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<document id="88F36405047609B92038D221D34140FD" ID-CLB-Dataset="42259" ID-DOI="10.11646/zootaxa.4103.6.1" ID-GBIF-Dataset="5d0b6265-8f97-4dd5-8e5d-b5c64db66ddc" ID-ISSN="1175-5326" ID-Zenodo-Dep="399676" ID-ZooBank="F4568761-B650-49D9-A94A-1B7E6B072663" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460959753691" checkinUser="plazi" docAuthor="Couri, Marcia S. &amp; Pont, Adrian C." docDate="2016" docId="038FA947C35A023579889350FDFFFC92" docLanguage="en" docName="zootaxa.4103.6.1.pdf.imf" docOrigin="Zootaxa 4103 (6)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coenosia flavohumeralis Couri &amp; Pont, 2016, sp. nov." docType="treatment" docVersion="12" lastPageNumber="507" masterDocId="FFB6D13FC35F0233791F976FFF9AFFAA" masterDocTitle="Coenosia Meigen (Diptera: Muscidae) from Angola: new species and records" masterLastPageNumber="512" masterPageNumber="501" pageNumber="506" updateTime="1698586785324" updateUser="ExternalLinkService">
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<mods:title id="EB0BD61D1C6537B5BCF19213B4C729E0">Coenosia Meigen (Diptera: Muscidae) from Angola: new species and records</mods:title>
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<mods:name id="D5F8F30E923B1762CA7D9456A24E14FE" type="personal">
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<mods:namePart id="6E119AF7B3350EC6B28A19363E080347">Couri, Marcia S.</mods:namePart>
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<mods:namePart id="73BB0B21974BD3E4ECEC968F8EDD2009">Pont, Adrian C.</mods:namePart>
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<mods:typeOfResource id="2AFC3F4F22CBB79D18DBA9314764BE45">text</mods:typeOfResource>
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<mods:title id="85F873787A24CD02FD1A9D2E0A9BE08C">Zootaxa</mods:title>
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<mods:date id="BBC786FB0F2499893E5A13D0F57DFC40">2016</mods:date>
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<mods:number id="6D5D838CC2A435C169F6C8DF182AF1F4">4103</mods:number>
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<treatment id="038FA947C35A023579889350FDFFFC92" ID-DOI="http://doi.org/10.5281/zenodo.6087572" ID-GBIF-Taxon="119592994" ID-Zenodo-Dep="6087572" LSID="urn:lsid:plazi:treatment:038FA947C35A023579889350FDFFFC92" httpUri="http://treatment.plazi.org/id/038FA947C35A023579889350FDFFFC92" lastPageId="6" lastPageNumber="507" pageId="5" pageNumber="506">
<subSubSection id="C33C4BDAC35A023679889350FEBDFBD1" pageId="5" pageNumber="506" type="nomenclature">
<paragraph id="8B991851C35A023679889350FDB3FBF0" blockId="5.[151,553,1087,1147]" box="[151,553,1087,1114]" pageId="5" pageNumber="506">
<heading id="D0D1AF3DC35A023679889350FDB3FBF0" bold="true" box="[151,553,1087,1114]" fontSize="11" level="1" pageId="5" pageNumber="506" reason="1">
<emphasis id="B952C443C35A023679889350FDB3FBF0" bold="true" box="[151,553,1087,1114]" pageId="5" pageNumber="506">
<taxonomicName id="4C2663D2C35A023679889350FE5EFBF3" ID-CoL="9HXWQ" box="[151,452,1087,1113]" class="Insecta" family="Muscidae" genus="Coenosia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="506" phylum="Arthropoda" rank="species" species="flavohumeralis" status="sp. nov.">
<emphasis id="B952C443C35A023679889350FE5EFBF3" bold="true" box="[151,452,1087,1113]" italics="true" pageId="5" pageNumber="506">Coenosia flavohumeralis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2617938C35A023678D3932FFDB3FBF0" box="[460,553,1088,1114]" pageId="5" pageNumber="506" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B991851C35A02367988930DFEBDFBD1" blockId="5.[151,553,1087,1147]" box="[151,295,1122,1147]" pageId="5" pageNumber="506">
(
<figureCitation id="131D04D4C35A02367980930DFE85FBD1" box="[159,287,1122,1147]" captionStart="FIGURES 1 9" captionStartId="3.[151,264,1918,1940]" captionTargetBox="[202,1376,201,1887]" captionTargetId="figure@3.[185,1390,193,1887]" captionTargetPageId="3" captionText="FIGURES 1 9. (1) Coenosia lucens sp. nov., male dorsal view; (2) Coenosia flavohumeralis sp. nov., male dorsal view; (3) Coenosia setosa sp. nov., male, dorsal view. (4 6) Coenosia lucens sp. nov., male: (4) sternite 5, dorsal view; (5) cercal plate, dorsal view; (6) aedeagal complex, lateral view; (7 9) Coenosia flavohumeralis sp. nov., male: (7) sternite 5, dorsal view; (8) cercal plate and surstyli, dorsal view; (9) aedeagal complex, lateral view." httpUri="https://zenodo.org/record/399836/files/figure.png" pageId="5" pageNumber="506">Figs 2, 79</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33C4BDAC35A0236798893C5FA9FFB4D" pageId="5" pageNumber="506" type="materials_examined">
<paragraph id="8B991851C35A0236798893C5FA9FFB4D" blockId="5.[151,1437,1194,2011]" pageId="5" pageNumber="506">
<emphasis id="B952C443C35A0236798893C5FE24FB69" bold="true" box="[151,446,1194,1219]" pageId="5" pageNumber="506">
<typeStatus id="549DA6F3C35A0236798893C5FF48FB69" box="[151,210,1194,1219]" pageId="5" pageNumber="506">Type</typeStatus>
material examined.
</emphasis>
<typeStatus id="549DA6F3C35A023678DB93C5FDB6FB69" box="[452,556,1194,1219]" pageId="5" pageNumber="506" type="holotype">Holotype</typeStatus>
male,
<collectingCountry id="F33158C1C35A02367B6993C4FD6BFB69" box="[630,753,1195,1219]" name="Angola" pageId="5" pageNumber="506">ANGOLA</collectingCountry>
: (A40), Tundavala, 810 mls. NW. Sa da Bandeira,
<date id="FF983E91C35A02367C2E93C5FF55FB4D" pageId="5" pageNumber="506" value="1972-03-27" valueMax="1972-03-29" valueMin="1972-03-27">2729.iii. 1972</date>
/ Southern African Exp., B.M.1972-1.
<typeStatus id="549DA6F3C35A02367B9993A0FD6CFB4D" box="[646,758,1231,1255]" pageId="5" pageNumber="506" type="paratype">Paratypes</typeStatus>
,
<specimenCount id="9D20D3D8C35A02367A1E93A0FCEAFB4D" box="[769,880,1230,1255]" pageId="5" pageNumber="506" type="female">2 females</specimenCount>
, same labels as
<typeStatus id="549DA6F3C35A02367D3D93A1FB1EFB4D" box="[1058,1156,1230,1255]" pageId="5" pageNumber="506" type="holotype">holotype</typeStatus>
(1
<collectionCode id="ED378094C35A02367DB593A0FB60FB4D" box="[1194,1274,1231,1255]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="5" pageNumber="506">MNRJ</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C33C4BDAC35A023679D8939DFBE5FAF9" pageId="5" pageNumber="506" type="diagnosis">
<paragraph id="8B991851C35A023679D8939DFBE5FAF9" blockId="5.[151,1437,1194,2011]" pageId="5" pageNumber="506">
<emphasis id="B952C443C35A023679D8939DFEDBFAA1" bold="true" box="[199,321,1266,1291]" pageId="5" pageNumber="506">Diagnosis.</emphasis>
Ground-colour brown with grey pollinosity; postpronotum yellow with a little grey pollinosity; legs yellow with a preapical brown ring on mid and hind femora; abdomen with brownish-grey pollinosity, syntergite 1+2 yellowish; sternite 5 large, triangular, with two concavities on apical margin (
<figureCitation id="131D04D4C35A02367D329255FBEBFAF9" box="[1069,1137,1338,1363]" captionStart="FIGURES 1 9" captionStartId="3.[151,264,1918,1940]" captionTargetBox="[202,1376,201,1887]" captionTargetId="figure@3.[185,1390,193,1887]" captionTargetPageId="3" captionText="FIGURES 1 9. (1) Coenosia lucens sp. nov., male dorsal view; (2) Coenosia flavohumeralis sp. nov., male dorsal view; (3) Coenosia setosa sp. nov., male, dorsal view. (4 6) Coenosia lucens sp. nov., male: (4) sternite 5, dorsal view; (5) cercal plate, dorsal view; (6) aedeagal complex, lateral view; (7 9) Coenosia flavohumeralis sp. nov., male: (7) sternite 5, dorsal view; (8) cercal plate and surstyli, dorsal view; (9) aedeagal complex, lateral view." httpUri="https://zenodo.org/record/399836/files/figure.png" pageId="5" pageNumber="506">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33C4BDAC35A023579D89231FB99FDCA" lastPageId="6" lastPageNumber="507" pageId="5" pageNumber="506" type="description">
<paragraph id="8B991851C35A023679D89231FE91F9E5" blockId="5.[151,1437,1194,2011]" pageId="5" pageNumber="506">
<emphasis id="B952C443C35A023679D89231FEC3FADD" bold="true" box="[199,345,1374,1399]" pageId="5" pageNumber="506">Description.</emphasis>
(
<figureCitation id="131D04D4C35A023678769231FE34FADD" box="[361,430,1374,1399]" captionStart="FIGURES 1 9" captionStartId="3.[151,264,1918,1940]" captionTargetBox="[202,1376,201,1887]" captionTargetId="figure@3.[185,1390,193,1887]" captionTargetPageId="3" captionText="FIGURES 1 9. (1) Coenosia lucens sp. nov., male dorsal view; (2) Coenosia flavohumeralis sp. nov., male dorsal view; (3) Coenosia setosa sp. nov., male, dorsal view. (4 6) Coenosia lucens sp. nov., male: (4) sternite 5, dorsal view; (5) cercal plate, dorsal view; (6) aedeagal complex, lateral view; (7 9) Coenosia flavohumeralis sp. nov., male: (7) sternite 5, dorsal view; (8) cercal plate and surstyli, dorsal view; (9) aedeagal complex, lateral view." httpUri="https://zenodo.org/record/399836/files/figure.png" pageId="5" pageNumber="506">Fig. 2</figureCitation>
) Ground-colour brown with grey pollinosity; head with frontal vitta dark brown; ocellar triangle yellow pollinose; fronto-orbital plate, face, parafacial and gena grey pollinose; antenna dark brown, tip of pedicel yellow; arista brown; palpus dark yellow on basal two-thirds and brown on apical third; scutum with one median brown vitta; scutellum uniformly grey pollinose; postpronotum yellow with a little grey pollinosity; pleura somewhat yellow with grey pollinosity; calypters white; haltere yellow; wing clear; legs yellow with a preapical brown ring on mid and hind femora; tarsi brown; abdomen with brownish-grey pollinosity, syntergite 1+2 yellowish.
</paragraph>
<paragraph id="8B991851C35A023679D89135FD67F9D8" blockId="5.[151,1437,1194,2011]" box="[199,765,1626,1651]" pageId="5" pageNumber="506">
<emphasis id="B952C443C35A023679D89135FE9BF9D9" box="[199,257,1626,1651]" italics="true" pageId="5" pageNumber="506">Male</emphasis>
. Body length:
<quantity id="4CDEB5B4C35A023678B89135FD9AF9D8" box="[423,512,1626,1651]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="506" unit="mm" value="2.5">2.5 mm</quantity>
; wing length:
<quantity id="4CDEB5B4C35A02367B809135FD62F9D8" box="[671,760,1626,1651]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="5" pageNumber="506" unit="mm" value="2.6">2.6 mm</quantity>
.
</paragraph>
<paragraph id="8B991851C35A023679D89111FEDCF8A9" blockId="5.[151,1437,1194,2011]" pageId="5" pageNumber="506">Head. Eye bare; frons at vertex about one-third of head-width; frontal row with 3 pairs of setae, and 1 pair of reclinate orbitals; inner and outer verticals and ocellars short and similar in size; antenna inserted at mid-level of eye, with postpedicel about 2.5 times the length of pedicel; arista with short and fine hairs along its whole length; palpus filiform.</paragraph>
<paragraph id="8B991851C35A023679D89061FBD7F838" blockId="5.[151,1437,1194,2011]" pageId="5" pageNumber="506">
<emphasis id="B952C443C35A023679D89061FE82F88D" box="[199,280,1806,1831]" italics="true" pageId="5" pageNumber="506">Thorax</emphasis>
. Acrostichal setulae in 2 rows, prescutellar pair not differentiated; dorsocentrals 1+3, all long; postpronotal 1, long; 1 presutural and 1 postsutural intra-alar, 1 postsutural supra-alar; 2 postalars; prealar absent; notopleuron with 2 setae, similar in size; anepisternum with series of 4 long setae; katepisternals 1+1+1, arranged in an equilateral triangle. Scutellum with 1 long basal and 1 long apical pair of setae.
</paragraph>
<paragraph id="8B991851C35A023579D890F0FA8FFECF" blockId="5.[151,1437,1194,2011]" lastBlockId="6.[151,1437,151,824]" lastPageId="6" lastPageNumber="507" pageId="5" pageNumber="506">
<emphasis id="B952C443C35A023679D890F0FF66F81C" box="[199,252,1951,1974]" italics="true" pageId="5" pageNumber="506">Legs</emphasis>
. Fore femur with 2 anterior setae on basal third and with 34 sparse setae on anterodorsal and posterodorsal surfaces; fore tibia with a long posterior median seta, anterodorsal, dorsal and posterodorsal preapicals, and 1 ventral apical; mid femur with 2 anterior setae on middle third, 45 well-spaced posteroventral and anteroventral setae on basal and middle thirds, and 2 posterior preapicals; mid tibia with 1 anteroventral and 1 median anterior, the anterior seta inserted a little below the other, with preapicals on dorsal and posterodorsal surfaces and 1 ventral apical; hind femur with an anterodorsal and an anteroventral row of well-spaced setae; hind tibia with 1 anterior and 1 posterodorsal median setae, inserted almost at the same level, 1 long submedian posterodorsal,1 anterodorsal and 1 dorsal on apical third, apicals on anteroventral and ventral surfaces.
</paragraph>
<paragraph id="8B991851C359023579D8961EFF5DFE06" blockId="6.[151,1437,151,824]" pageId="6" pageNumber="507">
<emphasis id="B952C443C359023579D8961EFF65FE22" box="[199,255,369,392]" italics="true" pageId="6" pageNumber="507">Wing</emphasis>
. Veins bare; veins M and R4+5 parallel at wing-tip; lower calypter about twice the length of the upper one.
</paragraph>
<paragraph id="8B991851C359023579D896D8FC4FFE5E" blockId="6.[151,1437,151,824]" pageId="6" pageNumber="507">
<emphasis id="B952C443C359023579D896D8FEABFE7A" box="[199,305,439,464]" italics="true" pageId="6" pageNumber="507">Abdomen</emphasis>
. Tergite 3 with 1 pair of lateral setae; tergites 45 each with a discal row of setae. Sternite 1 bare. Sternite 5 large, triangular, with two concavities on apical margin (
<figureCitation id="131D04D4C35902357A9D96B3FC5DFE5E" box="[898,967,476,501]" captionStart="FIGURES 1 9" captionStartId="3.[151,264,1918,1940]" captionTargetBox="[202,1376,201,1887]" captionTargetId="figure@3.[185,1390,193,1887]" captionTargetPageId="3" captionText="FIGURES 1 9. (1) Coenosia lucens sp. nov., male dorsal view; (2) Coenosia flavohumeralis sp. nov., male dorsal view; (3) Coenosia setosa sp. nov., male, dorsal view. (4 6) Coenosia lucens sp. nov., male: (4) sternite 5, dorsal view; (5) cercal plate, dorsal view; (6) aedeagal complex, lateral view; (7 9) Coenosia flavohumeralis sp. nov., male: (7) sternite 5, dorsal view; (8) cercal plate and surstyli, dorsal view; (9) aedeagal complex, lateral view." httpUri="https://zenodo.org/record/399836/files/figure.png" pageId="6" pageNumber="507">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="8B991851C359023579D89690FC19FD97" blockId="6.[151,1437,151,824]" pageId="6" pageNumber="507">
<emphasis id="B952C443C359023579D89690FEDBFDB2" box="[199,321,511,536]" italics="true" pageId="6" pageNumber="507">Terminalia</emphasis>
. Cercal plate and surstylus as in
<figureCitation id="131D04D4C35902357BCB9690FC87FDB2" box="[724,797,511,536]" captionStart="FIGURES 1 9" captionStartId="3.[151,264,1918,1940]" captionTargetBox="[202,1376,201,1887]" captionTargetId="figure@3.[185,1390,193,1887]" captionTargetPageId="3" captionText="FIGURES 1 9. (1) Coenosia lucens sp. nov., male dorsal view; (2) Coenosia flavohumeralis sp. nov., male dorsal view; (3) Coenosia setosa sp. nov., male, dorsal view. (4 6) Coenosia lucens sp. nov., male: (4) sternite 5, dorsal view; (5) cercal plate, dorsal view; (6) aedeagal complex, lateral view; (7 9) Coenosia flavohumeralis sp. nov., male: (7) sternite 5, dorsal view; (8) cercal plate and surstyli, dorsal view; (9) aedeagal complex, lateral view." httpUri="https://zenodo.org/record/399836/files/figure.png" pageId="6" pageNumber="507">Fig. 8</figureCitation>
; cercal plate with a tuft of setae on the ventral apex. Aedeagal complex as in
<figureCitation id="131D04D4C359023578B5954BFE74FD96" box="[426,494,548,573]" captionStart="FIGURES 1 9" captionStartId="3.[151,264,1918,1940]" captionTargetBox="[202,1376,201,1887]" captionTargetId="figure@3.[185,1390,193,1887]" captionTargetPageId="3" captionText="FIGURES 1 9. (1) Coenosia lucens sp. nov., male dorsal view; (2) Coenosia flavohumeralis sp. nov., male dorsal view; (3) Coenosia setosa sp. nov., male, dorsal view. (4 6) Coenosia lucens sp. nov., male: (4) sternite 5, dorsal view; (5) cercal plate, dorsal view; (6) aedeagal complex, lateral view; (7 9) Coenosia flavohumeralis sp. nov., male: (7) sternite 5, dorsal view; (8) cercal plate and surstyli, dorsal view; (9) aedeagal complex, lateral view." httpUri="https://zenodo.org/record/399836/files/figure.png" pageId="6" pageNumber="507">Fig. 9</figureCitation>
; distiphallus long and membranous.
