477 lines
56 KiB
XML
477 lines
56 KiB
XML
<document id="3179FED8CC41CF8AC5F18D96AC8BCEEF" ID-DOI="10.11646/zootaxa.4040.5.8" ID-GBIF-Dataset="eac6bd37-fc73-4150-a025-5dd9f7b8c4d4" ID-ISSN="1175-5326" ID-Zenodo-Dep="242468" ID-ZooBank="AF601781-0ABF-4321-A338-09A5ACC260EB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461337641809" checkinUser="plazi" docAuthor="Lackner, Tomáš" docDate="2015" docId="0382C7317557FF96BBAE9BDDFC971462" docLanguage="en" docName="zt04040p595.pdf" docOrigin="Zootaxa 4040 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hypocaccus Lackner, 2015, sp. nov." docType="treatment" docVersion="8" lastPageNumber="594" masterDocId="FFBBBF497556FF93BB399B01FFFF1137" masterDocTitle="Hypocaccus (s. l.) hirsutus sp. nov., an atypical new species of the genus Hypocaccus C. Thomson, 1867 from India (Coleoptera: Histeridae: Saprininae)" masterLastPageNumber="595" masterPageNumber="589" pageNumber="590" updateTime="1698619570719" updateUser="plazi">
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<mods:title id="43EBF680BFCEBB5CF6A44FC50B1BBBF9">Hypocaccus (s. l.) hirsutus sp. nov., an atypical new species of the genus Hypocaccus C. Thomson, 1867 from India (Coleoptera: Histeridae: Saprininae)</mods:title>
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<mods:name id="3F3BCBB1E0B7B8315A02C9C50FACDA14" type="personal">
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<mods:namePart id="5F50B81FB9A1A1BB960C21E61617CCD4">Lackner, Tomáš</mods:namePart>
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<mods:title id="85011275245B0B3E90334218F6092B6D">Zootaxa</mods:title>
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<mods:part id="D0C9B9A0FD18C3C5842B5428D2EE4677">
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<mods:date id="30F4A54C3213EC77FDD753BFEDAE4C11">2015</mods:date>
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<mods:number id="15FBECAF6A7D1F7B69BF6F11219FE912">4040</mods:number>
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<mods:number id="88EBF9CFF1D3AC9375F79D64C48FAF39">5</mods:number>
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<mods:classification id="3A39D26CC8277771019E9E8FA2A84326">journal article</mods:classification>
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<mods:identifier id="E4393531EED695FEA5CB8AC46855B569" type="DOI">10.11646/zootaxa.4040.5.8</mods:identifier>
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<treatment id="0382C7317557FF96BBAE9BDDFC971462" ID-DOI="http://doi.org/10.5281/zenodo.6095557" ID-GBIF-Taxon="119644671" ID-Zenodo-Dep="6095557" LSID="urn:lsid:plazi:treatment:0382C7317557FF96BBAE9BDDFC971462" httpUri="http://treatment.plazi.org/id/0382C7317557FF96BBAE9BDDFC971462" lastPageId="5" lastPageNumber="594" pageId="1" pageNumber="590">
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<subSubSection id="C33125AC7557FF92BBAE9BDDFD7011C0" box="[151,655,220,247]" pageId="1" pageNumber="590" type="nomenclature">
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<paragraph id="8B9476277557FF92BBAE9BDDFD7011C0" blockId="1.[151,655,220,247]" box="[151,655,220,247]" pageId="1" pageNumber="590">
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<heading id="D0DCC14B7557FF92BBAE9BDDFD7011C0" bold="true" box="[151,655,220,247]" fontSize="11" level="1" pageId="1" pageNumber="590" reason="1">
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<emphasis id="B95FAA357557FF92BBAE9BDDFD7011C0" bold="true" box="[151,655,220,247]" pageId="1" pageNumber="590">
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<taxonomicName id="4C2B0DA47557FF92BBAE9BDDFED611C1" ID-CoL="52M9" box="[151,297,220,246]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="590" phylum="Arthropoda" rank="genus">
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<emphasis id="B95FAA357557FF92BBAE9BDDFED611C1" bold="true" box="[151,297,220,246]" italics="true" pageId="1" pageNumber="590">Hypocaccus</emphasis>
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</taxonomicName>
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(sensu lato)
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<taxonomicName id="4C2B0DA47557FF92BAFF9BDDFDD611C1" box="[454,553,220,246]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="590" phylum="Arthropoda" rank="species" species="hirsutus" status="sp. nov.">
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<emphasis id="B95FAA357557FF92BAFF9BDDFDD611C1" bold="true" box="[454,553,220,246]" italics="true" pageId="1" pageNumber="590">hirsutus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26C174E7557FF92B9089BDCFD7011C0" box="[561,655,221,247]" pageId="1" pageNumber="590" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33125AC7557FF92BBAE9A25FE2E1703" pageId="1" pageNumber="590" type="materials_examined">
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<paragraph id="8B9476277557FF92BBAE9A25FDE010C7" blockId="1.[151,1437,292,496]" pageId="1" pageNumber="590">
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<emphasis id="B95FAA357557FF92BBAE9A25FE3E100A" bold="true" box="[151,449,292,317]" pageId="1" pageNumber="590">
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<typeStatus id="5490C8857557FF92BBAE9A25FF2D100A" box="[151,210,292,317]" pageId="1" pageNumber="590">Type</typeStatus>
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material examined.
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</emphasis>
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<typeStatus id="5490C8857557FF92BAF39A25FDCA100A" box="[458,565,292,317]" pageId="1" pageNumber="590" type="holotype">Holotype</typeStatus>
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, ♀, side-mounted on a triangular point, left antennal segments 3-11 missing, right protarsus and left mesotarsus broken off, with the following labels: “♀” (printed); followed by: “S
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<collectingCountry id="F33C36B77557FF92BE629A49FA681057" box="[1371,1431,328,352]" name="India" pageId="1" pageNumber="590">India</collectingCountry>
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, Kerala, 1994 / SHORANUR
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<geoCoordinate id="EE1F10E07557FF92BAE69A6DFDC110B2" box="[479,574,364,389]" direction="north" orientation="latitude" pageId="1" pageNumber="590" precision="925" value="10.766666">10º46’N</geoCoordinate>
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<geoCoordinate id="EE1F10E07557FF92B97C9A6DFD5F10B2" box="[581,672,364,389]" direction="east" orientation="longitude" pageId="1" pageNumber="590" precision="925" value="76.26667">76º16’E</geoCoordinate>
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/ bank of Ponnáni riv. /
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<date id="FF9550E77557FF92B8889A6DFC1910B3" box="[945,998,364,388]" pageId="1" pageNumber="590">31.I.</date>
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, Z. Kejval lgt.” (printed); followed by: “
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<emphasis id="B95FAA357557FF92BB9A9A90FE841090" box="[163,379,400,424]" italics="true" pageId="1" pageNumber="590">
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<taxonomicName id="4C2B0DA47557FF92BB9A9A90FED3109F" box="[163,300,401,424]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="590" phylum="Arthropoda" rank="genus">Hypocaccus</taxonomicName>
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(s.str.)
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</emphasis>
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? / spec. ?? / det. P. Kanaar 2002” (printed-written); followed by: “Gen. nov. / sp. nov. / P. Vienna det., 2003” (printed); followed by: “
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<taxonomicName id="4C2B0DA47557FF92B9B39AB5FCE910FB" box="[650,790,436,460]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="594" phylum="Arthropoda" rank="genus">Hypocaccus</taxonomicName>
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(s.l.) /
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<taxonomicName id="4C2B0DA47557FF92B8539AB5FC3C10FA" box="[874,963,436,461]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="590" phylum="Arthropoda" rank="species" species="hirsutus" status="sp. nov.">hirsutus</taxonomicName>
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<taxonomicNameLabel id="A26C174E7557FF92B8F59AB4FBDB10FB" box="[972,1060,437,460]" pageId="1" pageNumber="590" rank="species">sp. nov.</taxonomicNameLabel>
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/
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<typeStatus id="5490C8857557FF92BF049AB5FB2E10FB" box="[1085,1233,436,460]" pageId="1" pageNumber="590" type="holotype">HOLOTYPE</typeStatus>
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/ Det.
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<bibRefCitation id="EFBA0BD67557FF92BE1B9AB4FF2E10C7" author="Lackner" pageId="1" pageNumber="590" refString="Lackner, T. (2015) Revision of the genus Exaesiopus Reichardt, 1926. ZooKeys, 479, 65 - 108. http: // dx. doi. org / 10.3897 / zookeys. 479.8738" type="journal article" year="2015">T. Lackner 2015</bibRefCitation>
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” (red label, written) (
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<collectionCode id="ED3AEEE27557FF92BAFB9AD9FDEC10C7" box="[450,531,472,496]" pageId="1" pageNumber="590">TLAN</collectionCode>
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).
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</paragraph>
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<paragraph id="8B9476277557FF92BBAE9D01FE2E1703" blockId="1.[151,925,1536,1588]" pageId="1" pageNumber="590">
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FIGURE 1.
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<taxonomicName id="4C2B0DA47557FF92BA209D00FE6A1721" box="[281,405,1537,1558]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="590" phylum="Arthropoda" rank="genus">
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<emphasis id="B95FAA357557FF92BA209D00FE6A1721" box="[281,405,1537,1558]" italics="true" pageId="1" pageNumber="590">Hypocaccus</emphasis>
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</taxonomicName>
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(sensu lato)
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<taxonomicName id="4C2B0DA47557FF92B92F9D01FD971721" box="[534,616,1536,1558]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="590" phylum="Arthropoda" rank="species" species="hirsutus" status="sp. nov.">
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<emphasis id="B95FAA357557FF92B92F9D01FD971721" box="[534,616,1536,1558]" italics="true" pageId="1" pageNumber="590">hirsutus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26C174E7557FF92B9579D01FD3F1722" box="[622,704,1536,1557]" pageId="1" pageNumber="590" rank="species">sp. nov.</taxonomicNameLabel>
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, habitus, dorsal view. FIGURE 2.ditto, ventral view.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33125AC7557FF91BBFE9D5DFF0B167A" lastPageId="2" lastPageNumber="591" pageId="1" pageNumber="590" type="description">
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<paragraph id="8B9476277557FF91BBFE9D5DFCDF1077" blockId="1.[151,1437,1628,2015]" lastBlockId="2.[151,1437,151,2013]" lastPageId="2" lastPageNumber="591" pageId="1" pageNumber="590">
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<emphasis id="B95FAA357557FF92BBFE9D5DFEA61742" bold="true" box="[199,345,1628,1653]" pageId="1" pageNumber="590">Description.</emphasis>
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Body (Figs. 1–2) PEL:
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<quantity id="4CD3DBC27557FF92B95F9D5DFD311743" box="[614,718,1628,1653]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" pageId="1" pageNumber="590" unit="mm" value="2.55">2.55 mm</quantity>
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; APW:
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<quantity id="4CD3DBC27557FF92B81F9D5DFC721743" box="[806,909,1628,1653]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="1" pageNumber="590" unit="mm" value="1.25">1.25 mm</quantity>
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||
; PPW:
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<quantity id="4CD3DBC27557FF92B8D99D5DFBB81743" box="[992,1095,1628,1652]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="1" pageNumber="590" unit="mm" value="2.1">2.10 mm</quantity>
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; EW:
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<quantity id="4CD3DBC27557FF92BFB59D5DFB0B1743" box="[1164,1268,1628,1652]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="1" pageNumber="590" unit="mm" value="2.2">2.20 mm</quantity>
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; EL:
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<quantity id="4CD3DBC27557FF92BE089D5DFA671743" box="[1329,1432,1628,1653]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="590" unit="mm" value="1.5">1.50 mm</quantity>
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; cuticle light to rusty brown, pronotum somewhat darker than elytra, legs, antenna and mouthparts similarly colored. Head (Fig. 3): mandibles rather thin, pointed apically, left mandible with a large triangularly shaped subapical tooth; anterior margin of transverse labrum elevated, keel-like, other mouthparts not examined; clypeus transverse, with faint shallow punctures with keel-like median elevation, surface posterior to this elevation depressed, anterior to it clypeal disc even; frontal stria carinate, outwardly arcuate, supraorbital stria erased, absent; occipital stria complete, feebly carinate; frontal disc with scattered microscopic punctation, anteriorly with an almost straight transverse carina, posteriorly with a carinate chevron; between the two a short transverse carina present; frontal disc posterior to chevron with another transverse carina basically delimiting frontal disc from occiput; eyes flattened, well-visible from above. Antennal scape with numerous long amber setae; 8th antennomere shaped like a cupule surrounding antennal club up to one-third (Fig. 3), resembling that of
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<taxonomicName id="4C2B0DA47557FF91BF819CC6FF1F1187" class="Insecta" family="Histeridae" genus="Philothis" kingdom="Animalia" lastPageId="2" lastPageNumber="591" order="Coleoptera" pageId="1" pageNumber="590" phylum="Arthropoda" rank="species" species="atavus" subGenus="Atavinus">
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<emphasis id="B95FAA357557FF92BF819CC6FAE216E8" box="[1208,1309,1991,2015]" italics="true" pageId="1" pageNumber="590">Philothis</emphasis>
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(
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<emphasis id="B95FAA357557FF92BE089CC6FA6C16E9" box="[1329,1427,1991,2014]" italics="true" pageId="1" pageNumber="590">Atavinus</emphasis>
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)
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<emphasis id="B95FAA357554FF91BBAE9B98FF1F1187" box="[151,224,153,176]" italics="true" pageId="2" pageNumber="591">atavus</emphasis>
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</taxonomicName>
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(for fig. see
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<bibRefCitation id="EFBA0BD67554FF91BA4E9B96FDE81187" author="Lackner" box="[375,535,151,176]" pageId="2" pageNumber="591" refString="Lackner, T. (2010) Review of the Palaearctic genera of Saprininae (Coleoptera: Histeridae). Acta Entomologica Musei Nationalis Pragae, 50 (Supplementum), 1 - 254." type="journal article" year="2010">Lackner 2010</bibRefCitation>
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, fig. 6). Antennal club globular, conspicuously small; sensory structures of the antennal club examined only externally, due to the unique available specimen, which lacks one antenna. Although the antennal club has not been chemically cleared, a single stipe-shaped vesicle situated on internal distal side of the club was observable under great magnification. Basal half of antennal club glabrous, apical half with intermingled dense short and sparser longer erect sensilla.
