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<document id="67FF7295B9DD931C2B1909BC5C0D9AC9" ID-DOI="10.5281/zenodo.10788481" ID-ISSN="0253-116X" ID-Zenodo-Dep="10788481" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1709735275218" checkinUser="felipe" docAuthor="Riedel, Pénigot, Schwarz, Diller, Johansson &amp; Japoshvili, G." docDate="2023" docId="0384B524FFCBFFB0FF383D0EFE9D544A" docLanguage="en" docName="LinzerbiolBeitr.55.1.61-153.pdf" docOrigin="Linzer biologische Beiträge 55 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.10:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="10" docTitle="Cosmoconus (Alpicosmos) caucasicus RIEDEL 2023, nov.sp." docType="treatment" docVersion="2" lastPageNumber="141" masterDocId="FFBDCD5CFF84FFE1FF8A3E4EFFB15561" masterDocTitle="Darwin wasps (Hymenoptera, Ichneumonidae) of the Kintrishi National Park, Sakartvelo (Georgia), with descriptions of six new species" masterLastPageNumber="153" masterPageNumber="61" pageNumber="140" updateTime="1709739384998" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="7D9B26EF655CC90053B651736C993277">Darwin wasps (Hymenoptera, Ichneumonidae) of the Kintrishi National Park, Sakartvelo (Georgia), with descriptions of six new species</mods:title>
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<mods:namePart id="E36B2AEF0F103F428D870C41CE40B4A2">Schwarz</mods:namePart>
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<mods:namePart id="DEB02B78DCE24C92992CEAC3B9A9CA2B">Japoshvili, G.</mods:namePart>
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<mods:date id="BD53F4FD5B784897A3AB2184D2610067">2023</mods:date>
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<treatment id="0384B524FFCBFFB0FF383D0EFE9D544A" LSID="urn:lsid:plazi:treatment:0384B524FFCBFFB0FF383D0EFE9D544A" httpUri="http://treatment.plazi.org/id/0384B524FFCBFFB0FF383D0EFE9D544A" lastPageId="81" lastPageNumber="141" pageId="79" pageNumber="140">
<subSubSection id="C33757B9FFCBFFAEFF383D0EFDFF56EE" pageId="79" pageNumber="140" type="nomenclature">
<paragraph id="8B920432FFCBFFAEFF383D0EFCEE5637" blockId="79.[178,1085,832,911]" box="[178,863,832,854]" pageId="79" pageNumber="140">
<heading id="D0DAB35EFFCBFFAEFF383D0EFCEE5637" box="[178,863,832,854]" fontSize="8" level="2" pageId="79" pageNumber="140" reason="2">
<taxonomicName id="4C2D7FB1FFCBFFAEFF383D0EFD235634" authority="RIEDEL" authorityName="RIEDEL" authorityYear="2023" box="[178,658,832,853]" class="Insecta" family="Ichneumonidae" genus="Cosmoconus" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="148" phylum="Arthropoda" rank="species" species="caucasicus" status="sp. nov." subGenus="Alpicosmos">
<emphasis id="B959D820FFCBFFAEFF383D0EFE895634" bold="true" box="[178,312,832,853]" italics="true" pageId="79" pageNumber="140">Cosmoconus</emphasis>
(
<emphasis id="B959D820FFCBFFAEFECC3D0EFE0A5634" box="[326,443,832,853]" italics="true" pageId="79" pageNumber="140">Alpicosmos</emphasis>
)
<emphasis id="B959D820FFCBFFAEFE403D0EFD235634" bold="true" box="[458,658,832,853]" pageId="79" pageNumber="140">
<emphasis id="B959D820FFCBFFAEFE403D0EFD8D5634" bold="true" box="[458,572,832,853]" italics="true" pageId="79" pageNumber="140">caucasicus</emphasis>
RIEDEL
</emphasis>
</taxonomicName>
<emphasis id="B959D820FFCBFFAEFD133D0EFCEE5637" bold="true" box="[665,863,832,854]" pageId="79" pageNumber="140">
<taxonomicNameLabel id="A26A655BFFCBFFAEFD133D0EFD545634" box="[665,741,832,853]" pageId="79" pageNumber="140" rank="species">nov.sp.</taxonomicNameLabel>
(
<figureCitation id="131618B7FFCBFFAEFD783D0EFCE65637" box="[754,855,832,854]" captionStart="Figs" captionStartId="80.[178,220,1450,1472]" captionTargetBox="[179,1086,484,1438]" captionTargetId="figure-152@80.[179,1086,484,1438]" captionTargetPageId="80" captionText="Figs. 8-10: Cosmoconus (Alpicosmos) caucasicus RIEDEL nov.sp. HT: (8) habitus, (9) head from frontal, (10) head from dorsal." figureDoi="http://doi.org/10.5281/zenodo.10788487" httpUri="https://zenodo.org/record/10788487/files/figure.png" pageId="79" pageNumber="140">Figs 8-10</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="8B920432FFCBFFAEFF383D2CFDFF56EE" blockId="79.[178,1085,832,911]" pageId="79" pageNumber="140">
H o l o t y p e: (♁) SW
<collectingCountry id="F33A44A2FFCBFFAEFE2C3D2FFE5E5616" box="[422,495,865,887]" name="Georgia" pageId="79" pageNumber="140">Georgia</collectingCountry>
, Kintrishi NP,
<quantity id="4CD5A9D7FFCBFFAEFDFA3D2FFD1F5616" box="[624,686,865,887]" metricMagnitude="3" metricUnit="m" metricValue="1.634" pageId="79" pageNumber="140" unit="m" value="1634.0">1634m</quantity>
asl,
<geoCoordinate id="EE1962F5FFCBFFAEFD5C3D2FFCD05616" box="[726,865,865,887]" degrees="41.44535308" direction="north" orientation="latitude" pageId="79" pageNumber="140" precision="1" value="41.445354">41.44535308°N</geoCoordinate>
<geoCoordinate id="EE1962F5FFCBFFAEFCED3D2FFC435616" box="[871,1010,865,887]" degrees="41.05384144" direction="east" orientation="longitude" pageId="79" pageNumber="140" precision="1" value="41.05384">41.05384144°E</geoCoordinate>
, MT 12,
<date id="FF9322F2FFCBFFAEFF553D37FEEA56EE" box="[223,347,889,911]" pageId="79" pageNumber="140" value="2018-06-15" valueMax="2018-06-29" valueMin="2018-06-15">15-29.vi.2018</date>
, leg. G. Japoshvili (
<collectingRegion id="49E9CAD0FFCBFFAEFD8C3D37FDF256EE" box="[518,579,889,911]" country="Georgia" name="Tbilisi" pageId="79" pageNumber="140">Tbilisi</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C33757B9FFCBFFB0FF3B3DD0FE9D544A" lastPageId="81" lastPageNumber="142" pageId="79" pageNumber="140" type="description">
<paragraph id="8B920432FFCBFFAEFF3B3DD0FE455194" blockId="79.[177,1085,924,1269]" pageId="79" pageNumber="140">
D e s c r i p t i o n: ♁: Body length
<quantity id="4CD5A9D7FFCBFFAEFDBC3DD3FD2556D2" box="[566,660,925,947]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="79" pageNumber="140" unit="mm" value="12.5">12.5 mm</quantity>
. Fore wing
<quantity id="4CD5A9D7FFCBFFAEFC853DD3FCEF56D2" box="[783,862,925,947]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="79" pageNumber="140" unit="mm" value="8.5">8.5 mm</quantity>
long. Flagellum with 34 flagellomeres; 1
<superScript id="7C58A97AFFCBFFAEFEF13DF9FE3756A4" attach="left" box="[379,390,951,965]" fontSize="6" pageId="79" pageNumber="140">st</superScript>
flagellomere 3.7× longer than wide. Head covered with brownish setae. Temple slightly and roundly narrowed behind eye, dorsally 1.2× wider than eye, with rather dense fine punctures. OED 1.5× and OOD 1.1× ocellar diameter. Frons densely rugose-punctate, moderately impressed medially and with a small blunt tubercle between antennal sockets, without distinct horn or median ridge. Antennal sockets not modified. Face 1.6× wider than long, densely rugose-punctate. Clypeus with strong transverse central ridge, with scattered punctures basally and a row of larger punctures subapically. Mandible rather long, with two equally sized teeth. Malar space granulated, 0.7× as long as width of mandibular base. Genal carina reaching hypostomal carina far from mandibular base, hypostomal carina moderately elevated (about as wide as subdistal maxillary palpomere).
</paragraph>
<paragraph id="8B920432FFCBFFAEFF383B4AFD8C50B0" blockId="79.[178,1085,1284,1489]" pageId="79" pageNumber="140">
Mesosoma covered with brownish setae. Mesoscutum and mesopleuron densely punctate. Scutellum moderately elevated, almost pyramidal-shaped, with lateral carina in basal half. Propodeum with strong carina and large semi-oval spiracle. Area basalis and area superomedia confluent; anterior transverse carina (costula) present. Hind femur 3.9× longer than wide. 2
<superScript id="7C58A97AFFCBFFAEFEFF3B3BFE3450E2" attach="left" box="[373,389,1397,1411]" fontSize="6" pageId="79" pageNumber="140">nd</superScript>
hind tarsomere 1.15× longer than 5
<superScript id="7C58A97AFFCBFFAEFD7A3B3BFD4D50E2" attach="left" box="[752,764,1397,1411]" fontSize="6" pageId="79" pageNumber="140">th</superScript>
hind tarsomere. Claws simple. 1
<superScript id="7C58A97AFFCBFFAEFF343BD4FF7850C9" attach="left" box="[190,201,1434,1448]" fontSize="6" pageId="79" pageNumber="140">st</superScript>
tergite 2.5× longer than wide, with dense but fine punctures and distinct dorso-lateral carina. 2
<superScript id="7C58A97AFFCBFFAEFE833BF9FEA850A4" attach="left" box="[265,281,1463,1477]" fontSize="6" pageId="79" pageNumber="140">nd</superScript>
tergite 0.9× as long as wide.
</paragraph>
<paragraph id="8B920432FFCBFFB1FF383BAEFED7544A" blockId="79.[178,1085,1504,1585]" lastBlockId="80.[178,1085,219,299]" lastPageId="80" lastPageNumber="141" pageId="79" pageNumber="140">
Color: black. Flagellum black, ventrally ochreous in basal half and brownish in distal half. Mandible centrally reddish. Apical half of clypeus reddish-yellow. Apical margin of postpetiolus and 2
<superScript id="7C58A97AFFCBFFAEFEE73859FECC5344" attach="left" box="[365,381,1559,1573]" fontSize="6" pageId="79" pageNumber="140">nd</superScript>
to 4
<superScript id="7C58A97AFFCBFFAEFE273859FE085344" attach="left" box="[429,441,1559,1573]" fontSize="6" pageId="79" pageNumber="140">th</superScript>
tergites entirely yellowish. Coxae and trochanters black. Legs otherwise yellowish; fore femur in basal half, mid femur in basal 2/3 and hind femur entirely black; hind tibia black in apical 0.3. Wings with yellowish suffusion; pterostigma yellowish.
</paragraph>
<paragraph id="8B920432FFD4FFB1FF383F75FE805431" blockId="80.[178,305,314,336]" box="[178,305,314,336]" pageId="80" pageNumber="141">
<specimenCount id="9D2BCFBBFFD4FFB1FF383F75FF745431" box="[178,197,315,336]" count="1" pageId="80" pageNumber="141" type="female"></specimenCount>
unknown.
</paragraph>
<paragraph id="8B920432FFD4FFB1FF383F11FF4A54AC" blockId="80.[178,1085,350,461]" pageId="80" pageNumber="141">Taxonomical remark: This new ♁ is characterized by: distinct frontal horn absent, but small median tubercle between antennal grooves present; temple rather wide and only slightly and roundly narrowed behind eye; malar space rather long (0.7× mandibular width).</paragraph>
<caption id="DF5254BAFFD4FFB1FF383BE4FE7450B8" ID-DOI="http://doi.org/10.5281/zenodo.10788487" ID-Zenodo-Dep="10788487" httpUri="https://zenodo.org/record/10788487/files/figure.png" pageId="80" pageNumber="141" startId="80.[178,220,1450,1472]" targetBox="[179,1086,484,1438]" targetPageId="80" targetType="figure">
<paragraph id="8B920432FFD4FFB1FF383BE4FE7450B8" blockId="80.[178,1085,1450,1497]" pageId="80" pageNumber="141">
<emphasis id="B959D820FFD4FFB1FF383BE4FEA850DE" bold="true" box="[178,281,1450,1472]" pageId="80" pageNumber="141">Figs. 8-10:</emphasis>
<taxonomicName id="4C2D7FB1FFD4FFB1FEAB3BE5FD7C50DE" authority="RIEDEL" authorityName="RIEDEL" authorityYear="2023" box="[289,717,1451,1472]" class="Insecta" family="Ichneumonidae" genus="Cosmoconus" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="141" phylum="Arthropoda" rank="species" species="caucasicus" status="sp. nov." subGenus="Alpicosmos">
<emphasis id="B959D820FFD4FFB1FEAB3BE5FE2450A1" box="[289,405,1451,1472]" italics="true" pageId="80" pageNumber="141">Cosmoconus</emphasis>
(
<emphasis id="B959D820FFD4FFB1FE2E3BE5FDBC50A1" box="[420,525,1451,1472]" italics="true" pageId="80" pageNumber="141">Alpicosmos</emphasis>
)
<emphasis id="B959D820FFD4FFB1FD973BE5FD3150A1" box="[541,640,1451,1472]" italics="true" pageId="80" pageNumber="141">caucasicus</emphasis>
<smallCapsWord id="8D7492EEFFD4FFB1FD023BE5FD7C50DE" baselines="1467,1467" box="[648,717,1451,1472]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Riedel" pageId="80" pageNumber="141">RIEDEL</smallCapsWord>
</taxonomicName>
<taxonomicNameLabel id="A26A655BFFD4FFB1FD5F3BE5FCA650A1" box="[725,791,1451,1472]" pageId="80" pageNumber="141" rank="species">nov.sp.</taxonomicNameLabel>
HT: (
<emphasis id="B959D820FFD4FFB1FCD83BE4FCEC50DE" bold="true" box="[850,861,1450,1471]" pageId="80" pageNumber="141">8</emphasis>
) habitus, (
<emphasis id="B959D820FFD4FFB1FC493BE4FC7F50DE" bold="true" box="[963,974,1450,1471]" pageId="80" pageNumber="141">9</emphasis>
) head from frontal, (
<emphasis id="B959D820FFD4FFB1FE8B3B8CFEA950B6" bold="true" box="[257,280,1474,1495]" pageId="80" pageNumber="141">10</emphasis>
) head from dorsal.
</paragraph>
</caption>
<paragraph id="8B920432FFD4FFB0FF383BBCFE9D544A" blockId="80.[178,1085,1521,1602]" lastBlockId="81.[178,1085,219,299]" lastPageId="81" lastPageNumber="142" pageId="80" pageNumber="141">
It belongs to the subgenus
<taxonomicName id="4C2D7FB1FFD4FFB1FE6A3BBFFC975366" authority="KASPARYAN, 1999" authorityName="KASPARYAN" authorityYear="1999" box="[480,806,1521,1543]" pageId="80" pageNumber="141" rank="subGenus" subGenus="Alpicosmos">
<emphasis id="B959D820FFD4FFB1FE6A3BBFFDE45367" box="[480,597,1521,1542]" italics="true" pageId="80" pageNumber="141">Alpicosmos</emphasis>
<bibRefCitation id="EFBC79C3FFD4FFB1FDE83BBCFC975366" author="KASPARYAN D" box="[610,806,1522,1543]" pageId="80" pageNumber="141" refId="ref63930" refString="KASPARYAN D. R. &amp; V. I. TOLKANITZ (1999): Ichneumonidae subfamily Tryphoninae: tribes Sphinctini, Phytodietini, Oedemopsini, Tryphonini (Addendum), Idiogrammatini. Subfamilies Eucerotinae, Adelognathinae (addendum), Townesioninae. [Fauna of Russia and Neighbouring Countries. Insecta Hymenoptera.] Volume III. Number 3. (in Russian) Saint Petersburg. Nauka, 404 pp." type="book" year="1999">KASPARYAN, 1999</bibRefCitation>
</taxonomicName>
with the
<typeStatus id="5496BA90FFD4FFB1FC1D3BBCFC725366" box="[919,963,1522,1543]" pageId="80" pageNumber="141">type</typeStatus>
species
<taxonomicName id="4C2D7FB1FFD4FFB1FBAF3BBCFEC15345" authority="STROBL. It" authorityName="STROBL. It" class="Insecta" family="Ichneumonidae" genus="Ctenochira" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="141" phylum="Arthropoda" rank="species" species="genalis">
<emphasis id="B959D820FFD4FFB1FBAF3BBCFF4A5345" italics="true" pageId="80" pageNumber="141">C. genalis</emphasis>
STROBL. It
</taxonomicName>
differs from
<taxonomicName id="4C2D7FB1FFD4FFB1FE713841FDD25345" box="[507,611,1551,1572]" class="Insecta" family="Ichneumonidae" genus="Ctenochira" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="141" phylum="Arthropoda" rank="species" species="genalis">
<emphasis id="B959D820FFD4FFB1FE713841FDD25345" box="[507,611,1551,1572]" italics="true" pageId="80" pageNumber="141">C. genalis</emphasis>
</taxonomicName>
by its wider face, longer malar space, lack of median frontal ridge, and black face. In the key of Palaearctic
<taxonomicName id="4C2D7FB1FFD4FFB0FCA43862FF4A5590" authority="(KASPARYAN 1973 b)" baseAuthorityName="KASPARYAN" baseAuthorityYear="1973" class="Insecta" family="Ichneumonidae" genus="Cosmoconus" kingdom="Animalia" lastPageId="81" lastPageNumber="142" order="Hymenoptera" pageId="80" pageNumber="141" phylum="Arthropoda" rank="genus">
<emphasis id="B959D820FFD4FFB1FCA43862FC1E5320" box="[814,943,1580,1601]" italics="true" pageId="80" pageNumber="141">Cosmoconus</emphasis>
(
<bibRefCitation id="EFBC79C3FFD4FFB0FC373862FF445590" author="KASPARYAN D" lastPageId="81" lastPageNumber="142" pageId="80" pageNumber="141" refId="ref63325" refString="KASPARYAN D. R. (1973 b): Fauna of the USSR Hymenoptera Vol. III Number 1. Ichneumonidae (Subfamily Tryphoninae) Tribe Tryphonini. - Nauka Publishers, Leningrad. [Translated from Russian. Amerind Publishing Co. Ltd., New Delhi 1981, 414 pp.]" type="book" year="1973">KASPARYAN 1973b</bibRefCitation>
)
</taxonomicName>
, it runs to couplet 24 (23)
<taxonomicName id="4C2D7FB1FFD5FFB0FD9B3E95FCBD5591" authority="KASPARYAN" authorityName="KASPARYAN" box="[529,780,219,240]" class="Insecta" family="Ichneumonidae" genus="Cosmoconus" kingdom="Animalia" order="Hymenoptera" pageId="81" pageNumber="142" phylum="Arthropoda" rank="species" species="tibeticus">
<emphasis id="B959D820FFD5FFB0FD9B3E95FD375591" box="[529,646,219,240]" italics="true" pageId="81" pageNumber="142">C. tibeticus</emphasis>
KASPARYAN
</taxonomicName>
, but differs by its larger size, unmodified antennal sockets and presence of anterior transversal carina (costula) of propodeum.
</paragraph>
</subSubSection>
</treatment>
</document>