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<mods:title id="E01E7D33CC260FC59B19207265C08DAD">Taxonomic Studies on Australian Psammoecus Latreille (Coleoptera, Silvanidae, Brontinae)</mods:title>
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<mods:title id="178A1CC45FAE91DE84780CC8C34C64FE">European Journal of Taxonomy</mods:title>
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<mods:date id="02EE350F72221FFE5D396F229F916CCF">2020</mods:date>
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<paragraph id="8B9D1E2DFF8ABA44FDE4FD9381F1FD8D" blockId="7.[571,1016,581,643]" box="[571,1016,581,608]" pageId="7" pageNumber="142">
<heading id="D0D5A941FF8ABA44FDE4FD9381F1FD8D" box="[571,1016,581,608]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="142" reason="2">
<taxonomicName id="4C2265AEFF8ABA44FDE4FD9381F1FD8D" authority="Grouvelle, 1919" authorityName="Grouvelle" authorityYear="1919" box="[571,1016,581,608]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="142" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="B956C23FFF8ABA44FDE4FD93813BFD8D" bold="true" box="[571,818,581,608]" italics="true" pageId="7" pageNumber="142">Psammoecus obesus</emphasis>
<bibRefCitation id="EFB363DCFF8ABA44FCE5FD9081F1FD8D" author="Grouvelle A." box="[826,1016,581,608]" pageId="7" pageNumber="142" refId="ref13309" refString="Grouvelle A. 1919. Descriptions d'especes nouvelles du genre Psammoecus. Memoires entomologiques. Etudes sur les Coleopteres. Deuxieme Fascicule. Societe entomologique de France, Paris." type="book" year="1919">Grouvelle, 1919</bibRefCitation>
</taxonomicName>
</heading>
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<subSubSection id="C3384DA6FF8ABA44FD28FDBC8135FD6E" box="[759,828,617,643]" pageId="7" pageNumber="142" type="description">
<paragraph id="8B9D1E2DFF8ABA44FD28FDBC8135FD6E" blockId="7.[571,1016,581,643]" box="[759,828,617,643]" pageId="7" pageNumber="142">
<figureCitation id="131902A8FF8ABA44FD28FDBC8135FD6E" box="[759,828,617,643]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[257,1339,265,1820]" captionTargetId="figure-10@8.[257,1339,265,1854]" captionTargetPageId="8" captionText="Fig. 3. Psammoecus obesus Grouvelle, 1919. AD. Holotype, ♀ (NHMUK). A. Habitus. B. Head. C. Pronotum. D. Left antenna. EH. Specimen from Queensland, ♂ (ANIC). E. Aedeagus, ventral view. F. Aedeagus, lateral view. G. Parameres, ventral view. H. Median lobe, dorsal view. Scale bars: A, D = 1.0 mm; BC, EF = 0.5 mm; GH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4271758" httpUri="https://zenodo.org/record/4271758/files/figure.png" pageId="7" pageNumber="142">Fig. 3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3384DA6FF8ABA44FF62FD7E80A6FCEB" pageId="7" pageNumber="142" type="reference_group">
<paragraph id="8B9D1E2DFF8ABA44FF62FD7E80AFFD29" blockId="7.[189,678,682,709]" box="[189,678,682,709]" pageId="7" pageNumber="142">
<treatmentCitationGroup id="AB323903FF8ABA44FF62FD7E80AFFD29" box="[189,678,682,709]" pageId="7" pageNumber="142">
<treatmentCitation id="0A83383CFF8ABA44FF62FD7E80ABFD29" author="Grouvelle A." box="[189,674,682,709]" page="11" pageId="7" pageNumber="142" year="1919">
<taxonomicName id="4C2265AEFF8ABA44FF62FD7E80ABFD29" authority="Grouvelle, 1919: 11" authorityName="Grouvelle" authorityPageNumber="11" authorityYear="1919" box="[189,674,682,709]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="142" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="B956C23FFF8ABA44FF62FD7E83A4FD29" box="[189,429,682,708]" italics="true" pageId="7" pageNumber="142">Psammoecus obesus</emphasis>
<bibRefCitation id="EFB363DCFF8ABA44FE6AFD7F80ABFD29" author="Grouvelle A." box="[437,674,682,709]" pageId="7" pageNumber="142" refId="ref13309" refString="Grouvelle A. 1919. Descriptions d'especes nouvelles du genre Psammoecus. Memoires entomologiques. Etudes sur les Coleopteres. Deuxieme Fascicule. Societe entomologique de France, Paris." type="book" year="1919">Grouvelle, 1919: 11</bibRefCitation>
</taxonomicName>
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.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9D1E2DFF8ABA44FF62FD3980A6FCEB" blockId="7.[189,687,747,774]" box="[189,687,747,774]" pageId="7" pageNumber="142">
<treatmentCitationGroup id="AB323903FF8ABA44FF62FD3980A6FCEB" box="[189,687,747,774]" pageId="7" pageNumber="142">
<taxonomicName id="4C2265AEFF8ABA44FF62FD3983A4FCE8" authorityName="Grouvelle" authorityYear="1919" box="[189,429,747,773]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="142" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="B956C23FFF8ABA44FF62FD3983A4FCE8" box="[189,429,747,773]" italics="true" pageId="7" pageNumber="142">Psammoecus obesus</emphasis>
</taxonomicName>
<treatmentCitation id="0A83383CFF8ABA44FE14FD3E80A5FCEB" author="Hetschko A." box="[459,684,747,774]" page="83" pageId="7" pageNumber="142" year="1930">
<bibRefCitation id="EFB363DCFF8ABA44FE14FD3E80A5FCEB" author="Hetschko A." box="[459,684,747,774]" pageId="7" pageNumber="142" pagination="3 - 107" refId="ref13339" refString="Hetschko A. 1930. Fam. Cucujidae. In: Junk W. &amp; Schenkling S. (eds) Coleopterorum Catalogus 109: 3 - 107. W. Junk, Berlin." type="journal article" year="1930">Hetschko 1930: 83</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="C3384DA6FF8ABA44FF62FCF886A6FA68" pageId="7" pageNumber="142" type="diagnosis">
