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<document id="9DD4516345683FB2D1BCD74D1C6B5316" ID-CLB-Dataset="4031" ID-DOI="10.5852/ejt.2021.750.1365" ID-GBIF-Dataset="818d0bc4-31a6-4319-86a3-6e6b16ab72df" ID-ISSN="2118-9773" ID-Zenodo-Dep="5451747" ID-ZooBank="CE5D8517-AC04-4AF9-BD78-A2D6E21D33A8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630516667691" checkinUser="felipe" docAuthor="Pacheco, Thaynara L., Monné, Marcela L., Vaz-De, Fernando Z. &amp; Ahrens, Dirk" docDate="2021" docId="038B976CFF9DFFF6CA50C8911116CA05" docLanguage="en" docName="ejt-750_94-123.pdf" docOrigin="European Journal of Taxonomy 750 (750)" docSource="http://dx.doi.org/10.5852/ejt.2021.750.1365" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Sayloria abcora Pacheco &amp; Monné &amp; Vaz-De &amp; Ahrens 2021, comb. nov." docType="treatment" docVersion="9" lastPageNumber="118" masterDocId="FFB2EF14FF8BFFEEC84EC925155FCE04" masterDocTitle="Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor" masterLastPageNumber="123" masterPageNumber="94" pageNumber="116" updateTime="1699290068827" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="DCD08E91904C9ABD07E20044FE2BD440">Revision of type specimens of Astaena (Coleoptera: Scarabaeidae: Melolonthinae: Sericini) described by L. W. Saylor</mods:title>
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<mods:name id="3C60E9184FF5EBD20FE77F4CD8DC0486" type="personal">
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<mods:namePart id="FCC89B2D2AC1A5D1A89043F0569450A9">Pacheco, Thaynara L.</mods:namePart>
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<mods:affiliation id="10B2F1996E31E3BD2A22CFCEB6FC52F0">Laboratório de Coleoptera, Departamento de Entomologia, Museu Nacional, Universidade Federal do Rio de Janeiro, Rio de Janeiro, Brazil. &amp; Centre of Taxonomy and Evolutionary Research, Zoologisches Forschungsmuseum Alexander Koenig, Bonn, Germany.</mods:affiliation>
<mods:nameIdentifier id="A2155A79B8D4B42FE2C8D1CD244CB85E" type="email">thay.lpacheco@gmail.com</mods:nameIdentifier>
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<mods:roleTerm id="55F794E6BED4E29CC5088B0A1CC28C84">Author</mods:roleTerm>
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<mods:namePart id="7241A6509213257DCA92019A3F7D65E9">Monné, Marcela L.</mods:namePart>
<mods:nameIdentifier id="5493B00AE2F16C8AAC2EE203A7423B32" type="LSID">86B186B4-4297-4380-A98D-FC813B5D53A6&amp;2FF2B7D6-1A6B-43C1-9966-A1A949FB2B05</mods:nameIdentifier>
<mods:affiliation id="03B4BAD6A1CDB79FE071B26C548ED744">Laboratório de Coleoptera, Departamento de Entomologia, Museu Nacional, Universidade Federal do Rio de Janeiro, Rio de Janeiro, Brazil. &amp; Laboratório de Scarabaeoidologia, Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Cuiabá, Brazil.</mods:affiliation>
<mods:nameIdentifier id="11E3E0FB0A141DE8F00CF1143ACA558A" type="email">mlmonne2@gmail.com&amp;vazdemello@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="5D078EE5C80045091A43B81E65640267" type="personal">
<mods:role id="E12E1789354457A4A337B7425B970E2E">
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<mods:namePart id="7E662FB24954C15D5E6F72DC11E39FF9">Vaz-De, Fernando Z.</mods:namePart>
<mods:nameIdentifier id="0B1329DB87140177C1883131028ABA18" type="LSID">2FF2B7D6-1A6B-43C1-9966-A1A949FB2B05</mods:nameIdentifier>
<mods:affiliation id="805EB1E61885974C644774E124D494C4">Laboratório de Scarabaeoidologia, Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Cuiabá, Brazil.</mods:affiliation>
<mods:nameIdentifier id="924655147D28C92C049F93D39A045C65" type="email">vazdemello@gmail.com</mods:nameIdentifier>
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<mods:namePart id="25A816A2572D64C9EE8067DF492A9CB6">Ahrens, Dirk</mods:namePart>
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<mods:affiliation id="C22671EB73A59ABD476835D8B3CC0E75">Centre of Taxonomy and Evolutionary Research, Zoologisches Forschungsmuseum Alexander Koenig, Bonn, Germany.</mods:affiliation>
<mods:nameIdentifier id="31285C92F627FDEA530336D5E8586AF8" type="email">ahrens.dirk_col@gmx.de</mods:nameIdentifier>
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<mods:title id="C749573F31C4AAFAAD5A8E8EDD745A8B">European Journal of Taxonomy</mods:title>
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<mods:date id="608D97F720C1BBE40A0D1FA0421D5B7E">2021</mods:date>
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<mods:number id="2F90A7BD84EBD41C45CEF2E07F883A3B">2021-05-19</mods:number>
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<mods:identifier id="1B078DA979F664D63E72A507D5979452" type="CLB-Dataset">4031</mods:identifier>
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<treatment id="038B976CFF9DFFF6CA50C8911116CA05" ID-DOI="http://doi.org/10.5281/zenodo.5572738" ID-GBIF-Taxon="188647612" ID-Zenodo-Dep="5572738" LSID="urn:lsid:plazi:treatment:038B976CFF9DFFF6CA50C8911116CA05" httpUri="http://treatment.plazi.org/id/038B976CFF9DFFF6CA50C8911116CA05" lastPageId="24" lastPageNumber="118" pageId="22" pageNumber="116">
<subSubSection id="C33875F1FF9DFFF8CA50C8911149CFCB" box="[542,1046,436,463]" pageId="22" pageNumber="116" type="nomenclature">
<paragraph id="8B9D267AFF9DFFF8CA50C8911149CFCB" blockId="22.[542,1046,436,498]" box="[542,1046,436,463]" pageId="22" pageNumber="116">
<heading id="D0D59116FF9DFFF8CA50C8911149CFCB" box="[542,1046,436,463]" centered="true" fontSize="11" level="2" pageId="22" pageNumber="116" reason="2">
<taxonomicName id="4C225DF9FF9DFFF8CA50C89116D2CFCB" authority="(Saylor, 1946)" authorityName="Pacheco &amp; Monné &amp; Vaz-De &amp; Ahrens" authorityYear="2021" baseAuthorityName="Saylor" baseAuthorityYear="1946" box="[542,909,436,463]" class="Insecta" family="Melolonthidae" genus="Sayloria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="116" phylum="Arthropoda" rank="species" species="abcora" status="comb. nov.">
<emphasis id="B956FA68FF9DFFF8CA50C8911782CFCB" bold="true" box="[542,733,436,463]" italics="true" pageId="22" pageNumber="116">Sayloria abcora</emphasis>
(
<bibRefCitation id="EFB35B8BFF9DFFF8CAA3C89116DACFCB" author="Saylor L. W." box="[749,901,436,463]" pageId="22" pageNumber="116" pagination="215 - 232" refId="ref13932" refString="Saylor L. W. 1946. New South American scarab beetles of the genus Astaena. Revista de Entomologia 17: 215 - 232." type="journal article" year="1946">Saylor, 1946</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A2654713FF9DFFF8CBDAC8911149CFCB" box="[916,1046,436,463]" pageId="22" pageNumber="116" rank="species">comb. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33875F1FF9DFFF8CA88C8FD1632CFF6" box="[710,877,472,498]" pageId="22" pageNumber="116" type="description">
<paragraph id="8B9D267AFF9DFFF8CA88C8FD1632CFF6" blockId="22.[542,1046,436,498]" box="[710,877,472,498]" pageId="22" pageNumber="116">
<figureCitation id="13193AFFFF9DFFF8CA88C8FD1660CFF6" box="[710,831,472,498]" captionStart="Fig" captionStartId="23.[190,233,1858,1884]" captionTargetBox="[244,1344,265,1820]" captionTargetId="figure-16@23.[244,1344,265,1820]" captionTargetPageId="23" captionText="Fig. 6. A–E. Sayloria abcora (Saylor, 1946), holotype, ♂ (Type No. 7905; CAS). – F–I. Astaena apolinarmaria Saylor, 1946, holotype, ♀ (Type No. 7906; CAS). – J–K. Sayloria pottsi (Saylor, 1946), paratype, ♂ (CAS). – L–N. S. pottsi, holotype, ♀ (Type No. 7916; CAS). A, J. Parameres, dorsal view. B, K. Aedeagus, lateral view. C, G, L. Labels. D, H, M. Habitus, dorsal view. E, I, N. Habitus, lateral view. F. Head and pronotum, dorsal view. Scale bars: A–B, F, J–K = 0.5 mm; D–E, H–I, M–N = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5451759" httpUri="https://zenodo.org/record/5451759/files/figure.png" pageId="22" pageNumber="116">Figs 6A–I</figureCitation>
,
<figureCitation id="13193AFFFF9DFFF8CB05C8FD1632CFF6" box="[843,877,472,498]" captionStart="Fig" captionStartId="26.[189,232,1822,1848]" captionTargetBox="[261,1326,295,1750]" captionTargetId="figure-12@26.[241,1347,265,1784]" captionTargetPageId="26" captionText="Fig. 7. Distribution of type localities of the South American Sericini species described by Saylor. A. Astaena abaca Saylor, 1946, A. biciliata Saylor, 1946, A. bogotana Saylor, 1946, A. excisipes Saylor, 1947, A. explaniceps Saylor, 1947, A. incachaca Saylor, 1946, A. maqueta Saylor, 1947, and A. nigrona Saylor, 1947. B. A. fusagona Saylor, 1946, A. pygidia Saylor, 1946, A. rugithorax Saylor, 1946, A. salta Saylor, 1946, A. yungasa Saylor, 1946, and A. zyrota Saylor, 1946. C. Sayloria abcora (Saylor, 1946) comb. nov., S. bicoloripes (Saylor, 1946) comb. nov., and S. pottsi (Saylor, 1946) comb. nov.." figureDoi="http://doi.org/10.5281/zenodo.5451761" httpUri="https://zenodo.org/record/5451761/files/figure.png" pageId="22" pageNumber="116">7C</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33875F1FF9DFFF8C8F3CB3E163DCD2F" pageId="22" pageNumber="116" type="reference_group">
<paragraph id="8B9D267AFF9DFFF8C8F3CB3E170ACC30" blockId="22.[189,743,538,601]" box="[189,597,538,565]" pageId="22" pageNumber="116">
<treatmentCitationGroup id="AB320154FF9DFFF8C8F3CB3E170ACC30" box="[189,597,538,565]" pageId="22" pageNumber="116">
<treatmentCitation id="0A83006BFF9DFFF8C8F3CB3E170ECC30" author="Saylor L. W." box="[189,593,538,565]" page="224" pageId="22" pageNumber="116" year="1946">
<taxonomicName id="4C225DF9FF9DFFF8C8F3CB3E170ECC30" authority="Saylor, 1946: 224" authorityName="L. W. Saylor" authorityPageNumber="224" authorityYear="1946" box="[189,593,538,565]" class="Insecta" family="Melolonthidae" genus="Astaena" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="116" phylum="Arthropoda" rank="species" species="abcora">
<emphasis id="B956FA68FF9DFFF8C8F3CB3E1429CC30" box="[189,374,538,565]" italics="true" pageId="22" pageNumber="116">Astaena abcora</emphasis>
<bibRefCitation id="EFB35B8BFF9DFFF8C933CB3F170ECC30" author="Saylor L. W." box="[381,593,538,564]" pageId="22" pageNumber="116" pagination="215 - 232" refId="ref13932" refString="Saylor L. W. 1946. New South American scarab beetles of the genus Astaena. Revista de Entomologia 17: 215 - 232." type="journal article" year="1946">Saylor, 1946: 224</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9D267AFF9DFFF8C8F3CB1A17B8CC5C" blockId="22.[189,743,538,601]" box="[189,743,574,601]" pageId="22" pageNumber="116">
<taxonomicName id="4C225DF9FF9DFFF8C8F3CB1A172BCC5C" authority="Saylor, 1946" authorityName="L. W. Saylor" authorityYear="1946" box="[189,628,574,601]" class="Insecta" family="Melolonthidae" genus="Astaena" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="116" phylum="Arthropoda" rank="species" species="apolinar-maria">
<emphasis id="B956FA68FF9DFFF8C8F3CB1A1488CC5C" box="[189,471,574,601]" italics="true" pageId="22" pageNumber="116">Astaena apolinar-maria</emphasis>
<bibRefCitation id="EFB35B8BFF9DFFF8C990CB1B172BCC5C" author="Saylor L. W." box="[478,628,574,600]" pageId="22" pageNumber="116" pagination="215 - 232" refId="ref13932" refString="Saylor L. W. 1946. New South American scarab beetles of the genus Astaena. Revista de Entomologia 17: 215 - 232." type="journal article" year="1946">Saylor, 1946</bibRefCitation>
</taxonomicName>
<emphasis id="B956FA68FF9DFFF8CA35CB1B17B8CC5C" bold="true" box="[635,743,574,600]" pageId="22" pageNumber="116">
<taxonomicNameLabel id="A2654713FF9DFFF8CA35CB1B17B8CC5C" box="[635,743,574,600]" pageId="22" pageNumber="116">syn. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8B9D267AFF9DFFF8C8F3CBA41029CC9E" blockId="22.[189,1398,640,811]" box="[189,1398,640,667]" pageId="22" pageNumber="116">
<treatmentCitationGroup id="AB320154FF9DFFF8C8F3CBA41029CC9E" box="[189,1398,640,667]" pageId="22" pageNumber="116">
<taxonomicName id="4C225DF9FF9DFFF8C8F3CBA4142ACC9E" authorityName="L. W. Saylor" authorityYear="1946" box="[189,373,640,667]" class="Insecta" family="Melolonthidae" genus="Astaena" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="116" phylum="Arthropoda" rank="species" species="abcora">
<emphasis id="B956FA68FF9DFFF8C8F3CBA4142ACC9E" box="[189,373,640,667]" italics="true" pageId="22" pageNumber="116">Astaena abcora</emphasis>
</taxonomicName>
–
<treatmentCitation id="0A83006BFF9DFFF8C9DFCBA4171CCC9E" author="Frey G." box="[401,579,640,667]" page="348" pageId="22" pageNumber="116" year="1973">
<bibRefCitation id="EFB35B8BFF9DFFF8C9DFCBA4171CCC9E" author="Frey G." box="[401,579,640,667]" pageId="22" pageNumber="116" pagination="315 - 366" refId="ref13316" refString="Frey G. 1973. Synopsis der sudamerikanischen Sericinen. Entomologische Arbeiten aus dem Museum Frey 1973: 315 - 366." type="journal article" year="1973">Frey 1973: 348</bibRefCitation>
</treatmentCitation>
(identification key). —
<treatmentCitation id="0A83006BFF9DFFF8CB15CBA4114FCC9E" author="Evans A. V." box="[859,1040,640,667]" page="19" pageId="22" pageNumber="116" year="2003">
<bibRefCitation id="EFB35B8BFF9DFFF8CB15CBA4114FCC9E" author="Evans A. V." box="[859,1040,640,667]" pageId="22" pageNumber="116" pagination="1 - 458" refId="ref13003" refString="Evans A. V. 2003. A checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa 211: 1 - 458. https: // doi. org / 10.11646 / zootaxa. 211.1.1" type="journal article" year="2003">Evans 2003: 19</bibRefCitation>
</treatmentCitation>
(checklist). — Evans &amp;
<bibRefCitation id="EFB35B8BFF9DFFF8CD7ECBA51029CC9E" author="Smith A. B. T. &amp; Evans A. V." box="[1328,1398,640,666]" pageId="22" pageNumber="116" pagination="29 - 60" refId="ref14061" refString="Smith A. B. T. &amp; Evans A. V. 2005. A supplement to the checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae) with notes on their tribal classification. Zootaxa 1032: 29 - 60. https: // doi. org / 10.11646 / zootaxa. 1032.1.2" type="journal article" year="2005">Smith</bibRefCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9D267AFF9DFFF8C8A4CB811029CD02" blockId="22.[189,1398,640,811]" pageId="22" pageNumber="116">
2005: 29 (checklist); 2007: 30 (checklist); 2009: 30 (checklist).
<taxonomicName id="4C225DF9FF9DFFF8C8F3CBED1488CCE6" authorityName="L. W. Saylor" authorityYear="1946" box="[189,471,712,738]" class="Insecta" family="Melolonthidae" genus="Astaena" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="116" phylum="Arthropoda" rank="species" species="apolinar-maria">
<emphasis id="B956FA68FF9DFFF8C8F3CBED1488CCE6" box="[189,471,712,738]" italics="true" pageId="22" pageNumber="116">Astaena apolinar-maria</emphasis>
</taxonomicName>
–
<bibRefCitation id="EFB35B8BFF9DFFF8C9BACBEC17F7CCE7" author="Frey G." box="[500,680,712,739]" pageId="22" pageNumber="116" pagination="315 - 366" refId="ref13316" refString="Frey G. 1973. Synopsis der sudamerikanischen Sericinen. Entomologische Arbeiten aus dem Museum Frey 1973: 315 - 366." type="journal article" year="1973">Frey 1973: 352</bibRefCitation>
(identification key).
<taxonomicName id="4C225DF9FF9DFFF8C8F3CBC917C4CD02" authority="Evans 2003: 19" authorityName="Evans" authorityPageNumber="19" authorityYear="2003" box="[189,667,748,775]" class="Insecta" family="Melolonthidae" genus="Astaena" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="116" phylum="Arthropoda" rank="species" species="apolinarmaria">
<emphasis id="B956FA68FF9DFFF8C8F3CBC9148DCD02" box="[189,466,748,774]" italics="true" pageId="22" pageNumber="116">Astaena apolinarmaria</emphasis>
<bibRefCitation id="EFB35B8BFF9DFFF8C993CBC817C4CD02" author="Evans A. V." box="[477,667,748,775]" pageId="22" pageNumber="116" pagination="1 - 458" refId="ref13003" refString="Evans A. V. 2003. A checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa 211: 1 - 458. https: // doi. org / 10.11646 / zootaxa. 211.1.1" type="journal article" year="2003">Evans 2003: 19</bibRefCitation>
</taxonomicName>
(spelling corrected, checklist). — Evans &amp;
<bibRefCitation id="EFB35B8BFF9DFFF8CCF4CBC91029CD02" author="Smith A. B. T. &amp; Evans A. V." box="[1210,1398,748,775]" pageId="22" pageNumber="116" pagination="29 - 60" refId="ref14061" refString="Smith A. B. T. &amp; Evans A. V. 2005. A supplement to the checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae) with notes on their tribal classification. Zootaxa 1032: 29 - 60. https: // doi. org / 10.11646 / zootaxa. 1032.1.2" type="journal article" year="2005">Smith 2005: 29</bibRefCitation>
</paragraph>
<paragraph id="8B9D267AFF9DFFF8C8A4CA35163DCD2F" blockId="22.[189,1398,640,811]" box="[234,866,784,811]" pageId="22" pageNumber="116">(checklist); 2007: 30 (checklist); 2009: 30 (checklist).</paragraph>
</subSubSection>
<subSubSection id="C33875F1FF9DFFF8C8F3CA771005C818" pageId="22" pageNumber="116" type="materials_examined">
<paragraph id="8B9D267AFF9DFFF8C8F3CA7714B3CD68" blockId="22.[189,492,850,876]" box="[189,492,850,876]" pageId="22" pageNumber="116">
<heading id="D0D59116FF9DFFF8C8F3CA7714B3CD68" bold="true" box="[189,492,850,876]" fontSize="11" level="3" pageId="22" pageNumber="116" reason="3">
<emphasis id="B956FA68FF9DFFF8C8F3CA7714B3CD68" bold="true" box="[189,492,850,876]" pageId="22" pageNumber="116">Type material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B9D267AFF9DFFF8C8A4CAA411CBCDE7" blockId="22.[189,1399,897,996]" pageId="22" pageNumber="116">
<materialsCitation id="3B4A2C27FF9DFFF8C8A4CAA411CBCDE7" ID-GBIF-Occurrence="3383726319" collectingDate="ii-3" collectionCode="CAS" collectorName="L. W. Saylor Collection" country="Colombia" elevation="1143" location="Colombia" municipality="California Academy of Sciences Type" pageId="22" pageNumber="116" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B956FA68FF9DFFF8C8A4CAA41403CD9F" bold="true" box="[234,348,897,923]" pageId="22" pageNumber="116">
<typeStatus id="549998D8FF9DFFF8C8A4CAA41403CD9F" box="[234,348,897,923]" pageId="22" pageNumber="116" type="holotype">Holotype</typeStatus>
</emphasis>
(
<taxonomicName id="4C225DF9FF9DFFF8C920CAA71779CD9F" authorityName="L. W. Saylor" authorityYear="1946" box="[366,550,897,924]" class="Insecta" family="Melolonthidae" genus="Astaena" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="116" phylum="Arthropoda" rank="species" species="abcora">
<emphasis id="B956FA68FF9DFFF8C920CAA71779CD9F" box="[366,550,897,924]" italics="true" pageId="22" pageNumber="116">Astaena abcora</emphasis>
</taxonomicName>
)
<collectingCountry id="F33566EAFF9DFFF8C8F3CA801407CDBB" box="[189,344,933,959]" name="Colombia" pageId="22" pageNumber="116">COLOMBIA</collectingCountry>
•
<specimenCount id="9D24EDF3FF9DFFF8C921CA8314D7CDC4" box="[367,392,934,960]" pageId="22" pageNumber="116" type="male">♂</specimenCount>
; “Buenaventura alt.
