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<document id="956278B06B0B332BC67E461E419B5EE7" ID-DOI="10.11646/zootaxa.4810.2.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="3938177" ID-ZooBank="16464D12-3BF0-4345-B3CB-B2581E1C37B5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1594366343418" checkinUser="plazi" docAuthor="Tan, Ming Kai, Japir, Razy, Chung, Arthur Y. C. &amp; Wahab, Rodzay Bin Haji Abdul" docDate="2020" docId="0390913BA91BFF8BFF0C254CFD8BF92C" docLanguage="en" docName="zootaxa.4810.2.2.pdf" docOrigin="Zootaxa 4810 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Itara (Bornitara) spinosa Tan &amp; Japir &amp; Chung &amp; Wahab 2020, sp. nov." docType="treatment" docVersion="4" lastPageNumber="259" masterDocId="FFA9E943A910FF84FF9B227DFF8EFFC8" masterDocTitle="New taxa of crickets (Orthoptera: Grylloidea: Phaloriinae, Phalangopsinae Itarinae and Podoscirtinae) from Borneo (Brunei Darussalam and Sandakan)" masterLastPageNumber="270" masterPageNumber="244" pageNumber="255" updateTime="1698779676340" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="6F69B3705AFEED8F2BE8A222AB660AF4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6CE45737125C3F86B902B7AA1DBCF834">
<mods:title id="6C774A2B9171589E2D991B095298C655">New taxa of crickets (Orthoptera: Grylloidea: Phaloriinae, Phalangopsinae Itarinae and Podoscirtinae) from Borneo (Brunei Darussalam and Sandakan)</mods:title>
</mods:titleInfo>
<mods:name id="2F1D35C00A6AA03E7A3DC9BE2F6C9FEA" type="personal">
<mods:role id="63632D52EC687EE9D8937F21D436159B">
<mods:roleTerm id="DB175BA36ABA56297304F5D02D993B3C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1C5F60210DAA1957C01839EDC3EE392C">Tan, Ming Kai</mods:namePart>
<mods:affiliation id="00358CFD5D895B840B56FC46F8A1CA63">Institut de Systématique, Evolution et Biodiversité, Muséum national d’Histoire naturelle, CNRS, SU, EPHE, UA, 57 rue Cuvier, CP 50, 75231 Paris Cedex 05, France</mods:affiliation>
</mods:name>
<mods:name id="79C5876763E011446298EAE50265EE03" type="personal">
<mods:role id="F327E38291F7B723440B6FC0C0046968">
<mods:roleTerm id="E55F35DD971607FAF655DA63E8DD95F1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="72BCBC8FD429F07B69A5E5E8394C5C17">Japir, Razy</mods:namePart>
<mods:affiliation id="D3F3323965BAF796DD94C090F8982B7D">Forest Research Centre (Sepilok), Sabah Forestry Department, P. O. Box 1407, 90715 Sandakan, Sabah. Razy. Japir @ sabah. gov. my (RJ)</mods:affiliation>
</mods:name>
<mods:name id="59809587305DE59C4D2A36C19D243610" type="personal">
<mods:role id="9C3BB009F7080E48CF65675962EC326B">
<mods:roleTerm id="3931EF6845F34401B016737D5FAA7003">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1E19B453E7F2250301FEF067FF0D15E7">Chung, Arthur Y. C.</mods:namePart>
<mods:affiliation id="FD0892B3AF6E2550D0647DD6D8F1FB07">Forest Research Centre (Sepilok), Sabah Forestry Department, P. O. Box 1407, 90715 Sandakan, Sabah. Razy. Japir @ sabah. gov. my (RJ)</mods:affiliation>
</mods:name>
<mods:name id="F7301EF897727283E3ADC81A8284A552" type="personal">
<mods:role id="EFD7CC8EA46190F541A168ED52B4A31E">
<mods:roleTerm id="41C911D7775A5BC32F5C8E891165A0B2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C2F77E2A3A57E3AB80230CAC304451D6">Wahab, Rodzay Bin Haji Abdul</mods:namePart>
<mods:affiliation id="1B0BAEBB668F82D069A47F847B3285CC">Institute for Biodiversity and Environmental Research, Universiti Brunei Darussalam, Jalan Universiti, BE 1410, Brunei Darussalam. rodzay. wahab @ ubd. edu. bn; https: // orcid / org / 0000 - 0002 - 2151 - 7709</mods:affiliation>
</mods:name>
<mods:typeOfResource id="73101773E9AA5DDBEB4374A925A48215">text</mods:typeOfResource>
<mods:relatedItem id="6D90650A59C17502138022A0B9710EDD" type="host">
<mods:titleInfo id="27B1D6003B5359EF13B51393B2DD7601">
<mods:title id="BD424835D78E999A1F884C3B84F913F6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="6ABAE146C4A9EB7B8AF4E41B58830013">
<mods:date id="F19627EE0D2041C176C4B89316F5C000">2020</mods:date>
<mods:detail id="54A21D10D618E855A0C0E06A3A6FF049" type="pubDate">
<mods:number id="5B136327803BFAF09A18C9D56A964A61">2020-07-10</mods:number>
</mods:detail>
<mods:detail id="8A85CB4181CA9C87BBAB791525AF7681" type="volume">
<mods:number id="7135AECCA3C184B9AD3C3237947BA646">4810</mods:number>
</mods:detail>
<mods:detail id="2B1555CDDF42AA2D39C3F4952EA70AF0" type="issue">
<mods:number id="059A4948D3A90AEF7850931204CFD894">2</mods:number>
</mods:detail>
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<mods:start id="E99153CB7D6A853E2CFB5A03C916E575">244</mods:start>
<mods:end id="C4114E0FD0BBC304518CFBEB16398EF5">270</mods:end>
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<mods:classification id="42EC4A5479DB5D1BFC25650C22B88691">journal article</mods:classification>
<mods:identifier id="C17C59D4086AD9F117AC48AD48586F0F" type="DOI">10.11646/zootaxa.4810.2.2</mods:identifier>
<mods:identifier id="0D36D380DD86BB21C94336BF072F1AEB" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E5303BEE3B25E4C163B124AC74EA4A5D" type="Zenodo-Dep">3938177</mods:identifier>
<mods:identifier id="8431611DC4711CC88838C3675F008157" type="ZooBank">16464D12-3BF0-4345-B3CB-B2581E1C37B5</mods:identifier>
</mods:mods>
<treatment id="0390913BA91BFF8BFF0C254CFD8BF92C" ID-DOI="http://doi.org/10.5281/zenodo.4332663" ID-Zenodo-Dep="4332663" LSID="urn:lsid:plazi:treatment:0390913BA91BFF8BFF0C254CFD8BF92C" httpUri="http://treatment.plazi.org/id/0390913BA91BFF8BFF0C254CFD8BF92C" lastPageId="15" lastPageNumber="259" pageId="11" pageNumber="255">
<subSubSection id="C32373A6A91BFF8FFF0C254CFD65F882" box="[151,747,1840,1867]" pageId="11" pageNumber="255" type="nomenclature">
<paragraph id="8B86202DA91BFF8FFF0C254CFD65F882" blockId="11.[151,747,1840,1902]" box="[151,747,1840,1867]" pageId="11" pageNumber="255">
<heading id="D0CE9741A91BFF8FFF0C254CFD65F882" bold="true" box="[151,747,1840,1867]" fontSize="11" level="1" pageId="11" pageNumber="255" reason="1">
<emphasis id="B94DFC3FA91BFF8FFF0C254CFD65F882" bold="true" box="[151,747,1840,1867]" pageId="11" pageNumber="255">
<taxonomicName id="4C395BAEA91BFF8FFF0C254CFD0EF882" authority="Tan &amp; Wahab" authorityName="Tan &amp; Japir &amp; Chung &amp; Wahab" authorityYear="2020" box="[151,640,1840,1867]" class="Insecta" family="Haglotettigoniidae" genus="Itara" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="255" phylum="Arthropoda" rank="species" species="spinosa" status="sp. nov." subGenus="Bornitara">
<emphasis id="B94DFC3FA91BFF8FFF0C254CFF5DF883" bold="true" box="[151,211,1841,1867]" italics="true" pageId="11" pageNumber="255">Itara</emphasis>
(
<emphasis id="B94DFC3FA91BFF8FFF7E254DFED2F882" bold="true" box="[229,348,1840,1866]" italics="true" pageId="11" pageNumber="255">Bornitara</emphasis>
)
<emphasis id="B94DFC3FA91BFF8FFEF6254DFE46F882" bold="true" box="[365,456,1840,1866]" italics="true" pageId="11" pageNumber="255">spinosa</emphasis>
Tan &amp; Wahab
</taxonomicName>
,
<taxonomicNameLabel id="A27E4144A91BFF8FFD15254DFD65F882" box="[654,747,1840,1866]" pageId="11" pageNumber="255" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32373A6A91BFF8FFF0C2529FE91F8A6" box="[151,287,1876,1902]" pageId="11" pageNumber="255" type="description">
<paragraph id="8B86202DA91BFF8FFF0C2529FE91F8A6" blockId="11.[151,747,1840,1902]" box="[151,287,1876,1902]" pageId="11" pageNumber="255">
(
<figureCitation id="13023CA8A91BFF8FFF042529FE99F8A6" box="[159,279,1876,1902]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStart-2="FIGURE 11" captionStartId-0="12.[151,250,1945,1970]" captionStartId-1="13.[151,250,1944,1969]" captionStartId-2="14.[151,250,1403,1428]" captionTargetBox-0="[179,1408,208,1920]" captionTargetBox-1="[371,1210,181,1905]" captionTargetBox-2="[151,1436,181,1377]" captionTargetId-0="figure@12.[179,1408,181,1920]" captionTargetId-1="figure@13.[359,1229,181,1919]" captionTargetId-2="figure@14.[151,1436,181,1377]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="14" captionText-0="FIGURE 9. Itara (Bornitara) spinosa sp. nov. male holotype habitus in dorsal view (A), head and pronotum in dorsal (B) and lateral (C) views, face in anterior view (D) and metanotal gland in dorsal view (E). Scale bars: 1 mm." captionText-1="FIGURE 10. Itara (Bornitara) spinosa sp. nov. male holotype tegmen in dorsal view. Scale bar: 5 mm." captionText-2="FIGURE 11. Itara (Bornitara) spinosa sp. nov. male holotype abdominal apex in dorsal view (A), genitalia in lateral (B), dorsal (C) and ventral (D) views. Scale bar: 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3938197" figureDoi-1="http://doi.org/10.5281/zenodo.3938199" figureDoi-2="http://doi.org/10.5281/zenodo.3938201" httpUri-0="https://zenodo.org/record/3938197/files/figure.png" httpUri-1="https://zenodo.org/record/3938199/files/figure.png" httpUri-2="https://zenodo.org/record/3938201/files/figure.png" pageId="11" pageNumber="255">Figs. 9–11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32373A6A91BFF8FFF0C25E6FCAAF812" pageId="11" pageNumber="255" type="materials_examined">
<paragraph id="8B86202DA91BFF8FFF0C25E6FCAAF812" blockId="11.[151,1436,1947,2011]" pageId="11" pageNumber="255">
<emphasis id="B94DFC3FA91BFF8FFF0C25E6FE0EF87E" bold="true" box="[151,384,1947,1974]" pageId="11" pageNumber="255">Material examined.</emphasis>
<materialsCitation id="3B512A70A91BFF8FFE1D25E1FCAEF812" collectingDate="2019-07-12" collectionCode="UBDM" collectorName="M. K. Tan &amp; H. Yeo" location="Kuala Belalong Field Studies" municipality="Male" pageId="11" pageNumber="255" specimenCount="1" stateProvince="Temburong" typeStatus="holotype">
<collectingMunicipality id="6BE2BA57A91BFF8FFE1D25E1FE4EF87E" box="[390,448,1948,1974]" pageId="11" pageNumber="255">Male</collectingMunicipality>
<typeStatus id="54829E8FA91BFF8FFE5D25E1FDA6F87E" box="[454,552,1948,1974]" pageId="11" pageNumber="255" type="holotype">holotype</typeStatus>
(KB.19.9),
<collectingRegion id="49FDEECFA91BFF8FFD3625E1FC05F87E" box="[685,907,1948,1974]" country="Brunei" name="Temburong" pageId="11" pageNumber="255">Temburong District</collectingRegion>
,
<location id="8EE676F6A91BFF8FFC0D25E1FB53F87E" LSID="urn:lsid:plazi:treatment:0390913BA91BFF8BFF0C254CFD8BF92C:8EE676F6A91BFF8FFC0D25E1FB53F87E" box="[918,1245,1948,1974]" municipality="Male" name="Kuala Belalong Field Studies" pageId="11" pageNumber="255" stateProvince="Temburong">Kuala Belalong Field Studies</location>
<collectingRegion id="49FDEECFA91BFF8FFB7F25E1FABCF87E" box="[1252,1330,1948,1974]" country="France" name="Centre" pageId="11" pageNumber="255">Centre</collectingRegion>
, attracted to light,
<date id="FF8706EDA91BFF8FFF6E25BDFE07F812" box="[245,393,1984,2010]" pageId="11" pageNumber="255" value="2019-07-12">
<collectingDate id="EFC3FF05A91BFF8FFF6E25BDFE07F812" box="[245,393,1984,2010]" pageId="11" pageNumber="255" value="2019-07-12">12 July 2019</collectingDate>
</date>
, coll.
<collectorName id="26CC45FBA91BFF8FFE5725BCFDB1F813" box="[460,575,1985,2011]" pageId="11" pageNumber="255">M. K. Tan</collectorName>
&amp;
<collectorName id="26CC45FBA91BFF8FFDF925BCFD21F813" box="[610,687,1985,2011]" pageId="11" pageNumber="255">H. Yeo</collectorName>
(
<collectionCode id="ED28B8E8A91BFF8FFD2525BDFC96F812" box="[702,792,1984,2010]" pageId="11" pageNumber="255">UBDM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF4670A5A91CFF88FF0C25E4FB15F81D" ID-DOI="http://doi.org/10.5281/zenodo.3938197" ID-Zenodo-Dep="3938197" httpUri="https://zenodo.org/record/3938197/files/figure.png" pageId="12" pageNumber="256" startId="12.[151,250,1945,1970]" targetBox="[179,1408,208,1920]" targetPageId="12">
<paragraph id="8B86202DA91CFF88FF0C25E4FB15F81D" blockId="12.[151,1437,1945,2006]" pageId="12" pageNumber="256">
<emphasis id="B94DFC3FA91CFF88FF0C25E4FE9DF87A" bold="true" box="[151,275,1945,1970]" pageId="12" pageNumber="256">FIGURE 9.</emphasis>
<taxonomicName id="4C395BAEA91CFF88FE8125E7FD95F879" authority="Tan &amp; Japir &amp; Chung &amp; Wahab, 2020" authorityName="Tan &amp; Japir &amp; Chung &amp; Wahab" authorityYear="2020" box="[282,539,1946,1970]" class="Insecta" family="Haglotettigoniidae" genus="Itara" kingdom="Animalia" order="Orthoptera" pageId="12" pageNumber="256" phylum="Arthropoda" rank="species" species="spinosa" status="sp. nov." subGenus="Bornitara">
<emphasis id="B94DFC3FA91CFF88FE8125E7FEC3F879" box="[282,333,1946,1969]" italics="true" pageId="12" pageNumber="256">Itara</emphasis>
(
<emphasis id="B94DFC3FA91CFF88FEC025E7FE31F879" box="[347,447,1946,1969]" italics="true" pageId="12" pageNumber="256">Bornitara</emphasis>
)
<emphasis id="B94DFC3FA91CFF88FE5525E7FD95F879" box="[462,539,1946,1969]" italics="true" pageId="12" pageNumber="256">spinosa</emphasis>
</taxonomicName>
<emphasis id="B94DFC3FA91CFF88FDB925E7FDFDF87A" bold="true" box="[546,627,1946,1970]" pageId="12" pageNumber="256">
<taxonomicNameLabel id="A27E4144A91CFF88FDB925E7FDFDF87A" box="[546,627,1946,1970]" pageId="12" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male holotype habitus in dorsal view (A), head and pronotum in dorsal (B) and lateral (C) views, face in anterior view (D) and metanotal gland in dorsal view (E). Scale bars: 1 mm.
</paragraph>
</caption>
<caption id="DF4670A5A91DFF89FF0C25E5FB3AF878" ID-DOI="http://doi.org/10.5281/zenodo.3938199" ID-Zenodo-Dep="3938199" box="[151,1204,1944,1969]" httpUri="https://zenodo.org/record/3938199/files/figure.png" pageId="13" pageNumber="257" startId="13.[151,250,1944,1969]" targetBox="[371,1210,181,1905]" targetPageId="13">
<paragraph id="8B86202DA91DFF89FF0C25E5FB3AF878" blockId="13.[151,1204,1944,1969]" box="[151,1204,1944,1969]" pageId="13" pageNumber="257">
<emphasis id="B94DFC3FA91DFF89FF0C25E5FEAEF879" bold="true" box="[151,288,1944,1969]" pageId="13" pageNumber="257">FIGURE 10.</emphasis>
<taxonomicName id="4C395BAEA91DFF89FEBD25E4FDA9F878" authority="Tan &amp; Japir &amp; Chung &amp; Wahab, 2020" authorityName="Tan &amp; Japir &amp; Chung &amp; Wahab" authorityYear="2020" box="[294,551,1945,1969]" class="Insecta" family="Haglotettigoniidae" genus="Itara" kingdom="Animalia" order="Orthoptera" pageId="13" pageNumber="257" phylum="Arthropoda" rank="species" species="spinosa" status="sp. nov." subGenus="Bornitara">
<emphasis id="B94DFC3FA91DFF89FEBD25E4FED7F878" box="[294,345,1945,1968]" italics="true" pageId="13" pageNumber="257">Itara</emphasis>
(
<emphasis id="B94DFC3FA91DFF89FEFC25E4FE45F878" box="[359,459,1945,1968]" italics="true" pageId="13" pageNumber="257">Bornitara</emphasis>
)
<emphasis id="B94DFC3FA91DFF89FE4125E4FDA9F878" box="[474,551,1945,1968]" italics="true" pageId="13" pageNumber="257">spinosa</emphasis>
</taxonomicName>
<emphasis id="B94DFC3FA91DFF89FDB525E4FDF0F879" bold="true" box="[558,638,1945,1969]" pageId="13" pageNumber="257">
<taxonomicNameLabel id="A27E4144A91DFF89FDB525E4FDF0F879" box="[558,638,1945,1969]" pageId="13" pageNumber="257" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male holotype tegmen in dorsal view. Scale bar: 5 mm.
</paragraph>
</caption>
<caption id="DF4670A5A91EFF8AFF0C2706FDDBFA7F" ID-DOI="http://doi.org/10.5281/zenodo.3938201" ID-Zenodo-Dep="3938201" httpUri="https://zenodo.org/record/3938201/files/figure.png" pageId="14" pageNumber="258" startId="14.[151,250,1403,1428]" targetBox="[151,1436,181,1377]" targetPageId="14">
<paragraph id="8B86202DA91EFF8AFF0C2706FDDBFA7F" blockId="14.[151,1436,1403,1464]" pageId="14" pageNumber="258">
<emphasis id="B94DFC3FA91EFF8AFF0C2706FE93FA5C" bold="true" box="[151,285,1403,1428]" pageId="14" pageNumber="258">FIGURE 11.</emphasis>
<taxonomicName id="4C395BAEA91EFF8AFEB92701FDAFFA5B" authority="Tan &amp; Japir &amp; Chung &amp; Wahab, 2020" authorityName="Tan &amp; Japir &amp; Chung &amp; Wahab" authorityYear="2020" box="[290,545,1404,1428]" class="Insecta" family="Haglotettigoniidae" genus="Itara" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="258" phylum="Arthropoda" rank="species" species="spinosa" status="sp. nov." subGenus="Bornitara">
<emphasis id="B94DFC3FA91EFF8AFEB92701FEDBFA5B" box="[290,341,1404,1427]" italics="true" pageId="14" pageNumber="258">Itara</emphasis>
(
<emphasis id="B94DFC3FA91EFF8AFEF82701FE49FA5B" box="[355,455,1404,1427]" italics="true" pageId="14" pageNumber="258">Bornitara</emphasis>
)
<emphasis id="B94DFC3FA91EFF8AFE4F2701FDAFFA5B" box="[468,545,1404,1427]" italics="true" pageId="14" pageNumber="258">spinosa</emphasis>
</taxonomicName>
<emphasis id="B94DFC3FA91EFF8AFDBC2701FDF8FA5C" bold="true" box="[551,630,1404,1428]" pageId="14" pageNumber="258">
<taxonomicNameLabel id="A27E4144A91EFF8AFDBC2701FDF8FA5C" box="[551,630,1404,1428]" pageId="14" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
male holotype abdominal apex in dorsal view (A), genitalia in lateral (B), dorsal (C) and ventral (D) views. Scale bar: 1 mm.
