342 lines
39 KiB
XML
342 lines
39 KiB
XML
<document id="16F0EE97FFEB84AAB969D48AD75B7FD6" ID-DOI="10.11646/zootaxa.3999.1.2" ID-GBIF-Dataset="12d1122e-3201-4fb2-a139-cb58026f2146" ID-ISSN="1175-5326" ID-Zenodo-Dep="234596" ID-ZooBank="FBF20477-7DF3-4927-A415-D4BC98736B98" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461279299333" checkinUser="plazi" docAuthor="Heppner, John B. & Bae, Yang-Seop" docDate="2015" docId="8B2F87A12C15DF7EFF42D073FD8BFDEA" docLanguage="en" docName="zt03999p040.pdf" docOrigin="Zootaxa 3999 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nexosa Diakonoff 1977" docType="treatment" docVersion="8" masterDocId="7716FFD92C15DF7CFFD5D740FFACFFF2" masterDocTitle="Review of Nexosa Diakonoff in Vietnam, with a new species and a new subspecies, and transfer to the tribe Archipini (Lepidoptera: Tortricidae: Tortricinae: Archipini)" updateTime="1698613642155" updateUser="plazi">
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<mods:titleInfo id="B4D663CD1BF362C897FDBC4D3552E86C">
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<mods:title id="9818FA66DA4A8168DC95D3EA29737145">Review of Nexosa Diakonoff in Vietnam, with a new species and a new subspecies, and transfer to the tribe Archipini (Lepidoptera: Tortricidae: Tortricinae: Archipini)</mods:title>
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<mods:name id="5BE7E39DBFE173728FDC5838D13AA1AC" type="personal">
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<mods:namePart id="9256E0B2B428C384A5B9997C7BC7987C">Heppner, John B.</mods:namePart>
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<mods:name id="ECCFD2362F2D4B1379709FE1C087B27F" type="personal">
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<mods:roleTerm id="27AEDCA5FEF6276A596B9AF2860967C7">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2ECEE26F7BFD0EEAE785E9B4E97F933D">Bae, Yang-Seop</mods:namePart>
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<mods:typeOfResource id="D491BF4AA9303DFA97D1DE26F74411E7">text</mods:typeOfResource>
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<mods:title id="2A9E98A46CA7AA93FED71E939B33CC80">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="907367CBA529E555B9C3A13820F9D6D1">
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<mods:date id="69CC8F67B1917EF9D072F1E0644EFAF5">2015</mods:date>
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<mods:detail id="EA1D7513713979D7A73B597AD93150B9" type="volume">
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<mods:number id="699EBAA23FC7C41D67F30686109D16C9">3999</mods:number>
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<mods:detail id="4985EED966B3010D8128011C40D25C92" type="issue">
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<mods:number id="516E59789E83F4D1248B2E3D0D84732E">1</mods:number>
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<mods:classification id="E3B18005AE5DF40712C333D8903627C7">journal volume</mods:classification>
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<mods:identifier id="B1818AEB95FCEC78C24DFD63927801F8" type="DOI">10.11646/zootaxa.3999.1.2</mods:identifier>
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<mods:identifier id="82EF08A281A93472E7CA13007DF81AB7" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="5FDC937527F9141D61A4B5D1397B501C" type="Zenodo-Dep">234596</mods:identifier>
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<treatment id="8B2F87A12C15DF7EFF42D073FD8BFDEA" ID-DOI="http://doi.org/10.5281/zenodo.6094739" ID-GBIF-Taxon="119640305" ID-Zenodo-Dep="6094739" LSID="urn:lsid:plazi:treatment:8B2F87A12C15DF7EFF42D073FD8BFDEA" httpUri="http://treatment.plazi.org/id/8B2F87A12C15DF7EFF42D073FD8BFDEA" lastPageId="2" pageId="0">
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<subSubSection id="4B9C653C2C15DF7CFF42D073FB8CF846" pageId="0" type="nomenclature">
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<paragraph id="033936B72C15DF7CFF42D073FE13F8BF" blockId="0.[151,447,1843,1902]" box="[151,447,1843,1869]" pageId="0">
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<heading id="587181DB2C15DF7CFF42D073FE13F8BF" bold="true" box="[151,447,1843,1869]" fontSize="11" level="1" pageId="0" reason="1">
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<taxonomicName id="C4864D342C15DF7CFF42D073FE13F8BF" ID-CoL="92GNV" authority="Diakonoff, 1977" authorityName="Diakonoff" authorityYear="1977" box="[151,447,1843,1869]" class="Insecta" family="Tortricidae" genus="Nexosa" kingdom="Animalia" order="Lepidoptera" pageId="0" phylum="Arthropoda" rank="genus">
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<emphasis id="31F2EAA52C15DF7CFF42D073FE13F8BF" bold="true" box="[151,447,1843,1869]" pageId="0">
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<emphasis id="31F2EAA52C15DF7CFF42D073FF5DF8BF" bold="true" box="[151,241,1843,1869]" italics="true" pageId="0">Nexosa</emphasis>
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<bibRefCitation id="67174B462C15DF7CFF2DD073FE13F8BF" author="Diakonoff" box="[248,447,1843,1869]" pageId="0" refString="Diakonoff, A. N. (1977) Rearrangement of certain Glyphipterygidae sensu Meyrick, 1913, with descriptions of new taxa (Lepidoptera). