501 lines
63 KiB
XML
501 lines
63 KiB
XML
<document id="0B132F9E703A055D9D4DCBFE6F28DBA1" ID-DOI="10.5281/zenodo.275486" ID-GBIF-Dataset="0cbe8247-37a3-4243-bead-87a3bd2581c9" ID-ISSN="1175-5326" ID-Zenodo-Dep="275486" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460103532582" checkinUser="plazi" docAuthor="Ahyong, Shane T. & Chan, Tin-Yam" docDate="2010" docId="03828796FFC3C27F3595FEB2FB76F977" docLanguage="en" docName="zt02332p068.pdf" docOrigin="Zootaxa 2332" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Lithodes formosae Ahyong & Chan, 2010, sp. nov." docType="treatment" docVersion="9" lastPageNumber="67" masterDocId="FFBBFFEEFFC2C2793502FF95FFDAFFA5" masterDocTitle="Lithodes formosae, a new species of king crab from Taiwan (Crustacea: Decapoda: Lithodidae)" masterLastPageNumber="68" masterPageNumber="61" pageNumber="62" updateTime="1698234814124" updateUser="plazi">
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<mods:mods id="0B58A5CF1995F08F2CAB3B0D3F3640A8" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="28FFD16F054435532CD0F0A6DA40D044">
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<mods:title id="918826BC4F009B8AE97DF62C85E8866B">Lithodes formosae, a new species of king crab from Taiwan (Crustacea: Decapoda: Lithodidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="4283F809D6A5E276DC9C6B17E46BF304" type="personal">
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<mods:roleTerm id="F81EA733B4993C4B1A8E6B6958D7C50B">Author</mods:roleTerm>
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<mods:namePart id="A40FDCBFAFDA8E8A53ACC1A24C205F00">Ahyong, Shane T.</mods:namePart>
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</mods:name>
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<mods:name id="2A3F1D7B4F648E25D75F44EE015945A8" type="personal">
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<mods:role id="09E15D171CC6E8A217ECF1EC7D061D0C">
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<mods:roleTerm id="5B8ED85CC76D4AED894847F79EA293D6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4EC1A60F059BCCD8A6CBCDDEEB2A4B86">Chan, Tin-Yam</mods:namePart>
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<mods:typeOfResource id="8D9D2626B1C7E6547A50C18BA8344F76">text</mods:typeOfResource>
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<mods:title id="1178F9D50FCCBCEEE5FF452030F3A5A7">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="20E01B952B8D8890F366CCB14DD75F96">
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<mods:date id="0900858007CC95EB2E47DED4130B54A5">2010</mods:date>
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<mods:detail id="F62289CDD7BFACE2D8E96DE87A47F1FB" type="volume">
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<mods:number id="C57BAC27DA0D9A3AA258C2CEF6A4433B">2332</mods:number>
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<mods:start id="ABEA40C325EDF520E707D9405A2CAA11">61</mods:start>
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<mods:end id="514569A1B6F82ED770F4A0179FDCF1B2">68</mods:end>
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<mods:classification id="95D388561CFEF4C4DED100E8662E0E06">journal article</mods:classification>
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<mods:identifier id="056B7E087467CDE824578576F1BCF6D0" type="DOI">10.5281/zenodo.275486</mods:identifier>
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<mods:identifier id="FBE929110578597AEAB7790EA5B0E82A" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="2CA67C485C27A9CC7948267ACCD2A47E" type="Zenodo-Dep">275486</mods:identifier>
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<treatment id="03828796FFC3C27F3595FEB2FB76F977" ID-DOI="http://doi.org/10.5281/zenodo.3510878" ID-GBIF-Taxon="127654648" ID-Zenodo-Dep="3510878" LSID="urn:lsid:plazi:treatment:03828796FFC3C27F3595FEB2FB76F977" httpUri="http://treatment.plazi.org/id/03828796FFC3C27F3595FEB2FB76F977" lastPageId="6" lastPageNumber="67" pageId="1" pageNumber="62">
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<subSubSection id="C331650BFFC3C2783595FEB2FECDFEC1" pageId="1" pageNumber="62" type="nomenclature">
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<paragraph id="8B943680FFC3C2783595FEB2FE01FEE4" blockId="1.[151,475,295,356]" box="[151,475,295,321]" pageId="1" pageNumber="62">
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<heading id="D0DC81ECFFC3C2783595FEB2FE01FEE4" bold="true" box="[151,475,295,321]" fontSize="11" level="1" pageId="1" pageNumber="62" reason="1">
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<emphasis id="B95FEA92FFC3C2783595FEB2FE01FEE4" bold="true" box="[151,475,295,321]" pageId="1" pageNumber="62">
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<taxonomicName id="4C2B4D03FFC3C2783595FEB2FEACFEE4" ID-CoL="86ed675f-2aef-4ded-8319-e4f154ab923d" box="[151,374,295,321]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="62" phylum="Arthropoda" rank="species" species="formosae" status="sp. nov.">
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<emphasis id="B95FEA92FFC3C2783595FEB2FEACFEE4" bold="true" box="[151,374,295,321]" italics="true" pageId="1" pageNumber="62">Lithodes formosae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26C57E9FFC3C278347FFEB2FE01FEE4" box="[381,475,295,321]" pageId="1" pageNumber="62" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B943680FFC3C2783595FEDFFECDFEC1" blockId="1.[151,475,295,356]" box="[151,279,330,356]" pageId="1" pageNumber="62">
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(
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<figureCitation id="13102A05FFC3C278359DFEDFFED5FEC1" box="[159,271,330,356]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[151,255,1957,1981]" captionStartId-1="3.[151,255,1650,1674]" captionStartId-2="4.[151,255,1645,1669]" captionStartId-3="5.[151,255,1973,1997]" captionTargetBox-0="[151,1436,197,1934]" captionTargetBox-1="[151,1436,194,1628]" captionTargetBox-2="[151,1436,182,1619]" captionTargetBox-3="[151,1434,1093,1948]" captionTargetId-0="figure@2.[151,1436,197,1934]" captionTargetId-1="figure@3.[151,1436,194,1628]" captionTargetId-2="figure@4.[151,1436,182,1621]" captionTargetId-3="figure@5.[151,1436,1093,1949]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 1. Lithodes formosae sp. nov., ovigerous female holotype (cl 78.4 mm, pcl 55.9 mm, cw 54.7 mm), Dasi, Ilan County, NE Taiwan. A, dorsal habitus. B, cephalothorax, right lateral view." captionText-1="FIGURE 2. Lithodes formosae sp. nov., ovigerous female holotype (cl 78.4 mm, pcl 55.9 mm, cw 54.7 mm), Dasi, Ilan County, NE Taiwan. A, carapace. B, right pereopod 4. C, right pereopod 4 merus and carpus. D, right pereopod 4 dactylus." captionText-2="FIGURE 3. Lithodes formosae sp. nov., ovigerous female holotype (cl 78.4 mm, pcl 55.9 mm, cw 54.7 mm), Dasi, Ilan County, NE Taiwan. A, right chela. B, left chela. C, anterior cephalothorax, dorsal view. D, anterior cephalothorax including antenna, right oblique view. E, posterior carapace and abdomen. F, abdomen." captionText-3="FIGURE 4. Lithodes formosae sp. nov., ovigerous female holotype (cl 78.4 mm, pcl 55.9 mm, cw 54.7 mm), Dasi, Ilan County, NE Taiwan. Colour in life." httpUri-0="https://zenodo.org/record/275487/files/figure.png" httpUri-1="https://zenodo.org/record/275488/files/figure.png" httpUri-2="https://zenodo.org/record/275489/files/figure.png" httpUri-3="https://zenodo.org/record/275490/files/figure.png" pageId="1" pageNumber="62">Figs. 1–4</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C331650BFFC3C2783595FE00FD62FE09" box="[151,696,405,428]" pageId="1" pageNumber="62" type="description">
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<paragraph id="8B943680FFC3C2783595FE00FD62FE09" blockId="1.[151,696,405,428]" box="[151,696,405,428]" pageId="1" pageNumber="62">
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<treatmentCitationGroup id="AB3B11AEFFC3C2783595FE00FD62FE09" box="[151,696,405,428]" pageId="1" pageNumber="62">
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<taxonomicName id="4C2B4D03FFC3C2783595FE00FF29FE09" ID-CoL="3b7554cf-217a-4b1b-a13e-6125ec346ed4" box="[151,243,405,428]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="62" phylum="Arthropoda" rank="genus">
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<emphasis id="B95FEA92FFC3C2783595FE00FF29FE09" box="[151,243,405,428]" italics="true" pageId="1" pageNumber="62">Lithodes</emphasis>
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</taxonomicName>
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sp. —
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<treatmentCitation id="0A8A1091FFC3C2783442FE00FD50FE09" ID-DOI="http://doi.org/10.5281/zenodo.3502362" ID-Zenodo-Dep="3502362" author="Macpherson" box="[320,650,405,428]" httpUri="http://treatment.plazi.org/id/437B87C4991CFFDECCF63A70FEEAFE17" page="47" pageId="1" pageNumber="62" year="2008">
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<bibRefCitation id="EFBA4B71FFC3C2783442FE00FDBEFE09" author="Macpherson" box="[320,612,405,428]" pageId="1" pageNumber="68" refString="Macpherson, E. & Chan, T. Y. (2008) Some lithodid crabs (Crustacea: Decapoda: Lithodidae) from Taiwan and adjacent waters, with the description of one new species from Guam. Zootaxa, 1924, 43 - 52." type="journal article" year="2008">Macpherson & Chan, 2008</bibRefCitation>
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: 47
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</treatmentCitation>
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–48.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C331650BFFC3C2783595FE4FFB35FDBF" pageId="1" pageNumber="62" type="materials_examined">
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<paragraph id="8B943680FFC3C2783595FE4FFB35FDBF" blockId="1.[151,1437,474,1956]" pageId="1" pageNumber="62">
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<emphasis id="B95FEA92FFC3C2783595FE4FFE90FE51" bold="true" box="[151,330,474,500]" pageId="1" pageNumber="62">
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<typeStatus id="54908822FFC3C2783595FE4FFF08FE51" box="[151,210,474,500]" pageId="1" pageNumber="62">Type</typeStatus>
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material.
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</emphasis>
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<collectionCode id="ED3AAE45FFC3C2783451FE4FFE7EFE51" box="[339,420,474,500]" httpUri="http://grbio.org/cool/be15-j6k8" name="Institute of Marine Biology, National Taiwan Ocean University" pageId="1" pageNumber="62">NTOU</collectionCode>
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A01090, ovigerous female
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<typeStatus id="54908822FFC3C27837E5FE4FFC97FE51" box="[743,845,474,500]" pageId="1" pageNumber="62" type="holotype">holotype</typeStatus>
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(cl
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<quantity id="4CD39B65FFC3C278367BFE4FFC3EFE51" box="[889,996,474,500]" metricMagnitude="-2" metricUnit="m" metricValue="7.84" pageId="1" pageNumber="62" unit="mm" value="78.4">78.4 mm</quantity>
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, pcl
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<quantity id="4CD39B65FFC3C278311BFE4FFB5EFE51" box="[1049,1156,474,500]" metricMagnitude="-2" metricUnit="m" metricValue="5.59" pageId="1" pageNumber="62" unit="mm" value="55.9">55.9 mm</quantity>
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, cw
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<quantity id="4CD39B65FFC3C27831B5FE4FFAF9FE51" box="[1207,1315,474,500]" metricMagnitude="-2" metricUnit="m" metricValue="5.47" pageId="1" pageNumber="62" unit="mm" value="54.7">54.7 mm</quantity>
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), off Dasi, Ilan County, northeastern
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<collectingCountry id="F33C7610FFC3C27834C4FD95FDC5FDBF" box="[454,543,512,538]" name="Taiwan" pageId="1" pageNumber="62">Taiwan</collectingCountry>
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, commercial trawl,
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<quantity id="4CD39B65FFC3C2783605FD95FC55FDBF" box="[775,911,512,538]" metricMagnitude="2" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="1" pageNumber="62" unit="m" value="550.0" valueMax="600.0" valueMin="500.0">500–600 m</quantity>
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, coll. T.
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<collectingCountry id="F33C7610FFC3C27836EEFD95FBD8FDBF" box="[1004,1026,512,538]" name="Italy" pageId="1" pageNumber="62">Y</collectingCountry>
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. Chan,
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<date id="FF951040FFC3C278315BFD95FB31FDBF" box="[1113,1259,512,538]" pageId="1" pageNumber="62" value="1998-06-06">6 June 1998</date>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C331650BFFC3C27835C4FDB0FE83FC5A" pageId="1" pageNumber="62" type="diagnosis">
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<paragraph id="8B943680FFC3C27835C4FDB0FE83FC5A" blockId="1.[151,1437,474,1956]" pageId="1" pageNumber="62">
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<emphasis id="B95FEA92FFC3C27835C4FDB0FE9CFD9A" bold="true" box="[198,326,549,575]" pageId="1" pageNumber="62">Diagnosis.</emphasis>
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Carapace dorsal surface with long, slender spines; gastric region with 4 upright spines, anterior pair longer than half length of hepatic spine, posterior pair half length of hepatic spine; cardiac and intestinal regions each with 2 long upright spines; branchial surface with 2 long upright spines. Branchial margins with 2 primary, laterodorsally directed spines positioned at a level between pereopods 2 and 3, and between pereopods 3 and 4, respectively (anterior spine 1/3 length of hepatic spine, posterior spine 1/6 length of hepatic spine); margin between posterior primary branchial spine and intestinal spine with 5 protrusions of which 2 or 3 are developed as short spines. Adult rostrum shorter than half pcl; proximal half strongly upraised by about 45°. Abdominal somite 2 comprising 3 separate plates in adults. Antennal peduncle segment 2 with slender outer spine reaching midlength of segment 4. Chelipeds unequal; palms generally smooth, with low tubercles on dorsal surface. Ambulatory legs spinose; surface between major spines smooth or with very few, scattered spines. Merus about as long as pcl, length less than 7 times height. Propodus length 11–12 times height. Dactylus half length of propodus; flexor margin with 6 or 7 minute, widely spaced corneous spines.
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</paragraph>
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</subSubSection>
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<subSubSection id="C331650BFFC3C27C35C4FB9FFC05FBA9" lastPageId="5" lastPageNumber="66" pageId="1" pageNumber="62" type="description">
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<paragraph id="8B943680FFC3C27835C4FB9FFC2DFA41" blockId="1.[151,1437,474,1956]" pageId="1" pageNumber="62">
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<emphasis id="B95FEA92FFC3C27835C4FB9FFD89FB81" bold="true" box="[198,595,1034,1060]" pageId="1" pageNumber="62">
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Description of female
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<typeStatus id="54908822FFC3C27834E2FB9FFD94FB81" box="[480,590,1034,1060]" pageId="1" pageNumber="62" type="holotype">holotype</typeStatus>
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.
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</emphasis>
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<emphasis id="B95FEA92FFC3C278375EFB9FFD00FB86" box="[604,730,1034,1059]" italics="true" pageId="1" pageNumber="62">Carapace:</emphasis>
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Pyriform, about as long as wide; regions indicated; dorsal surface armed with long, slender spines and scattered, widely separated granules. Hepatic spine longest of carapace spines (0.4 pcl), directed anterolaterally. Gastric region convex, with 2 pairs of long, upright spines, anterior pair longer than half length of hepatic spine, posterior pair half length of hepatic spine. Cardiac region and intestinal regions each with pair of long upright spines, as long as anterior gastric spines. Branchial surface with 2 long upright spines: 1 spine at level of pereopod 3 coxa (almost as long as hepatic spine); 1 spine at level of pereopod 4 coxa (slightly longer than anterior gastric spines), without conical tubercle at base of posterior spine. Branchial margins with 2 primary, laterodorsally directed spines positioned at a level between pereopods 2 and 3, and between pereopods 3 and 4, respectively (anterior spine 1/3 length of hepatic spine, posterior spine 1/6 length of hepatic spine); with 2 low tubercles anterior to first primary branchial spine, and 2 low tubercles between primary branchial spines; margin between posterior primary branchial spine and intestinal spine with 5 protrusions of which 2 or 3 are developed as short spines. Pterygostomian region granulate or tuberculate, with small, anterior, submarginal granule.
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</paragraph>
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<paragraph id="8B943680FFC3C27835C4FA65FF37F961" blockId="1.[151,1437,474,1956]" pageId="1" pageNumber="62">
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<emphasis id="B95FEA92FFC3C27835C4FA65FE7AF9AC" box="[198,416,1520,1545]" italics="true" pageId="1" pageNumber="62">Rostrum and orbit</emphasis>
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: Rostrum 0.4 pcl, comprising proximal and distal portions; proximal portion dorsally angled by about 45°, with pair of divergent dorsal spines reaching to midlength of distal portion of rostrum; distal portion subhorizontal, distally bifurcate for half length, forming pair of divergent spines; basal subrostral spine elongate, extending anteriorly beyond apices of outer orbital spine. Posterior orbital margin concave; outer orbital spine directed anteriorly, slightly shorter than anterolateral spine, reaching to end of cornea.
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</paragraph>
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<paragraph id="8B943680FFC3C27835C4F945FD26F94F" blockId="1.[151,1437,474,1956]" box="[198,764,1744,1770]" pageId="1" pageNumber="62">
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<emphasis id="B95FEA92FFC3C27835C4F945FE51F94C" box="[198,395,1744,1769]" italics="true" pageId="1" pageNumber="62">Ocular peduncle</emphasis>
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: Longer than cornea; unarmed.
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</paragraph>
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<paragraph id="8B943680FFC3C27835C4F960FE2EF891" blockId="1.[151,1437,474,1956]" pageId="1" pageNumber="62">
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<emphasis id="B95FEA92FFC3C27835C4F960FEE7F8AB" box="[198,317,1781,1806]" italics="true" pageId="1" pageNumber="62">Antennule</emphasis>
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: Peduncle unarmed, reaching anteriorly beyond antennal peduncle by about 2/3 length of distal antennular peduncle segment.
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</paragraph>
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<paragraph id="8B943680FFC3C27835C4F8D5FDA3F801" blockId="1.[151,1437,474,1956]" pageId="1" pageNumber="62">
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<emphasis id="B95FEA92FFC3C27835C4F8D5FEF2F8FC" box="[198,296,1856,1881]" italics="true" pageId="1" pageNumber="62">Antenna</emphasis>
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: Basal segment unarmed; outer margin of segment 2 with slender spine, reaching midlength of segment 4; segment 3 unarmed; scaphocerite minute, obsolete, blunt, shorter than segment 4; segment 4 unarmed, about half length of segment 5.
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</paragraph>
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<caption id="DF546608FFC0C27B3595F830FC62F878" httpUri="https://zenodo.org/record/275487/files/figure.png" pageId="2" pageNumber="63" targetBox="[151,1436,197,1934]" targetPageId="2">
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<paragraph id="8B943680FFC0C27B3595F830FC62F878" blockId="2.[151,1436,1957,2013]" pageId="2" pageNumber="63">
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<emphasis id="B95FEA92FFC0C27B3595F830FEC3F818" bold="true" box="[151,281,1957,1981]" pageId="2" pageNumber="63">FIGURE 1.</emphasis>
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<taxonomicName id="4C2B4D03FFC0C27B3422F833FE3CF818" box="[288,486,1958,1981]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="formosae" status="sp. nov.">
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<emphasis id="B95FEA92FFC0C27B3422F833FE3CF818" box="[288,486,1958,1981]" italics="true" pageId="2" pageNumber="63">Lithodes formosae</emphasis>
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</taxonomicName>
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<emphasis id="B95FEA92FFC0C27B34EFF830FD99F818" bold="true" box="[493,579,1957,1981]" pageId="2" pageNumber="63">
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<taxonomicNameLabel id="A26C57E9FFC0C27B34EFF830FD99F818" box="[493,579,1957,1981]" pageId="2" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, ovigerous female holotype (cl 78.4 mm, pcl 55.9 mm, cw 54.7 mm), Dasi, Ilan County, NE Taiwan. A, dorsal habitus. B, cephalothorax, right lateral view.
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</paragraph>
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</caption>
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<caption id="DF546608FFC1C27A3595F9E7FF2DF96C" httpUri="https://zenodo.org/record/275488/files/figure.png" pageId="3" pageNumber="64" targetBox="[151,1436,194,1628]" targetPageId="3">
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||
<paragraph id="8B943680FFC1C27A3595F9E7FF2DF96C" blockId="3.[151,1437,1650,1737]" pageId="3" pageNumber="64">
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<emphasis id="B95FEA92FFC1C27A3595F9E7FEC3F92F" bold="true" box="[151,281,1650,1674]" pageId="3" pageNumber="64">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C2B4D03FFC1C27A3422F9E7FE3CF92C" box="[288,486,1650,1673]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="64" phylum="Arthropoda" rank="species" species="formosae" status="sp. nov.">
|
||
<emphasis id="B95FEA92FFC1C27A3422F9E7FE3CF92C" box="[288,486,1650,1673]" italics="true" pageId="3" pageNumber="64">Lithodes formosae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95FEA92FFC1C27A34EFF9E7FD99F92F" bold="true" box="[493,579,1650,1674]" pageId="3" pageNumber="64">
|
||
<taxonomicNameLabel id="A26C57E9FFC1C27A34EFF9E7FD99F92F" box="[493,579,1650,1674]" pageId="3" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, ovigerous female holotype (cl 78.4 mm, pcl 55.9 mm, cw 54.7 mm), Dasi, Ilan County, NE Taiwan. A, carapace. B, right pereopod 4. C, right pereopod 4 merus and carpus. D, right pereopod 4 dactylus.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B943680FFC1C27A35C4F88DFC1BF849" blockId="3.[151,1436,1815,2028]" pageId="3" pageNumber="64">
|
||
<emphasis id="B95FEA92FFC1C27A35C4F88DFEEFF894" box="[198,309,1816,1841]" italics="true" pageId="3" pageNumber="64">Abdomen</emphasis>
|
||
: Somite 1 with low blunt triangular teeth on posterior margin. Somite 2 rugose; composed of 3 distinct plates separated by narrow suture; median surface with pair of low submedian teeth; posterior margin with 4 low prominences; lateral margins of lateral plates irregularly dentate. Somites 3–5 with nodular median plates; surface of lateral plates pitted, irregular, rugose; marginal plates with left margin irregularly crenulate (fused with laterals), right marginals subdivided, with angular to pointed apices. Somite 6 longer than wide, distal margin with 2 small distal tubercles. Telson rounded, unarmed.
|
||
</paragraph>
|
||
<caption id="DF546608FFC6C27D3595F9F8FBE2F96E" httpUri="https://zenodo.org/record/275489/files/figure.png" pageId="4" pageNumber="65" targetBox="[151,1436,182,1619]" targetPageId="4">
|
||
<paragraph id="8B943680FFC6C27D3595F9F8FBE2F96E" blockId="4.[151,1436,1645,1739]" pageId="4" pageNumber="65">
|
||
<emphasis id="B95FEA92FFC6C27D3595F9F8FEC3F920" bold="true" box="[151,281,1645,1669]" pageId="4" pageNumber="65">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C2B4D03FFC6C27D3422F9F8FE3CF921" box="[288,486,1645,1668]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="formosae" status="sp. nov.">
|
||
<emphasis id="B95FEA92FFC6C27D3422F9F8FE3CF921" box="[288,486,1645,1668]" italics="true" pageId="4" pageNumber="65">Lithodes formosae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95FEA92FFC6C27D34EFF9F8FD99F920" bold="true" box="[493,579,1645,1669]" pageId="4" pageNumber="65">
|
||
<taxonomicNameLabel id="A26C57E9FFC6C27D34EFF9F8FD99F920" box="[493,579,1645,1669]" pageId="4" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, ovigerous female holotype (cl 78.4 mm, pcl 55.9 mm, cw 54.7 mm), Dasi, Ilan County, NE Taiwan. A, right chela. B, left chela. C, anterior cephalothorax, dorsal view. D, anterior cephalothorax including antenna, right oblique view. E, posterior carapace and abdomen. F, abdomen.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B943680FFC6C27D35C4F883FCFAF84F" blockId="4.[151,1436,1813,2026]" pageId="4" pageNumber="65">
|
||
<emphasis id="B95FEA92FFC6C27D35C4F883FEE1F88A" box="[198,315,1814,1839]" italics="true" pageId="4" pageNumber="65">Chelipeds</emphasis>
|
||
: Slightly dimorphic, unequal; spination similar. Surfaces of both chelipeds smooth or with scattered granules in addition to major spines. Coxa with tufts of setae, unarmed; ischiobasis with 3 or 4 stout ventral spines and about 6 blunt tubercles. Merus inner margin, with stout subdistal spine and 2 or 3 smaller spines or acute tubercles; ventral margin with 2 or 3 low conical spines; dorsal and lateral surface spinose, spines largest distally. Carpus with 2 or 3 short, prominent spines on dorsal and lateral surfaces. Palms generally smooth, with low tubercles on dorsal surface.
|
||
</paragraph>
|
||
<paragraph id="8B943680FFC7C27C35C4FF0DFB42FF59" blockId="5.[151,1437,152,1036]" pageId="5" pageNumber="66">Major cheliped 1.50 pcl; upper palm length 1.76 times height; occlusal margins of fingers corneous for distal third, proximally with 3 calcareous nodules; dactylus dorsal margin broadly convex, with rows of tufts of golden setae and blunt proximal tooth, 1.17 times longer than dorsal margin of palm.</paragraph>
|
||
<paragraph id="8B943680FFC7C27C35C4FE9DFCB1FEC9" blockId="5.[151,1437,152,1036]" pageId="5" pageNumber="66">Minor cheliped 1.51 pcl; upper palm length 2.04 times height; occlusal margin corneous for distal quarter, proximally crenulate; dactylus dorsal margin broadly convex, with rows of tufts of golden setae and blunt proximal tooth, 1.41 times longer than dorsal margin of palm.</paragraph>
|
||
<paragraph id="8B943680FFC7C27C35C4FEEDFDB2FCA2" blockId="5.[151,1437,152,1036]" pageId="5" pageNumber="66">
|
||
<emphasis id="B95FEA92FFC7C27C35C4FEEDFE53FE34" box="[198,393,376,401]" italics="true" pageId="5" pageNumber="66">Ambulatory legs</emphasis>
|
||
: Similar, slender, elongate; segments spinose, surface between major spines smooth or with very few, scattered spines. Pereopod 4 longest. Distal margins of coxae unarmed; surface smooth. Ischiobasis with 2 non-setose distal spines. Merus subcircular to ovate in cross section; extensor margin with 3 or 4 larger spines intervened by 1 or 2 small spines in addition to long, prominent, distal spine; distal spine directed oblique to meral axis; flexor margin with 2 rows of 4 or 5 small spines; upper surface with irregular row of 6 or 7 small spines. Carpus about half merus length; extensor margin with distal and second proximal spines longest, exceeding half carpus length; surface with scattered small spines. Propodus compressed; with 2 rows of 5–8 small spines on extensor margin and 2–4 small spines on dorsal surface; flexor margin with 6 or 7 small spines. Dactylus distally curved, rounded in cross section distally; with 4 corneous spines proximally; extensor margin with small corneous spine at midlength; flexor margin with about 6 or 7 minute, widely spaced corneous spines; apex corneous.
|
||
</paragraph>
|
||
<paragraph id="8B943680FFC7C27C35C4FC87FBC4FCF7" blockId="5.[151,1437,152,1036]" pageId="5" pageNumber="66">Pereopod 2 length 2.64 pcl. Merus 0.95 pcl; length: height ratio 6.40. Carpus 0.52 merus length. Propodus 0.86 merus length; length: height ratio 11.04. Dactylus 0.50 propodus length.</paragraph>
|
||
<paragraph id="8B943680FFC7C27C35C4FCC8FBC4FC39" blockId="5.[151,1437,152,1036]" pageId="5" pageNumber="66">Pereopod 3 length 2.83 pcl. Merus 1.02 pcl; length: height ratio 6.85. Carpus 0.53 merus length. Propodus 0.88 merus length, length: height ratio 12.04. Dactylus 0.53 propodus length.</paragraph>
|
||
<paragraph id="8B943680FFC7C27C35C4FC3DFBC7FC42" blockId="5.[151,1437,152,1036]" pageId="5" pageNumber="66">Pereopod 4 length 2.91 pcl. Merus 0.99 pcl; length: height ratio 6.90. Carpus 0.58 merus length. Propodus 0.94 merus length, length: height ratio 11.64. Dactylus 0.48 propodus length.</paragraph>
|
||
<paragraph id="8B943680FFC7C27C35C4FC67FC05FBA9" blockId="5.[151,1437,152,1036]" box="[198,991,1010,1036]" pageId="5" pageNumber="66">
|
||
<emphasis id="B95FEA92FFC7C27C35C4FC67FEA9FBA9" bold="true" box="[198,371,1010,1036]" pageId="5" pageNumber="66">Colour in life.</emphasis>
|
||
(
|
||
<figureCitation id="13102A05FFC7C27C3486FC67FE10FBA9" box="[388,458,1010,1036]" captionStart="FIGURE 4" captionStartId="5.[151,255,1973,1997]" captionTargetBox="[151,1434,1093,1948]" captionTargetId="figure@5.[151,1436,1093,1949]" captionTargetPageId="5" captionText="FIGURE 4. Lithodes formosae sp. nov., ovigerous female holotype (cl 78.4 mm, pcl 55.9 mm, cw 54.7 mm), Dasi, Ilan County, NE Taiwan. Colour in life." httpUri="https://zenodo.org/record/275490/files/figure.png" pageId="5" pageNumber="66">Fig. 4</figureCitation>
|
||
) Body red. Cornea dark brown. Eggs orange.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF546608FFC7C27C3595F820FDD4F849" httpUri="https://zenodo.org/record/275490/files/figure.png" pageId="5" pageNumber="66" targetBox="[151,1434,1093,1948]" targetPageId="5">
|
||
<paragraph id="8B943680FFC7C27C3595F820FDD4F849" blockId="5.[151,1436,1973,2028]" pageId="5" pageNumber="66">
|
||
<emphasis id="B95FEA92FFC7C27C3595F820FEC3F868" bold="true" box="[151,281,1973,1997]" pageId="5" pageNumber="66">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C2B4D03FFC7C27C3423F820FE3CF869" box="[289,486,1973,1996]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="formosae" status="sp. nov.">
|
||
<emphasis id="B95FEA92FFC7C27C3423F820FE3CF869" box="[289,486,1973,1996]" italics="true" pageId="5" pageNumber="66">Lithodes formosae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26C57E9FFC7C27C34ECF820FD99F869" box="[494,579,1973,1996]" pageId="5" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, ovigerous female holotype (cl 78.4 mm, pcl 55.9 mm, cw 54.7 mm), Dasi, Ilan County, NE Taiwan. Colour in life.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C331650BFFC4C27F35C4FF02FEC4FF72" pageId="6" pageNumber="67" type="etymology">
|
||
<paragraph id="8B943680FFC4C27F35C4FF02FEC4FF72" blockId="6.[151,1437,151,1746]" pageId="6" pageNumber="67">
|
||
<emphasis id="B95FEA92FFC4C27F35C4FF02FE89FF14" bold="true" box="[198,339,151,177]" pageId="6" pageNumber="67">Etymology.</emphasis>
|
||
This new species is named after a former name for
|
||
<collectingCountry id="F33C7610FFC4C27F36B9FF0DFBCBFF17" box="[955,1041,152,178]" name="Taiwan" pageId="6" pageNumber="67">Taiwan</collectingCountry>
|
||
(“
|
||
<collectingCountry id="F33C7610FFC4C27F312EFF0DFB47FF17" box="[1068,1181,152,178]" name="Taiwan" pageId="6" pageNumber="67">Formosa</collectingCountry>
|
||
”), the
|
||
<typeStatus id="54908822FFC4C27F31EBFF0DFAC1FF17" box="[1257,1307,152,178]" pageId="6" pageNumber="67">type</typeStatus>
|
||
locality of the species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C331650BFFC4C27F35C4FF77FE51FA27" pageId="6" pageNumber="67" type="discussion">
|
||
<paragraph id="8B943680FFC4C27F35C4FF77FE51FA27" blockId="6.[151,1437,151,1746]" pageId="6" pageNumber="67">
|
||
<emphasis id="B95FEA92FFC4C27F35C4FF77FEE5FF59" bold="true" box="[198,319,226,252]" pageId="6" pageNumber="67">Remarks.</emphasis>
|
||
<bibRefCitation id="EFBA4B71FFC4C27F344EFF77FDE3FF59" author="Macpherson" box="[332,569,226,252]" pageId="6" pageNumber="68" refString="Macpherson, E. (1988) Revision of the family Lithodidae Samouelle, 1819 (Crustacea, Decapoda, Anomura) in the Atlantic Ocean. Monografias de Zoologia Marina, 2, 9 - 153." type="journal article" year="1988">Macpherson (1988)</bibRefCitation>
|
||
listed 15 species of
|
||
<taxonomicName id="4C2B4D03FFC4C27F3640FF77FC7DFF5E" box="[834,935,226,251]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FEA92FFC4C27F3640FF77FC7DFF5E" box="[834,935,226,251]" italics="true" pageId="6" pageNumber="67">Lithodes</emphasis>
|
||
</taxonomicName>
|
||
worldwide of which seven occur in the Atlantic Ocean and eight in the Indo-Pacific. Since 1988, three species of
|
||
<taxonomicName id="4C2B4D03FFC4C27F310CFE9DFBAEFE84" box="[1038,1140,264,289]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FEA92FFC4C27F310CFE9DFBAEFE84" box="[1038,1140,264,289]" italics="true" pageId="6" pageNumber="67">Lithodes</emphasis>