</paragraph>
</caption>
<subSubSection id="C32373A6A91EFF8AFF5C2798FB93F831" pageId="14" pageNumber="258" type="diagnosis">
<paragraph id="8B86202DA91EFF8AFF5C2798FCFBF9A4" blockId="14.[151,1437,1509,2041]" pageId="14" pageNumber="258">
<emphasis id="B94DFC3FA91EFF8AFF5C2798FECCF9C8" bold="true" box="[199,322,1509,1536]" pageId="14" pageNumber="258">Diagnosis.</emphasis>
This new species differs from congeners by epiphallus fairly broad, with a distinct dorsal spine (ds) at middle of epiphallus, with large ventral spine (vs) and fairly long and slightly hooked apex (ap), ectoparameres (ec) with distinctly shaped anterior half and long and cylindrical posterior half; and tegminal venation with welldeveloped additional vein connecting all oblique veins near CuA.
</paragraph>
<paragraph id="8B86202DA91EFF8AFF5C2408FB93F831" blockId="14.[151,1437,1509,2041]" pageId="14" pageNumber="258">
<emphasis id="B94DFC3FA91EFF8AFF5C2408FD91F958" bold="true" box="[199,543,1653,1680]" pageId="14" pageNumber="258">Comparison with congeners.</emphasis>
The new species resembles
<taxonomicName id="4C395BAEA91EFF8AFCFF240AFB66F958" authority="Gorochov, 1997" authorityName="Gorochov" authorityYear="1997" box="[868,1256,1654,1680]" class="Insecta" family="Haglotettigoniidae" genus="Itara" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="255" phylum="Arthropoda" rank="species" species="sabahensis" subGenus="Bornitara">
<emphasis id="B94DFC3FA91EFF8AFCFF240AFCE3F958" box="[868,877,1655,1680]" italics="true" pageId="14" pageNumber="258">I</emphasis>
. (
<emphasis id="B94DFC3FA91EFF8AFC1D240AFC19F958" box="[902,919,1655,1680]" italics="true" pageId="14" pageNumber="258">B</emphasis>
.)
<emphasis id="B94DFC3FA91EFF8AFC2B240BFBA5F958" box="[944,1067,1654,1680]" italics="true" pageId="14" pageNumber="258">sabahensis</emphasis>
<bibRefCitation id="EFA85DDCA91EFF8AFBA8240BFB66F958" author="Gorochov, A. V." box="[1075,1256,1654,1680]" pageId="14" pageNumber="258" pagination="47 - 75" refId="ref8541" refString="Gorochov, A. V. (1997) Partial revision of the subfamily Itarinae (Orthoptera: Gryllidae). Zoosystematica Rossica, 6 (1 - 2), 47 - 75." type="journal article" year="1997">Gorochov, 1997</bibRefCitation>
</taxonomicName>
from Sandakan (
<collectingRegion id="49FDEECFA91EFF8AFF0424E7FF67F97D" box="[159,233,1690,1717]" country="Malaysia" name="Sabah" pageId="14" pageNumber="258">Sabah</collectingRegion>
), as well as
<taxonomicName id="4C395BAEA91EFF8AFEEB24E6FD5BF97C" authority="Chopard, 1930" authorityName="Chopard" authorityYear="1930" box="[368,725,1690,1716]" class="Insecta" family="Haglotettigoniidae" genus="Itara" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="258" phylum="Arthropoda" rank="species" species="latipennis" subGenus="Bornitara">
<emphasis id="B94DFC3FA91EFF8AFEEB24E6FE0FF97C" box="[368,385,1691,1716]" italics="true" pageId="14" pageNumber="258">I.</emphasis>
(
<emphasis id="B94DFC3FA91EFF8AFE0A24E6FE27F97C" box="[401,425,1691,1716]" italics="true" pageId="14" pageNumber="258">B.</emphasis>
)
<emphasis id="B94DFC3FA91EFF8AFE2224E7FDA9F97C" box="[441,551,1690,1716]" italics="true" pageId="14" pageNumber="258">latipennis</emphasis>
Chopard, 1930
</taxonomicName>
and
<taxonomicName id="4C395BAEA91EFF8AFC9024E6FBE5F97C" authority="Gorochov, 2012" authorityName="Gorochov" authorityYear="2012" box="[779,1131,1690,1716]" class="Insecta" family="Haglotettigoniidae" genus="Itara" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="258" phylum="Arthropoda" rank="species" species="matangi" subGenus="Bornitara">
<emphasis id="B94DFC3FA91EFF8AFC9024E6FC92F97C" box="[779,796,1691,1716]" italics="true" pageId="14" pageNumber="258">I.</emphasis>
(
<emphasis id="B94DFC3FA91EFF8AFCB724E6FCCAF97C" box="[812,836,1691,1716]" italics="true" pageId="14" pageNumber="258">B.</emphasis>
)
<emphasis id="B94DFC3FA91EFF8AFCCF24E6FC3EF97C" box="[852,944,1691,1716]" italics="true" pageId="14" pageNumber="258">matangi</emphasis>
Gorochov, 2012
</taxonomicName>
most, both from the southern part of
<collectingCountry id="F32E60BDA91EFF8AFE8D24C3FEF8F910" box="[278,374,1726,1752]" name="Malaysia" pageId="14" pageNumber="258">Sarawak</collectingCountry>
relatively far from
<collectingCountry id="F32E60BDA91EFF8AFDCE24C3FCBEF910" box="[597,816,1726,1752]" name="Brunei" pageId="14" pageNumber="258">Brunei Darussalam</collectingCountry>
, by shape of male genitalia and tegminal venation; but differs from
<taxonomicName id="4C395BAEA91EFF8AFEBD249EFE51F934" authorityName="Chopard" authorityYear="1930" box="[294,479,1762,1788]" class="Insecta" family="Haglotettigoniidae" genus="Itara" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="258" phylum="Arthropoda" rank="species" species="latipennis" subGenus="Bornitara">
<emphasis id="B94DFC3FA91EFF8AFEBD249EFEB9F934" box="[294,311,1763,1788]" italics="true" pageId="14" pageNumber="258">I.</emphasis>
(
<emphasis id="B94DFC3FA91EFF8AFED3249EFEEEF934" box="[328,352,1763,1788]" italics="true" pageId="14" pageNumber="258">B.</emphasis>
)
<emphasis id="B94DFC3FA91EFF8AFEEA249FFE51F934" box="[369,479,1762,1788]" italics="true" pageId="14" pageNumber="258">latipennis</emphasis>
</taxonomicName>
by ventral spine (vs) on epiphallus longer in profile (more similar to
<taxonomicName id="4C395BAEA91EFF8AFB6D249EFA12F934" authorityName="Gorochov" authorityYear="2012" box="[1270,1436,1762,1788]" class="Insecta" family="Haglotettigoniidae" genus="Itara" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="258" phylum="Arthropoda" rank="species" species="matangi" subGenus="Bornitara">
<emphasis id="B94DFC3FA91EFF8AFB6D249EFA89F934" box="[1270,1287,1763,1788]" italics="true" pageId="14" pageNumber="258">I.</emphasis>
(
<emphasis id="B94DFC3FA91EFF8AFA83249EFABEF934" box="[1304,1328,1763,1788]" italics="true" pageId="14" pageNumber="258">B.</emphasis>
)
<emphasis id="B94DFC3FA91EFF8AFADB249EFA12F934" box="[1344,1436,1763,1788]" italics="true" pageId="14" pageNumber="258">matangi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C395BAEA91EFF8AFF53257AFE1EF8E8" authorityName="Gorochov" authorityYear="1997" box="[200,400,1798,1824]" class="Insecta" family="Haglotettigoniidae" genus="Itara" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="258" phylum="Arthropoda" rank="species" species="sabahensis" subGenus="Bornitara">
<emphasis id="B94DFC3FA91EFF8AFF53257AFE1EF8E8" box="[200,400,1798,1824]" italics="true" pageId="14" pageNumber="258">I. (B.) sabahensis</emphasis>
</taxonomicName>
), apex of epiphallus more strongly hooked (ap) in profile (similar to
<taxonomicName id="4C395BAEA91EFF8AFB30257AFADCF8E8" authorityName="Gorochov" authorityYear="2012" box="[1195,1362,1798,1824]" class="Insecta" family="Haglotettigoniidae" genus="Itara" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="258" phylum="Arthropoda" rank="species" species="matangi" subGenus="Bornitara">
<emphasis id="B94DFC3FA91EFF8AFB30257AFB32F8E8" box="[1195,1212,1799,1824]" italics="true" pageId="14" pageNumber="258">I.</emphasis>
(
<emphasis id="B94DFC3FA91EFF8AFB55257AFB68F8E8" box="[1230,1254,1799,1824]" italics="true" pageId="14" pageNumber="258">B.</emphasis>
)
<emphasis id="B94DFC3FA91EFF8AFB6D257AFADCF8E8" box="[1270,1362,1799,1824]" italics="true" pageId="14" pageNumber="258">matangi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C395BAEA91EFF8AFA17257AFECDF88C" authorityName="Gorochov" authorityYear="1997" class="Insecta" family="Haglotettigoniidae" genus="Itara" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="258" phylum="Arthropoda" rank="species" species="sabahensis" subGenus="Bornitara">
<emphasis id="B94DFC3FA91EFF8AFA17257AFECDF88C" italics="true" pageId="14" pageNumber="258">I. (B.) sabahensis</emphasis>
</taxonomicName>
) and anterior emargination (ae) narrower; from
<taxonomicName id="4C395BAEA91EFF8AFCC12556FC70F88C" authorityName="Gorochov" authorityYear="2012" box="[858,1022,1834,1860]" class="Insecta" family="Haglotettigoniidae" genus="Itara" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="258" phylum="Arthropoda" rank="species" species="matangi" subGenus="Bornitara">
<emphasis id="B94DFC3FA91EFF8AFCC12556FCE5F88C" box="[858,875,1835,1860]" italics="true" pageId="14" pageNumber="258">I.</emphasis>
(
<emphasis id="B94DFC3FA91EFF8AFCE12556FC1CF88C" box="[890,914,1835,1860]" italics="true" pageId="14" pageNumber="258">B.</emphasis>
)
<emphasis id="B94DFC3FA91EFF8AFC392556FC70F88C" box="[930,1022,1835,1860]" italics="true" pageId="14" pageNumber="258">matangi</emphasis>
</taxonomicName>
by posterior half of epiphallus not so strongly concave and narrow in dorsal view; differs from
<taxonomicName id="4C395BAEA91EFF8AFC832532FC55F8A0" authorityName="Gorochov" authorityYear="1997" box="[792,987,1870,1896]" class="Insecta" family="Haglotettigoniidae" genus="Itara" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="258" phylum="Arthropoda" rank="species" species="sabahensis" subGenus="Bornitara">
<emphasis id="B94DFC3FA91EFF8AFC832532FCAFF8A0" box="[792,801,1871,1896]" italics="true" pageId="14" pageNumber="258">I</emphasis>
. (
<emphasis id="B94DFC3FA91EFF8AFCA22532FCC4F8A0" box="[825,842,1871,1896]" italics="true" pageId="14" pageNumber="258">B</emphasis>
.)
<emphasis id="B94DFC3FA91EFF8AFCFB2533FC55F8A0" box="[864,987,1870,1896]" italics="true" pageId="14" pageNumber="258">sabahensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C395BAEA91EFF8AFC722532FB11F8A0" authorityName="Chopard" authorityYear="1930" box="[1001,1183,1870,1896]" class="Insecta" family="Haglotettigoniidae" genus="Itara" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="258" phylum="Arthropoda" rank="species" species="latipennis" subGenus="Bornitara">
<emphasis id="B94DFC3FA91EFF8AFC722532FC74F8A0" box="[1001,1018,1871,1896]" italics="true" pageId="14" pageNumber="258">I.</emphasis>
(
<emphasis id="B94DFC3FA91EFF8AFB922532FBAFF8A0" box="[1033,1057,1871,1896]" italics="true" pageId="14" pageNumber="258">B.</emphasis>
)
<emphasis id="B94DFC3FA91EFF8AFBAA2533FB11F8A0" box="[1073,1183,1870,1896]" italics="true" pageId="14" pageNumber="258">latipennis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C395BAEA91EFF8AFB4E2532FAF4F8A0" authorityName="Gorochov" authorityYear="2012" box="[1237,1402,1870,1896]" class="Insecta" family="Haglotettigoniidae" genus="Itara" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="258" phylum="Arthropoda" rank="species" species="matangi" subGenus="Bornitara">
<emphasis id="B94DFC3FA91EFF8AFB4E2532FB68F8A0" box="[1237,1254,1871,1896]" italics="true" pageId="14" pageNumber="258">I.</emphasis>
(
<emphasis id="B94DFC3FA91EFF8AFB6D2532FA80F8A0" box="[1270,1294,1871,1896]" italics="true" pageId="14" pageNumber="258">B.</emphasis>
)
<emphasis id="B94DFC3FA91EFF8AFA852532FAF4F8A0" box="[1310,1402,1871,1896]" italics="true" pageId="14" pageNumber="258">matangi</emphasis>
</taxonomicName>
by epiphallus with distinct dorsal spine (ds) at middle of epiphallus and ectoparameres (ec) posterior process slenderer and more elongated, touching each other at apex in ventral view. This new species also differs from
<taxonomicName id="4C395BAEA91EFF8AFB6025EAFEE9F81C" authority="Gorochov, 1997" authorityName="Gorochov" authorityYear="1997" class="Insecta" family="Haglotettigoniidae" genus="Itara" kingdom="Animalia" order="Orthoptera" pageId="14" pageNumber="258" phylum="Arthropoda" rank="species" species="chopardi" subGenus="Bornitara">
<emphasis id="B94DFC3FA91EFF8AFB6025EAFA82F878" box="[1275,1292,1943,1968]" italics="true" pageId="14" pageNumber="258">I.</emphasis>
(
<emphasis id="B94DFC3FA91EFF8AFA8725EAFABAF878" box="[1308,1332,1943,1968]" italics="true" pageId="14" pageNumber="258">B.</emphasis>
)
<emphasis id="B94DFC3FA91EFF8AFADF25EBFF23F81C" italics="true" pageId="14" pageNumber="258">chopardi</emphasis>
<bibRefCitation id="EFA85DDCA91EFF8AFF2F25C7FEE9F81C" author="Gorochov, A. V." box="[180,359,1978,2004]" pageId="14" pageNumber="258" pagination="47 - 75" refId="ref8541" refString="Gorochov, A. V. (1997) Partial revision of the subfamily Itarinae (Orthoptera: Gryllidae). Zoosystematica Rossica, 6 (1 - 2), 47 - 75." type="journal article" year="1997">Gorochov, 1997</bibRefCitation>
</taxonomicName>
from Murud (not too far from
<collectingCountry id="F32E60BDA91EFF8AFD2625C7FC85F81D" box="[701,779,1978,2005]" name="Brunei" pageId="14" pageNumber="258">Brunei</collectingCountry>
) by presence of well-developed additional vein connecting all oblique veins near CuA (instead of interrupted near the longest oblique vein).
