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<document id="8A56CDDA96BCB7D7BC3AF0CB4FAAEE98" ID-DOI="10.1080/00222930802277616" ID-ISSN="1464-5262" ID-Zenodo-Dep="5223730" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629391701861" checkinUser="felipe" docAuthor="Csuzdi, Csaba" docDate="2008" docId="038287A1F317FFA76A83FBD3FD83CC90" docLanguage="en" docName="JNatHist.42.33-34.2227-2241.pdf" docOrigin="Journal of Natural History 42 (33 - 34)" docSource="http://dx.doi.org/10.1080/00222930802277616" docStyle="DocumentStyle:FCA295E821436EF9307F50C7AC261B53.5:JNatHist.2008-2012.journal_article.1cover" docStyleId="FCA295E821436EF9307F50C7AC261B53" docStyleName="JNatHist.2008-2012.journal_article.1cover" docStyleVersion="5" docTitle="Dichogaster (Diplothecodrilus) paulclarki Csuzdi 2008, sp. nov." docType="treatment" docVersion="3" lastPageNumber="2235" masterDocId="FFBBFFD9F310FFAE6B16FF8BFFA6CE6E" masterDocTitle="Earthworms from the Kivu Region, D. R. Congo with description of two new Dichogaster Beddard, 1888 (Oligochaeta: Acanthodrilidae, Ocnerodrilidae)" masterLastPageNumber="2241" masterPageNumber="2227" pageNumber="2233" updateTime="1699071437077" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="C975A34DB0F8FD4C60C57D4276D41B8A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CD55BBCFB36DCC2AEECEA0113474449F">
<mods:title id="C27146800BA5EEB6CE4005972A6E7DD7">Earthworms from the Kivu Region, D. R. Congo with description of two new Dichogaster Beddard, 1888 (Oligochaeta: Acanthodrilidae, Ocnerodrilidae)</mods:title>
</mods:titleInfo>
<mods:name id="94AD3711845661F0871AE443D6BAE9CD" type="personal">
<mods:role id="A6C78160A7C54D71C9A0B7A7BFDB8AE6">
<mods:roleTerm id="E2611953D9236D9B848B86605D54F58B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6440749B3C5D58BDBB624AA173720348">Csuzdi, Csaba</mods:namePart>
</mods:name>
<mods:typeOfResource id="2F7A82771D4BC7F72A696F9386CB2A9E">text</mods:typeOfResource>
<mods:relatedItem id="21AFCBAB085B94EF48C9B7EA613FEBCF" type="host">
<mods:titleInfo id="D980B3322FFE6DB2E4E94C475F68D711">
<mods:title id="5FE99E700B8D58426A0B225162BD0CD7">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="2C2D21DC5567FFB5D9B667A8D1D6BD22">
<mods:date id="89742D71AF6E6CEBA6741E13059D1714">2008</mods:date>
<mods:detail id="7C559A5D3E433D33E8ECA084DF504E2C" type="pubDate">
<mods:number id="05E369BA7CC6A171DB765ADA08A5D6E4">2008-09-30</mods:number>
</mods:detail>
<mods:detail id="08FE7A4462CE708C2387988D8C70C0A0" type="volume">
<mods:number id="25F07964BB1E604D33BAF08A0BA31F24">42</mods:number>
</mods:detail>
<mods:detail id="97B059FC2DFF07E4FC467F05CF84C9FB" type="issue">
<mods:number id="7D3EA9C1170EA7C53318D0025F6F548B">33 - 34</mods:number>
</mods:detail>
<mods:extent id="00F7B429B61ED28BB80017CD0D8305BF" unit="page">
<mods:start id="7EDC8025AC51FB17FC8918B9230BC284">2227</mods:start>
<mods:end id="221B5C2CFFA79C20E277DFCEF28D5278">2241</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="716EDA5E10BF25DB370F54401AC82295">
<mods:url id="E905D4C007881E7ECC8F6FD7B306ADB8">http://dx.doi.org/10.1080/00222930802277616</mods:url>
</mods:location>
<mods:classification id="10184FC63D0CEAB53689D8ABEE92FF3F">journal article</mods:classification>
<mods:identifier id="6F63D7C711AC836859047ED2023F5827" type="DOI">10.1080/00222930802277616</mods:identifier>
<mods:identifier id="6D1343FE7061F6434D88D2DEBC3EDB6B" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="59F8A035F8E3EEFE9C9324604BB94A86" type="Zenodo-Dep">5223730</mods:identifier>
</mods:mods>
<treatment id="038287A1F317FFA76A83FBD3FD83CC90" LSID="urn:lsid:plazi:treatment:038287A1F317FFA76A83FBD3FD83CC90" httpUri="http://treatment.plazi.org/id/038287A1F317FFA76A83FBD3FD83CC90" lastPageId="9" lastPageNumber="2235" pageId="7" pageNumber="2233">
<subSubSection id="C331653CF317FFA96A83FBD3FC77CA1E" box="[405,977,1112,1136]" pageId="7" pageNumber="2233" type="nomenclature">
<paragraph id="8B9436B7F317FFA96A83FBD3FC77CA1E" blockId="7.[405,977,1112,1168]" box="[405,977,1112,1136]" pageId="7" pageNumber="2233">
<heading id="D0DC81DBF317FFA96A83FBD3FC77CA1E" box="[405,977,1112,1136]" centered="true" fontSize="36" level="2" pageId="7" pageNumber="2233" reason="7">
<taxonomicName id="4C2B4D34F317FFA96A83FBD3FCCACA1E" authority="Csuzdi, 2008" authorityName="Csuzdi" authorityYear="2008" box="[405,876,1112,1136]" class="Clitellata" family="Octochaetidae" genus="Dichogaster" kingdom="Animalia" order="Haplotaxida" pageId="7" pageNumber="2233" phylum="Annelida" rank="species" species="paulclarki" status="sp. nov." subGenus="Diplothecodrilus">
<emphasis id="B95FEAA5F317FFA96A83FBD3FCCACA1E" bold="true" box="[405,876,1112,1136]" italics="true" pageId="7" pageNumber="2233">Dichogaster (Diplothecodrilus) paulclarki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C57DEF317FFA96863FBD3FC77CA1E" box="[885,977,1112,1136]" pageId="7" pageNumber="2233" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C331653CF317FFA76976FBF3FD83CC90" lastPageId="9" lastPageNumber="2235" pageId="7" pageNumber="2233" type="description">
<paragraph id="8B9436B7F317FFA96976FBF3FCA0CAFE" blockId="7.[405,977,1112,1168]" box="[608,774,1144,1168]" pageId="7" pageNumber="2233">
(
<figureCitation id="13102A32F317FFA96971FBF3FD58CAFE" box="[615,766,1144,1168]" captionStart="Figure 4" captionStartId="8.[146,213,809,831]" captionTargetBox="[146,1098,165,773]" captionTargetId="figure-176@8.[146,1104,165,773]" captionTargetPageId="8" captionText="Figure 4. Dichogaster (Dt.) paulclarki sp. nov. (A) Ventral part of the clitellar region; (B) penial seta; (C) ornamentation of the penial seta; (D) spermatheca. St, spermathecal pores; Pr, prostate pores." figureDoi="http://doi.org/10.5281/zenodo.5223738" httpUri="https://zenodo.org/record/5223738/files/figure.png" pageId="7" pageNumber="2233">Figure 4A–D</figureCitation>
)
</paragraph>
<paragraph id="8B9436B7F317FFA96BC3FB42FE0FCA8F" blockId="7.[213,425,1225,1249]" box="[213,425,1225,1249]" pageId="7" pageNumber="2233">
<emphasis id="B95FEAA5F317FFA96BC3FB42FE0FCA8F" box="[213,425,1225,1249]" italics="true" pageId="7" pageNumber="2233">Material examined</emphasis>
</paragraph>
<paragraph id="8B9436B7F317FFA96BC3FB7AFE6BCB47" blockId="7.[213,1169,1265,1321]" pageId="7" pageNumber="2233">
<materialsCitation id="3B433CEAF317FFA96BC3FB7AFE6BCB47" collectingDate="1973-07-25" collectionCode="R" collectorName="Gasana N'Doli" country="Democratic Republic of the Congo" location="Kivu Region" pageId="7" pageNumber="2233" specimenCode="BMNH 1997.1628" specimenCount="1" typeStatus="holotype">
<emphasis id="B95FEAA5F317FFA96BC3FB7AFE9ACB67" box="[213,316,1265,1289]" italics="true" pageId="7" pageNumber="2233">
<typeStatus id="54908815F317FFA96BC3FB7AFE9ACB67" box="[213,316,1265,1289]" pageId="7" pageNumber="2233" type="holotype">Holotype</typeStatus>
</emphasis>
.
<specimenCode id="DB8D9ECCF317FFA96A44FB7AFD8DCB67" box="[338,555,1265,1289]" collectionCode="BMNH" pageId="7" pageNumber="2233">BMNH 1997.1628</specimenCode>
D.
<collectionCode id="ED3AAE72F317FFA96940FB7AFDC8CB67" box="[598,622,1265,1289]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="2233">R</collectionCode>
.
<collectingCountry id="F33C7627F317FFA96997FB7AFD76CB67" box="[641,720,1265,1289]" name="Democratic Republic of the Congo" pageId="7" pageNumber="2233">Congo</collectingCountry>
,
<location id="8EF4606CF317FFA969F2FB7AFC24CB67" LSID="urn:lsid:plazi:treatment:038287A1F317FFA76A83FBD3FD83CC90:8EF4606CF317FFA969F2FB7AFC24CB67" box="[740,898,1265,1289]" country="Democratic Republic of the Congo" name="Kivu Region" pageId="7" pageNumber="2233">Kivu Region</location>
, Bugabo, leg.
<collectorName id="26DE5361F317FFA96F2CFB7AFE8ECB47" pageId="7" pageNumber="2233">Gasana N’Doli</collectorName>
,
<date id="FF951077F317FFA96A23FA9AFE6FCB47" box="[309,457,1297,1321]" pageId="7" pageNumber="2233" value="1973-07-25">
<collectingDate id="EFD1E99FF317FFA96A23FA9AFE6FCB47" box="[309,457,1297,1321]" pageId="7" pageNumber="2233" value="1973-07-25">25 July 1973</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8B9436B7F317FFA96BC3FAD1FB37CBDF" blockId="7.[213,1169,1369,1457]" pageId="7" pageNumber="2233">
<materialsCitation id="3B433CEAF317FFA96BC3FAD1FD10CB1F" box="[213,694,1369,1394]" collectionCode="HNHM" pageId="7" pageNumber="2233" specimenCount="1" typeStatus="paratype">
<emphasis id="B95FEAA5F317FFA96BC3FAD1FEE3CB1C" box="[213,325,1370,1394]" italics="true" pageId="7" pageNumber="2233">
<typeStatus id="54908815F317FFA96BC3FAD1FEE3CB1C" box="[213,325,1370,1394]" pageId="7" pageNumber="2233" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<collectionCode id="ED3AAE72F317FFA96A45FAD2FE08CB1F" box="[339,430,1369,1393]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="7" pageNumber="2233">HNHM</collectionCode>
AF/3544, one exemplar
</materialsCitation>
,
<materialsCitation id="3B433CEAF317FFA969A9FAD2FDB4CBFF" collectionCode="BMNH" pageId="7" pageNumber="2233" specimenCode="BMNH 1997.1629" specimenCount="1" typeStatus="holotype">
<specimenCode id="DB8D9ECCF317FFA969A9FAD2FC2ACB1F" box="[703,908,1369,1393]" collectionCode="BMNH" pageId="7" pageNumber="2233">BMNH 1997.1629</specimenCode>
, one exemplar. Locality same as that of the
<typeStatus id="54908815F317FFA96ABBFAF2FDB4CBFF" box="[429,530,1401,1425]" pageId="7" pageNumber="2233" type="holotype">holotype</typeStatus>
</materialsCitation>
. HNHM AF/3571, one exemplar,
<materialsCitation id="3B433CEAF317FFA96899FAF2FEF9CBDF" collectionCode="BMNH" pageId="7" pageNumber="2233" specimenCode="BMNH 1997.1593" specimenCount="1" typeStatus="paratype">
<specimenCode id="DB8D9ECCF317FFA96899FAF2FBF8CBFF" box="[911,1118,1401,1425]" collectionCode="BMNH" pageId="7" pageNumber="2233">BMNH 1997.1593</specimenCode>
, one exemplar. D
</materialsCitation>
.
<materialsCitation id="3B433CEAF317FFA96A75FA12FB2BCBDF" box="[355,1165,1433,1457]" collectingDate="1973-07-15" collectionCode="R" collectorName="Gasana N'Doli" country="Democratic Republic of the Congo" location="Kivu Region" municipality="River Mululu" pageId="7" pageNumber="2233" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED3AAE72F317FFA96A75FA12FEDCCBDF" box="[355,378,1433,1457]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="2233">R</collectionCode>
.
<collectingCountry id="F33C7627F317FFA96A93FA12FE74CBDF" box="[389,466,1433,1457]" name="Democratic Republic of the Congo" pageId="7" pageNumber="2233">Congo</collectingCountry>
,
<location id="8EF4606CF317FFA96ACBFA12FDC9CBDF" LSID="urn:lsid:plazi:treatment:038287A1F317FFA76A83FBD3FD83CC90:8EF4606CF317FFA96ACBFA12FDC9CBDF" box="[477,623,1433,1457]" country="Democratic Republic of the Congo" municipality="River Mululu" name="Kivu Region" pageId="7" pageNumber="2233">Kivu Region</location>
,
<collectingMunicipality id="6BF0ACCDF317FFA9696FFA12FCB4CBDF" box="[633,786,1433,1457]" pageId="7" pageNumber="2233">River Mululu</collectingMunicipality>
, leg.
<collectorName id="26DE5361F317FFA9685CFA12FC53CBDF" box="[842,1013,1433,1457]" pageId="7" pageNumber="2233">Gasana N’Doli</collectorName>
,
<date id="FF951077F317FFA96F17FA12FB2BCBDF" box="[1025,1165,1433,1457]" pageId="7" pageNumber="2233" value="1973-07-15">
<collectingDate id="EFD1E99FF317FFA96F17FA12FB2BCBDF" box="[1025,1165,1433,1457]" pageId="7" pageNumber="2233" value="1973-07-15">15 July 1973</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8B9436B7F317FFA96BC3FA61FEE4C86C" blockId="7.[213,322,1514,1538]" box="[213,322,1514,1538]" pageId="7" pageNumber="2233">
<emphasis id="B95FEAA5F317FFA96BC3FA61FEE4C86C" box="[213,322,1514,1538]" italics="true" pageId="7" pageNumber="2233">Diagnosis</emphasis>
</paragraph>
<paragraph id="8B9436B7F317FFA96BC3F999FB37C8C4" blockId="7.[213,1169,1554,1706]" pageId="7" pageNumber="2233">
Length
<quantity id="4CD39B52F317FFA96A3AF999FE04C844" box="[300,418,1554,1578]" metricMagnitude="-2" metricUnit="m" metricValue="4.55" metricValueMax="5.0" metricValueMin="4.1" pageId="7" pageNumber="2233" unit="mm" value="45.5" valueMax="50.0" valueMin="41.0">41–50 mm</quantity>
, D.
<quantity id="4CD39B52F317FFA96AD9F999FD99C844" box="[463,575,1554,1578]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.3" metricValueMin="2.0" pageId="7" pageNumber="2233" unit="mm" value="2.15" valueMax="2.3" valueMin="2.0">2–2.3 mm</quantity>
, no. of segments 95–115. Pigmentation lacking. First dorsal pore in 5/6. Clitellum 13–20.
<specimenCount id="9D2DFD3EF317FFA96979F9B9FDDBC825" box="[623,637,1586,1611]" pageId="7" pageNumber="2233" type="female">♀</specimenCount>
14 paired before setae a–a. Prostatic pores 17, 19. Spermathecal pores 7/8, 8/9, spermathecae with a subdivided irregular ampoule, and an equally long duct. Gizzard in 5/6, last pair of hearts in 13. Calciferous glands in 15–17 of equal size. Excretory system meroic with four meronephridia per side.
</paragraph>
<caption id="DF54663FF318FFA66B84FCA2FE8ACD14" ID-DOI="http://doi.org/10.5281/zenodo.5223738" ID-Zenodo-Dep="5223738" httpUri="https://zenodo.org/record/5223738/files/figure.png" pageId="8" pageNumber="2234" startId="8.[146,213,809,831]" targetBox="[146,1098,165,773]" targetPageId="8">
<paragraph id="8B9436B7F318FFA66B84FCA2FE8ACD14" blockId="8.[146,1102,809,890]" pageId="8" pageNumber="2234">
Figure 4.
<taxonomicName id="4C2B4D34F318FFA66BEBFCA2FD83CD50" authority="Csuzdi, 2008" authorityName="Csuzdi" authorityYear="2008" box="[253,549,809,830]" class="Clitellata" family="Octochaetidae" genus="Dichogaster" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="2234" phylum="Annelida" rank="species" species="paulclarki" status="sp. nov." subGenus="Diplothecodrilus">
<emphasis id="B95FEAA5F318FFA66BEBFCA2FD83CD50" box="[253,549,809,830]" italics="true" pageId="8" pageNumber="2234">Dichogaster (Dt.) paulclarki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C57DEF318FFA66938FCA2FD24CD51" box="[558,642,809,831]" pageId="8" pageNumber="2234" rank="species">sp. nov.</taxonomicNameLabel>
(A) Ventral part of the clitellar region; (B) penial seta; (C) ornamentation of the penial seta; (D) spermatheca. St, spermathecal pores; Pr, prostate pores.
</paragraph>
</caption>
<paragraph id="8B9436B7F318FFA66B84FC38FF45CD85" blockId="8.[146,1103,947,1003]" pageId="8" pageNumber="2234">
Penial seta L.
<quantity id="4CD39B52F318FFA66A2DFC38FE19CDA5" box="[315,447,947,971]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="8" pageNumber="2234" unit="mm" value="1.75" valueMax="1.8" valueMin="1.7">1.7–1.8 mm</quantity>
, D.
<quantity id="4CD39B52F318FFA66AE2FC38FDC2CDA5" box="[500,612,947,971]" metricMagnitude="-5" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="2234" unit="mm" value="0.015">0.015 mm</quantity>
, tip sharply pointed, ornamentation small thorns.
</paragraph>
<paragraph id="8B9436B7F318FFA66B84FBAFFEFECA52" blockId="8.[146,344,1060,1084]" box="[146,344,1060,1084]" pageId="8" pageNumber="2234">
<emphasis id="B95FEAA5F318FFA66B84FBAFFEFECA52" box="[146,344,1060,1084]" italics="true" pageId="8" pageNumber="2234">Derivatio nominis</emphasis>
</paragraph>
<paragraph id="8B9436B7F318FFA66B84FBC7FCB9CAEB" blockId="8.[146,1103,1100,1157]" pageId="8" pageNumber="2234">The specific epithet refers to Dr Paul Clark (The Natural History Museum, London) for always being helpful throughout my stay in London.</paragraph>
<paragraph id="8B9436B7F318FFA66B84FB35FEC8CAB8" blockId="8.[146,366,1214,1238]" box="[146,366,1214,1238]" pageId="8" pageNumber="2234">
<emphasis id="B95FEAA5F318FFA66B84FB35FEC8CAB8" box="[146,366,1214,1238]" italics="true" pageId="8" pageNumber="2234">External characters</emphasis>
</paragraph>
<paragraph id="8B9436B7F318FFA66B84FB6CFCC3CA90" blockId="8.[146,869,1254,1279]" box="[146,869,1254,1279]" pageId="8" pageNumber="2234">
<emphasis id="B95FEAA5F318FFA66B84FB6CFF5FCA91" box="[146,249,1255,1279]" italics="true" pageId="8" pageNumber="2234">
<typeStatus id="54908815F318FFA66B84FB6CFF5FCA91" box="[146,249,1255,1279]" pageId="8" pageNumber="2234" type="holotype">Holotype</typeStatus>
</emphasis>
. Length
<quantity id="4CD39B52F318FFA66A75FB6DFE17CA90" box="[355,433,1254,1278]" metricMagnitude="-2" metricUnit="m" metricValue="4.6" pageId="8" pageNumber="2234" unit="mm" value="46.0">46 mm</quantity>
, diameter
<quantity id="4CD39B52F318FFA6693CFB6DFDCDCA90" box="[554,619,1254,1278]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="2234" unit="mm" value="2.0">2 mm</quantity>
, no. of segments 115.
</paragraph>
<paragraph id="8B9436B7F318FFA66B84FABCFC53C87E" blockId="8.[146,1103,1335,1552]" pageId="8" pageNumber="2234">
<emphasis id="B95FEAA5F318FFA66B84FABCFEA4CB21" box="[146,258,1335,1359]" italics="true" pageId="8" pageNumber="2234">
<typeStatus id="54908815F318FFA66B84FABCFEA4CB21" box="[146,258,1335,1359]" pageId="8" pageNumber="2234" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<quantity id="4CD39B52F318FFA66A04FABCFE23CB21" box="[274,389,1335,1359]" metricMagnitude="-2" metricUnit="m" metricValue="4.55" metricValueMax="5.0" metricValueMin="4.1" pageId="8" pageNumber="2234" unit="mm" value="45.5" valueMax="50.0" valueMin="41.0">41–50 mm</quantity>
in length,
<quantity id="4CD39B52F318FFA66917FABCFDC8CB21" box="[513,622,1335,1359]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.3" metricValueMin="2.0" pageId="8" pageNumber="2234" unit="mm" value="2.15" valueMax="2.3" valueMin="2.0">2–2.3 mm</quantity>
in diameter, number of segments 95–115. Colour pale, pigmentation lacking. Prostomium schizolobous. First dorsal pore in 5/ 6. Setae all ventral, setal arrangement after the clitellum aa:ab:bc:cd:dd54:1.1:4:1:34. Clitellum extends over segments 13–20, male field oblong. Prostatic pores paired on 17, 19 in the setal line b. Male pores externally not visible, they are on 18, in the seminal groves. Female pores paired on 14 just before the setae aa. Spermathecal pores paired in 7/8, 8/9 near to the setae b. Porophores absent (
<figureCitation id="13102A32F318FFA66865FA73FC4FC87E" box="[883,1001,1528,1552]" captionStart="Figure 4" captionStartId="8.[146,213,809,831]" captionTargetBox="[146,1098,165,773]" captionTargetId="figure-176@8.[146,1104,165,773]" captionTargetPageId="8" captionText="Figure 4. Dichogaster (Dt.) paulclarki sp. nov. (A) Ventral part of the clitellar region; (B) penial seta; (C) ornamentation of the penial seta; (D) spermatheca. St, spermathecal pores; Pr, prostate pores." figureDoi="http://doi.org/10.5281/zenodo.5223738" httpUri="https://zenodo.org/record/5223738/files/figure.png" pageId="8" pageNumber="2234">Figure 4A</figureCitation>
).
</paragraph>
<paragraph id="8B9436B7F318FFA66B84F9C2FEC3C80F" blockId="8.[146,357,1609,1633]" box="[146,357,1609,1633]" pageId="8" pageNumber="2234">
<emphasis id="B95FEAA5F318FFA66B84F9C2FEC3C80F" box="[146,357,1609,1633]" italics="true" pageId="8" pageNumber="2234">Internal characters</emphasis>
</paragraph>
<paragraph id="8B9436B7F318FFA76B84F9F9FBE3CC4D" blockId="8.[146,1103,1650,1706]" lastBlockId="9.[213,1169,168,547]" lastPageId="9" lastPageNumber="2235" pageId="8" pageNumber="2234">
The first visible septum 6/7, thickened septa lacking. Oesophageal gizzards small, in segment 5 and 6. Three pairs of calciferous glands in segment 15–17 of equal size. Excretory system meroic, with four meronephridia on each side. Paired hearts are present in segments 10–13. Typhlosole very small, begins in segment 21, auxiliary typhlosole lacking. Testes in 10 and 11 enclosed in perioesophageal testis sacs. Seminal vesicles in 11, 12, and a pair of large ovaries in segment 13. Seminal duct convoluted, discharging in segment 18. Two pairs of prostatic glands are present in 17 and 19, slightly intruding into the neighbouring segments and each with a penial setal sac containing a mature and several immature penial setae. The mature penial seta is undulated at the ectal third, about
<quantity id="4CD39B52F319FFA7694EFE01FD7ECFCC" box="[600,728,394,418]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="9" pageNumber="2235" unit="mm" value="1.75" valueMax="1.8" valueMin="1.7">1.7–1.8 mm</quantity>
long and
<quantity id="4CD39B52F319FFA76851FE01FC14CFCC" box="[839,946,394,418]" metricMagnitude="-5" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="2235" unit="mm" value="0.015">0.015 mm</quantity>
wide with a sharply pointed tip. Ornamentation consists of scattered thorns (
<figureCitation id="13102A32F319FFA76842FE21FC78CFAC" box="[852,990,426,450]" captionStart="Figure 4" captionStartId="8.[146,213,809,831]" captionTargetBox="[146,1098,165,773]" captionTargetId="figure-176@8.[146,1104,165,773]" captionTargetPageId="8" captionText="Figure 4. Dichogaster (Dt.) paulclarki sp. nov. (A) Ventral part of the clitellar region; (B) penial seta; (C) ornamentation of the penial seta; (D) spermatheca. St, spermathecal pores; Pr, prostate pores." figureDoi="http://doi.org/10.5281/zenodo.5223738" httpUri="https://zenodo.org/record/5223738/files/figure.png" pageId="9" pageNumber="2235">Figure 4B,C</figureCitation>
). There are two pairs of spermathecae in segment 8 and 9. The ampoule is subdivided. The ental part small and turns back to the pear-shaped ectal part which bears a small rounded diverticulum. The duct is narrow and half as long as the ampoule (
<figureCitation id="13102A32F319FFA768D5FD80FB9FCC4D" box="[963,1081,523,547]" captionStart="Figure 4" captionStartId="8.[146,213,809,831]" captionTargetBox="[146,1098,165,773]" captionTargetId="figure-176@8.[146,1104,165,773]" captionTargetPageId="8" captionText="Figure 4. Dichogaster (Dt.) paulclarki sp. nov. (A) Ventral part of the clitellar region; (B) penial seta; (C) ornamentation of the penial seta; (D) spermatheca. St, spermathecal pores; Pr, prostate pores." figureDoi="http://doi.org/10.5281/zenodo.5223738" httpUri="https://zenodo.org/record/5223738/files/figure.png" pageId="9" pageNumber="2235">Figure 4D</figureCitation>
).
</paragraph>
<paragraph id="8B9436B7F319FFA76BC3FDD7FE9ECC1A" blockId="9.[213,312,604,628]" box="[213,312,604,628]" pageId="9" pageNumber="2235">
<emphasis id="B95FEAA5F319FFA76BC3FDD7FE9ECC1A" box="[213,312,604,628]" italics="true" pageId="9" pageNumber="2235">Remarks</emphasis>
</paragraph>
<paragraph id="8B9436B7F319FFA76BC3FD0EFD83CC90" blockId="9.[213,1169,645,766]" pageId="9" pageNumber="2235">
This species belongs to the
<emphasis id="B95FEAA5F319FFA76912FD0EFDEFCCF3" box="[516,585,645,669]" italics="true" pageId="9" pageNumber="2235">affinis</emphasis>
species group and, by the penial setae, resembles
<taxonomicName id="4C2B4D34F319FFA76F63FD0EFDC9CCD3" authority="Stephenson, 1933" authorityName="Stephenson" authorityYear="1933" class="Clitellata" family="Octochaetidae" genus="Dichogaster" kingdom="Animalia" order="Haplotaxida" pageId="9" pageNumber="2235" phylum="Annelida" rank="species" species="kigogoana" subGenus="Diplothecodrilus">
<emphasis id="B95FEAA5F319FFA76F63FD0EFE35CCD3" italics="true" pageId="9" pageNumber="2235">D. (Dt.) kigogoana</emphasis>
Stephenson, 1933
</taxonomicName>
. Nevertheless it differs from the latter in the position of the last pair of hearts, in the shape of the spermathecae and in the absence of a thickened septa.
</paragraph>
</subSubSection>
</treatment>
</document>
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