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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.213.96546" ID-Pensoft-Pub="1314-2003-213-131" ID-Pensoft-UUID="05698862B038562DA7D4512233C53B11" ModsDocID="1314-2003-213-131" checkinTime="1668554621434" checkinUser="pensoft" docAuthor="Wang, Yiheng, Sun, Jiahui, Wang, Jingyi, Mao, Qiang, Dong, Wenpan, Yuan, Qingjun, Guo, Lanping &amp; Huang, Luqi" docDate="2022" docId="9E55CA0A36A05B18AD138B887F7FC1D3" docLanguage="en" docName="PhytoKeys 213: 131-141" docOrigin="PhytoKeys 213" docPubDate="2022-11-15" docSource="http://dx.doi.org/10.3897/phytokeys.213.96546" docTitle="Coptis huanjiangensis L. Q. Huang, Q. J. Yuan &amp; Y. H. Wang 2022, sp. nov." docType="treatment" docVersion="1" id="05698862B038562DA7D4512233C53B11" lastPageNumber="131" masterDocId="05698862B038562DA7D4512233C53B11" masterDocTitle="Coptis huanjiangensis, a new species of Ranunculaceae from Guangxi, China" masterLastPageNumber="141" masterPageNumber="131" pageNumber="131" updateTime="1668554621434" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Coptis huanjiangensis, a new species of Ranunculaceae from Guangxi, China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wang, Yiheng</mods:namePart>
<mods:affiliation>State Key Laboratory Breeding Base of Dao-di Herbs, National Resource Center for Chinese Materia Medica, China Academy of Chinese Medical Sciences, Beijing 100700, China &amp; Key Laboratory of Biology and Cultivation of Herb Medicine, Ministry of Agriculture and Rural Affairs, Beijing 100700, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sun, Jiahui</mods:namePart>
<mods:affiliation>State Key Laboratory Breeding Base of Dao-di Herbs, National Resource Center for Chinese Materia Medica, China Academy of Chinese Medical Sciences, Beijing 100700, China &amp; Key Laboratory of Biology and Cultivation of Herb Medicine, Ministry of Agriculture and Rural Affairs, Beijing 100700, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wang, Jingyi</mods:namePart>
<mods:affiliation>State Key Laboratory Breeding Base of Dao-di Herbs, National Resource Center for Chinese Materia Medica, China Academy of Chinese Medical Sciences, Beijing 100700, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mao, Qiang</mods:namePart>
<mods:affiliation>Crop Research Institute, Sichuan Academy of Agricultural Sciences, Chengdu 610023, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dong, Wenpan</mods:namePart>
<mods:affiliation>Laboratory of Systematic Evolution and Biogeography of Woody Plants, School of Ecology and Nature Conservation, Beijing Forestry University, Beijing 100083, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yuan, Qingjun</mods:namePart>
<mods:affiliation>State Key Laboratory Breeding Base of Dao-di Herbs, National Resource Center for Chinese Materia Medica, China Academy of Chinese Medical Sciences, Beijing 100700, China &amp; Key Laboratory of Biology and Cultivation of Herb Medicine, Ministry of Agriculture and Rural Affairs, Beijing 100700, China</mods:affiliation>
<mods:nameIdentifier type="email">yuanqingjun@icmm.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Guo, Lanping</mods:namePart>
<mods:affiliation>State Key Laboratory Breeding Base of Dao-di Herbs, National Resource Center for Chinese Materia Medica, China Academy of Chinese Medical Sciences, Beijing 100700, China &amp; Key Laboratory of Biology and Cultivation of Herb Medicine, Ministry of Agriculture and Rural Affairs, Beijing 100700, China</mods:affiliation>
<mods:nameIdentifier type="email">glp01@126.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Huang, Luqi</mods:namePart>
<mods:affiliation>State Key Laboratory Breeding Base of Dao-di Herbs, National Resource Center for Chinese Materia Medica, China Academy of Chinese Medical Sciences, Beijing 100700, China</mods:affiliation>
<mods:nameIdentifier type="email">huangluqi01@126.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-11-15</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>213</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>131</mods:start>