<paragraph id="8B9D1E2DFF8ABA44FF62FCF8833FFCAA" blockId="7.[189,310,813,839]" box="[189,310,813,839]" pageId="7" pageNumber="142">
<heading id="D0D5A941FF8ABA44FF62FCF8833FFCAA" bold="true" box="[189,310,813,839]" fontSize="11" level="3" pageId="7" pageNumber="142" reason="3">
<emphasis id="B956C23FFF8ABA44FF62FCF8833FFCAA" bold="true" box="[189,310,813,839]" pageId="7" pageNumber="142">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B9D1E2DFF8ABA44FF62FC8E86A6FA68" blockId="7.[188,1401,859,1414]" pageId="7" pageNumber="142">
The following combination of character states distinguishes this species: body elongate-oval (
<figureCitation id="131902A8FF8ABA44FAD5FC8E8761FC9B" box="[1290,1384,859,886]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[257,1339,265,1820]" captionTargetId="figure-10@8.[257,1339,265,1854]" captionTargetPageId="8" captionText="Fig. 3. Psammoecus obesus Grouvelle, 1919. AD. Holotype, ♀ (NHMUK). A. Habitus. B. Head. C. Pronotum. D. Left antenna. EH. Specimen from Queensland, ♂ (ANIC). E. Aedeagus, ventral view. F. Aedeagus, lateral view. G. Parameres, ventral view. H. Median lobe, dorsal view. Scale bars: A, D = 1.0 mm; BC, EF = 0.5 mm; GH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4271758" httpUri="https://zenodo.org/record/4271758/files/figure.png" pageId="7" pageNumber="142">Fig. 3A</figureCitation>
), length
<quantity id="4CDAB3C8FF8ABA44FECDFCAA83CAFC74" box="[274,451,895,921]" metricMagnitude="-3" metricUnit="m" metricValue="3.185" metricValueMax="3.29" metricValueMin="3.08" pageId="7" pageNumber="142" unit="mm" value="3.185" valueMax="3.29" valueMin="3.08">3.083.29 mm</quantity>
; eyes large (
<figureCitation id="131902A8FF8ABA44FDBFFCAA80B6FC74" box="[608,703,895,921]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[257,1339,265,1820]" captionTargetId="figure-10@8.[257,1339,265,1854]" captionTargetPageId="8" captionText="Fig. 3. Psammoecus obesus Grouvelle, 1919. AD. Holotype, ♀ (NHMUK). A. Habitus. B. Head. C. Pronotum. D. Left antenna. EH. Specimen from Queensland, ♂ (ANIC). E. Aedeagus, ventral view. F. Aedeagus, lateral view. G. Parameres, ventral view. H. Median lobe, dorsal view. Scale bars: A, D = 1.0 mm; BC, EF = 0.5 mm; GH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4271758" httpUri="https://zenodo.org/record/4271758/files/figure.png" pageId="7" pageNumber="142">Fig. 3B</figureCitation>
), irregularly rounded with stronger curvature posteriorly; temples distinct, short, strongly curved; frontal grooves distinct, diverging anteriorly, parallel near end; reaching anterior
<emphasis id="B956C23FFF8ABA44FE51FC1783AEFC0D" bold="true" box="[398,423,962,992]" pageId="7" pageNumber="142">²5</emphasis>
of eyes; antennae stout (
<figureCitation id="131902A8FF8ABA44FD12FC108122FC32" box="[717,811,965,991]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[257,1339,265,1820]" captionTargetId="figure-10@8.[257,1339,265,1854]" captionTargetPageId="8" captionText="Fig. 3. Psammoecus obesus Grouvelle, 1919. AD. Holotype, ♀ (NHMUK). A. Habitus. B. Head. C. Pronotum. D. Left antenna. EH. Specimen from Queensland, ♂ (ANIC). E. Aedeagus, ventral view. F. Aedeagus, lateral view. G. Parameres, ventral view. H. Median lobe, dorsal view. Scale bars: A, D = 1.0 mm; BC, EF = 0.5 mm; GH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4271758" httpUri="https://zenodo.org/record/4271758/files/figure.png" pageId="7" pageNumber="142">Fig. 3D</figureCitation>
), antennomere 8 slightly darkened, antennomeres 910 darkened, antennomere 11 bright yellowish-brown; punctation on vertex moderate, punctures distinctly elongate, as large as eye facet diameter; microsculpture on vertex absent, pubescence moderate; pronotum widest just in front of middle (
<figureCitation id="131902A8FF8ABA44FCF7FBFA818CFBA4" box="[808,901,1071,1097]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[257,1339,265,1820]" captionTargetId="figure-10@8.[257,1339,265,1854]" captionTargetPageId="8" captionText="Fig. 3. Psammoecus obesus Grouvelle, 1919. AD. Holotype, ♀ (NHMUK). A. Habitus. B. Head. C. Pronotum. D. Left antenna. EH. Specimen from Queensland, ♂ (ANIC). E. Aedeagus, ventral view. F. Aedeagus, lateral view. G. Parameres, ventral view. H. Median lobe, dorsal view. Scale bars: A, D = 1.0 mm; BC, EF = 0.5 mm; GH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4271758" httpUri="https://zenodo.org/record/4271758/files/figure.png" pageId="7" pageNumber="142">Fig. 3C</figureCitation>
), lateral pronotal margins with five teeth; tooth I slightly and tooth V distinctly bigger than IIIV, distance between I and II wider than distances between other teeth, anterior and posterior denticles present, small; pronotal punctation dense, punctures as large as on vertex, distinctly widened, microsculpture absent, pubescence moderate, setae directed medially; elytra widest at middle (
<figureCitation id="131902A8FF8ABA44FD87FB6E80BEFB3B" box="[600,695,1211,1238]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[257,1339,265,1820]" captionTargetId="figure-10@8.[257,1339,265,1854]" captionTargetPageId="8" captionText="Fig. 3. Psammoecus obesus Grouvelle, 1919. AD. Holotype, ♀ (NHMUK). A. Habitus. B. Head. C. Pronotum. D. Left antenna. EH. Specimen from Queensland, ♂ (ANIC). E. Aedeagus, ventral view. F. Aedeagus, lateral view. G. Parameres, ventral view. H. Median lobe, dorsal view. Scale bars: A, D = 1.0 mm; BC, EF = 0.5 mm; GH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4271758" httpUri="https://zenodo.org/record/4271758/files/figure.png" pageId="7" pageNumber="142">Fig. 3A</figureCitation>