<quantity id="4CDA8B9FFF9DFFF8CA38CA801649CDBB" box="[630,790,933,960]" metricMagnitude="3" metricUnit="m" metricValue="1.143" metricValueMax="1.2192" metricValueMin="1.0668" pageId="22" pageNumber="116" unit="ft" value="3750.0" valueMax="4000.0" valueMin="3500.0">
<elevation id="000FC149FF9DFFF8CA38CA801649CDBB" box="[630,790,933,960]" metricMagnitude="3" metricUnit="m" metricValue="1.143" metricValueMax="1.2192" metricValueMin="1.0668" pageId="22" pageNumber="116" unit="ft" value="3750.0" valueMax="4000.0" valueMin="3500.0">3500-4000 ft.</elevation>
</quantity>
<location id="8EFD70A1FF9DFFF8CB52CA8016CCCDBB" LSID="urn:lsid:plazi:treatment:038B976CFF9DFFF6CA50C8911116CA05:8EFD70A1FF9DFFF8CB52CA8016CCCDBB" box="[796,915,933,959]" country="Colombia" municipality="California Academy of Sciences Type" name="Colombia" pageId="22" pageNumber="116">Colombia</location>
,
<date id="FF9C00BAFF9DFFF8CBD0CA801698CDBB" box="[926,967,933,959]" pageId="22" pageNumber="116">
<collectingDate id="EFD8F952FF9DFFF8CBD0CA801698CDBB" box="[926,967,933,959]" pageId="22" pageNumber="116">ii-3</collectingDate>
</date>
/ L.W. Saylor Collection / California Academy of Sciences Type nº 7905 / Holotype
<taxonomicName id="4C225DF9FF9DFFF8CAA7CAEC1162CDE7" authority="L. W. Saylor" authorityName="L. W. Saylor" authorityYear="1946" box="[745,1085,969,996]" class="Insecta" family="Melolonthidae" genus="Astaena" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="116" phylum="Arthropoda" rank="species" species="abcora">
<emphasis id="B956FA68FF9DFFF8CAA7CAEC16FECDE7" box="[745,929,969,995]" italics="true" pageId="22" pageNumber="116">Astaena abcora</emphasis>
L.W. Saylor
</taxonomicName>
”;
<collectionCode id="ED33BEBFFF9DFFF8CC1DCAEC11CFCDE7" box="[1107,1168,969,995]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="22" pageNumber="116" type="Museum">CAS</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8B9D267AFF9DFFF8C8A4CD2E1758CA95" blockId="22.[187,1399,1035,1169]" pageId="22" pageNumber="116">
<materialsCitation id="3B4A2C27FF9DFFF8C8A4CD2E1029CA4D" ID-GBIF-Occurrence="3383726310" collectingDate="ii-3" collectorName="L. W. Saylor Collection &amp; Collection" country="Colombia" elevation="1143" location="COLOMBIA" pageId="22" pageNumber="116" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<emphasis id="B956FA68FF9DFFF8C8A4CD2E1403CA21" bold="true" box="[234,348,1035,1061]" pageId="22" pageNumber="116">
<typeStatus id="549998D8FF9DFFF8C8A4CD2E1403CA21" box="[234,348,1035,1061]" pageId="22" pageNumber="116" type="paratype">Paratype</typeStatus>
</emphasis>
(
<taxonomicName id="4C225DF9FF9DFFF8C923CD2E1779CA21" authorityName="L. W. Saylor" authorityYear="1946" box="[365,550,1035,1061]" class="Insecta" family="Melolonthidae" genus="Astaena" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="116" phylum="Arthropoda" rank="species" species="abcora">
<emphasis id="B956FA68FF9DFFF8C923CD2E1779CA21" box="[365,550,1035,1061]" italics="true" pageId="22" pageNumber="116">Astaena abcora</emphasis>
</taxonomicName>
)
<collectingCountry id="F33566EAFF9DFFF8C8F3CD0A1407CA4D" box="[189,344,1071,1097]" name="Colombia" pageId="22" pageNumber="116">COLOMBIA</collectingCountry>
•
<specimenCount id="9D24EDF3FF9DFFF8C922CD0A14C8CA4E" box="[364,407,1071,1098]" pageId="22" pageNumber="116" type="female">1 ♀</specimenCount>
; “Buenaventura alt.
<quantity id="4CDA8B9FFF9DFFF8CACCCD0A167FCA4D" box="[642,800,1071,1098]" metricMagnitude="3" metricUnit="m" metricValue="1.143" metricValueMax="1.2192" metricValueMin="1.0668" pageId="22" pageNumber="116" unit="ft" value="3750.0" valueMax="4000.0" valueMin="3500.0">
<elevation id="000FC149FF9DFFF8CACCCD0A167FCA4D" box="[642,800,1071,1098]" metricMagnitude="3" metricUnit="m" metricValue="1.143" metricValueMax="1.2192" metricValueMin="1.0668" pageId="22" pageNumber="116" unit="ft" value="3750.0" valueMax="4000.0" valueMin="3500.0">3500-4000 ft.</elevation>
</quantity>
Colombia,
<date id="FF9C00BAFF9DFFF8CBEACD0A1692CA4D" box="[932,973,1071,1097]" pageId="22" pageNumber="116">
<collectingDate id="EFD8F952FF9DFFF8CBEACD0A1692CA4D" box="[932,973,1071,1097]" pageId="22" pageNumber="116">ii-3</collectingDate>
</date>
/ L.W. Saylor Collection / Collection
</materialsCitation>
<materialsCitation id="3B4A2C27FF9DFFF8C8F3CD761758CA95" ID-GBIF-Occurrence="3383726303" collectionCode="CAS" location="San Francisco" municipality="Academy of Sciences" pageId="22" pageNumber="116" specimenCode="CASENT 8438114" specimenCount="1" stateProvince="California" typeStatus="allotype">
of the California Academy of Sciences, San Francisco, California / Allotype
<taxonomicName id="4C225DF9FF9DFFF8CC62CD761028CA69" authority="L. W. Saylor" authorityName="L. W. Saylor" authorityYear="1946" box="[1068,1399,1107,1133]" class="Insecta" family="Melolonthidae" genus="Astaena" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="116" phylum="Arthropoda" rank="species" species="abcora">
<emphasis id="B956FA68FF9DFFF8CC62CD7611BCCA69" box="[1068,1251,1107,1133]" italics="true" pageId="22" pageNumber="116">Astaena abcora</emphasis>
L.W. Saylor
</taxonomicName>
/
<specimenCode id="DB848E01FF9DFFF8C885CD5214EFCA95" box="[203,432,1143,1169]" pageId="22" pageNumber="116">CASENT 8438114</specimenCode>
”;
<collectionCode id="ED33BEBFFF9DFFF8C988CD52175CCA95" box="[454,515,1143,1169]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="22" pageNumber="116" type="Museum">CAS</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8B9D267AFF9DFFF8C8A4CD9C1124CB1F" blockId="22.[189,1398,1209,1307]" pageId="22" pageNumber="116">
<materialsCitation id="3B4A2C27FF9DFFF8C8A4CD9C1124CB1F" ID-GBIF-Occurrence="3383726320" collectionCode="CAS" collectorName="L. W. Saylor Collection" country="Colombia" location="COLOMBIA" pageId="22" pageNumber="116" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="B956FA68FF9DFFF8C8A4CD9C1403CAD7" bold="true" box="[234,348,1209,1235]" pageId="22" pageNumber="116">
<typeStatus id="549998D8FF9DFFF8C8A4CD9C1403CAD7" box="[234,348,1209,1235]" pageId="22" pageNumber="116" type="holotype">Holotype</typeStatus>
</emphasis>
(
<taxonomicName id="4C225DF9FF9DFFF8C920CD9C1721CAD7" authorityName="Evans" authorityYear="2003" box="[366,638,1209,1235]" class="Insecta" family="Melolonthidae" genus="Astaena" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="116" phylum="Arthropoda" rank="species" species="apolinarmaria">
<emphasis id="B956FA68FF9DFFF8C920CD9C1721CAD7" box="[366,638,1209,1235]" italics="true" pageId="22" pageNumber="116">Astaena apolinarmaria</emphasis>
</taxonomicName>
)
<collectingCountry id="F33566EAFF9DFFF8C8F3CDF81407CAF3" box="[189,344,1245,1271]" name="Colombia" pageId="22" pageNumber="116">COLOMBIA</collectingCountry>
•
<specimenCount id="9D24EDF3FF9DFFF8C920CDF814DACAF3" box="[366,389,1245,1271]" pageId="22" pageNumber="116" type="female">♀</specimenCount>
; “Pensilvania Colomb / L.W. Saylor Collection / California Academy of Sciences Type nº 7906 / Apolinar-Maria / Holotype
<taxonomicName id="4C225DF9FF9DFFF8CA21CC24117BCB1F" authority="L. W. Saylor" authorityName="L. W. Saylor" authorityYear="1946" box="[623,1060,1281,1307]" class="Insecta" family="Melolonthidae" genus="Astaena" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="116" phylum="Arthropoda" rank="species" species="apolinar-maria">
<emphasis id="B956FA68FF9DFFF8CA21CC2416D6CB1F" box="[623,905,1281,1307]" italics="true" pageId="22" pageNumber="116">Astaena apolinar-maria</emphasis>
L.W. Saylor
</taxonomicName>
”;
<collectionCode id="ED33BEBFFF9DFFF8CC74CC241128CB1F" box="[1082,1143,1281,1307]" collectionName="USA, California, San Francisco, California Academy of Sciences" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="22" pageNumber="116" type="Museum">CAS</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8B9D267AFF9DFFF8C8F3CC67176BCB58" blockId="22.[189,564,1346,1372]" box="[189,564,1346,1372]" pageId="22" pageNumber="116">
<heading id="D0D59116FF9DFFF8C8F3CC67176BCB58" bold="true" box="[189,564,1346,1372]" fontSize="11" level="3" pageId="22" pageNumber="116" reason="3">
<emphasis id="B956FA68FF9DFFF8C8F3CC67176BCB58" bold="true" box="[189,564,1346,1372]" pageId="22" pageNumber="116">Additional material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B9D267AFF9DFFF8C8F3CC571005C818" blockId="22.[189,1399,1394,1564]" pageId="22" pageNumber="116">
<materialsCitation id="3B4A2C27FF9DFFF8C8F3CC571029CB88" ID-GBIF-Occurrence="3383726333" box="[189,1398,1394,1420]" collectingDate="1973-04-20" collectionCode="CMNC" collectorName="J. Helava" country="Colombia" elevation="1800" location="Antioquia" pageId="22" pageNumber="116" specimenCount="5" specimenCount-female="1" specimenCount-male="4" stateProvince="Antioquia">
<collectingCountry id="F33566EAFF9DFFF8C8F3CC571407CB88" box="[189,344,1394,1420]" name="Colombia" pageId="22" pageNumber="116">COLOMBIA</collectingCountry>
•
<specimenCount id="9D24EDF3FF9DFFF8C937CC57149ECB88" box="[377,449,1394,1420]" pageId="22" pageNumber="116" type="male">4 ♂♂</specimenCount>
,
<specimenCount id="9D24EDF3FF9DFFF8C99CCC57175BCB88" box="[466,516,1394,1420]" pageId="22" pageNumber="116" type="female">1 ♀</specimenCount>
; “COLOM:
<collectingRegion id="49E6E898FF9DFFF8CAD1CC57164BCB88" box="[671,788,1394,1420]" country="Colombia" name="Antioquia" pageId="22" pageNumber="116">Antioquia</collectingRegion>
<quantity id="4CDA8B9FFF9DFFF8CB6ECC57162BCB88" box="[800,884,1394,1420]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="22" pageNumber="116" unit="m" value="1800.0">
<elevation id="000FC149FF9DFFF8CB6ECC57162BCB88" box="[800,884,1394,1420]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="22" pageNumber="116" unit="m" value="1800.0">1800m</elevation>
</quantity>
, nr. Yarumai
<date id="FF9C00BAFF9DFFF8CC53CC571122CB88" box="[1053,1149,1394,1420]" pageId="22" pageNumber="116" value="1973-04-20">
<collectingDate id="EFD8F952FF9DFFF8CC53CC571122CB88" box="[1053,1149,1394,1420]" pageId="22" pageNumber="116" value="1973-04-20">20.iv.73</collectingDate>
</date>
,
<collectorName id="26D743ACFF9DFFF8CCC0CC57105ACB88" box="[1166,1285,1394,1420]" pageId="22" pageNumber="116">J. Helava</collectorName>
”;
<collectionCode id="ED33BEBFFF9DFFF8CD6ECC571029CB88" box="[1312,1398,1394,1420]" collectionName="Canada, Ottawa, Canadian Museum of Nature" pageId="22" pageNumber="116">CMNC</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B4A2C27FF9DFFF8C89DCCB3103ECBB4" ID-GBIF-Occurrence="3383726312" box="[211,1377,1430,1457]" collectionCode="ZFMK, MNHN" collectorName="Manizales A. M. Patino" country="Colombia" pageId="22" pageNumber="116" specimenCount="30" specimenCount-female="11" specimenCount-male="19">
<specimenCount id="9D24EDF3FF9DFFF8C89DCCB31476CBB4" box="[211,297,1430,1456]" pageId="22" pageNumber="116" type="male">19 ♂♂</specimenCount>
,
<specimenCount id="9D24EDF3FF9DFFF8C974CCB314D1CBB4" box="[314,398,1430,1456]" pageId="22" pageNumber="116" type="female">11 ♀♀</specimenCount>
; “Manizales A.M. Patino / Slg. R. Oberthür Eing. Nr.4, 1956”;
<collectionCode id="ED33BEBFFF9DFFF8CCEDCCB311A7CBB4" box="[1187,1272,1430,1456]" collectionName="Germany, Bonn, Zoologische Forschungsinstitut und Museum &quot;Alexander Koenig&quot;" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="22" pageNumber="116" type="Museum">ZFMK</collectionCode>
,
<collectionCode id="ED33BEBFFF9DFFF8CD49CCB3103ECBB4" box="[1287,1377,1430,1456]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="22" pageNumber="116">MNHN</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B4A2C27FF9DFFF8C8F3CC9F168FCBD0" ID-GBIF-Occurrence="3383726328" box="[189,976,1466,1492]" collectionCode="SMTD" collectorName="Coll. C. Felsche Kauf" country="Colombia" pageId="22" pageNumber="116" specimenCount="4" specimenCount-female="1" specimenCount-male="3">
<specimenCount id="9D24EDF3FF9DFFF8C8F3CC9F145ECBD0" box="[189,257,1466,1492]" pageId="22" pageNumber="116" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9D24EDF3FF9DFFF8C941CC9F1463CBD0" box="[271,316,1466,1492]" pageId="22" pageNumber="116" type="female">1 ♀</specimenCount>
; “S. America /
<collectorName id="26D743ACFF9DFFF8C9BCCC9F17ACCBD0" box="[498,755,1466,1492]" pageId="22" pageNumber="116">Coll. C. Felsche Kauf</collectorName>
20, 1918”;
<collectionCode id="ED33BEBFFF9DFFF8CB31CC9F168FCBD0" box="[895,976,1466,1492]" collectionName="Germany, Dresden, Museum fuer Tierkunde" pageId="22" pageNumber="116">SMTD</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B4A2C27FF9DFFF8CBA4CC9F14B1CBFC" ID-GBIF-Occurrence="3383726332" collectionCode="SMTD" collectorName="Gehr. W. Muller Vermacht." country="Colombia" location="Columbia" pageId="22" pageNumber="116" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D24EDF3FF9DFFF8CBA4CC9F1148CBD0" box="[1002,1047,1466,1492]" pageId="22" pageNumber="116" type="female">1 ♀</specimenCount>
; “Columbia / Gehr. W.Müller Vermächt., 1909”;
<collectionCode id="ED33BEBFFF9DFFF8C9D3CCFB14B1CBFC" box="[413,494,1502,1528]" collectionName="Germany, Dresden, Museum fuer Tierkunde" pageId="22" pageNumber="116">SMTD</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B4A2C27FF9DFFF8CA44CCFB11C4CBFC" ID-GBIF-Occurrence="3383726301" box="[522,1179,1501,1528]" collectionCode="SMTD" collectorName="Coll. C. Felsche Kauf" country="Colombia" location="Cauca" pageId="22" pageNumber="116" specimenCount="1" specimenCount-female="1" stateProvince="Cauca">
<specimenCount id="9D24EDF3FF9DFFF8CA44CCFB1766CBFC" box="[522,569,1502,1528]" pageId="22" pageNumber="116" type="female">1 ♀</specimenCount>
; “
<collectingRegion id="49E6E898FF9DFFF8CA1DCCFB17C0CBFC" box="[595,671,1502,1528]" country="Colombia" name="Cauca" pageId="22" pageNumber="116">Cauca</collectingRegion>
/
<collectorName id="26D743ACFF9DFFF8CAF7CCF816E3CBF3" box="[697,956,1501,1528]" pageId="22" pageNumber="116">Coll. C. Felsche Kauf</collectorName>
20, 1918”;
<collectionCode id="ED33BEBFFF9DFFF8CC04CCFB11C4CBFC" box="[1098,1179,1502,1528]" collectionName="Germany, Dresden, Museum fuer Tierkunde" pageId="22" pageNumber="116">SMTD</collectionCode>
</materialsCitation>
•
<materialsCitation id="3B4A2C27FF9DFFF8CCF9CCFB1008C818" ID-GBIF-Occurrence="3383726307" collectionCode="NHMB" country="Colombia" determinerName="Det. G. Frey" location="Columbia Canon de Gallo" pageId="22" pageNumber="116" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D24EDF3FF9DFFF8CCF9CCFB11B9CBFC" box="[1207,1254,1502,1528]" pageId="22" pageNumber="116" type="female">1 ♀</specimenCount>
; “
<location id="8EFD70A1FF9DFFF8CD4ECCF8142AC81F" LSID="urn:lsid:plazi:treatment:038B976CFF9DFFF6CA50C8911116CA05:8EFD70A1FF9DFFF8CD4ECCF8142AC81F" country="Colombia" name="Columbia Canon de Gallo" pageId="22" pageNumber="116">Columbia Cañon de Gallo</location>
/
<taxonomicName id="4C225DF9FF9DFFF8C9C4CF2714B7C818" authorityName="Erichson" authorityYear="1847" box="[394,488,1538,1564]" class="Insecta" family="Melolonthidae" genus="Astaena" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="116" phylum="Arthropoda" rank="genus">Astaena</taxonomicName>
apollinarmaria Sayl. Det. G. Frey, 1972 / cum typo comparatum”;
<collectionCode id="ED33BEBFFF9DFFF8CCB5CF271008C818" box="[1275,1367,1538,1564]" collectionName="Switzerland, Basel, Naturhistorisches Museum" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="22" pageNumber="116">NHMB</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33875F1FF9DFFF6C8F3CF611494CD69" lastPageId="24" lastPageNumber="118" pageId="22" pageNumber="116" type="description">
<paragraph id="8B9D267AFF9DFFF8C8F3CF611483C859" blockId="22.[189,476,1603,1630]" box="[189,476,1603,1630]" pageId="22" pageNumber="116">
<heading id="D0D59116FF9DFFF8C8F3CF611483C859" bold="true" box="[189,476,1603,1630]" fontSize="11" level="3" pageId="22" pageNumber="116" reason="3">
<emphasis id="B956FA68FF9DFFF8C8F3CF611483C859" bold="true" box="[189,476,1603,1630]" pageId="22" pageNumber="116">
<typeStatus id="549998D8FF9DFFF8C8F3CF611470C85A" box="[189,303,1604,1630]" pageId="22" pageNumber="116" type="holotype">Holotype</typeStatus>
redescription
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9D267AFF9DFFF8C8F3CF5617A0C889" blockId="22.[189,767,1651,1678]" box="[189,767,1651,1678]" pageId="22" pageNumber="116">
MEASUREMENTS. Length:
<quantity id="4CDA8B9FFF9DFFF8C9A8CF561772C889" box="[486,557,1651,1677]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="22" pageNumber="116" unit="mm" value="7.0">7 mm</quantity>
. Width:
<quantity id="4CDA8B9FFF9DFFF8CAC1CF5617A4C889" box="[655,763,1651,1678]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" pageId="22" pageNumber="116" unit="mm" value="3.75">3.75 mm</quantity>
.
</paragraph>
<paragraph id="8B9D267AFF9DFFF8C8F3CF901194C95B" blockId="22.[189,1399,1717,1887]" pageId="22" pageNumber="116">HEAD. Black, surface with simple setae. Labroclypeus, shape of anterior margin weakly sinuated medially, posteriorly with sparse punctation. Ocular canthus with terminal setae, ratio to eye: 1/3. Eyes, ratio diameter/interocular width: 0.40. Antennae with eight antennomeres; antennal club with three antennomeres; ratio antennal club/remaining antennomeres: 1/1.5; ratio third vs fourth antennomere: 1/2; fourth antennomere, lateral projection absent; ratio fifth vs fourth antennomere: 1/5.</paragraph>
<paragraph id="8B9D267AFF9DFFF6C8F3CEA21746CF89" blockId="22.[189,1399,1926,2025]" lastBlockId="24.[189,1399,265,397]" lastPageId="24" lastPageNumber="118" pageId="22" pageNumber="116">PRONOTUM AND ELYTRA. Surface of pronotum and elytra without setae. Pronotum bicolored, widest at base, anterior marginal line present, posterior marginal line absent, lateral margin weakly convex and serrated, antero-lateral angle dorso-ventrally flattened, postero-lateral angle dorso-ventrally not flattened, punctation on disc sparse, setae of lateral margin dense, median longitudinal excavation absent. Scutellum, punctures absent medially. Elytra unicolored, brownish-orange, surface opaque, microsculpture absent, punctation on intervals present, sparse, diameter of strial and interval punctation equal, posterior edge straight.</paragraph>
<caption id="DF5D76F2FF9CFFF9C8F0CE671027C9EE" ID-DOI="http://doi.org/10.5281/zenodo.5451759" ID-Zenodo-Dep="5451759" httpUri="https://zenodo.org/record/5451759/files/figure.png" pageId="23" pageNumber="117" startId="23.[190,233,1858,1884]" targetBox="[244,1344,265,1820]" targetPageId="23">
<paragraph id="8B9D267AFF9CFFF9C8F0CE671027C9EE" blockId="23.[190,1400,1858,2026]" pageId="23" pageNumber="117">
<emphasis id="B956FA68FF9CFFF9C8F0CE67140CC958" bold="true" box="[190,339,1858,1884]" pageId="23" pageNumber="117">Fig. 6. A–E</emphasis>
.
<taxonomicName id="4C225DF9FF9CFFF9C929CE6617B9C959" authority="(Saylor, 1946)" authorityName="Pacheco &amp; Monné &amp; Vaz-De &amp; Ahrens" authorityYear="2021" baseAuthorityName="Saylor" baseAuthorityYear="1946" box="[359,742,1858,1885]" class="Insecta" family="Melolonthidae" genus="Sayloria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="117" phylum="Arthropoda" rank="species" species="abcora">
<emphasis id="B956FA68FF9CFFF9C929CE661776C959" box="[359,553,1859,1885]" italics="true" pageId="23" pageNumber="117">Sayloria abcora</emphasis>
(
<bibRefCitation id="EFB35B8BFF9CFFF9CA70CE661783C959" author="Saylor L. W." box="[574,732,1858,1885]" pageId="23" pageNumber="117" pagination="215 - 232" refId="ref13932" refString="Saylor L. W. 1946. New South American scarab beetles of the genus Astaena. Revista de Entomologia 17: 215 - 232." type="journal article" year="1946">Saylor, 1946</bibRefCitation>
)
</taxonomicName>
, holotype, ♂ (Type No. 7905; CAS). –
<emphasis id="B956FA68FF9CFFF9CC97CE661059C959" bold="true" box="[1241,1286,1859,1885]" pageId="23" pageNumber="117">F–I</emphasis>
.
<taxonomicName id="4C225DF9FF9CFFF9CD57CE661758C984" authority="Saylor, 1946" authorityName="Saylor" authorityYear="1946" class="Insecta" family="Melolonthidae" genus="Astaena" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="117" phylum="Arthropoda" rank="species" species="apolinarmaria">
<emphasis id="B956FA68FF9CFFF9CD57CE661436C984" italics="true" pageId="23" pageNumber="117">Astaena apolinarmaria</emphasis>
<bibRefCitation id="EFB35B8BFF9CFFF9C93ECE431758C984" author="Saylor L. W." box="[368,519,1894,1920]" pageId="23" pageNumber="117" pagination="215 - 232" refId="ref13932" refString="Saylor L. W. 1946. New South American scarab beetles of the genus Astaena. Revista de Entomologia 17: 215 - 232." type="journal article" year="1946">Saylor, 1946</bibRefCitation>
</taxonomicName>
, holotype, ♀ (Type No. 7906; CAS). –
<emphasis id="B956FA68FF9CFFF9CB9DCE431158C984" bold="true" box="[979,1031,1894,1920]" pageId="23" pageNumber="117">J–K</emphasis>
.
<taxonomicName id="4C225DF9FF9CFFF9CC5BCE43102CC985" authority="(Saylor, 1946)" authorityName="Pacheco &amp; Monné &amp; Vaz-De &amp; Ahrens" authorityYear="2021" baseAuthorityName="Saylor" baseAuthorityYear="1946" box="[1045,1395,1894,1921]" class="Insecta" family="Melolonthidae" genus="Sayloria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="117" phylum="Arthropoda" rank="species" species="pottsi">
<emphasis id="B956FA68FF9CFFF9CC5BCE43119EC984" box="[1045,1217,1894,1920]" italics="true" pageId="23" pageNumber="117">Sayloria pottsi</emphasis>
(
<bibRefCitation id="EFB35B8BFF9CFFF9CC9ECE431036C985" author="Saylor L. W." box="[1232,1385,1894,1921]" pageId="23" pageNumber="117" pagination="215 - 232" refId="ref13932" refString="Saylor L. W. 1946. New South American scarab beetles of the genus Astaena. Revista de Entomologia 17: 215 - 232." type="journal article" year="1946">Saylor, 1946</bibRefCitation>
)
</taxonomicName>
, paratype, ♂ (CAS). –
<emphasis id="B956FA68FF9CFFF9C98ECEAC14A7C9A7" bold="true" box="[448,504,1929,1955]" pageId="23" pageNumber="117">L–N</emphasis>
.
<taxonomicName id="4C225DF9FF9CFFF9CA48CEAC173AC9A7" authorityName="Pacheco &amp; Monné &amp; Vaz-De &amp; Ahrens" authorityYear="2021" baseAuthorityName="Saylor" baseAuthorityYear="1946" box="[518,613,1929,1955]" class="Insecta" family="Melolonthidae" genus="Sayloria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="117" phylum="Arthropoda" rank="species" species="pottsi">
<emphasis id="B956FA68FF9CFFF9CA48CEAC173AC9A7" box="[518,613,1929,1955]" italics="true" pageId="23" pageNumber="117">S. pottsi</emphasis>
</taxonomicName>
, holotype, ♀ (Type No. 7916; CAS).
<emphasis id="B956FA68FF9CFFF9CC6ECEAC110CC9A7" bold="true" box="[1056,1107,1929,1955]" pageId="23" pageNumber="117">A, J</emphasis>
. Parameres, dorsal view.
<emphasis id="B956FA68FF9CFFF9C8F0CE8915A7C9C2" bold="true" box="[190,248,1964,1990]" pageId="23" pageNumber="117">B, K</emphasis>
. Aedeagus, lateral view.
<emphasis id="B956FA68FF9CFFF9CA6ECE891720C9C2" bold="true" box="[544,639,1964,1990]" pageId="23" pageNumber="117">C, G, L</emphasis>
. Labels.
<emphasis id="B956FA68FF9CFFF9CAA2CE89160EC9C2" bold="true" box="[748,849,1964,1990]" pageId="23" pageNumber="117">D, H, M</emphasis>
. Habitus, dorsal view.
<emphasis id="B956FA68FF9CFFF9CC2CCE8911EAC9C2" bold="true" box="[1122,1205,1964,1990]" pageId="23" pageNumber="117">E, I, N</emphasis>
. Habitus, lateral view.
<emphasis id="B956FA68FF9CFFF9C94ACEF51449C9EE" bold="true" box="[260,278,2000,2026]" pageId="23" pageNumber="117">F</emphasis>
. Head and pronotum, dorsal view. Scale bars: A–B, F, J–K = 0.5 mm; D–E, H–I, M–N = 1 mm.
</paragraph>
</caption>
<paragraph id="8B9D267AFF93FFF6C8F3C89017BDCFF6" blockId="24.[189,1398,437,499]" pageId="24" pageNumber="118">ABDOMEN. Lateral carina absent. Second visible ventrite without row of setae in medial region. Apical visible ventrite without suture to apical tergite.</paragraph>
<paragraph id="8B9D267AFF93FFF6C8F3CB3F17A7CD28" blockId="24.[186,1399,538,812]" pageId="24" pageNumber="118">LEGS. Ratio of length of metepisternum/metacoxa: 1/0.77. Metacoxa without longer lateral bristles, a laterally extended longitudinal shelf present. Metatibia, widest at apex, ratio of width/length: 1/3.34, dorsal margin carinated, ventral margin not serrated, distal spines groups disposed at two thirds of metatibial length. First metatarsomere, ratio to dorsal metatibial spur: 1/2. Dorsal surface of tarsi glabrous, without punctures. Metatarsi, length of first protarsomere subequal to second and third combined, ventral margin serrated and glabrous. Protibia with three teeth. Protarsi, length of first protarsomere subequal to second and third combined, second protarsomere narrow, claws symmetrical, inner protarsal claw bifid, apex of basal tooth of inner protarsal claw acute.</paragraph>
<paragraph id="8B9D267AFF93FFF6C8F3CA761494CD69" blockId="24.[189,459,851,877]" box="[189,459,851,877]" pageId="24" pageNumber="118">
AEDEAGUS.
<figureCitation id="13193AFFFF93FFF6C906CA761498CD69" box="[328,455,851,877]" captionStart="Fig" captionStartId="23.[190,233,1858,1884]" captionTargetBox="[244,1344,265,1820]" captionTargetId="figure-16@23.[244,1344,265,1820]" captionTargetPageId="23" captionText="Fig. 6. A–E. Sayloria abcora (Saylor, 1946), holotype, ♂ (Type No. 7905; CAS). – F–I. Astaena apolinarmaria Saylor, 1946, holotype, ♀ (Type No. 7906; CAS). – J–K. Sayloria pottsi (Saylor, 1946), paratype, ♂ (CAS). – L–N. S. pottsi, holotype, ♀ (Type No. 7916; CAS). A, J. Parameres, dorsal view. B, K. Aedeagus, lateral view. C, G, L. Labels. D, H, M. Habitus, dorsal view. E, I, N. Habitus, lateral view. F. Head and pronotum, dorsal view. Scale bars: A–B, F, J–K = 0.5 mm; D–E, H–I, M–N = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5451759" httpUri="https://zenodo.org/record/5451759/files/figure.png" pageId="24" pageNumber="118">Fig. 6A–B</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33875F1FF93FFF6C8F3CAB01116CA05" pageId="24" pageNumber="118" type="discussion">
<paragraph id="8B9D267AFF93FFF6C8F3CAB01470CDAB" blockId="24.[189,303,917,943]" box="[189,303,917,943]" pageId="24" pageNumber="118">
<heading id="D0D59116FF93FFF6C8F3CAB01470CDAB" bold="true" box="[189,303,917,943]" fontSize="11" level="3" pageId="24" pageNumber="118" reason="3">
<emphasis id="B956FA68FF93FFF6C8F3CAB01470CDAB" bold="true" box="[189,303,917,943]" pageId="24" pageNumber="118">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B9D267AFF93FFF6C8F3CAE61116CA05" blockId="24.[189,1399,963,1025]" pageId="24" pageNumber="118">
There is no difference between female
<typeStatus id="549998D8FF93FFF6CAE9CAE11785CDDA" box="[679,730,964,990]" pageId="24" pageNumber="118">type</typeStatus>
specimens of
<taxonomicName id="4C225DF9FF93FFF6CBDACAE11155CDD9" authorityName="Pacheco &amp; Monné &amp; Vaz-De &amp; Ahrens" authorityYear="2021" baseAuthorityName="Saylor" baseAuthorityYear="1946" box="[916,1034,963,990]" class="Insecta" family="Melolonthidae" genus="Sayloria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="118" phylum="Arthropoda" rank="species" species="abcora">
<emphasis id="B956FA68FF93FFF6CBDACAE11155CDD9" box="[916,1034,963,990]" italics="true" pageId="24" pageNumber="118">S. abcora</emphasis>
</taxonomicName>
and
<taxonomicName id="4C225DF9FF93FFF6CC01CAE11042CDD9" authorityName="Saylor" authorityYear="1946" box="[1103,1309,963,990]" class="Insecta" family="Melolonthidae" genus="Astaena" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="118" phylum="Arthropoda" rank="species" species="apolinarmaria">
<emphasis id="B956FA68FF93FFF6CC01CAE11042CDD9" box="[1103,1309,963,990]" italics="true" pageId="24" pageNumber="118">S. apolinarmaria</emphasis>
</taxonomicName>
except coloration, which is very variable. Therefore, we consider them synonymous.
</paragraph>
</subSubSection>
</treatment>
</document>
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