</paragraph>
</subSubSection>
<subSubSection id="C32373A6A91FFF8BFF5C22E5FE6FF953" pageId="15" pageNumber="259" type="description">
<paragraph id="8B86202DA91FFF8BFF5C22E5FBCBFD4F" blockId="15.[151,1437,152,1764]" pageId="15" pageNumber="259">
<emphasis id="B94DFC3FA91FFF8BFF5C22E5FED7FF7B" bold="true" box="[199,345,152,179]" pageId="15" pageNumber="259">Description.</emphasis>
Habitus very typical of the genus, relatively large and somewhat dorsoventrally compressed (
<figureCitation id="13023CA8A91FFF8BFAF422E4FF33FF1F" captionStart="FIGURE 9" captionStartId="12.[151,250,1945,1970]" captionTargetBox="[179,1408,208,1920]" captionTargetId="figure@12.[179,1408,181,1920]" captionTargetPageId="12" captionText="FIGURE 9. Itara (Bornitara) spinosa sp. nov. male holotype habitus in dorsal view (A), head and pronotum in dorsal (B) and lateral (C) views, face in anterior view (D) and metanotal gland in dorsal view (E). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3938197" httpUri="https://zenodo.org/record/3938197/files/figure.png" pageId="15" pageNumber="259">Fig. 9A</figureCitation>
). Head rounded, very finely pubescent; rostrum 1.7 times as wide as scapus, with apex truncated (in dorsal view) (
<figureCitation id="13023CA8A91FFF8BFF3B229CFF76FF33" box="[160,248,225,251]" captionStart="FIGURE 9" captionStartId="12.[151,250,1945,1970]" captionTargetBox="[179,1408,208,1920]" captionTargetId="figure@12.[179,1408,181,1920]" captionTargetPageId="12" captionText="FIGURE 9. Itara (Bornitara) spinosa sp. nov. male holotype habitus in dorsal view (A), head and pronotum in dorsal (B) and lateral (C) views, face in anterior view (D) and metanotal gland in dorsal view (E). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3938197" httpUri="https://zenodo.org/record/3938197/files/figure.png" pageId="15" pageNumber="259">Fig. 9B</figureCitation>
); scapus stout; maxillary palpi slender and elongated, with apical segment longest, obliquely clavate and apex truncated (
<figureCitation id="13023CA8A91FFF8BFED12378FE2CFED7" box="[330,418,261,287]" captionStart="FIGURE 9" captionStartId="12.[151,250,1945,1970]" captionTargetBox="[179,1408,208,1920]" captionTargetId="figure@12.[179,1408,181,1920]" captionTargetPageId="12" captionText="FIGURE 9. Itara (Bornitara) spinosa sp. nov. male holotype habitus in dorsal view (A), head and pronotum in dorsal (B) and lateral (C) views, face in anterior view (D) and metanotal gland in dorsal view (E). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3938197" httpUri="https://zenodo.org/record/3938197/files/figure.png" pageId="15" pageNumber="259">Fig. 9C</figureCitation>
); eyes oval in dorsal view (
<figureCitation id="13023CA8A91FFF8BFD492378FCA7FED7" box="[722,809,261,287]" captionStart="FIGURE 9" captionStartId="12.[151,250,1945,1970]" captionTargetBox="[179,1408,208,1920]" captionTargetId="figure@12.[179,1408,181,1920]" captionTargetPageId="12" captionText="FIGURE 9. Itara (Bornitara) spinosa sp. nov. male holotype habitus in dorsal view (A), head and pronotum in dorsal (B) and lateral (C) views, face in anterior view (D) and metanotal gland in dorsal view (E). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3938197" httpUri="https://zenodo.org/record/3938197/files/figure.png" pageId="15" pageNumber="259">Fig. 9B</figureCitation>
), oblong in lateral view (
<figureCitation id="13023CA8A91FFF8BFBDB2378FB16FED7" box="[1088,1176,261,287]" captionStart="FIGURE 9" captionStartId="12.[151,250,1945,1970]" captionTargetBox="[179,1408,208,1920]" captionTargetId="figure@12.[179,1408,181,1920]" captionTargetPageId="12" captionText="FIGURE 9. Itara (Bornitara) spinosa sp. nov. male holotype habitus in dorsal view (A), head and pronotum in dorsal (B) and lateral (C) views, face in anterior view (D) and metanotal gland in dorsal view (E). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3938197" httpUri="https://zenodo.org/record/3938197/files/figure.png" pageId="15" pageNumber="259">Fig. 9C</figureCitation>
); median ocellus round and small; lateral ocelli oblong and large, located near eyes (
<figureCitation id="13023CA8A91FFF8BFCDB2354FC15FE8B" box="[832,923,297,323]" captionStart="FIGURE 9" captionStartId="12.[151,250,1945,1970]" captionTargetBox="[179,1408,208,1920]" captionTargetId="figure@12.[179,1408,181,1920]" captionTargetPageId="12" captionText="FIGURE 9. Itara (Bornitara) spinosa sp. nov. male holotype habitus in dorsal view (A), head and pronotum in dorsal (B) and lateral (C) views, face in anterior view (D) and metanotal gland in dorsal view (E). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3938197" httpUri="https://zenodo.org/record/3938197/files/figure.png" pageId="15" pageNumber="259">Fig. 9D</figureCitation>
). Pronotal disc 1.8 times as wide as long, distinctly widening posteriorly (anterior margin 0.6 times as wide as posterior margin); finely pubescent with strong hairs along anterior and posterior margins; anterior margin of disc straight to slightly concave (when dried); posterior margin of disc slightly convex (
<figureCitation id="13023CA8A91FFF8BFDDA23E8FD14FE67" box="[577,666,405,431]" captionStart="FIGURE 9" captionStartId="12.[151,250,1945,1970]" captionTargetBox="[179,1408,208,1920]" captionTargetId="figure@12.[179,1408,181,1920]" captionTargetPageId="12" captionText="FIGURE 9. Itara (Bornitara) spinosa sp. nov. male holotype habitus in dorsal view (A), head and pronotum in dorsal (B) and lateral (C) views, face in anterior view (D) and metanotal gland in dorsal view (E). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3938197" httpUri="https://zenodo.org/record/3938197/files/figure.png" pageId="15" pageNumber="259">Fig. 9B</figureCitation>
). Pronotal lateral lobe 1.6 times as long as high, with ventral margin straight (