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</paragraph>
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<paragraph id="8B9476277554FF91BBFE9A4CFAA610C2" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="591">Pronotum sub-trapezoidal, antennal angles acute, pronotal sides narrowing anteriorly; marginal pronotal stria carinate, complete, becoming rather thin and sub-carinate in posterior angles. Entire pronotal disc with punctation, punctures on most of disc forming elongate confluent rugae; only on ante-scutellar area punctures free, separated by their own to several times their diameter. Pronotal base with a double row of dense punctures; ante-scutellar area with a faint depression; pronotal hypomeron with dense long amber setae. Scutellum rather small, triangular.</paragraph>
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<paragraph id="8B9476277554FF91BBFE9AFEFA8813C7" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="591">Elytra: marginal epipleural stria well-impressed; elytral epipleura punctate; marginal elytral stria complete, carinate, continuous with complete apical elytral stria that is continuous with complete and carinate sutural elytral stria. Oblique humeral stria short, double; inner subhumeral stria present as short median fragment; elytral disc with four discal elytral stria 1–4, first stria the longest, surpassing elytral half apically; striae 2–4 somewhat shortened, reaching approximately half of elytral length apically; fourth stria weakly developed, interrupted on its half-length. All striae rather weak and obscured by dense elytral punctation; entire elytral disc densely punctuate, punctures separated by less than their diameter; punctation becomes sparser near elytral flanks and elytral base.</paragraph>
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<paragraph id="8B9476277554FF91BBFE99FDFEB3126A" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="591">Propygidium transverse, its punctation even denser than that of elytra, punctures almost confluent; pygidium with similar, but sparser punctation, punctures here separated by their own to twice their diameter, becoming sparser apically.</paragraph>
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<paragraph id="8B9476277554FF91BBFE9869FDF512DA" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="591">Prosternum (Fig. 4): entire prosternal surface with alutaceous microsculpture, prosternal process with punctures; surface between carinate and complete carinal prosternal striae densely punctate; lateral prosternal striae complete and carinate, convergent anteriorly, united in front of united carinal prosternal striae, passing mesad to small but deep prosternal foveae.</paragraph>
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<paragraph id="8B9476277554FF91BBFE98F6FD5E1560" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="591">Mesoventrite (Fig. 4): transverse, punctate, punctures separated by about their own to twice their diameter; marginal stria of mesoventrite carinate laterally, anteriorly obsolete, antero-medially absent; meso-metaventral stria undulate, concealing meso-metaventral suture.</paragraph>
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<paragraph id="8B9476277554FF91BBFE9F65FCB0143A" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="591">Metaventrite (Fig. 4) covered with scattered fine punctures separated by several times their diameter, surface along median longitudinal line slightly depressed; lateral metaventral stria straight, carinate, not reaching metacoxa. Lateral disc of metaventrite with rather deep punctures of various sizes, punctures without setae; metepisternum with similar punctation becoming weaker and sparser on fused metepimeron; lateral metepisternal stria present as a short vague fragment on fused metepimeron.</paragraph>
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<paragraph id="8B9476277554FF91BBFE9E16FCD21462" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="591">First visible abdominal ventrite with punctation similar to that of metaventrite, completely striate laterally; punctation becoming microscopic and very sparse apically.</paragraph>
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<paragraph id="8B9476277554FF91BBFE9E60FBBC178A" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="591">Legs: protibia: outer margin apically with two rather approximate low teeth topped by large round denticle; this double-tooth rather widely separated from the following 6 proximal low teeth each topped by round denticle diminishing size in proximal direction. Protarsal groove shallow, protibial spur large, bent, growing out from apical protibial margin; posterior face of protibia rugulose-lacunose; apical margin ventrally with two tiny denticles, posterior protibial stria carinate and complete, separating rugulose-lacunose outer part of protibia from smooth median part of prosternal surface; inner row of setae sparse; inner posterior denticles absent. Mesotibia and metatibia similar, outer margin of both with dense row of long thick denticles supplemented by another, sparser row of shorter denticles on anterior face; both meso-and metatibial spurs well developed and rather long, straight; each meso-and metatarsomere with two long strongly sclerotized setae; tarsal claws in both cases approximately as long as half-length of their respective apical-most tarsomeres, slightly bent apically.</paragraph>
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<paragraph id="8B9476277554FF91BBFE9DC6FE8817D7" blockId="2.[151,1437,151,2013]" box="[199,375,1735,1760]" pageId="2" pageNumber="591">Male unknown.</paragraph>
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<paragraph id="8B9476277554FF91BBFE9DEDFF0B167A" blockId="2.[151,1437,151,2013]" pageId="2" pageNumber="591">
|
||
<emphasis id="B95FAA357554FF91BBFE9DEDFE281632" bold="true" box="[199,471,1772,1797]" pageId="2" pageNumber="591">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="4C2B0DA47554FF91BADF9DECFD261633" box="[486,729,1772,1796]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="591" phylum="Arthropoda" rank="species" species="hirsutus" status="sp. nov.">
|
||
<emphasis id="B95FAA357554FF91BADF9DECFD261633" box="[486,729,1772,1796]" italics="true" pageId="2" pageNumber="591">Hypocaccus hirsutus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26C174E7554FF91B9D09DEDFCB41632" box="[745,843,1772,1797]" pageId="2" pageNumber="591" rank="species">
|
||