), with transverse brown maculae behind middle, interstices distinctly wider than striae, pubescence moderate, microsculpture absent; male genitalia (
<figureCitation id="131902A8FF8ABA44FB2FFB0A873FFB14" box="[1264,1334,1247,1273]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[257,1339,265,1820]" captionTargetId="figure-10@8.[257,1339,265,1854]" captionTargetPageId="8" captionText="Fig. 3. Psammoecus obesus Grouvelle, 1919. AD. Holotype, ♀ (NHMUK). A. Habitus. B. Head. C. Pronotum. D. Left antenna. EH. Specimen from Queensland, ♂ (ANIC). E. Aedeagus, ventral view. F. Aedeagus, lateral view. G. Parameres, ventral view. H. Median lobe, dorsal view. Scale bars: A, D = 1.0 mm; BC, EF = 0.5 mm; GH = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4271758" httpUri="https://zenodo.org/record/4271758/files/figure.png" pageId="7" pageNumber="142">Fig. 3</figureCitation>
EH) strongly sclerotised, median lobe in dorsal view wide, narrowed abruptly near ostium, apical end well rounded, in lateral view also narrowed abruptly, with sharp apical end; parameres in ventral view very wide, inner margins, apex and apical third of lateral margins with distinct setae; in lateral view with very wide basis, narrowed abruptly towards apex with dorsal margin almost forming angle.
</paragraph>
</subSubSection>
<subSubSection id="C3384DA6FF8ABA44FF62FA7883F9F834" pageId="7" pageNumber="142" type="materials_examined">
<paragraph id="8B9D1E2DFF8ABA44FF62FA7883A3FA2A" blockId="7.[189,426,1453,1479]" box="[189,426,1453,1479]" pageId="7" pageNumber="142">
<heading id="D0D5A941FF8ABA44FF62FA7883A3FA2A" bold="true" box="[189,426,1453,1479]" fontSize="11" level="3" pageId="7" pageNumber="142" reason="3">
<emphasis id="B956C23FFF8ABA44FF62FA7883A3FA2A" bold="true" box="[189,426,1453,1479]" pageId="7" pageNumber="142">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B9D1E2DFF8ABA44FF35FA098355FA1B" blockId="7.[189,1399,1500,1632]" box="[234,348,1500,1526]" pageId="7" pageNumber="142">
<emphasis id="B956C23FFF8ABA44FF35FA098355FA1B" bold="true" box="[234,348,1500,1526]" pageId="7" pageNumber="142">
<typeStatus id="5499A08FFF8ABA44FF35FA098355FA1B" box="[234,348,1500,1526]" pageId="7" pageNumber="142" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B9D1E2DFF8ABA44FF62FA2A86CEF98D" blockId="7.[189,1399,1500,1632]" pageId="7" pageNumber="142">
<materialsCitation id="3B4A1470FF8ABA44FF62FA2A86CEF98D" collectionCode="NHMUK" collectorName="Pascoe" country="Indonesia" determinerName="R. G. Booth" location="Maluku" pageId="7" pageNumber="142" specimenCount="1" specimenCount-female="1" stateProvince="Maluku" typeStatus="holotype">
<collectingCountry id="F3355EBDFF8ABA44FF62FA2A8352F9F4" box="[189,347,1535,1561]" name="Indonesia" pageId="7" pageNumber="142">INDONESIA</collectingCountry>
<specimenCount id="9D24D5A4FF8ABA44FEAEFA2A8381F9F4" box="[369,392,1535,1561]" pageId="7" pageNumber="142" type="female"></specimenCount>
;
<collectingRegion id="49E6D0CFFF8ABA44FE4AFA2A83FAF9F4" box="[405,499,1535,1561]" country="Indonesia" name="Maluku Utara" pageId="7" pageNumber="142">Maluku</collectingRegion>
, Seram;
<collectorName id="26D77BFBFF8ABA44FD89FA2A80AEF9F4" box="[598,679,1535,1561]" pageId="7" pageNumber="142">Pascoe</collectorName>
leg.; “HOLO - |
<typeStatus id="5499A08FFF8ABA44FCBAFA2A81ABF9F4" box="[869,930,1535,1561]" pageId="7" pageNumber="142">Type</typeStatus>
” [round label with red border]; “Ceram” [blue, elliptic label]; “Pascoe | Coll. | 9360”; “
<taxonomicName id="4C2265AEFF8ABA44FD2CF9F68185F9D1" authorityName=", Pal" authorityYear="1985" box="[755,908,1571,1596]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="142" phylum="Arthropoda" rank="genus">
<emphasis id="B956C23FFF8ABA44FD2CF9F68185F9D1" box="[755,908,1571,1596]" italics="true" pageId="7" pageNumber="142">Psammoecus</emphasis>
</taxonomicName>
|
<taxonomicName id="4C2265AEFF8ABA44FC7AF9F781FCF9D1" authorityName="Grouvelle" authorityYear="1919" box="[933,1013,1570,1596]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="142" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="B956C23FFF8ABA44FC7AF9F781FCF9D1" box="[933,1013,1570,1596]" italics="true" pageId="7" pageNumber="142">obesus</emphasis>
</taxonomicName>
| ty. Grouv” [Grouvelles hand]; “
<typeStatus id="5499A08FFF8ABA44FF17F9908373F9B2" box="[200,378,1605,1631]" pageId="7" pageNumber="142" type="holotype">HOLOTYPUS</typeStatus>
|
<taxonomicName id="4C2265AEFF8ABA44FE52F993802FF9B2" authorityName=", Pal" authorityYear="1985" box="[397,550,1606,1631]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="142" phylum="Arthropoda" rank="genus">
<emphasis id="B956C23FFF8ABA44FE52F993802FF9B2" box="[397,550,1606,1631]" italics="true" pageId="7" pageNumber="142">Psammoecus</emphasis>
</taxonomicName>
|
<taxonomicName id="4C2265AEFF8ABA44FDE4F990812CF9B2" authority="Grouv. 1919" authorityName="Grouvelle" authorityYear="1919" box="[571,805,1605,1631]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="142" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="B956C23FFF8ABA44FDE4F9908082F9B2" box="[571,651,1605,1631]" italics="true" pageId="7" pageNumber="142">obesus</emphasis>
Grouv. 1919
</taxonomicName>
| det.
<determinerName id="7E41F2E9FF8ABA44FCB4F99081FBF9B2" box="[875,1010,1605,1631]" pageId="7" pageNumber="142">R.G. Booth</determinerName>
2013”;
<collectionCode id="ED3386E8FF8ABA44FB8FF99386CAF98D" box="[1104,1219,1606,1632]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="7" pageNumber="142" type="Museum">NHMUK</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8B9D1E2DFF8ABA44FF35F95383AFF94D" blockId="7.[188,1399,1670,1908]" box="[234,422,1670,1696]" pageId="7" pageNumber="142">
<materialsCitation id="3B4A1470FF8ABA44FF35F95383AFF94D" box="[234,422,1670,1696]" collectionCode="NHMUK" collectorName="Perkins" country="Australia" latitude="-16.916666" location="Cairns" longLatPrecision="1280" longitude="145.76666" pageId="7" pageNumber="142" specimenCount="1" stateProvince="Queensland">
<emphasis id="B956C23FFF8ABA44FF35F95383AFF94D" bold="true" box="[234,422,1670,1696]" pageId="7" pageNumber="142">Other material</emphasis>
</materialsCitation>
</paragraph>
<paragraph id="8B9D1E2DFF8ABA44FF62F97F8723F899" blockId="7.[188,1399,1670,1908]" pageId="7" pageNumber="142">
<collectingCountry id="F3355EBDFF8ABA44FF62F97F836BF929" box="[189,354,1706,1732]" name="Australia" pageId="7" pageNumber="142">AUSTRALIA</collectingCountry>
<specimenCount id="9D24D5A4FF8ABA44FEA7F97F83C2F929" box="[376,459,1706,1732]" pageId="7" pageNumber="142" type="generic">1 spec.</specimenCount>
;
<collectingRegion id="49E6D0CFFF8ABA44FE09F97F806DF929" box="[470,612,1706,1732]" country="Australia" name="Queensland" pageId="7" pageNumber="142">Queensland</collectingRegion>
,
<location id="8EFD48F6FF8ABA44FDB2F97F80B4F929" LSID="urn:lsid:plazi:treatment:038BAF3BFF8ABA4AFDE4FD9386BFFE09:8EFD48F6FF8ABA44FDB2F97F80B4F929" box="[621,701,1706,1732]" country="Australia" latitude="-16.916666" longLatPrecision="1280" longitude="145.76666" name="Cairns" pageId="7" pageNumber="142" stateProvince="Queensland">Cairns</location>
;
<geoCoordinate id="EE1678EAFF8ABA44FD17F97D8124F929" box="[712,813,1704,1732]" degrees="16" direction="south" minutes="55" orientation="latitude" pageId="7" pageNumber="142" precision="925" value="-16.916666">16°55 S</geoCoordinate>
,
<geoCoordinate id="EE1678EAFF8ABA44FCE7F97D81A6F929" box="[824,943,1704,1732]" degrees="145" direction="east" minutes="46" orientation="longitude" pageId="7" pageNumber="142" precision="925" value="145.76666">145°46 E</geoCoordinate>
; 1942;
<collectorName id="26D77BFBFF8ABA44FBDDF97F8653F929" box="[1026,1114,1706,1732]" pageId="7" pageNumber="142">Perkins</collectorName>
leg.;
<collectionCode id="ED3386E8FF8ABA44FB46F97F8701F929" box="[1177,1288,1706,1732]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="7" pageNumber="142" type="Museum">NHMUK</collectionCode>
<materialsCitation id="3B4A1470FF8ABA44FAC1F97F877EF90A" collectedFrom="rainforest" collectingDate="1994-11" collectionCode="ANIC" collectorName="C. Reid" country="Australia" location="Cassowary Coast Region" pageId="7" pageNumber="142" specimenCount="1" stateProvince="Queensland">
<specimenCount id="9D24D5A4FF8ABA44FAC1F97F877BF929" box="[1310,1394,1706,1732]" pageId="7" pageNumber="142" type="generic">1 spec.</specimenCount>
;
<collectingRegion id="49E6D0CFFF8ABA44FF62F9188342F90A" box="[189,331,1741,1767]" country="Australia" name="Queensland" pageId="7" pageNumber="142">Queensland</collectingRegion>
,
<location id="8EFD48F6FF8ABA44FE88F918808DF90A" LSID="urn:lsid:plazi:treatment:038BAF3BFF8ABA4AFDE4FD9386BFFE09:8EFD48F6FF8ABA44FE88F918808DF90A" box="[343,644,1741,1767]" country="Australia" name="Cassowary Coast Region" pageId="7" pageNumber="142" stateProvince="Queensland">Cassowary Coast Region</location>
,
<locationDeviation id="9A2F070FFF8ABA44FD4EF918817EF90A" box="[657,887,1741,1768]" location="Tully" pageId="7" pageNumber="142">
<quantity id="4CDAB3C8FF8ABA44FD4EF91880D5F90A" box="[657,732,1741,1767]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="142" unit="km" value="10.0">10 km</quantity>
NE of Tully
</locationDeviation>
;
<date id="FF9C38EDFF8ABA44FC5BF91B860AF90A" box="[900,1027,1741,1768]" pageId="7" pageNumber="142" value="1994-11">
<collectingDate id="EFD8C105FF8ABA44FC5BF91B860AF90A" box="[900,1027,1741,1768]" pageId="7" pageNumber="142" value="1994-11">Nov. 1994</collectingDate>
</date>
;
<collectorName id="26D77BFBFF8ABA44FBCFF9188663F90A" box="[1040,1130,1741,1767]" pageId="7" pageNumber="142">C. Reid</collectorName>
leg.;
<collectedFrom id="383E5DD0FF8ABA44FB70F918872AF90A" box="[1199,1315,1741,1767]" pageId="7" pageNumber="142">rainforest</collectedFrom>
;
<collectionCode id="ED3386E8FF8ABA44FAF0F918877EF90A" box="[1327,1399,1741,1767]" collectionName="Australia, Australian Capital Territory, Canberra City, CSIRO, Australian National Insect Collection" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="7" pageNumber="142">ANIC</collectionCode>
</materialsCitation>
<materialsCitation id="3B4A1470FF8ABA44FF0EF925876CF8E7" box="[209,1381,1776,1803]" collectingDate="1966-12-01" collectingDateMax="1966-12-02" collectingDateMin="1966-12-01" collectionCode="ANIC" collectorName="J. G. Brooks" country="Australia" location="Cardstone" municipality="Cassowary Coast Region" pageId="7" pageNumber="142" specimenCount="1" stateProvince="Queensland">
<specimenCount id="9D24D5A4FF8ABA44FF0EF925832EF8E6" box="[209,295,1776,1803]" pageId="7" pageNumber="142" type="generic">1 spec.</specimenCount>
;
<collectingRegion id="49E6D0CFFF8ABA44FEEBF92583CBF8E7" box="[308,450,1776,1802]" country="Australia" name="Queensland" pageId="7" pageNumber="142">Queensland</collectingRegion>
,
<collectingMunicipality id="6BF98457FF8ABA44FE11F92580F2F8E7" box="[462,763,1776,1802]" pageId="7" pageNumber="142">Cassowary Coast Region</collectingMunicipality>
,
<location id="8EFD48F6FF8ABA44FCD7F925818AF8E7" LSID="urn:lsid:plazi:treatment:038BAF3BFF8ABA4AFDE4FD9386BFFE09:8EFD48F6FF8ABA44FCD7F925818AF8E7" box="[776,899,1776,1802]" country="Australia" municipality="Cassowary Coast Region" name="Cardstone" pageId="7" pageNumber="142" stateProvince="Queensland">Cardstone</location>
;
<date id="FF9C38EDFF8ABA44FC50F9258648F8E7" box="[911,1089,1776,1803]" pageId="7" pageNumber="142" value="1966-12-01" valueMax="1966-12-02" valueMin="1966-12-01">
<collectingDate id="EFD8C105FF8ABA44FC50F9258648F8E7" box="[911,1089,1776,1803]" pageId="7" pageNumber="142" value="1966-12-01" valueMax="1966-12-02" valueMin="1966-12-01">12 Dec. 1966</collectingDate>
</date>
;
<collectorName id="26D77BFBFF8ABA44FB91F92586D2F8E7" box="[1102,1243,1776,1802]" pageId="7" pageNumber="142">J.G. Brooks</collectorName>
leg.;
<collectionCode id="ED3386E8FF8ABA44FAC2F925876CF8E7" box="[1309,1381,1776,1802]" collectionName="Australia, Australian Capital Territory, Canberra City, CSIRO, Australian National Insect Collection" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="7" pageNumber="142">ANIC</collectionCode>
</materialsCitation>
<materialsCitation id="3B4A1470FF8ABA44FF61F8C68087F8BC" collectingDate="1989-06-27" collectingDateMax="1989-07-12" collectingDateMin="1989-06-27" collectionCode="ANIC" collectorName="T. A. Weir" country="Australia" elevation="500" latitude="-13.75" location="McIlwraith Range" longLatPrecision="1289" longitude="143.36667" municipality="Shire of Cook" pageId="7" pageNumber="142" specimenCount="1" stateProvince="Queensland">
<specimenCount id="9D24D5A4FF8ABA44FF61F8C6831AF8C3" box="[190,275,1811,1838]" pageId="7" pageNumber="142" type="generic">1 spec.</specimenCount>
;
<collectingRegion id="49E6D0CFFF8ABA44FEFEF8C683A6F8C0" box="[289,431,1811,1837]" country="Australia" name="Queensland" pageId="7" pageNumber="142">Queensland</collectingRegion>
,
<collectingMunicipality id="6BF98457FF8ABA44FE64F8C6806FF8C0" box="[443,614,1811,1837]" pageId="7" pageNumber="142">Shire of Cook</collectingMunicipality>
,
<location id="8EFD48F6FF8ABA44FDACF8C68146F8C3" LSID="urn:lsid:plazi:treatment:038BAF3BFF8ABA4AFDE4FD9386BFFE09:8EFD48F6FF8ABA44FDACF8C68146F8C3" box="[627,847,1811,1838]" country="Australia" latitude="-13.75" longLatPrecision="1289" longitude="143.36667" municipality="Shire of Cook" name="McIlwraith Range" pageId="7" pageNumber="142" stateProvince="Queensland">McIlwraith Range</location>
;
<elevation id="000FF91EFF8ABA44FC84F8C681E1F8C0" box="[859,1000,1811,1838]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="142" unit="m" value="500.0">
<quantity id="4CDAB3C8FF8ABA44FC84F8C681AEF8C3" box="[859,935,1811,1838]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="142" unit="m" value="500.0">500 m</quantity>
a.s.l.
</elevation>
;
<geoCoordinate id="EE1678EAFF8ABA44FC28F8C78656F8C0" box="[1015,1119,1810,1838]" degrees="13" direction="south" minutes="45" orientation="latitude" pageId="7" pageNumber="142" precision="925" value="-13.75">13°45 S</geoCoordinate>
,
<geoCoordinate id="EE1678EAFF8ABA44FBB2F8C786EFF8C3" box="[1133,1254,1810,1838]" degrees="143" direction="east" minutes="22" orientation="longitude" pageId="7" pageNumber="142" precision="925" value="143.36667">143°22 E</geoCoordinate>
;
<date id="FF9C38EDFF8ABA44FB2BF8C68322F8BC" pageId="7" pageNumber="142" value="1989-06-27" valueMax="1989-07-12" valueMin="1989-06-27">
<collectingDate id="EFD8C105FF8ABA44FB2BF8C68322F8BC" pageId="7" pageNumber="142" value="1989-06-27" valueMax="1989-07-12" valueMin="1989-06-27">27 Jun.12 Jul. 1989</collectingDate>
</date>
;
<collectorName id="26D77BFBFF8ABA44FEE9F8E283AEF8BD" box="[310,423,1846,1873]" pageId="7" pageNumber="142">T.A. Weir</collectorName>
leg.; at light;
<collectionCode id="ED3386E8FF8ABA44FD99F8E28087F8BC" box="[582,654,1847,1873]" collectionName="Australia, Australian Capital Territory, Canberra City, CSIRO, Australian National Insect Collection" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="7" pageNumber="142">ANIC</collectionCode>
</materialsCitation>
<materialsCitation id="3B4A1470FF8ABA44FD7AF8E28124F899" collectedFrom="dead leaves, edge of rainforest" collectingDate="1966-10-29" collectionCode="ANIC" collectorName="E. Britton" country="Australia" location="Daintree" municipality="Shire of Douglas" pageId="7" pageNumber="142" specimenCount="1" stateProvince="Queensland">
<specimenCount id="9D24D5A4FF8ABA44FD7AF8E280E7F8BC" box="[677,750,1847,1873]" pageId="7" pageNumber="142">1 spm</specimenCount>
;
<collectingRegion id="49E6D0CFFF8ABA44FD26F8E3818EF8BD" box="[761,903,1846,1872]" country="Australia" name="Queensland" pageId="7" pageNumber="142">Queensland</collectingRegion>
,
<collectingMunicipality id="6BF98457FF8ABA44FC4DF8E38653F8BD" box="[914,1114,1846,1872]" pageId="7" pageNumber="142">Shire of Douglas</collectingMunicipality>
,
<location id="8EFD48F6FF8ABA44FBBBF8E386C4F8BD" LSID="urn:lsid:plazi:treatment:038BAF3BFF8ABA4AFDE4FD9386BFFE09:8EFD48F6FF8ABA44FBBBF8E386C4F8BD" box="[1124,1229,1846,1872]" country="Australia" municipality="Shire of Douglas" name="Daintree" pageId="7" pageNumber="142" stateProvince="Queensland">Daintree</location>
;
<date id="FF9C38EDFF8ABA44FB07F8E2877BF8BD" box="[1240,1394,1846,1873]" pageId="7" pageNumber="142" value="1966-10-29">
<collectingDate id="EFD8C105FF8ABA44FB07F8E2877BF8BD" box="[1240,1394,1846,1873]" pageId="7" pageNumber="142" value="1966-10-29">29 Oct. 1966</collectingDate>
</date>
;
<collectorName id="26D77BFBFF8ABA44FF63F88F8339F899" box="[188,304,1882,1908]" pageId="7" pageNumber="142">E. Britton</collectorName>
leg.;
<collectedFrom id="383E5DD0FF8ABA44FEADF88F80D2F899" box="[370,731,1882,1908]" pageId="7" pageNumber="142">dead leaves, edge of rainforest</collectedFrom>
;
<collectionCode id="ED3386E8FF8ABA44FD3AF88F8124F899" box="[741,813,1882,1908]" collectionName="Australia, Australian Capital Territory, Canberra City, CSIRO, Australian National Insect Collection" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="7" pageNumber="142">ANIC</collectionCode>
</materialsCitation>
<materialsCitation id="3B4A1470FF8ABA44FC99F88F872CF899" box="[838,1317,1882,1908]" collectedFrom="dead leaves, edge of rainforest" collectingDate="1966-10-29" collectionCode="MKF" collectorName="E. Britton" country="Australia" location="Daintree" municipality="Shire of Douglas" pageId="7" pageNumber="142" specimenCount="1" stateProvince="Queensland">
<specimenCount id="9D24D5A4FF8ABA44FC99F88F8199F899" box="[838,912,1882,1908]" pageId="7" pageNumber="142">1 spm</specimenCount>
; same data as for preceding;
<collectionCode id="ED3386E8FF8ABA44FB3BF88F872CF899" box="[1252,1317,1882,1908]" pageId="7" pageNumber="142">MKF</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B9D1E2DFF8ABA44FF62F84E83F9F834" blockId="7.[189,1399,1946,2009]" pageId="7" pageNumber="142">
<materialsCitation id="3B4A1470FF8ABA44FF62F84E83E5F834" collectingDate="1938-04" collectionCode="NHMUK" collectorName="Cheesman" country="Indonesia" elevation="762" latitude="-0.2" location="Camp Nok" longLatPrecision="1308" longitude="130.83333" municipality="Waigeo" pageId="7" pageNumber="142" specimenCount="2" stateProvince="West Papua">
<collectingCountry id="F3355EBDFF8ABA44FF62F84E8352F858" box="[189,347,1947,1973]" name="Indonesia" pageId="7" pageNumber="142">INDONESIA</collectingCountry>
<specimenCount id="9D24D5A4FF8ABA44FEAAF84E83C6F85B" box="[373,463,1947,1974]" pageId="7" pageNumber="142">2 specs</specimenCount>
;
<collectingRegion id="49E6D0CFFF8ABA44FE04F8498061F85B" box="[475,616,1948,1974]" country="Indonesia" name="Papua Barat" pageId="7" pageNumber="142">West Papua</collectingRegion>
,
<collectingMunicipality id="6BF98457FF8ABA44FDACF84E80C6F858" box="[627,719,1947,1973]" pageId="7" pageNumber="142">Waigeo</collectingMunicipality>
,
<location id="8EFD48F6FF8ABA44FD04F84E8156F858" LSID="urn:lsid:plazi:treatment:038BAF3BFF8ABA4AFDE4FD9386BFFE09:8EFD48F6FF8ABA44FD04F84E8156F858" box="[731,863,1947,1973]" country="Indonesia" latitude="-0.2" longLatPrecision="1308" longitude="130.83333" municipality="Waigeo" name="Camp Nok" pageId="7" pageNumber="142" stateProvince="West Papua">Camp Nok</location>
;
<elevation id="000FF91EFF8ABA44FCB3F84E8608F858" box="[876,1025,1947,1974]" metricMagnitude="2" metricUnit="m" metricValue="7.62" pageId="7" pageNumber="142" unit="ft" value="2500.0">
<quantity id="4CDAB3C8FF8ABA44FCB3F84E81C9F858" box="[876,960,1947,1974]" metricMagnitude="2" metricUnit="m" metricValue="7.62" pageId="7" pageNumber="142" unit="ft" value="2500.0">2500 ft</quantity>
a.s.l.
</elevation>
;
<geoCoordinate id="EE1678EAFF8ABA44FBD0F84F866EF858" box="[1039,1127,1946,1974]" degrees="0" direction="south" minutes="12" orientation="latitude" pageId="7" pageNumber="142" precision="925" value="-0.2">0°12 S</geoCoordinate>
,
<geoCoordinate id="EE1678EAFF8ABA44FBAAF84F86E4F85B" box="[1141,1261,1946,1974]" degrees="130" direction="east" minutes="50" orientation="longitude" pageId="7" pageNumber="142" precision="925" value="130.83333">130°50 E</geoCoordinate>
;
<date id="FF9C38EDFF8ABA44FB27F84E877BF858" box="[1272,1394,1947,1973]" pageId="7" pageNumber="142" value="1938-04">
<collectingDate id="EFD8C105FF8ABA44FB27F84E877BF858" box="[1272,1394,1947,1973]" pageId="7" pageNumber="142" value="1938-04">Apr. 1938</collectingDate>
</date>
;
<collectorName id="26D77BFBFF8ABA44FF62F86B833EF835" box="[189,311,1982,2008]" pageId="7" pageNumber="142">Cheesman</collectorName>
leg.;
<collectionCode id="ED3386E8FF8ABA44FEA6F86A83E5F834" box="[377,492,1983,2009]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="7" pageNumber="142" type="Museum">NHMUK</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF5D4EA5FF85BA4BFF62F8B180D7F804" ID-DOI="http://doi.org/10.5281/zenodo.4271758" ID-Zenodo-Dep="4271758" httpUri="https://zenodo.org/record/4271758/files/figure.png" pageId="8" pageNumber="143" startId="8.[189,232,1892,1918]" targetBox="[257,1339,265,1820]" targetPageId="8">
<paragraph id="8B9D1E2DFF85BA4BFF62F8B180D7F804" blockId="8.[188,1399,1892,2025]" pageId="8" pageNumber="143">
<emphasis id="B956C23FFF85BA4BFF62F8B18306F892" bold="true" box="[189,271,1892,1919]" pageId="8" pageNumber="143">Fig. 3.</emphasis>
<taxonomicName id="4C2265AEFF85BA4BFEC5F8B380E8F892" authority="Grouvelle, 1919" authorityName="Grouvelle" authorityYear="1919" box="[282,737,1893,1919]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="143" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="B956C23FFF85BA4BFEC5F8B38006F892" box="[282,527,1893,1919]" italics="true" pageId="8" pageNumber="143">Psammoecus obesus</emphasis>
<bibRefCitation id="EFB363DCFF85BA4BFDC4F8B080E8F892" author="Grouvelle A." box="[539,737,1893,1919]" pageId="8" pageNumber="143" refId="ref13309" refString="Grouvelle A. 1919. Descriptions d'especes nouvelles du genre Psammoecus. Memoires entomologiques. Etudes sur les Coleopteres. Deuxieme Fascicule. Societe entomologique de France, Paris." type="book" year="1919">Grouvelle, 1919</bibRefCitation>
</taxonomicName>
.
<emphasis id="B956C23FFF85BA4BFD2EF8B18123F893" bold="true" box="[753,810,1892,1918]" pageId="8" pageNumber="143">AD</emphasis>
. Holotype, ♀ (NHMUK).
<emphasis id="B956C23FFF85BA4BFBABF8B18680F893" bold="true" box="[1140,1161,1892,1918]" pageId="8" pageNumber="143">A</emphasis>
. Habitus.
<emphasis id="B956C23FFF85BA4BFAD4F8B08716F892" bold="true" box="[1291,1311,1893,1919]" pageId="8" pageNumber="143">B</emphasis>
. Head.
<emphasis id="B956C23FFF85BA4BFF62F85D82DBF84F" bold="true" box="[189,210,1928,1954]" pageId="8" pageNumber="143">C</emphasis>
. Pronotum.
<emphasis id="B956C23FFF85BA4BFEBFF85D837CF84F" bold="true" box="[352,373,1928,1954]" pageId="8" pageNumber="143">D</emphasis>
. Left antenna.
<emphasis id="B956C23FFF85BA4BFDFDF85D8052F84F" bold="true" box="[546,603,1928,1954]" pageId="8" pageNumber="143">EH</emphasis>
. Specimen from Queensland, ♂ (ANIC).
<emphasis id="B956C23FFF85BA4BFB9FF85D865DF84F" bold="true" box="[1088,1108,1928,1954]" pageId="8" pageNumber="143">E</emphasis>
. Aedeagus, ventral view.
<emphasis id="B956C23FFF85BA4BFF63F87E82C7F828" bold="true" box="[188,206,1963,1989]" pageId="8" pageNumber="143">F</emphasis>
. Aedeagus, lateral view.
<emphasis id="B956C23FFF85BA4BFE2AF87E8005F828" bold="true" box="[501,524,1963,1989]" pageId="8" pageNumber="143">G</emphasis>
. Parameres, ventral view.
<emphasis id="B956C23FFF85BA4BFC9CF87E8153F828" bold="true" box="[835,858,1963,1989]" pageId="8" pageNumber="143">H</emphasis>
. Median lobe, dorsal view. Scale bars: A, D = 1.0 mm; BC, EF = 0.5 mm; GH = 0.2 mm.
</paragraph>
</caption>
<subSubSection id="C3384DA6FF84BA4AFF62FEDD81EEFEBC" box="[189,999,264,338]" pageId="9" pageNumber="144" type="distribution">
<paragraph id="8B9D1E2DFF84BA4AFF62FEDD835FFECF" blockId="9.[189,342,264,290]" box="[189,342,264,290]" pageId="9" pageNumber="144">
<heading id="D0D5A941FF84BA4AFF62FEDD835FFECF" bold="true" box="[189,342,264,290]" fontSize="11" level="3" pageId="9" pageNumber="144" reason="3">
<emphasis id="B956C23FFF84BA4AFF62FEDD835FFECF" bold="true" box="[189,342,264,290]" pageId="9" pageNumber="144">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B9D1E2DFF84BA4AFF62FEE281EEFEBC" blockId="9.[189,999,311,338]" box="[189,999,311,338]" pageId="9" pageNumber="144">
The species has been found in
<collectingCountry id="F3355EBDFF84BA4AFDFCFEE28086FEBC" box="[547,655,311,337]" name="Australia" pageId="9" pageNumber="144">Australia</collectingCountry>
(
<collectingRegion id="49E6D0CFFF84BA4AFD41FEE28125FEBF" box="[670,812,311,338]" country="Australia" name="Queensland" pageId="9" pageNumber="144">Queensland</collectingRegion>
) and
<collectingCountry id="F3355EBDFF84BA4AFCB2FEE281EAFEBC" box="[877,995,311,337]" name="Indonesia" pageId="9" pageNumber="144">Indonesia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3384DA6FF84BA4AFF62FEAC86BFFE09" pageId="9" pageNumber="144" type="discussion">
<paragraph id="8B9D1E2DFF84BA4AFF62FEAC8326FE7E" blockId="9.[189,303,377,403]" box="[189,303,377,403]" pageId="9" pageNumber="144">
<heading id="D0D5A941FF84BA4AFF62FEAC8326FE7E" bold="true" box="[189,303,377,403]" fontSize="11" level="3" pageId="9" pageNumber="144" reason="3">
<emphasis id="B956C23FFF84BA4AFF62FEAC8326FE7E" bold="true" box="[189,303,377,403]" pageId="9" pageNumber="144">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B9D1E2DFF84BA4AFF62FE7D86BFFE09" blockId="9.[188,1398,423,485]" pageId="9" pageNumber="144">
<taxonomicName id="4C2265AEFF84BA4AFF62FE7D83A4FE2C" authorityName="Grouvelle" authorityYear="1919" box="[189,429,423,449]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="species" species="obesus">
<emphasis id="B956C23FFF84BA4AFF62FE7D83A4FE2C" box="[189,429,423,449]" italics="true" pageId="9" pageNumber="144">Psammoecus obesus</emphasis>
</taxonomicName>
is closely related to
<taxonomicName id="4C2265AEFF84BA4AFD44FE7D81D7FE2C" authority="Grouvelle, 1883" authorityName="Grouvelle" authorityYear="1883" box="[667,990,423,449]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="species" species="hirsutus">
<emphasis id="B956C23FFF84BA4AFD44FE7D811FFE2C" box="[667,790,423,449]" italics="true" pageId="9" pageNumber="144">P. hirsutus</emphasis>
Grouvelle, 1883
</taxonomicName>
,
<taxonomicName id="4C2265AEFF84BA4AFC36FE7D8749FE2C" authority="Grouvelle, 1919" authorityName="Grouvelle" authorityYear="1919" box="[1001,1344,423,449]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="species" species="decoratus">
<emphasis id="B956C23FFF84BA4AFC36FE7D8670FE2C" box="[1001,1145,423,449]" italics="true" pageId="9" pageNumber="144">P. decoratus</emphasis>
<bibRefCitation id="EFB363DCFF84BA4AFB5FFE728749FE2C" author="Grouvelle A." box="[1152,1344,423,449]" pageId="9" pageNumber="144" refId="ref13309" refString="Grouvelle A. 1919. Descriptions d'especes nouvelles du genre Psammoecus. Memoires entomologiques. Etudes sur les Coleopteres. Deuxieme Fascicule. Societe entomologique de France, Paris." type="book" year="1919">Grouvelle, 1919</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C2265AEFF84BA4AFF63FE1E800CFE09" authority="Grouvelle, 1919" authorityName="Grouvelle" authorityYear="1919" box="[188,517,458,484]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="144" phylum="Arthropoda" rank="species" species="signatus">
<emphasis id="B956C23FFF84BA4AFF63FE1E8332FE09" box="[188,315,459,484]" italics="true" pageId="9" pageNumber="144">P. signatus</emphasis>
<bibRefCitation id="EFB363DCFF84BA4AFE9CFE1F800CFE09" author="Grouvelle A." box="[323,517,458,484]" pageId="9" pageNumber="144" refId="ref13309" refString="Grouvelle A. 1919. Descriptions d'especes nouvelles du genre Psammoecus. Memoires entomologiques. Etudes sur les Coleopteres. Deuxieme Fascicule. Societe entomologique de France, Paris." type="book" year="1919">Grouvelle, 1919</bibRefCitation>
</taxonomicName>
. It differs from all three by the shape of the male genitalia.
</paragraph>
</subSubSection>
</treatment>
</document>