<figureCitation id="13023CA8A91FFF8BFF6223C4FEDEFE1B" box="[249,336,441,467]" captionStart="FIGURE 9" captionStartId="12.[151,250,1945,1970]" captionTargetBox="[179,1408,208,1920]" captionTargetId="figure@12.[179,1408,181,1920]" captionTargetPageId="12" captionText="FIGURE 9. Itara (Bornitara) spinosa sp. nov. male holotype habitus in dorsal view (A), head and pronotum in dorsal (B) and lateral (C) views, face in anterior view (D) and metanotal gland in dorsal view (E). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3938197" httpUri="https://zenodo.org/record/3938197/files/figure.png" pageId="15" pageNumber="259">Fig. 9C</figureCitation>
). Fore and middle legs generally pubescent (especially along ventral margin), with a few stout setae usually located along dorsal margin; tarsus with a row of stout setae on ventral surface. Fore tibia with distinctly swollen on inner side, with inner tympanum very elongated slit-like, with outer tympanum open and having large, elongate, oblong tympanal membrane. Hind femur finely pubescent, without ventral spines; hind tibia also pubescent, with about 8 small and stout denticules at anterior half on each dorsal side, with 5 long dorsal subapical spurs on inner and outer margins; hind basitarsus with about 6 inner and 6 outer denticles.
</paragraph>
<paragraph id="8B86202DA91FFF8BFF5C20ECFE03FCDF" blockId="15.[151,1437,152,1764]" pageId="15" pageNumber="259">
Metanotal gland (
<figureCitation id="13023CA8A91FFF8BFE0B20ECFE69FD63" box="[400,487,657,683]" captionStart="FIGURE 9" captionStartId="12.[151,250,1945,1970]" captionTargetBox="[179,1408,208,1920]" captionTargetId="figure@12.[179,1408,181,1920]" captionTargetPageId="12" captionText="FIGURE 9. Itara (Bornitara) spinosa sp. nov. male holotype habitus in dorsal view (A), head and pronotum in dorsal (B) and lateral (C) views, face in anterior view (D) and metanotal gland in dorsal view (E). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3938197" httpUri="https://zenodo.org/record/3938197/files/figure.png" pageId="15" pageNumber="259">Fig. 9E</figureCitation>
) with roundly curved transverse concavity in anterior part of metanotum, large and low median convexity on middle part, rather large and shallow concavity (approximately coincident with much lighter area at centre of metanotum) at apex of this concavity, and an oval whitish tubercle and keel-like structures behind this concavity.
</paragraph>
<paragraph id="8B86202DA91FFF8BFF5C215CFB18FC27" blockId="15.[151,1437,152,1764]" pageId="15" pageNumber="259">
Tegmen (
<figureCitation id="13023CA8A91FFF8BFEB4215CFE0EFCF3" box="[303,384,801,827]" captionStart="FIGURE 10" captionStartId="13.[151,250,1944,1969]" captionTargetBox="[371,1210,181,1905]" captionTargetId="figure@13.[359,1229,181,1919]" captionTargetPageId="13" captionText="FIGURE 10. Itara (Bornitara) spinosa sp. nov. male holotype tegmen in dorsal view. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3938199" httpUri="https://zenodo.org/record/3938199/files/figure.png" pageId="15" pageNumber="259">Fig. 10</figureCitation>
) very finely pubescent, extending beyond abdominal apex, with dorsal field longer than lateral field; with four oblique veins in harp area, and well developed additional vein connecting all oblique veins near CuA; tegminal mirror large, 1.5 times as wide as long, with anterior margin angular, with posterior margin widely rounded, with a sinuous dividing; anal area of tegmen longer than length of mirror; lateral field wide, with 10 parallel strong branches of Sc and about 7 cross-veins (often indistinct) between R and M (
<figureCitation id="13023CA8A91FFF8BFBE121CCFB41FC03" box="[1146,1231,945,971]" captionStart="FIGURE 10" captionStartId="13.[151,250,1944,1969]" captionTargetBox="[371,1210,181,1905]" captionTargetId="figure@13.[359,1229,181,1919]" captionTargetPageId="13" captionText="FIGURE 10. Itara (Bornitara) spinosa sp. nov. male holotype tegmen in dorsal view. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3938199" httpUri="https://zenodo.org/record/3938199/files/figure.png" pageId="15" pageNumber="259">Fig. 10</figureCitation>
); R and M mostly parallel, strongly diverging towards apex; hind wings slightly surpassing tegmina (
<figureCitation id="13023CA8A91FFF8BFBAE21A8FB07FC27" box="[1077,1161,981,1007]" captionStart="FIGURE 10" captionStartId="13.[151,250,1944,1969]" captionTargetBox="[371,1210,181,1905]" captionTargetId="figure@13.[359,1229,181,1919]" captionTargetPageId="13" captionText="FIGURE 10. Itara (Bornitara) spinosa sp. nov. male holotype tegmen in dorsal view. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3938199" httpUri="https://zenodo.org/record/3938199/files/figure.png" pageId="15" pageNumber="259">Fig. 10</figureCitation>
).
</paragraph>
<paragraph id="8B86202DA91FFF8BFF5C2184FD4EFBFF" blockId="15.[151,1437,152,1764]" pageId="15" pageNumber="259">
Anal plate tongue-shaped, with lateral keel distinct in anterior half (
<figureCitation id="13023CA8A91FFF8BFC782184FBC0FBDB" box="[995,1102,1017,1043]" captionStart="FIGURE 11" captionStartId="14.[151,250,1403,1428]" captionTargetBox="[151,1436,181,1377]" captionTargetId="figure@14.[151,1436,181,1377]" captionTargetPageId="14" captionText="FIGURE 11. Itara (Bornitara) spinosa sp. nov. male holotype abdominal apex in dorsal view (A), genitalia in lateral (B), dorsal (C) and ventral (D) views. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3938201" httpUri="https://zenodo.org/record/3938201/files/figure.png" pageId="15" pageNumber="259">Fig. 11A</figureCitation>
). Subgenital plate elongated, gradually narrowing to subacute apex (
<figureCitation id="13023CA8A91FFF8BFDD72660FD3DFBFF" box="[588,691,1053,1079]" captionStart="FIGURE 11" captionStartId="14.[151,250,1403,1428]" captionTargetBox="[151,1436,181,1377]" captionTargetId="figure@14.[151,1436,181,1377]" captionTargetPageId="14" captionText="FIGURE 11. Itara (Bornitara) spinosa sp. nov. male holotype abdominal apex in dorsal view (A), genitalia in lateral (B), dorsal (C) and ventral (D) views. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3938201" httpUri="https://zenodo.org/record/3938201/files/figure.png" pageId="15" pageNumber="259">Fig. 11A</figureCitation>
).