<emphasis id="B95FAA357554FF91B9D09DEDFCB41632" bold="true" box="[745,843,1772,1797]" pageId="2" pageNumber="591">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs markedly from all species of the genus
|
||
<taxonomicName id="4C2B0DA47554FF91BBAE9C10FEDF161F" box="[151,288,1809,1832]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="591" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FAA357554FF91BBAE9C10FEDF161F" box="[151,288,1809,1832]" italics="true" pageId="2" pageNumber="591">Hypocaccus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F33C36B77554FF91BA5E9C0EFE5E161F" box="[359,417,1807,1832]" name="India" pageId="2" pageNumber="591">India</collectingCountry>
|
||
by densely ciliate pronotal hypomeron as well as by strongly elevated anterior margin of clypeus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33125AC7554FF91BBFE9C56FB961647" box="[199,1129,1879,1905]" pageId="2" pageNumber="591" type="biology_ecology">
|
||
<paragraph id="8B9476277554FF91BBFE9C56FB961647" blockId="2.[151,1437,151,2013]" box="[199,1129,1879,1905]" pageId="2" pageNumber="591">
|
||
<emphasis id="B95FAA357554FF91BBFE9C56FED81647" bold="true" box="[199,295,1879,1904]" pageId="2" pageNumber="591">Biology.</emphasis>
|
||
Unknown, the
|
||
<typeStatus id="5490C8857554FF91BAEA9C58FDFC1646" box="[467,515,1881,1905]" pageId="2" pageNumber="591">type</typeStatus>
|
||
specimen was collected on the bank of Ponnáni River.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33125AC7554FF91BBFE9C7DFA6316A2" box="[199,1436,1916,1941]" pageId="2" pageNumber="591" type="distribution">
|
||
<paragraph id="8B9476277554FF91BBFE9C7DFA6316A2" blockId="2.[151,1437,151,2013]" box="[199,1436,1916,1941]" pageId="2" pageNumber="591">
|
||
<emphasis id="B95FAA357554FF91BBFE9C7DFE9F16A2" bold="true" box="[199,352,1916,1941]" pageId="2" pageNumber="591">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="5490C8857554FF91B9669C7CFD7016A2" box="[607,655,1917,1941]" pageId="2" pageNumber="591">type</typeStatus>
|
||
locality, village of Shoranur in the Indian province of Kerala (Fig. 5).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33125AC7554FF96BBFE9CA1FC971462" lastPageId="5" lastPageNumber="594" pageId="2" pageNumber="591" type="discussion">
|
||
<paragraph id="8B9476277554FF90BBFE9CA1FC7C1680" blockId="2.[151,1437,151,2013]" lastBlockId="3.[151,1045,1953,2006]" lastPageId="3" lastPageNumber="592" pageId="2" pageNumber="591">
|
||
<emphasis id="B95FAA357554FF91BBFE9CA1FEB3168E" bold="true" box="[199,332,1952,1977]" pageId="2" pageNumber="591">Discussion.</emphasis>
|
||
The genus
|
||
<taxonomicName id="4C2B0DA47554FF91BAD89CA0FCAA168E" authority="C. Thomson, 1867" authorityName="C. Thomson" authorityYear="1867" box="[481,853,1952,1977]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="591" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FAA357554FF91BAD89CA0FD95168F" box="[481,618,1953,1976]" italics="true" pageId="2" pageNumber="591">Hypocaccus</emphasis>
|
||
C. Thomson, 1867
|
||
</taxonomicName>
|
||
contains currently three subgenera:
|
||
<taxonomicName id="4C2B0DA47554FF91BE359CA0FA6A168F" box="[1292,1429,1953,1976]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="591" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FAA357554FF91BE359CA0FA6A168F" box="[1292,1429,1953,1976]" italics="true" pageId="2" pageNumber="591">Hypocaccus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2B0DA47554FF91BBAE9CC5FDE816EA" authority="Reichardt, 1926" authorityName="Reichardt" authorityYear="1926" box="[151,535,1988,2013]" class="Insecta" family="Histeridae" genus="Baeckmanniolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="591" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FAA357554FF91BBAE9CC5FEAF16EB" box="[151,336,1988,2012]" italics="true" pageId="2" pageNumber="591">Baeckmanniolus</emphasis>
|
||
Reichardt, 1926
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2B0DA47554FF91B9659CC4FC9116EB" authority="Reichardt, 1932" authorityName="Reichardt" authorityYear="1932" box="[604,878,1988,2013]" class="Insecta" family="Histeridae" genus="Nessus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="591" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FAA357554FF91B9659CC4FD5516EB" box="[604,682,1989,2012]" italics="true" pageId="2" pageNumber="591">Nessus</emphasis>
|
||
Reichardt, 1932
|
||
</taxonomicName>
|
||
. The primarily Holarctic nominotypical subgenus FIGURE 3.
|
||
<taxonomicName id="4C2B0DA47555FF90BA209CA3FE6A1680" box="[281,405,1954,1975]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="592" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FAA357555FF90BA209CA3FE6A1680" box="[281,405,1954,1975]" italics="true" pageId="3" pageNumber="592">Hypocaccus</emphasis>
|
||
</taxonomicName>
|
||
(sensu lato)
|
||
<taxonomicName id="4C2B0DA47555FF90B92F9CA0FD971680" box="[534,616,1953,1975]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="592" phylum="Arthropoda" rank="species" species="hirsutus" status="sp. nov.">
|
||
<emphasis id="B95FAA357555FF90B92F9CA0FD971680" box="[534,616,1953,1975]" italics="true" pageId="3" pageNumber="592">hirsutus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26C174E7555FF90B9579CA3FD3F1680" box="[622,704,1954,1975]" pageId="3" pageNumber="592" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, head, dorsal view.
|
||
</paragraph>
|
||
<paragraph id="8B9476277555FF90BBAE9CC1FBEA16E1" blockId="3.[151,1045,1953,2006]" box="[151,1045,1984,2006]" pageId="3" pageNumber="592">
|
||
FIGURE 4.
|
||
<taxonomicName id="4C2B0DA47555FF90BA209CC0FE6A16E1" box="[281,405,1985,2006]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="592" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FAA357555FF90BA209CC0FE6A16E1" box="[281,405,1985,2006]" italics="true" pageId="3" pageNumber="592">Hypocaccus</emphasis>
|
||
</taxonomicName>
|
||
(sensu lato)
|
||
<taxonomicName id="4C2B0DA47555FF90B92F9CC1FD9716E1" box="[534,616,1984,2006]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="592" phylum="Arthropoda" rank="species" species="hirsutus" status="sp. nov.">
|
||
<emphasis id="B95FAA357555FF90B92F9CC1FD9716E1" box="[534,616,1984,2006]" italics="true" pageId="3" pageNumber="592">hirsutus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26C174E7555FF90B9579CC0FD3F16E1" box="[622,704,1985,2006]" pageId="3" pageNumber="592" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, prosternum, meso+metaventrite.
|
||
</paragraph>
|
||
<paragraph id="8B9476277552FF97BBAE9DFAFC371627" blockId="4.[151,968,1786,1808]" box="[151,968,1786,1808]" pageId="4" pageNumber="593">
|
||
FIGURE 5.
|
||
<typeStatus id="5490C8857552FF97BA209DFAFEB41627" box="[281,331,1787,1808]" pageId="4" pageNumber="593">Type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="4C2B0DA47552FF97BAFC9DFAFDBE1627" box="[453,577,1787,1808]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="593" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FAA357552FF97BAFC9DFAFDBE1627" box="[453,577,1787,1808]" italics="true" pageId="4" pageNumber="593">Hypocaccus</emphasis>
|
||
</taxonomicName>
|
||
(sensu lato)
|
||
<taxonomicName id="4C2B0DA47552FF97B9FA9DFBFCEA1627" box="[707,789,1786,1808]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="593" phylum="Arthropoda" rank="species" species="hirsutus" status="sp. nov.">
|
||
<emphasis id="B95FAA357552FF97B9FA9DFBFCEA1627" box="[707,789,1786,1808]" italics="true" pageId="4" pageNumber="593">hirsutus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26C174E7552FF97B8229DFAFC931627" box="[795,876,1787,1808]" pageId="4" pageNumber="593" rank="species">sp. nov.</taxonomicNameLabel>
|
||
in
|
||
<collectingCountry id="F33C36B77552FF97B8B59DFBFC3C1627" box="[908,963,1786,1808]" name="India" pageId="4" pageNumber="593">India</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9476277552FF97BBAE9C3FFE6A16D1" blockId="4.[151,1436,1853,2022]" pageId="4" pageNumber="593">
|
||
<taxonomicName id="4C2B0DA47552FF97BBAE9C3FFEDF1662" box="[151,288,1854,1877]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="593" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FAA357552FF97BBAE9C3FFEDF1662" box="[151,288,1854,1877]" italics="true" pageId="4" pageNumber="593">Hypocaccus</emphasis>
|
||
</taxonomicName>
|
||
is normally found on sandy soils, banks of rivers, and seashores, and includes 44 described species (
|
||
<bibRefCitation id="EFBA0BD67552FF97BBA79C60FEAF164D" author="Lackner" box="[158,336,1889,1914]" pageId="4" pageNumber="593" refString="Lackner, T. (2014 d) Phylogeny of the Saprininae subfamily reveals interesting ecological shifts in the history of the subfamily (Coleoptera: Histeridae). Zoological Journal of the Linnean Society, 172, 521 - 555. http: // dx. doi. org / 10.1111 / zoj. 12182" type="journal article" year="2014" yearSuffix="d">Lackner 2014d</bibRefCitation>
|
||
). The pronotal hypomeron of
|
||
<taxonomicName id="4C2B0DA47552FF97B9959C63FCCA164E" box="[684,821,1890,1913]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="593" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="B95FAA357552FF97B9959C63FCCA164E" box="[684,821,1890,1913]" italics="true" pageId="4" pageNumber="593">Hypocaccus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26C174E7552FF97B8069C63FC82164E" box="[831,893,1890,1913]" pageId="4" pageNumber="593" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
is normally asetose; however, several Nearctic psammophilous representatives (e.g.,
|
||
<taxonomicName id="4C2B0DA47552FF97B9789C87FCBA16A9" authority="Mazur, 1991" authorityName="Mazur" authorityYear="1991" box="[577,837,1926,1950]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="593" phylum="Arthropoda" rank="species" species="texaco">
|
||
<emphasis id="B95FAA357552FF97B9789C87FD5216AA" box="[577,685,1926,1949]" italics="true" pageId="4" pageNumber="593">H. texaco</emphasis>
|
||
Mazur, 1991
|
||
</taxonomicName>
|
||
) do possess ciliate pronotal hypomeron. From
|
||
<collectingCountry id="F33C36B77552FF97BE629C84FA6816A9" box="[1371,1431,1925,1950]" name="India" pageId="4" pageNumber="593">India</collectingCountry>
|
||
, three species have been recorded so far:
|
||
<taxonomicName id="4C2B0DA47552FF97B9569CABFC2816F5" authority="Marseul, 1864" authorityName="Marseul" authorityYear="1864" box="[623,983,1961,1986]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="593" phylum="Arthropoda" rank="species" species="brahminius">
|
||
<emphasis id="B95FAA357552FF97B9569CABFCEC16F6" box="[623,787,1961,1985]" italics="true" pageId="4" pageNumber="593">H. brahminius</emphasis>
|
||
(Marseul, 1864)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2B0DA47552FF97B8DC9CABFABA16F6" authority="Paykull, 1811" authorityName="Paykull" authorityYear="1811" box="[997,1349,1961,1986]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="593" phylum="Arthropoda" rank="species" species="brasiliensis">
|
||
<emphasis id="B95FAA357552FF97B8DC9CABFB7416F6" box="[997,1163,1961,1985]" italics="true" pageId="4" pageNumber="593">H. brasiliensis</emphasis>
|
||
(Paykull, 1811)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2B0DA47552FF97BEB89CABFE6F16D1" authority="Marseul, 1862" authorityName="Marseul" authorityYear="1862" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="593" phylum="Arthropoda" rank="species" species="sinae">
|
||
<emphasis id="B95FAA357552FF97BEB89CABFF2D16D2" italics="true" pageId="4" pageNumber="593">H. sinae</emphasis>
|
||
(Marseul, 1862)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9476277553FF96BBFE9B96FE29109A" blockId="5.[151,1437,151,1365]" pageId="5" pageNumber="594">
|
||
Another subgenus, the strictly beach-dwelling
|
||
<taxonomicName id="4C2B0DA47553FF96B9E79B99FC681187" box="[734,919,152,176]" class="Insecta" family="Histeridae" genus="Baeckmanniolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FAA357553FF96B9E79B99FC681187" box="[734,919,152,176]" italics="true" pageId="5" pageNumber="594">Baeckmanniolus</emphasis>
|
||
</taxonomicName>
|
||
, with 11 described species (
|
||
<bibRefCitation id="EFBA0BD67553FF96BFE79B96FA701187" author="Lackner" box="[1246,1423,151,176]" pageId="5" pageNumber="594" refString="Lackner, T. (2014 d) Phylogeny of the Saprininae subfamily reveals interesting ecological shifts in the history of the subfamily (Coleoptera: Histeridae). Zoological Journal of the Linnean Society, 172, 521 - 555. http: // dx. doi. org / 10.1111 / zoj. 12182" type="journal article" year="2014" yearSuffix="d">Lackner 2014d</bibRefCitation>
|
||
), occurs mainly on seashores of all continents (except
|
||
<collectingCountry id="F33C36B77553FF96B9DA9BBDFCA611E2" box="[739,857,188,213]" name="Antarctica" pageId="5" pageNumber="594">Antarctica</collectingCountry>
|
||
) but has not yet been recorded from
|
||
<collectingCountry id="F33C36B77553FF96BFCA9BBDFAD011E2" box="[1267,1327,188,213]" name="India" pageId="5" pageNumber="594">India</collectingCountry>
|
||
, although one species,
|
||
<taxonomicName id="4C2B0DA47553FF96BA149BE0FD5C11CF" authority="Schmidt, 1890" authorityName="Schmidt" authorityYear="1890" box="[301,675,223,248]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="species" species="varians" subGenus="B.">
|
||
<emphasis id="B95FAA357553FF96BA149BE0FEBE11CF" box="[301,321,225,248]" italics="true" pageId="5" pageNumber="594">H</emphasis>
|
||
. (
|
||
<emphasis id="B95FAA357553FF96BA649BE0FE9111CF" box="[349,366,225,248]" italics="true" pageId="5" pageNumber="594">B</emphasis>
|
||
.)
|
||
<emphasis id="B95FAA357553FF96BAB39BE1FE2111C0" box="[394,478,224,247]" italics="true" pageId="5" pageNumber="594">varians</emphasis>
|
||
(Schmidt, 1890)
|
||
</taxonomicName>
|
||
is widely spread in Indo-Malayan as well as Australian regions.
