130" phylum="Arthropoda" rank="genus">
<emphasis id="B94E0B52F6EAE0D37BBBFD9AFDF7FDF9" box="[412,524,538,564]" italics="true" pageId="129" pageNumber="130">Crassolus</emphasis>
</taxonomicName>
, from which
<taxonomicName id="4C3AACC3F6EAE0D37882FD9AFBC6FDF9" authority="Kluge, 2021" authorityName="Kluge" authorityYear="2021" box="[677,1085,538,564]" class="Insecta" family="Baetidae" genus="Centroptella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="129" pageNumber="130" phylum="Arthropoda" rank="species" species="papuanica" status="sp. nov." subGenus="Chopralla">
<emphasis id="B94E0B52F6EAE0D37882FD9AFCC9FDF9" box="[677,818,538,564]" italics="true" pageId="129" pageNumber="130">Centroptella</emphasis>
(
<emphasis id="B94E0B52F6EAE0D37964FD9AFC4DFDF9" box="[835,950,538,564]" italics="true" pageId="129" pageNumber="130">Chopralla</emphasis>
)
<emphasis id="B94E0B52F6EAE0D379E0FD9BFBC6FDF9" box="[967,1085,539,564]" italics="true" pageId="129" pageNumber="130">papuanica</emphasis>
</taxonomicName>
<emphasis id="B94E0B52F6EAE0D37E62FD9AFB7EFDF9" bold="true" box="[1093,1157,538,564]" pageId="129" pageNumber="130">
<taxonomicNameLabel id="A27DB629F6EAE0D37E62FD9AFB7EFDF9" box="[1093,1157,538,564]" pageId="129" pageNumber="130" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
differs by the characters listed in
<tableCitation id="C6B8E2FBF6EAE0D37AD0FDBDFEAAFD9A" box="[247,337,573,599]" captionStart-0="TABLE 2" captionStart-1="TABLE 3" captionStartId-0="14.[151,239,1232,1256]" captionStartId-1="16.[151,239,151,175]" captionTargetPageId-0="14" captionTargetPageId-1="16" captionText-0="TABLE 2. Characters of Centroptella s. str., Crassolus and Chopralla (autapomorphies are marked with bold font and asterisk." captionText-1="TABLE 3. Some species-specific characters of Centroptella s. l." httpUri-0="http://table.plazi.org/id/DF4587C8F665E05C7AB0FB50FF0BFAC1" httpUri-1="http://table.plazi.org/id/DF4587C8F67BE0427AB0FF17FCBEFF62" pageId="129" pageNumber="130" tableUuid-0="DF4587C8F665E05C7AB0FB50FF0BFAC1" tableUuid-1="DF4587C8F67BE0427AB0FF17FCBEFF62">Table 2.</tableCitation>
</paragraph>
</subSubSection>
</treatment>
</document>
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