<mods:end>141</mods:end>
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</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.213.96546</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.213.96546</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-213-131</mods:identifier>
<mods:identifier type="Pensoft-UUID">05698862B038562DA7D4512233C53B11</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:9E55CA0A36A05B18AD138B887F7FC1D3" httpUri="http://treatment.plazi.org/id/9E55CA0A36A05B18AD138B887F7FC1D3" lastPageNumber="131" pageId="0" pageNumber="131">
<subSubSection pageId="0" pageNumber="131" type="nomenclature">
<paragraph pageId="0" pageNumber="131">
<taxonomicName LSID="9E55CA0A-36A0-5B18-AD13-8B887F7FC1D3" authority="L. Q. Huang, Q. J. Yuan &amp; Y. H. Wang" authorityName="L. Q. Huang, Q. J. Yuan &amp; Y. H. Wang" authorityYear="2022" class="Magnoliopsida" family="Ranunculaceae" genus="Coptis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coptis huanjiangensis" order="Ranunculales" pageId="0" pageNumber="131" phylum="Tracheophyta" rank="species" species="huanjiangensis" status="sp. nov.">Coptis huanjiangensis L.Q.Huang, Q.J.Yuan &amp; Y.H.Wang</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="131">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="131" type="description">
<paragraph pageId="0" pageNumber="131">
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Coptis huanjiangensis L. Q. Huang, Q. J. Yuan &amp; Y. H. Wang, sp. nov. A habit B flower, frontal view C flower, back view D opened corolla E petals F sepals G, H follicles I inflorescence J root. Drawn by Yingbao Sun." figureDoi="10.3897/phytokeys.213.96546.figure2" httpUri="https://binary.pensoft.net/fig/769793" pageId="0" pageNumber="131">Figs 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Coptis huanjiangensis L. Q. Huang, Q. J. Yuan &amp; Y. H. Wang, sp. nov. A species habitat (Jiuwanshan National Natural Reserve, Huanjiang County, Guangxi, China) B plant in florescence stage C plant in fruiting stage D leaf, frontal and back view E margin with sparsely upturned spiny hairs F-H inflorescence and flowers I-K follicles and seeds L root. Photos by Yiheng Wang, Jingyi Wang &amp; Qiang Mao." figureDoi="10.3897/phytokeys.213.96546.figure3" httpUri="https://binary.pensoft.net/fig/769794" pageId="0" pageNumber="131">, 3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="131" type="diagnosis">
<paragraph pageId="0" pageNumber="131">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="131">
<taxonomicName authorityName="L. Q. Huang, Q. J. Yuan &amp; Y. H. Wang" authorityYear="2022" class="Magnoliopsida" family="Ranunculaceae" genus="Coptis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coptis huanjiangensis" order="Ranunculales" pageId="0" pageNumber="131" phylum="Tracheophyta" rank="species" species="huanjiangensis">
<emphasis italics="true" pageId="0" pageNumber="131">Coptis huanjiangensis</emphasis>
</taxonomicName>
is morphologically similar to
<taxonomicName lsidName="C. chinensis" pageId="0" pageNumber="131" rank="species" species="chinensis">
<emphasis italics="true" pageId="0" pageNumber="131">C. chinensis</emphasis>
</taxonomicName>
,
<taxonomicName lsidName="C. deltoidei" pageId="0" pageNumber="131" rank="species" species="deltoidei">
<emphasis italics="true" pageId="0" pageNumber="131">C. deltoidei</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="C. omeiensis" pageId="0" pageNumber="131" rank="species" species="omeiensis">
<emphasis italics="true" pageId="0" pageNumber="131">C. omeiensis</emphasis>
</taxonomicName>
, but it differs from these species by having notably longer petioles (15-40 cm), scapes (20-32 cm), and bigger leaf blades with lobes remote obviously.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="131" type="materials_examined">
<paragraph pageId="0" pageNumber="131">
<typeStatus>Type</typeStatus>
.