</paragraph>
<paragraph id="8B86202DA91FFF8BFF5C263CFC99FA9F" blockId="15.[151,1437,152,1764]" pageId="15" pageNumber="259">
Male genitalia as shown in
<figureCitation id="13023CA8A91FFF8BFD9B263CFDD7FB93" box="[512,601,1089,1115]" captionStart="FIGURE 11" captionStartId="14.[151,250,1403,1428]" captionTargetBox="[151,1436,181,1377]" captionTargetId="figure@14.[151,1436,181,1377]" captionTargetPageId="14" captionText="FIGURE 11. Itara (Bornitara) spinosa sp. nov. male holotype abdominal apex in dorsal view (A), genitalia in lateral (B), dorsal (C) and ventral (D) views. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3938201" httpUri="https://zenodo.org/record/3938201/files/figure.png" pageId="15" pageNumber="259">Figs. 11</figureCitation>
B–D: Epiphallus in dorsal view elongated but fairly broad, gradually narrowing to apex, lateral margin slightly concave after middle, apex somewhat truncated; in ventral view with platelike structure in apical half, this plate with anterior margin faintly sinuous and narrowly emarginated in middle, laterally produced into ventrad-pointing spine (vs); in profile with characteristics S-shaped dorsal edge, with a distinct dorsal spine at middle of epiphallus (ds), lacks row of denticles on ventral surface, apex strongly curved upwards and faintly hooked (ap), ventral margin with a relatively large spine in middle, anterior emargination (ae) narrow. Ectoparameres (ec) with anterior half wide, inner lobe rounded; posterior half cylindrical, elongated (as long as length of anterior half), slightly curved with apex obtuse.
</paragraph>
<paragraph id="8B86202DA91FFF8BFF5C271CFE10FAB3" blockId="15.[151,1437,152,1764]" box="[199,414,1377,1403]" pageId="15" pageNumber="259">Female. Unknown.</paragraph>
<paragraph id="8B86202DA91FFF8BFF5C27F8FAF6F99B" blockId="15.[151,1437,152,1764]" pageId="15" pageNumber="259">
Colouration. Body generally brown (
<figureCitation id="13023CA8A91FFF8BFDCF27F8FD24FA57" box="[596,682,1413,1439]" captionStart="FIGURE 9" captionStartId="12.[151,250,1945,1970]" captionTargetBox="[179,1408,208,1920]" captionTargetId="figure@12.[179,1408,181,1920]" captionTargetPageId="12" captionText="FIGURE 9. Itara (Bornitara) spinosa sp. nov. male holotype habitus in dorsal view (A), head and pronotum in dorsal (B) and lateral (C) views, face in anterior view (D) and metanotal gland in dorsal view (E). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3938197" httpUri="https://zenodo.org/record/3938197/files/figure.png" pageId="15" pageNumber="259">Fig. 9A</figureCitation>
). Head and pronotum dark brown (
<figureCitation id="13023CA8A91FFF8BFBBA27F8FBFBFA57" box="[1057,1141,1413,1439]" captionStart="FIGURE 9" captionStartId="12.[151,250,1945,1970]" captionTargetBox="[179,1408,208,1920]" captionTargetId="figure@12.[179,1408,181,1920]" captionTargetPageId="12" captionText="FIGURE 9. Itara (Bornitara) spinosa sp. nov. male holotype habitus in dorsal view (A), head and pronotum in dorsal (B) and lateral (C) views, face in anterior view (D) and metanotal gland in dorsal view (E). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3938197" httpUri="https://zenodo.org/record/3938197/files/figure.png" pageId="15" pageNumber="259">Fig. 9B</figureCitation>
); scapus and pedicel slightly lighter in colouration; frons yellow brown; gena whitish immediately ventral of eye and then dark brown; maxillary palpi greyish. Tegminal dorsal field mostly transparent with tint of grey, veins usually darker (but not always), anal area infumated yellow brown; area between M, R and Sc infumated dark brown; lateral tegminal field pale mostly transparent with darker brown veins (
<figureCitation id="13023CA8A91FFF8BFE4D2468FDA8F9E7" box="[470,550,1557,1583]" captionStart="FIGURE 10" captionStartId="13.[151,250,1944,1969]" captionTargetBox="[371,1210,181,1905]" captionTargetId="figure@13.[359,1229,181,1919]" captionTargetPageId="13" captionText="FIGURE 10. Itara (Bornitara) spinosa sp. nov. male holotype tegmen in dorsal view. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3938199" httpUri="https://zenodo.org/record/3938199/files/figure.png" pageId="15" pageNumber="259">Fig. 10</figureCitation>
). Hind wings infumated dark. Legs generally with coxae pale, femora pale brown except near knees which are darker, tibiae and tarsi brown. Abdominal tergites grey brown, sternites yellow brown.
</paragraph>
<paragraph id="8B86202DA91FFF8BFF5C2420FE6FF953" blockId="15.[151,1437,152,1764]" pageId="15" pageNumber="259">
<emphasis id="B94DFC3FA91FFF8BFF5C2420FE6DF9BF" bold="true" box="[199,483,1628,1655]" pageId="15" pageNumber="259">Measurements (in mm).</emphasis>
Male
<typeStatus id="54829E8FA91FFF8BFDB12420FD02F9BF" box="[554,652,1629,1655]" pageId="15" pageNumber="259" type="holotype">holotype</typeStatus>
BL = 15.9; HL = 1.9; PL = 2.4; PW = 4.3; TL = 14.8; TW = 6.4; HWL = 4.1; HFL = 7.0; HTL = 5.8.
</paragraph>
</subSubSection>
<subSubSection id="C32373A6A91FFF8BFF5C24D8FD8BF92C" pageId="15" pageNumber="259" type="etymology">
<paragraph id="8B86202DA91FFF8BFF5C24D8FD8BF92C" blockId="15.[151,1437,152,1764]" pageId="15" pageNumber="259">
<emphasis id="B94DFC3FA91FFF8BFF5C24D8FEC0F977" bold="true" box="[199,334,1701,1727]" pageId="15" pageNumber="259">Etymology.</emphasis>
The species name refers to the characteristic distinct dorsal spine (ds) at middle of epiphallus (spinosa = point or spike, feminine).
</paragraph>
</subSubSection>
</treatment>
</document>
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