</paragraph>
<paragraph id="8B991851C359023579D89528FB99FDCA" blockId="6.[151,1437,151,824]" box="[199,1027,583,608]" pageId="6" pageNumber="507">
<emphasis id="B952C443C359023579D89528FE81FDCA" box="[199,283,583,608]" italics="true" pageId="6" pageNumber="507">Female</emphasis>
: Body length:
<quantity id="4CDEB5B4C359023578DD9528FD81FDF5" box="[450,539,583,608]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="6" pageNumber="507" unit="mm" value="2.6">2.6 mm</quantity>
; wing length:
<quantity id="4CDEB5B4C35902357BA59528FCDEFDF5" box="[698,836,583,608]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.7" metricValueMin="2.6" pageId="6" pageNumber="507" unit="mm" value="2.65" valueMax="2.7" valueMin="2.6">2.62.7 mm</quantity>
. Similar to male.
</paragraph>
</subSubSection>
<subSubSection id="C33C4BDAC359023579D89503FEC7FD0D" pageId="6" pageNumber="507" type="etymology">
<paragraph id="8B991851C359023579D89503FEC7FD0D" blockId="6.[151,1437,151,824]" pageId="6" pageNumber="507">
<emphasis id="B952C443C359023579D89503FED4FD2F" bold="true" box="[199,334,620,645]" pageId="6" pageNumber="507">Etymology.</emphasis>
The specific name refers to the yellow colour of the postpronotal lobes, contrasting with the grey pollinose scutum.
</paragraph>
</subSubSection>
<subSubSection id="C33C4BDAC359023579D895DBFDFFFC92" pageId="6" pageNumber="507" type="discussion">
<paragraph id="8B991851C359023579D895DBFE2CFCBF" blockId="6.[151,1437,151,824]" pageId="6" pageNumber="507">
<emphasis id="B952C443C359023579D895DBFED6FD67" bold="true" box="[199,332,692,717]" pageId="6" pageNumber="507">Discussion.</emphasis>
This species runs in Emdens (1940) key to the
<taxonomicName id="4C2663D2C35902357A7D95DAFC50FD66" box="[866,970,693,716]" class="Insecta" family="Muscidae" genus="Coenosia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="507" phylum="Arthropoda" rank="species" species="vittata">
<emphasis id="B952C443C35902357A7D95DAFC50FD66" box="[866,970,693,716]" italics="true" pageId="6" pageNumber="507">C. vittata</emphasis>
</taxonomicName>
-group, and within this group to
<taxonomicName id="4C2663D2C35902357C2D95DAFE2BFD5A" authority="Emden, 1940" authorityName="Emden" authorityYear="1940" class="Insecta" family="Muscidae" genus="Coenosia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="507" phylum="Arthropoda" rank="species" species="edwardsi">
<emphasis id="B952C443C35902357C2D95DAFF66FD5A" italics="true" pageId="6" pageNumber="507">Coenosia edwardsi</emphasis>
(
<bibRefCitation id="EFB765A0C3590235781295B8FE33FD5A" author="Emden" box="[269,425,727,752]" pageId="6" pageNumber="507" refString="Emden, F. I. van (1940) Muscidae: B - Coenosiinae. In: Ruwenzori Expedition 1934 - 5. Vol. 2. British Museum (Natural History), London, pp. 91 - 255." type="book chapter" year="1940">Emden, 1940</bibRefCitation>
)
</taxonomicName>
, but in this species the basal scutellar setae is much shorter than the apical one, and the fore femur is dark brown.
</paragraph>
<paragraph id="8B991851C359023579D89470FDFFFC92" blockId="6.[151,1437,151,824]" box="[199,613,799,824]" pageId="6" pageNumber="507">
<emphasis id="B952C443C359023579D89470FD9EFC92" bold="true" box="[199,516,799,824]" pageId="6" pageNumber="507">Geographical distribution.</emphasis>
<collectingCountry id="F33158C1C35902357B149470FDFBFC92" box="[523,609,799,824]" name="Angola" pageId="6" pageNumber="507">Angola</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>