|
||
</taxonomicName>
|
||
have been described, all from the Pacific Ocean:
|
||
<taxonomicName id="4C2B4D03FFC4C27F34CAFEB8FCB7FEE2" authority="Macpherson, 1991" authorityName="Macpherson" authorityYear="1991" box="[456,877,301,327]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="megacantha">
|
||
<emphasis id="B95FEA92FFC4C27F34CAFEB8FD5AFEE3" box="[456,640,301,326]" italics="true" pageId="6" pageNumber="67">L. megacantha</emphasis>
|
||
<bibRefCitation id="EFBA4B71FFC4C27F378AFEB8FCB7FEE2" author="Macpherson" box="[648,877,301,327]" pageId="6" pageNumber="68" refString="Macpherson, E. (1991) A new species of the genus Lithodes (Crustacea, Decapoda, Lithodidae) from the French Polynesia. Bulletin du Museum national d'Histoire naturelle (Paris), 4 (13), A (1 - 2), 153 - 158." type="journal article" year="1991">Macpherson, 1991</bibRefCitation>
|
||
</taxonomicName>
|
||
[
|
||
<typeStatus id="54908822FFC4C27F367CFEB8FC6CFEE2" box="[894,950,301,327]" pageId="6" pageNumber="67">type</typeStatus>
|
||
locality:
|
||
<collectingCountry id="F33C7610FFC4C27F312FFEB8FAD0FEE2" box="[1069,1290,301,327]" name="French Polynesia" pageId="6" pageNumber="67">French Polynesia</collectingCountry>
|
||
],
|
||
<taxonomicName id="4C2B4D03FFC4C27F301CFEB8FE04FEC9" authority="Macpherson & Chan, 2008" authorityName="Macpherson & Chan" authorityYear="2008" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="paulayi">
|
||
<emphasis id="B95FEA92FFC4C27F301CFEB8FA47FEE3" box="[1310,1437,301,326]" italics="true" pageId="6" pageNumber="67">L. paulayi</emphasis>
|
||
<bibRefCitation id="EFBA4B71FFC4C27F3595FEC7FE04FEC9" author="Macpherson" box="[151,478,338,364]" pageId="6" pageNumber="68" refString="Macpherson, E. & Chan, T. Y. (2008) Some lithodid crabs (Crustacea: Decapoda: Lithodidae) from Taiwan and adjacent waters, with the description of one new species from Guam. Zootaxa, 1924, 43 - 52." type="journal article" year="2008">Macpherson & Chan, 2008</bibRefCitation>
|
||
</taxonomicName>
|
||
[
|
||
<typeStatus id="54908822FFC4C27F34ECFEC7FDFFFEC9" box="[494,549,338,364]" pageId="6" pageNumber="67">type</typeStatus>
|
||
locality:
|
||
<collectingCountry id="F33C7610FFC4C27F379AFEC7FD3DFEC9" box="[664,743,338,364]" name="Guam" pageId="6" pageNumber="67">Guam</collectingCountry>
|
||
], and
|
||
<taxonomicName id="4C2B4D03FFC4C27F362CFEC7FB31FEC9" authority="Hall & Thatje, 2009" authorityName="Hall & Thatje" authorityYear="2009" box="[814,1259,338,364]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="galapagensis">
|
||
<emphasis id="B95FEA92FFC4C27F362CFEC7FC37FECE" box="[814,1005,338,363]" italics="true" pageId="6" pageNumber="67">L. galapagensis</emphasis>
|
||
<bibRefCitation id="EFBA4B71FFC4C27F36F4FEC7FB31FEC9" author="Hall" box="[1014,1259,338,364]" pageId="6" pageNumber="68" refString="Hall, S. & Thatje, S. (2009) Four new species of the family Lithodidae (Decapoda: Anomura) from the collections of the National Museum of Natural History, Smithsonian Institution. Zootaxa, 2302, 31 - 47." type="journal article" year="2009">Hall & Thatje, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
[
|
||
<typeStatus id="54908822FFC4C27F31F8FEC7FAEBFEC9" box="[1274,1329,338,364]" pageId="6" pageNumber="67">type</typeStatus>
|
||
locality: Galapagos Islands].
|
||
<taxonomicName id="4C2B4D03FFC4C27F348EFEEDFDB0FE34" box="[396,618,376,401]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="formosae" status="sp. nov.">
|
||
<emphasis id="B95FEA92FFC4C27F348EFEEDFDB0FE34" box="[396,618,376,401]" italics="true" pageId="6" pageNumber="67">Lithodes formosae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95FEA92FFC4C27F3770FEE2FD08FE34" bold="true" box="[626,722,375,401]" pageId="6" pageNumber="67">
|
||
<taxonomicNameLabel id="A26C57E9FFC4C27F3770FEE2FD08FE34" box="[626,722,375,401]" pageId="6" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is the nineteenth known species of
|
||
<taxonomicName id="4C2B4D03FFC4C27F3180FEEDFB33FE34" box="[1154,1257,376,401]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95FEA92FFC4C27F3180FEEDFB33FE34" box="[1154,1257,376,401]" italics="true" pageId="6" pageNumber="67">Lithodes</emphasis>
|
||
</taxonomicName>
|
||
and resembles other long-spined species of the genus in which the carapace and walking legs bear long spines even as early adults —
|
||
<taxonomicName id="4C2B4D03FFC4C27F3408FE57FDE1FE79" authority="Sakai, 1971" authorityName="Sakai" authorityYear="1971" box="[266,571,450,476]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="longispina">
|
||
<emphasis id="B95FEA92FFC4C27F3408FE57FE7FFE7E" box="[266,421,450,475]" italics="true" pageId="6" pageNumber="67">L. longispina</emphasis>
|
||
<bibRefCitation id="EFBA4B71FFC4C27F34AFFE57FDE1FE79" author="Sakai" box="[429,571,450,476]" pageId="6" pageNumber="68" refString="Sakai, T. (1971) Illustrations of 15 species of crabs of the family Lithodidae, two of which are new to science. Researches on Crustacea, 4 - 5, 1 - 49." type="journal article" year="1971">Sakai, 1971</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2B4D03FFC4C27F374AFE57FD2DFE7E" box="[584,759,450,475]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="megacantha">
|
||
<emphasis id="B95FEA92FFC4C27F374AFE57FD2DFE7E" box="[584,759,450,475]" italics="true" pageId="6" pageNumber="67">L. megacantha</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C2B4D03FFC4C27F363BFE57FC6AFE7E" box="[825,944,450,475]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="paulayi">
|
||
<emphasis id="B95FEA92FFC4C27F363BFE57FC6AFE7E" box="[825,944,450,475]" italics="true" pageId="6" pageNumber="67">L. paulayi</emphasis>
|
||
</taxonomicName>
|
||
. The new species is readily distinguished from
|
||
<taxonomicName id="4C2B4D03FFC4C27F35D5FE7DFEA9FDA4" box="[215,371,488,513]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="longispina">
|
||
<emphasis id="B95FEA92FFC4C27F35D5FE7DFEA9FDA4" box="[215,371,488,513]" italics="true" pageId="6" pageNumber="67">L. longispina</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2B4D03FFC4C27F34AEFE7DFD81FDA4" box="[428,603,488,513]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="megacantha">
|
||
<emphasis id="B95FEA92FFC4C27F34AEFE7DFD81FDA4" box="[428,603,488,513]" italics="true" pageId="6" pageNumber="67">L. megacantha</emphasis>
|
||
</taxonomicName>
|
||
by having two dorsal branchial spines instead of one; the first primary marginal branchial spine is considerably shorter than, rather than subequal to the hepatic spine; the second primary marginal branchial spine is considerably shorter than the first, rather than subequal; and the distolateral spine of the basal antennal segment reaches the midlength rather than end of the second segment. The new species is most similar to
|
||
<taxonomicName id="4C2B4D03FFC4C27F3735FDE8FD75FD33" box="[567,687,637,662]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="paulayi">
|
||
<emphasis id="B95FEA92FFC4C27F3735FDE8FD75FD33" box="[567,687,637,662]" italics="true" pageId="6" pageNumber="67">L. paulayi</emphasis>
|
||
</taxonomicName>
|
||
in sharing two dorsal branchial carapace spines, proportionally short primary marginal branchial spines, and similar basal antennal segment ornamentation. The two species differ in the number of marginal branchial spines between the posterior primary branchial spine and intestinal spines (5 protrusions of which 2 or 3 are developed as short spines in