</paragraph>
<paragraph pageId="0" pageNumber="131">
<materialsCitation accessionNumber="HJ220124" collectingDate="2022-01-24" country="China" county="Huanjiang County" elevation="1082" latitude="25.200298" location="Jiuwanshan National Natural Reserve" longLatPrecision="1" longitude="108.6412" specimenCount="1" stateProvince="Guangxi" typeStatus="holotype">
<collectingCountry name="China">China</collectingCountry>
.
<collectingRegion country="China" name="Guangxi">Guangxi</collectingRegion>
:
<collectingCounty>Huanjiang County</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:9E55CA0A36A05B18AD138B887F7FC1D3:512D58474677DCAFC4876D51D349723B" country="China" county="Huanjiang County" latitude="25.200298" longLatPrecision="1" longitude="108.6412" name="Jiuwanshan National Natural Reserve" stateProvince="Guangxi">Jiuwanshan National Natural Reserve</location>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.082" unit="m" value="1082.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.082" unit="m" value="1082.0">1082 m</elevation>
</quantity>
,
<geoCoordinate degrees="25" direction="north" minutes="12" orientation="latitude" precision="1" seconds="1.07" value="25.200298">25°12'1.07&quot;N</geoCoordinate>
,
<geoCoordinate degrees="108" direction="east" minutes="38" orientation="longitude" precision="1" seconds="28.32" value="108.6412">108°38'28.32&quot;E</geoCoordinate>
, valleys,
<collectingDate value="2022-01-24">24 January 2022</collectingDate>
, Yiheng Wang
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/HJ220124">HJ220124</accessionNumber>
I02 (
<typeStatus>holotype</typeStatus>
CMMI!, isotype CMMI!) (Suppl. materials 2, 3)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="131" type="description">
<paragraph pageId="0" pageNumber="131">Description.</paragraph>
<paragraph pageId="0" pageNumber="131">
Herbs perennial, rhizomes branched, without stolons. Leaves basal, petiole 15-40 cm, glabrous. Leaf blade ovate-triangular, 12-22
<normalizedToken originalValue="×">x</normalizedToken>
9-22 cm, three-sect, papery to subleathery, abaxially glabrous, adaxially nearly glabrous on veins, base cordate, margin with sparsely upturned spiny hairs; central segment petiolulate (petiole 2.5-4 cm), ovate-rhombic, 11-18
<normalizedToken originalValue="×">x</normalizedToken>
7-14 cm, deeply four-ten-lobed, lobes remote, ultimate lobes margin acute serrate, apex acute or obtuse; lateral segments similar to or slightly shorter than the central one, obliquely ovate, unequally two-parted. Scapes one to several, erect, longer or shorter than the leaves, 20-32 cm tall, glabrous, sulcate. Inflorescences terminal, often monochasial, five-ten-flowered; flowers small, actinomorphic, bisexual; bracts lanceolate, palmately divided. Sepals five or six, greenish or redish yellow, long ellipsoid or lanceolate, 5.5-9.0
<normalizedToken originalValue="×">x</normalizedToken>
1.8-3.5 mm, sparsely puberulous. Petals spatulate, 2-5 mm long, glabrous, apex rounded to obtuse, 1/3-1/2 as long as sepals. Stamens numerous, glabrous, 2-4 mm-long, outer ones slightly shorter than petals. Pistils 8-14, 3-5 mm long; follicles 4.5-9.0 mm long, stipitate; seeds ellipsoid, ca. 1-2 mm long, brown.
</paragraph>
<caption doi="10.3897/phytokeys.213.96546.figure2" httpUri="https://binary.pensoft.net/fig/769793" pageId="0" pageNumber="131" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="131">
<emphasis bold="true" pageId="0" pageNumber="131">Figure 2.</emphasis>
<taxonomicName authorityName="L. Q. Huang, Q. J. Yuan &amp; Y. H. Wang" authorityYear="2022" class="Magnoliopsida" family="Ranunculaceae" genus="Coptis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coptis huanjiangensis" order="Ranunculales" pageId="0" pageNumber="131" phylum="Tracheophyta" rank="species" species="huanjiangensis">
<emphasis italics="true" pageId="0" pageNumber="131">Coptis huanjiangensis</emphasis>
</taxonomicName>
L.Q.Huang, Q.J.Yuan &amp; Y.H.Wang, sp. nov.
<emphasis bold="true" pageId="0" pageNumber="131">A</emphasis>
habit
<emphasis bold="true" pageId="0" pageNumber="131">B</emphasis>
flower, frontal view
<emphasis bold="true" pageId="0" pageNumber="131">C</emphasis>
flower, back view
<emphasis bold="true" pageId="0" pageNumber="131">D</emphasis>
opened corolla
<emphasis bold="true" pageId="0" pageNumber="131">E</emphasis>
petals
<emphasis bold="true" pageId="0" pageNumber="131">F</emphasis>
sepals
<emphasis bold="true" pageId="0" pageNumber="131">G, H</emphasis>
follicles
<emphasis bold="true" pageId="0" pageNumber="131">I</emphasis>
inflorescence
<emphasis bold="true" pageId="0" pageNumber="131">J</emphasis>
root. Drawn by Yingbao Sun.
</paragraph>
</caption>
<caption doi="10.3897/phytokeys.213.96546.figure3" httpUri="https://binary.pensoft.net/fig/769794" pageId="0" pageNumber="131" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="131">
<emphasis bold="true" pageId="0" pageNumber="131">Figure 3.</emphasis>
<taxonomicName authorityName="L. Q. Huang, Q. J. Yuan &amp; Y. H. Wang" authorityYear="2022" class="Magnoliopsida" family="Ranunculaceae" genus="Coptis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coptis huanjiangensis" order="Ranunculales" pageId="0" pageNumber="131" phylum="Tracheophyta" rank="species" species="huanjiangensis">
<emphasis italics="true" pageId="0" pageNumber="131">Coptis huanjiangensis</emphasis>
</taxonomicName>
L.Q.Huang, Q.J.Yuan &amp; Y.H.Wang, sp. nov.
<emphasis bold="true" pageId="0" pageNumber="131">A</emphasis>
species habitat (Jiuwanshan National Natural Reserve, Huanjiang County, Guangxi, China)
<emphasis bold="true" pageId="0" pageNumber="131">B</emphasis>
plant in florescence stage
<emphasis bold="true" pageId="0" pageNumber="131">C</emphasis>
plant in fruiting stage
<emphasis bold="true" pageId="0" pageNumber="131">D</emphasis>
leaf, frontal and back view
<emphasis bold="true" pageId="0" pageNumber="131">E</emphasis>
margin with sparsely upturned spiny hairs
<emphasis bold="true" pageId="0" pageNumber="131">F-H</emphasis>
inflorescence and flowers
<emphasis bold="true" pageId="0" pageNumber="131">I-K</emphasis>
follicles and seeds
<emphasis bold="true" pageId="0" pageNumber="131">L</emphasis>
root. Photos by Yiheng Wang, Jingyi Wang &amp; Qiang Mao.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="131" type="distribution">
<paragraph pageId="0" pageNumber="131">Distribution and habitat.</paragraph>
<paragraph pageId="0" pageNumber="131">This species has only been found in the valleys of Jiuwanshan National Natural Reserve, Huanjiang County up until now. It grows in shaded places in valleys at 800-1200 m. a. s. l.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="131" type="etymology">
<paragraph pageId="0" pageNumber="131">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="131">The specific epithet is derived from the type locality, Huanjiang County, Guangxi.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="131" type="phenology">
<paragraph pageId="0" pageNumber="131">Phenology.</paragraph>
<paragraph pageId="0" pageNumber="131">The species was observed flowering in February - March and fruiting in April-June.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="131" type="notes">
<paragraph pageId="0" pageNumber="131">Note.</paragraph>
<paragraph pageId="0" pageNumber="131">
There are seven species and one variant of
<taxonomicName class="Magnoliopsida" family="Ranunculaceae" genus="Coptis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Coptis" order="Ranunculales" pageId="0" pageNumber="131" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="131">Coptis</emphasis>
</taxonomicName>
distributed in China. An identification key is presented below.
</paragraph>
</subSubSection>
</treatment>
</document>