|
||
<bibRefCitation id="EFBA0BD67553FF96BBAE9A04FECB102B" author="Mazur" box="[151,308,260,284]" pageId="5" pageNumber="594" refString="Mazur, S. (2011) A concise catalogue of the Histeridae. Warsaw University of Life Sciences, SGGW Press, Warsaw, 332 pp." type="book" year="2011">Mazur (2011)</bibRefCitation>
|
||
reports it from neighboring
|
||
<collectingCountry id="F33C36B77553FF96B9B89A05FD09102A" box="[641,758,260,285]" name="Sri Lanka" pageId="5" pageNumber="594">Sri Lanka</collectingCountry>
|
||
. Species of
|
||
<taxonomicName id="4C2B0DA47553FF96B8BD9A05FBC2102B" box="[900,1085,260,284]" class="Insecta" family="Histeridae" genus="Baeckmanniolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FAA357553FF96B8BD9A05FBC2102B" box="[900,1085,260,284]" italics="true" pageId="5" pageNumber="594">Baeckmanniolus</emphasis>
|
||
</taxonomicName>
|
||
do not have a ciliate pronotal hypomeron. The two subgenera,
|
||
<taxonomicName id="4C2B0DA47553FF96B9329A28FD6B1077" box="[523,660,297,320]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FAA357553FF96B9329A28FD6B1077" box="[523,660,297,320]" italics="true" pageId="5" pageNumber="594">Hypocaccus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B95FAA357553FF96B9F59A29FC881077" box="[716,887,296,320]" italics="true" pageId="5" pageNumber="594">Baeckmaniolus</emphasis>
|
||
are separated based on their pronotal punctation (species in the subgenus
|
||
<taxonomicName id="4C2B0DA47553FF96BA899A4DFD961053" box="[432,617,332,356]" class="Insecta" family="Histeridae" genus="Baeckmanniolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FAA357553FF96BA899A4DFD961053" box="[432,617,332,356]" italics="true" pageId="5" pageNumber="594">Baeckmanniolus</emphasis>
|
||
</taxonomicName>
|
||
possess impunctate pronotum
|
||
<emphasis id="B95FAA357553FF96B8FC9A4CFC1B1053" box="[965,996,333,356]" italics="true" pageId="5" pageNumber="594">vs.</emphasis>
|
||
punctate in the subgenus
|
||
<taxonomicName id="4C2B0DA47553FF96BE339A4CFA6C1053" box="[1290,1427,333,356]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FAA357553FF96BE339A4CFA6C1053" box="[1290,1427,333,356]" italics="true" pageId="5" pageNumber="594">Hypocaccus</emphasis>
|
||
</taxonomicName>
|
||
) and number of rows of denticles on the metatibia (two in
|
||
<taxonomicName id="4C2B0DA47553FF96B82E9A70FC5F10BF" box="[791,928,369,392]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FAA357553FF96B82E9A70FC5F10BF" box="[791,928,369,392]" italics="true" pageId="5" pageNumber="594">Hypocaccus</emphasis>
|
||
</taxonomicName>
|
||
vs. three in
|
||
<taxonomicName id="4C2B0DA47553FF96BF119A71FB1E10BF" box="[1064,1249,368,392]" class="Insecta" family="Histeridae" genus="Baeckmanniolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FAA357553FF96BF119A71FB1E10BF" box="[1064,1249,368,392]" italics="true" pageId="5" pageNumber="594">Baeckmanniolus</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<bibRefCitation id="EFBA0BD67553FF96BFC09A6EFA6710BF" author="Lackner" box="[1273,1432,367,392]" pageId="5" pageNumber="594" refString="Lackner, T. (2010) Review of the Palaearctic genera of Saprininae (Coleoptera: Histeridae). Acta Entomologica Musei Nationalis Pragae, 50 (Supplementum), 1 - 254." type="journal article" year="2010">Lackner 2010</bibRefCitation>
|
||
, Bousquet & Laplante 2006).
|
||
</paragraph>
|
||
<paragraph id="8B9476277553FF96BBFE9AB6FDE61292" blockId="5.[151,1437,151,1365]" pageId="5" pageNumber="594">
|
||
The third subgenus is the species-rich
|
||
<taxonomicName id="4C2B0DA47553FF96B9459AB8FD3510E7" box="[636,714,441,464]" class="Insecta" family="Histeridae" genus="Nessus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FAA357553FF96B9459AB8FD3510E7" box="[636,714,441,464]" italics="true" pageId="5" pageNumber="594">Nessus</emphasis>
|
||
</taxonomicName>
|
||
containing 53 species distributed almost exclusively in the Old World and occurring chiefly on carcasses, mammal excrement, and in rodent nests, with some specialized Middle- Asian psammophilous forms (
|
||
<bibRefCitation id="EFBA0BD67553FF96BACF9AFEFD55132F" author="Lackner" box="[502,682,511,536]" pageId="5" pageNumber="594" refString="Lackner, T. (2014 d) Phylogeny of the Saprininae subfamily reveals interesting ecological shifts in the history of the subfamily (Coleoptera: Histeridae). Zoological Journal of the Linnean Society, 172, 521 - 555. http: // dx. doi. org / 10.1111 / zoj. 12182" type="journal article" year="2014" yearSuffix="d">Lackner 2014d</bibRefCitation>
|
||
). Four species:
|
||
<taxonomicName id="4C2B0DA47553FF96B85E9900FB33132F" authority="Marseul, 1857" authorityName="Marseul" authorityYear="1857" box="[871,1228,511,536]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="species" species="fugax" subGenus="N.">
|
||
<emphasis id="B95FAA357553FF96B85E9900FC84132F" box="[871,891,513,536]" italics="true" pageId="5" pageNumber="594">H</emphasis>
|
||
. (
|
||
<emphasis id="B95FAA357553FF96B8AE9900FC55132F" box="[919,938,513,536]" italics="true" pageId="5" pageNumber="594">N</emphasis>
|
||
.)
|
||
<emphasis id="B95FAA357553FF96B8FF9901FBFB132F" box="[966,1028,512,536]" italics="true" pageId="5" pageNumber="594">fugax</emphasis>
|
||
(Marseul, 1857)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2B0DA47553FF96BFE59900FE96130A" authority="Desbordes, 1925" authorityName="Desbordes" authorityYear="1925" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="species" species="fugitivus" subGenus="N.">
|
||
<emphasis id="B95FAA357553FF96BFE59900FB0F132F" box="[1244,1264,513,536]" italics="true" pageId="5" pageNumber="594">H</emphasis>
|
||
. (
|
||
<emphasis id="B95FAA357553FF96BE359900FAE0132F" box="[1292,1311,513,536]" italics="true" pageId="5" pageNumber="594">N</emphasis>
|
||
.)
|
||
<emphasis id="B95FAA357553FF96BE059901FA63132F" box="[1340,1436,512,536]" italics="true" pageId="5" pageNumber="594">fugitivus</emphasis>
|
||
(Desbordes, 1925)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2B0DA47553FF96BA4F9924FCD6130B" authority="Reichardt, 1932" authorityName="Reichardt" authorityYear="1932" box="[374,809,548,573]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="species" species="malabaricus" subGenus="N.">
|
||
<emphasis id="B95FAA357553FF96BA4F9924FE75130B" box="[374,394,549,572]" italics="true" pageId="5" pageNumber="594">H</emphasis>
|
||
. (
|
||
<emphasis id="B95FAA357553FF96BA9A9924FE49130B" box="[419,438,549,572]" italics="true" pageId="5" pageNumber="594">N</emphasis>
|
||
.)
|
||
<emphasis id="B95FAA357553FF96BAF79925FDA5130B" box="[462,602,548,572]" italics="true" pageId="5" pageNumber="594">malabaricus</emphasis>
|
||
(Reichardt, 1932)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2B0DA47553FF96B85B9924FB20130B" authority="Erichson, 1834" authorityName="Erichson" authorityYear="1834" box="[866,1247,548,573]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="species" species="rubripes" subGenus="N.">
|
||
<emphasis id="B95FAA357553FF96B85B9924FC89130B" box="[866,886,549,572]" italics="true" pageId="5" pageNumber="594">H</emphasis>
|
||
. (
|
||
<emphasis id="B95FAA357553FF96B8B69924FC5D130B" box="[911,930,549,572]" italics="true" pageId="5" pageNumber="594">N</emphasis>
|
||
.)
|
||
<emphasis id="B95FAA357553FF96B8839925FBE6130B" box="[954,1049,548,572]" italics="true" pageId="5" pageNumber="594">rubripes</emphasis>
|
||
(Erichson, 1834)
|
||
</taxonomicName>
|
||
are known from
|
||
<collectingCountry id="F33C36B77553FF96BBAE9946FF2E1357" box="[151,209,583,608]" name="India" pageId="5" pageNumber="594">India</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFBA0BD67553FF96BBE69949FE911357" author="Mazur" box="[223,366,584,608]" pageId="5" pageNumber="594" refString="Mazur, S. (2011) A concise catalogue of the Histeridae. Warsaw University of Life Sciences, SGGW Press, Warsaw, 332 pp." type="book" year="2011">Mazur 2011</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C2B0DA47553FF96BAB99948FE311357" box="[384,462,585,608]" class="Insecta" family="Histeridae" genus="Nessus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FAA357553FF96BAB99948FE311357" box="[384,462,585,608]" italics="true" pageId="5" pageNumber="594">Nessus</emphasis>
|
||
</taxonomicName>
|
||
was historically treated as a subgenus of the genus
|
||
<taxonomicName id="4C2B0DA47553FF96BF2C9949FA8E1357" authority="Bickhardt, 1914" authorityName="Bickhardt" authorityYear="1914" box="[1045,1393,583,608]" class="Insecta" family="Histeridae" genus="Hypocacculus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FAA357553FF96BF2C9949FB4C1357" box="[1045,1203,584,608]" italics="true" pageId="5" pageNumber="594">Hypocacculus</emphasis>
|
||
Bickhardt, 1914
|
||
</taxonomicName>
|
||
but was moved into
|
||
<taxonomicName id="4C2B0DA47553FF96BA64996CFE1913B3" box="[349,486,621,644]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FAA357553FF96BA64996CFE1913B3" box="[349,486,621,644]" italics="true" pageId="5" pageNumber="594">Hypocaccus</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFBA0BD67553FF96B920996CFD4813B3" author="Mazur" box="[537,695,620,644]" pageId="5" pageNumber="594" refString="Mazur, S. (2011) A concise catalogue of the Histeridae. Warsaw University of Life Sciences, SGGW Press, Warsaw, 332 pp." type="book" year="2011">Mazur (2011)</bibRefCitation>
|
||
without explanation. Although
|
||
<bibRefCitation id="EFBA0BD67553FF96BF16996CFB3213B3" author="Mazur" box="[1071,1229,620,644]" pageId="5" pageNumber="594" refString="Mazur, S. (2011) A concise catalogue of the Histeridae. Warsaw University of Life Sciences, SGGW Press, Warsaw, 332 pp." type="book" year="2011">Mazur (2011)</bibRefCitation>
|
||
did not give any reasons for inclusion of
|
||
<taxonomicName id="4C2B0DA47553FF96BA9D9990FE0D139F" box="[420,498,657,680]" class="Insecta" family="Histeridae" genus="Nessus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FAA357553FF96BA9D9990FE0D139F" box="[420,498,657,680]" italics="true" pageId="5" pageNumber="594">Nessus</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<taxonomicName id="4C2B0DA47553FF96B92C9990FD61139F" box="[533,670,657,680]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FAA357553FF96B92C9990FD61139F" box="[533,670,657,680]" italics="true" pageId="5" pageNumber="594">Hypocaccus</emphasis>
|
||
</taxonomicName>
|
||
, he apparently did so based on few differences between the two taxa (the densely punctate-variolate structure of frons, occasionally with transverse rugae is often shared between the two taxa), with the exception of perhaps the body size, that is normally much smaller in
|
||
<taxonomicName id="4C2B0DA47553FF96BF5599D8FB4513C7" box="[1132,1210,729,752]" class="Insecta" family="Histeridae" genus="Nessus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FAA357553FF96BF5599D8FB4513C7" box="[1132,1210,729,752]" italics="true" pageId="5" pageNumber="594">Nessus</emphasis>
|
||
</taxonomicName>
|
||
than in
|
||
<taxonomicName id="4C2B0DA47553FF96BE2A99D8FA6313C7" box="[1299,1436,729,752]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FAA357553FF96BE2A99D8FA6313C7" box="[1299,1436,729,752]" italics="true" pageId="5" pageNumber="594">Hypocaccus</emphasis>
|
||
</taxonomicName>
|
||
(Mazur pers. comm.). There are, however, also rather ‘small’
|
||
<taxonomicName id="4C2B0DA47553FF96B85F99FCFC101223" box="[870,1007,765,788]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FAA357553FF96B85F99FCFC101223" box="[870,1007,765,788]" italics="true" pageId="5" pageNumber="594">Hypocaccus</emphasis>
|
||
</taxonomicName>
|
||
species, e.g., an undescribed species from
|
||
<collectingCountry id="F33C36B77553FF96BBE1981EFEB7120F" box="[216,328,799,824]" name="Mongolia" pageId="5" pageNumber="594">Mongolia</collectingCountry>
|
||
, which only reaches 2.00 mm, a length ‘normal’ for species of
|
||
<taxonomicName id="4C2B0DA47553FF96BF039820FB77120F" box="[1082,1160,801,824]" class="Insecta" family="Histeridae" genus="Nessus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FAA357553FF96BF039820FB77120F" box="[1082,1160,801,824]" italics="true" pageId="5" pageNumber="594">Nessus</emphasis>
|
||
</taxonomicName>
|
||
(Lackner unpublished). There are also ‘large’ species of
|
||
<taxonomicName id="4C2B0DA47553FF96B91D9844FD8D126B" box="[548,626,837,860]" class="Insecta" family="Histeridae" genus="Nessus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FAA357553FF96B91D9844FD8D126B" box="[548,626,837,860]" italics="true" pageId="5" pageNumber="594">Nessus</emphasis>
|
||
</taxonomicName>
|
||
, e.g., the psammophilous
|
||
<taxonomicName id="4C2B0DA47553FF96B8949844FAA6126A" authority="Kryzhanovskij, 1966" authorityName="Kryzhanovskij" authorityYear="1966" box="[941,1369,836,861]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="species" species="vlasovi" subGenus="N.">
|
||
<emphasis id="B95FAA357553FF96B8949844FC3E126B" box="[941,961,837,860]" italics="true" pageId="5" pageNumber="594">H</emphasis>
|
||
. (
|
||
<emphasis id="B95FAA357553FF96B8E79844FC0E126B" box="[990,1009,837,860]" italics="true" pageId="5" pageNumber="594">N</emphasis>
|
||
.)
|
||
<emphasis id="B95FAA357553FF96BF349845FBA3126B" box="[1037,1116,836,860]" italics="true" pageId="5" pageNumber="594">vlasovi</emphasis>
|
||
Kryzhanovskij, 1966
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F33C36B77553FF96BBAE9866FECA12B7" box="[151,309,871,896]" name="Turkmenistan" pageId="5" pageNumber="594">Turkmenistan</collectingCountry>
|
||
, that reaches up to 3.00 mm, a body length ‘normal’ for most species of
|
||
<taxonomicName id="4C2B0DA47553FF96BFFD9868FAB212B7" box="[1220,1357,873,896]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="B95FAA357553FF96BFFD9868FAB212B7" box="[1220,1357,873,896]" italics="true" pageId="5" pageNumber="594">Hypocaccus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26C174E7553FF96BE629869FA631248" box="[1371,1436,872,895]" pageId="5" pageNumber="594" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
(Kryzhnovskij & Reichardt 1976).
|
||
</paragraph>
|
||
<paragraph id="8B9476277553FF96BBFE98AEFC971462" blockId="5.[151,1437,151,1365]" pageId="5" pageNumber="594">
|
||
The newly described species
|
||
<taxonomicName id="4C2B0DA47553FF96B91798B0FD4D12FF" box="[558,690,944,968]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="species" species="hirsutus">
|
||
<emphasis id="B95FAA357553FF96B91798B0FD4D12FF" box="[558,690,944,968]" italics="true" pageId="5" pageNumber="594">H. hirsutus</emphasis>
|
||
</taxonomicName>
|
||
does not fit neatly into any current subgenus of Old World
|
||
<taxonomicName id="4C2B0DA47553FF96BBAE98D4FEDF12DB" box="[151,288,981,1004]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FAA357553FF96BBAE98D4FEDF12DB" box="[151,288,981,1004]" italics="true" pageId="5" pageNumber="594">Hypocaccus</emphasis>
|
||
</taxonomicName>
|
||
based on the setose pronotal hypomeron, as well as strongly elevated anterior clypeal margin. It is morphologically most similar to members of
|
||
<taxonomicName id="4C2B0DA47553FF96B99998F8FCD61527" box="[672,809,1017,1040]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="B95FAA357553FF96B99998F8FCD61527" box="[672,809,1017,1040]" italics="true" pageId="5" pageNumber="594">Hypocaccus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26C174E7553FF96B80B98F9FC901538" box="[818,879,1016,1039]" pageId="5" pageNumber="594" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
On the other hand, the elevated anterior clypeal margin resembles the monotypic African genus
|
||
<taxonomicName id="4C2B0DA47553FF96B98D9F1DFBEF1502" authority="Vienna, 1995" authorityName="Vienna" authorityYear="1995" box="[692,1040,1052,1077]" class="Insecta" family="Histeridae" genus="Parahypocaccus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FAA357553FF96B98D9F1DFC901503" box="[692,879,1052,1076]" italics="true" pageId="5" pageNumber="594">Parahypocaccus</emphasis>
|
||
Vienna, 1995
|
||
</taxonomicName>
|
||
, but differs from it by the presence of prosternal foveae. Apparently, the external morphological characters (body size, pronotal pilosity, etc.) vary between species of both
|
||
<taxonomicName id="4C2B0DA47553FF96BA8F9F64FDC0154B" box="[438,575,1125,1148]" class="Insecta" family="Histeridae" genus="Hypocaccus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="B95FAA357553FF96BA8F9F64FDC0154B" box="[438,575,1125,1148]" italics="true" pageId="5" pageNumber="594">Hypocaccus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26C174E7553FF96B9709F64FD78154B" box="[585,647,1125,1148]" pageId="5" pageNumber="594" sensu="stricto">s. str.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C2B0DA47553FF96B9FA9F64FCEE154B" box="[707,785,1125,1148]" class="Insecta" family="Histeridae" genus="Nessus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FAA357553FF96B9FA9F64FCEE154B" box="[707,785,1125,1148]" italics="true" pageId="5" pageNumber="594">Nessus</emphasis>
|
||
</taxonomicName>
|
||
, most likely convergent adaptations to the environment. Recent phylogenetic analysis of the
|
||
<taxonomicName id="4C2B0DA47553FF96B9059F86FD4B1597" box="[572,692,1159,1184]" class="Insecta" family="Histeridae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="subFamily" subFamily="Saprininae">Saprininae</taxonomicName>
|
||
genera and subgenera (
|
||
<bibRefCitation id="EFBA0BD67553FF96B8F19F86FB881597" author="Lackner" box="[968,1143,1159,1184]" pageId="5" pageNumber="594" refString="Lackner, T. (2014 d) Phylogeny of the Saprininae subfamily reveals interesting ecological shifts in the history of the subfamily (Coleoptera: Histeridae). Zoological Journal of the Linnean Society, 172, 521 - 555. http: // dx. doi. org / 10.1111 / zoj. 12182" type="journal article" year="2014" yearSuffix="d">Lackner 2014d</bibRefCitation>
|
||
) based on morphological characters failed to clarify the relationships between the taxa
|
||
<taxonomicName id="4C2B0DA47553FF96B8769FACFBCA15F3" authority="Hypocaccus" authorityName="Hypocaccus" box="[847,1077,1197,1220]" class="Insecta" family="Histeridae" genus="Nessus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FAA357553FF96B8769FACFC6215F3" box="[847,925,1197,1220]" italics="true" pageId="5" pageNumber="594">Nessus</emphasis>
|
||
,
|
||
<emphasis id="B95FAA357553FF96B8959FACFBCA15F3" box="[940,1077,1197,1220]" italics="true" pageId="5" pageNumber="594">Hypocaccus</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="4C2B0DA47553FF96BF659FADFAEA15F3" box="[1116,1301,1196,1220]" class="Insecta" family="Histeridae" genus="Baeckmanniolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="594" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FAA357553FF96BF659FADFAEA15F3" box="[1116,1301,1196,1220]" italics="true" pageId="5" pageNumber="594">Baeckmanniolus</emphasis>
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</taxonomicName>
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||
, recovering them in a large unresolved clade of taxa sharing a single synapomorphy of single pear or stipe-shaped vesicle inside their antennal club. The newly-described species lacks numerous characters (e.g. male genitalia, mouthparts) that would be needed for inclusion in a cladistic analysis for proper (sub)generic placement. A subsequent phylogenetic study based on molecular characters is in progress.
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||
</paragraph>
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||
</subSubSection>
|
||
</treatment>
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||
</document> |