|
||
<taxonomicName id="4C2B4D03FFC4C27F36DBFD78FBBDFCA3" box="[985,1127,749,774]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="formosae" status="sp. nov.">
|
||
<emphasis id="B95FEA92FFC4C27F36DBFD78FBBDFCA3" box="[985,1127,749,774]" italics="true" pageId="6" pageNumber="67">L. formosae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95FEA92FFC4C27F316DFD78FB14FCA2" bold="true" box="[1135,1230,749,775]" pageId="6" pageNumber="67">
|
||
<taxonomicNameLabel id="A26C57E9FFC4C27F316DFD78FB14FCA2" box="[1135,1230,749,775]" pageId="6" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
; 6–8 spines in
|
||
<taxonomicName id="4C2B4D03FFC4C27F3086FD78FF35FC8E" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="paulayi">
|
||
<emphasis id="B95FEA92FFC4C27F3086FD78FF35FC8E" italics="true" pageId="6" pageNumber="67">L. paulayi</emphasis>
|
||
</taxonomicName>
|
||
), proportional length of the rostrum and dorsal spines (markedly longer in
|
||
<taxonomicName id="4C2B4D03FFC4C27F316EFC87FB3EFC8E" box="[1132,1252,786,811]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="paulayi">
|
||
<emphasis id="B95FEA92FFC4C27F316EFC87FB3EFC8E" box="[1132,1252,786,811]" italics="true" pageId="6" pageNumber="67">L. paulayi</emphasis>
|
||
</taxonomicName>
|
||
), and spination of the chelipeds (a few dorsal tubercles on the palms in
|
||
<taxonomicName id="4C2B4D03FFC4C27F3658FCADFC2AFCF4" box="[858,1008,824,849]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="formosae" status="sp. nov.">
|
||
<emphasis id="B95FEA92FFC4C27F3658FCADFC2AFCF4" box="[858,1008,824,849]" italics="true" pageId="6" pageNumber="67">L. formosae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26C57E9FFC4C27F36FBFCADFB84FCF7" box="[1017,1118,824,850]" pageId="6" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; dorsal spines and lateral tubercles in
|
||
<taxonomicName id="4C2B4D03FFC4C27F3427FCC8FE46FCD3" box="[293,412,861,886]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="paulayi">
|
||
<emphasis id="B95FEA92FFC4C27F3427FCC8FE46FCD3" box="[293,412,861,886]" italics="true" pageId="6" pageNumber="67">L. paulayi</emphasis>
|
||
</taxonomicName>
|
||
). The rostrum of
|
||
<taxonomicName id="4C2B4D03FFC4C27F3768FCC8FD3AFCD3" box="[618,736,861,886]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="paulayi">
|
||
<emphasis id="B95FEA92FFC4C27F3768FCC8FD3AFCD3" box="[618,736,861,886]" italics="true" pageId="6" pageNumber="67">L. paulayi</emphasis>
|
||
</taxonomicName>
|
||
is about 0.8 pcl compared to 0.4 pcl in
|
||
<taxonomicName id="4C2B4D03FFC4C27F31AEFCC8FAE2FCD3" box="[1196,1336,861,886]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="formosae" status="sp. nov.">
|
||
<emphasis id="B95FEA92FFC4C27F31AEFCC8FAE2FCD3" box="[1196,1336,861,886]" italics="true" pageId="6" pageNumber="67">L. formosae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95FEA92FFC4C27F303DFCC8FA41FCD2" bold="true" box="[1343,1435,861,887]" pageId="6" pageNumber="67">
|
||
<taxonomicNameLabel id="A26C57E9FFC4C27F303DFCC8FA41FCD2" box="[1343,1435,861,887]" pageId="6" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Proportional spine and rostral length decreases with increasing size in lithodids, so the difference in relative rostral length between
|
||
<taxonomicName id="4C2B4D03FFC4C27F34A6FC3DFDEBFC64" box="[420,561,936,961]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="formosae" status="sp. nov.">
|
||
<emphasis id="B95FEA92FFC4C27F34A6FC3DFDEBFC64" box="[420,561,936,961]" italics="true" pageId="6" pageNumber="67">L. formosae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95FEA92FFC4C27F373BFC32FD4DFC64" bold="true" box="[569,663,935,961]" pageId="6" pageNumber="67">
|
||
<taxonomicNameLabel id="A26C57E9FFC4C27F373BFC32FD4DFC64" box="[569,663,935,961]" pageId="6" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C2B4D03FFC4C27F37D3FC3DFC92FC64" box="[721,840,936,961]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="paulayi">
|
||
<emphasis id="B95FEA92FFC4C27F37D3FC3DFC92FC64" box="[721,840,936,961]" italics="true" pageId="6" pageNumber="67">L. paulayi</emphasis>
|
||
</taxonomicName>
|
||
is all the more significant because the
|
||
<typeStatus id="54908822FFC4C27F3017FC3DFAA1FC67" box="[1301,1403,936,962]" pageId="6" pageNumber="67" type="holotype">holotype</typeStatus>
|
||
of the former is about half the size of that of the latter. The proportional dorsal spine length in the
|
||
<typeStatus id="54908822FFC4C27F31F4FC58FA86FC42" box="[1270,1372,973,999]" pageId="6" pageNumber="67" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C2B4D03FFC4C27F3086FC58FEDDFBAE" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="formosae" status="sp. nov.">
|
||
<emphasis id="B95FEA92FFC4C27F3086FC58FEDDFBAE" italics="true" pageId="6" pageNumber="67">L. formosae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95FEA92FFC4C27F340DFC67FEAAFBA9" bold="true" box="[271,368,1010,1036]" pageId="6" pageNumber="67">
|
||
<taxonomicNameLabel id="A26C57E9FFC4C27F340DFC67FEAAFBA9" box="[271,368,1010,1036]" pageId="6" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to that of the much larger
|
||
<typeStatus id="54908822FFC4C27F3633FC67FC41FBA9" box="[817,923,1010,1036]" pageId="6" pageNumber="67" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C2B4D03FFC4C27F36C6FC67FB9AFBAE" box="[964,1088,1010,1035]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="paulayi">
|
||
<emphasis id="B95FEA92FFC4C27F36C6FC67FB9AFBAE" box="[964,1088,1010,1035]" italics="true" pageId="6" pageNumber="67">L. paulayi</emphasis>
|
||
</taxonomicName>
|
||
. The
|
||
<typeStatus id="54908822FFC4C27F3185FC67FB35FBA9" box="[1159,1263,1010,1036]" pageId="6" pageNumber="67" type="paratype">paratype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C2B4D03FFC4C27F301BFC67FA4FFBAE" box="[1305,1429,1010,1035]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="paulayi">
|
||
<emphasis id="B95FEA92FFC4C27F301BFC67FA4FFBAE" box="[1305,1429,1010,1035]" italics="true" pageId="6" pageNumber="67">L. paulayi</emphasis>
|
||
</taxonomicName>
|
||
, however, is a similar size to the
|
||
<typeStatus id="54908822FFC4C27F370DFB8DFDAFFB97" box="[527,629,1048,1074]" pageId="6" pageNumber="67" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C2B4D03FFC4C27F379EFB8DFCF2FB94" box="[668,808,1048,1073]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="formosae" status="sp. nov.">
|
||
<emphasis id="B95FEA92FFC4C27F379EFB8DFCF2FB94" box="[668,808,1048,1073]" italics="true" pageId="6" pageNumber="67">L. formosae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95FEA92FFC4C27F362DFB82FC56FB94" bold="true" box="[815,908,1047,1073]" pageId="6" pageNumber="67">
|
||
<taxonomicNameLabel id="A26C57E9FFC4C27F362DFB82FC56FB94" box="[815,908,1047,1073]" pageId="6" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, but in the latter, the spines are less than half the proportional length as those of the former (compare
|
||
<figureCitation id="13102A05FFC4C27F362CFBA8FCA2FBF2" box="[814,888,1085,1111]" captionStart="FIGURE 1" captionStartId="2.[151,255,1957,1981]" captionTargetBox="[151,1436,197,1934]" captionTargetId="figure@2.[151,1436,197,1934]" captionTargetPageId="2" captionText="FIGURE 1. Lithodes formosae sp. nov., ovigerous female holotype (cl 78.4 mm, pcl 55.9 mm, cw 54.7 mm), Dasi, Ilan County, NE Taiwan. A, dorsal habitus. B, cephalothorax, right lateral view." httpUri="https://zenodo.org/record/275487/files/figure.png" pageId="6" pageNumber="67">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="13102A05FFC4C27F3685FBA8FC4FFBF2" box="[903,917,1085,1111]" captionStart="FIGURE 2" captionStartId="3.[151,255,1650,1674]" captionTargetBox="[151,1436,194,1628]" captionTargetId="figure@3.[151,1436,194,1628]" captionTargetPageId="3" captionText="FIGURE 2. Lithodes formosae sp. nov., ovigerous female holotype (cl 78.4 mm, pcl 55.9 mm, cw 54.7 mm), Dasi, Ilan County, NE Taiwan. A, carapace. B, right pereopod 4. C, right pereopod 4 merus and carpus. D, right pereopod 4 dactylus." httpUri="https://zenodo.org/record/275488/files/figure.png" pageId="6" pageNumber="67">2</figureCitation>
|
||
with
|
||
<bibRefCitation id="EFBA4B71FFC4C27F36D8FBA8FACCFBF2" author="Macpherson" box="[986,1302,1085,1111]" pageId="6" pageNumber="68" refString="Macpherson, E. & Chan, T. Y. (2008) Some lithodid crabs (Crustacea: Decapoda: Lithodidae) from Taiwan and adjacent waters, with the description of one new species from Guam. Zootaxa, 1924, 43 - 52." type="journal article" year="2008">Macpherson & Chan 2008</bibRefCitation>
|
||
: fig. 2a–d). Comparison of partial sequences of the cytochrome oxidase subunit I gene (COI) (658 bp) from five specimens of
|
||
<taxonomicName id="4C2B4D03FFC4C27F3442FB1DFE38FB04" box="[320,482,1160,1185]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="longispina">
|
||
<emphasis id="B95FEA92FFC4C27F3442FB1DFE38FB04" box="[320,482,1160,1185]" italics="true" pageId="6" pageNumber="67">L. longispina</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F33C7610FFC4C27F372CFB1DFDA9FB07" box="[558,627,1160,1186]" name="Japan" pageId="6" pageNumber="67">Japan</collectingCountry>
|
||
(GenBank accession nos
|
||
<accessionNumber id="9478AB63FFC4C27F36B0FB1DFBEDFB07" box="[946,1079,1160,1186]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AB476813" pageId="6" pageNumber="67">AB476813</accessionNumber>
|
||
–476817) with the respective
|
||
<typeStatus id="54908822FFC4C27F3595FB38FED1FB62" box="[151,267,1197,1223]" pageId="6" pageNumber="67" type="holotype">holotypes</typeStatus>
|
||
of
|
||
<taxonomicName id="4C2B4D03FFC4C27F3436FB38FE18FB63" box="[308,450,1197,1222]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="formosae" status="sp. nov.">
|
||
<emphasis id="B95FEA92FFC4C27F3436FB38FE18FB63" box="[308,450,1197,1222]" italics="true" pageId="6" pageNumber="67">L. formosae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95FEA92FFC4C27F34C9FB38FDF0FB62" bold="true" box="[459,554,1197,1223]" pageId="6" pageNumber="67">
|
||
<taxonomicNameLabel id="A26C57E9FFC4C27F34C9FB38FDF0FB62" box="[459,554,1197,1223]" pageId="6" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(GenBank accession no.
|
||
<accessionNumber id="9478AB63FFC4C27F365FFB38FC38FB62" box="[861,994,1197,1223]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/GU289678" pageId="6" pageNumber="67">GU289678</accessionNumber>
|
||
) and
|
||
<taxonomicName id="4C2B4D03FFC4C27F3124FB38FB45FB63" box="[1062,1183,1197,1222]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="paulayi">
|
||
<emphasis id="B95FEA92FFC4C27F3124FB38FB45FB63" box="[1062,1183,1197,1222]" italics="true" pageId="6" pageNumber="67">L. paulayi</emphasis>
|
||
</taxonomicName>
|
||
(GenBank accession no.
|
||
<accessionNumber id="9478AB63FFC4C27F35C1FB47FE9DFB49" box="[195,327,1234,1260]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/GU289677" pageId="6" pageNumber="67">GU289677</accessionNumber>
|
||
) corroborates the specific status of the new species. Pairwise nucleotide divergences are 6.2% between
|
||
<taxonomicName id="4C2B4D03FFC4C27F3401FB6DFE4EFAB4" box="[259,404,1272,1297]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="formosae" status="sp. nov.">
|
||
<emphasis id="B95FEA92FFC4C27F3401FB6DFE4EFAB4" box="[259,404,1272,1297]" italics="true" pageId="6" pageNumber="67">L. formosae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95FEA92FFC4C27F349EFB62FE26FAB4" bold="true" box="[412,508,1271,1297]" pageId="6" pageNumber="67">
|
||
<taxonomicNameLabel id="A26C57E9FFC4C27F349EFB62FE26FAB4" box="[412,508,1271,1297]" pageId="6" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C2B4D03FFC4C27F373AFB6DFD69FAB4" box="[568,691,1272,1297]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="paulayi">
|
||
<emphasis id="B95FEA92FFC4C27F373AFB6DFD69FAB4" box="[568,691,1272,1297]" italics="true" pageId="6" pageNumber="67">L. paulayi</emphasis>
|
||
</taxonomicName>
|
||
, 6.3–6.8% (mean 6.5%) between
|
||
<taxonomicName id="4C2B4D03FFC4C27F314CFB6DFB05FAB4" box="[1102,1247,1272,1297]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="formosae" status="sp. nov.">
|
||
<emphasis id="B95FEA92FFC4C27F314CFB6DFB05FAB4" box="[1102,1247,1272,1297]" italics="true" pageId="6" pageNumber="67">L. formosae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95FEA92FFC4C27F31E5FB62FA92FAB4" bold="true" box="[1255,1352,1271,1297]" pageId="6" pageNumber="67">
|
||
<taxonomicNameLabel id="A26C57E9FFC4C27F31E5FB62FA92FAB4" box="[1255,1352,1271,1297]" pageId="6" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C2B4D03FFC4C27F3086FB6DFECEFA93" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="longispina">
|
||
<emphasis id="B95FEA92FFC4C27F3086FB6DFECEFA93" italics="true" pageId="6" pageNumber="67">L. longispina</emphasis>
|
||
</taxonomicName>
|
||
, and 6.3–6.6% (mean 6.4%) between
|
||
<taxonomicName id="4C2B4D03FFC4C27F37DFFA88FC8CFA93" box="[733,854,1309,1334]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="paulayi">
|
||
<emphasis id="B95FEA92FFC4C27F37DFFA88FC8CFA93" box="[733,854,1309,1334]" italics="true" pageId="6" pageNumber="67">L. paulayi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2B4D03FFC4C27F3693FA88FBF4FA93" box="[913,1070,1309,1334]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="longispina">
|
||
<emphasis id="B95FEA92FFC4C27F3693FA88FBF4FA93" box="[913,1070,1309,1334]" italics="true" pageId="6" pageNumber="67">L. longispina</emphasis>
|
||
</taxonomicName>
|
||
. The interspecific divergences between the three species contrasts strongly with the intraspecific divergence of 0.2–0.8% (mean 0.4%) within
|
||
<taxonomicName id="4C2B4D03FFC4C27F35EBFAFDFE59FA24" box="[233,387,1384,1409]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="longispina">
|
||
<emphasis id="B95FEA92FFC4C27F35EBFAFDFE59FA24" box="[233,387,1384,1409]" italics="true" pageId="6" pageNumber="67">L. longispina</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C331650BFFC4C27F35C4FA18FD08F909" pageId="6" pageNumber="67" type="reference_group">
|
||
<paragraph id="8B943680FFC4C27F35C4FA18FD08F909" blockId="6.[151,1437,151,1746]" pageId="6" pageNumber="67">
|
||
<taxonomicName id="4C2B4D03FFC4C27F35C4FA18FE7FFA03" ID-CoL="86ed675f-2aef-4ded-8319-e4f154ab923d" box="[198,421,1421,1446]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="formosae" status="sp. nov.">
|
||
<emphasis id="B95FEA92FFC4C27F35C4FA18FE7FFA03" box="[198,421,1421,1446]" italics="true" pageId="6" pageNumber="67">Lithodes formosae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95FEA92FFC4C27F34B3FA18FDC9FA02" bold="true" box="[433,531,1421,1447]" pageId="6" pageNumber="67">
|
||
<taxonomicNameLabel id="A26C57E9FFC4C27F34B3FA18FDC9FA02" box="[433,531,1421,1447]" pageId="6" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is the second species of the genus known from
|
||
<collectingCountry id="F33C7610FFC4C27F3174FA18FB15FA02" box="[1142,1231,1421,1447]" name="Taiwan" pageId="6" pageNumber="67">Taiwan</collectingCountry>
|
||
, with
|
||
<taxonomicName id="4C2B4D03FFC4C27F3022FA18FE8AFA69" ID-CoL="22f1dfdc-6ed5-46c3-8ed3-301ad667f63d" authority="Ortmann, 1892" authorityName="Ortmann" authorityYear="1892" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="turritus">
|
||
<emphasis id="B95FEA92FFC4C27F3022FA18FA46FA03" box="[1312,1436,1421,1446]" italics="true" pageId="6" pageNumber="67">L. turritus</emphasis>
|
||
<bibRefCitation id="EFBA4B71FFC4C27F3595FA27FE8AFA69" author="Ortmann" box="[151,336,1458,1484]" pageId="6" pageNumber="68" refString="Ortmann, A. E. (1892) Die Decapoden-Krebse des Strassburger Museums, mit besonderer Berucksichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und zur Zeit im Strassburger Museum aufbewahrten Formen. IV. Die Abtheilungen Galatheidea und Paguridea. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 6, 241 - 326." type="journal article" year="1892">Ortmann, 1892</bibRefCitation>
|
||
</taxonomicName>
|
||
, recorded by
|
||
<bibRefCitation id="EFBA4B71FFC4C27F34F7FA27FD64FA69" box="[501,702,1458,1484]" pageId="6" pageNumber="68" refString="Wu, S. H., Chan, T. Y. & Yu, H. P. (1998) First record of the king crab Lithodes turritus Ortmann, 1892 (Decapoda, Lithodidae) in Taiwanese waters. Crustaceana, 71 (7), 818 - 823." type="journal article">
|
||
Wu
|
||
<emphasis id="B95FEA92FFC4C27F372AFA26FDBFFA69" box="[552,613,1459,1484]" italics="true" pageId="6" pageNumber="67">et al.</emphasis>
|
||
(1998)
|
||
</bibRefCitation>
|
||
and more recently by
|
||
<bibRefCitation id="EFBA4B71FFC4C27F36CDFA27FAFDFA69" author="Macpherson" box="[975,1319,1458,1484]" pageId="6" pageNumber="68" refString="Macpherson, E. & Chan, T. Y. (2008) Some lithodid crabs (Crustacea: Decapoda: Lithodidae) from Taiwan and adjacent waters, with the description of one new species from Guam. Zootaxa, 1924, 43 - 52." type="journal article" year="2008">Macpherson & Chan (2008)</bibRefCitation>
|
||
. The new species is readily distinguished from
|
||
<taxonomicName id="4C2B4D03FFC4C27F374BFA4DFD1AFA54" ID-CoL="22f1dfdc-6ed5-46c3-8ed3-301ad667f63d" box="[585,704,1496,1521]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="turritus">
|
||
<emphasis id="B95FEA92FFC4C27F374BFA4DFD1AFA54" box="[585,704,1496,1521]" italics="true" pageId="6" pageNumber="67">L. turritus</emphasis>
|
||
</taxonomicName>
|
||
(of similar size to the
|
||
<typeStatus id="54908822FFC4C27F36C4FA4DFBF6FA57" box="[966,1068,1496,1522]" pageId="6" pageNumber="67" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C2B4D03FFC4C27F3151FA4DFB04FA54" ID-CoL="86ed675f-2aef-4ded-8319-e4f154ab923d" box="[1107,1246,1496,1521]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="formosae" status="sp. nov.">
|
||
<emphasis id="B95FEA92FFC4C27F3151FA4DFB04FA54" box="[1107,1246,1496,1521]" italics="true" pageId="6" pageNumber="67">L. formosae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95FEA92FFC4C27F31E7FA42FA99FA54" bold="true" box="[1253,1347,1495,1521]" pageId="6" pageNumber="67">
|
||
<taxonomicNameLabel id="A26C57E9FFC4C27F31E7FA42FA99FA54" box="[1253,1347,1495,1521]" pageId="6" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
) by the strongly upraised proximal half of the rostrum (45° versus less than 30°); the length of the primary marginal branchial spines, which are considerably shorter than half the length of the hepatic spine, compared to twothirds to as long as the hepatic spine; and the armature of the posterior branchial margin, which bears only low tubercles or very short conical spines compared to long slender spines in
|
||
<taxonomicName id="4C2B4D03FFC4C27F36F3F9F8FBB2F923" ID-CoL="22f1dfdc-6ed5-46c3-8ed3-301ad667f63d" box="[1009,1128,1645,1670]" class="Malacostraca" family="Lithodidae" genus="Lithodes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="turritus">
|
||
<emphasis id="B95FEA92FFC4C27F36F3F9F8FBB2F923" box="[1009,1128,1645,1670]" italics="true" pageId="6" pageNumber="67">L. turritus</emphasis>
|
||
</taxonomicName>
|
||
(compare
|
||
<figureCitation id="13102A05FFC4C27F31E7F9F8FAF4F922" box="[1253,1326,1645,1671]" captionStart="FIGURE 1" captionStartId="2.[151,255,1957,1981]" captionTargetBox="[151,1436,197,1934]" captionTargetId="figure@2.[151,1436,197,1934]" captionTargetPageId="2" captionText="FIGURE 1. Lithodes formosae sp. nov., ovigerous female holotype (cl 78.4 mm, pcl 55.9 mm, cw 54.7 mm), Dasi, Ilan County, NE Taiwan. A, dorsal habitus. B, cephalothorax, right lateral view." httpUri="https://zenodo.org/record/275487/files/figure.png" pageId="6" pageNumber="67">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="13102A05FFC4C27F303EF9F8FA90F922" box="[1340,1354,1645,1671]" captionStart="FIGURE 2" captionStartId="3.[151,255,1650,1674]" captionTargetBox="[151,1436,194,1628]" captionTargetId="figure@3.[151,1436,194,1628]" captionTargetPageId="3" captionText="FIGURE 2. Lithodes formosae sp. nov., ovigerous female holotype (cl 78.4 mm, pcl 55.9 mm, cw 54.7 mm), Dasi, Ilan County, NE Taiwan. A, carapace. B, right pereopod 4. C, right pereopod 4 merus and carpus. D, right pereopod 4 dactylus." httpUri="https://zenodo.org/record/275488/files/figure.png" pageId="6" pageNumber="67">2</figureCitation>
|
||
A with
|
||
<bibRefCitation id="EFBA4B71FFC4C27F3595F907FEC4F909" author="Ikeda" box="[151,286,1682,1708]" pageId="6" pageNumber="68" refString="Ikeda, H. (1998) The Deep-Sea Crabs of Sagami Bay. Hayama Shiosai Museum, Kanagawa, Japan. 180 pp." type="book" year="1998">Ikeda 1998</bibRefCitation>
|
||
: pl. 66, fig. 1,
|
||
<bibRefCitation id="EFBA4B71FFC4C27F34C5F907FDA3F909" author="Wu" box="[455,633,1682,1708]" pageId="6" pageNumber="68" refString="Wu, S. H., Chan, T. Y. & Yu, H. P. (1998) First record of the king crab Lithodes turritus Ortmann, 1892 (Decapoda, Lithodidae) in Taiwanese waters. Crustaceana, 71 (7), 818 - 823." type="journal article" year="1998">
|
||
Wu
|
||
<emphasis id="B95FEA92FFC4C27F34FAF906FDE8F909" box="[504,562,1683,1708]" italics="true" pageId="6" pageNumber="67">et al.</emphasis>
|
||
1998
|
||
</bibRefCitation>
|
||
: fig. 1).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C331650BFFC4C27F35C4F922FB76F977" box="[198,1196,1719,1746]" pageId="6" pageNumber="67" type="distribution">
|
||
<paragraph id="8B943680FFC4C27F35C4F922FB76F977" blockId="6.[151,1437,151,1746]" box="[198,1196,1719,1746]" pageId="6" pageNumber="67">
|
||
<emphasis id="B95FEA92FFC4C27F35C4F922FEBDF974" bold="true" box="[198,359,1719,1745]" pageId="6" pageNumber="67">Distribution.</emphasis>
|
||
Presently known only from off Dasi, northeastern
|
||
<collectingCountry id="F33C7610FFC4C27F36B8F92DFBC9F977" box="[954,1043,1720,1746]" name="Taiwan" pageId="6" pageNumber="67">Taiwan</collectingCountry>
|
||
;
|
||
<quantity id="4CD39B65FFC4C27F3122F92DFB72F977" box="[1056,1192,1720,1746]" metricMagnitude="2" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="6" pageNumber="67" unit="m" value="550.0" valueMax="600.0" valueMin="500.0">
|
||
500–
|
||
<specimenCount id="9D2DFD09FFC4C27F3159F92DFB72F977" box="[1115,1192,1720,1746]" pageId="6" pageNumber="67" type="generic">600 m</specimenCount>
|
||
</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |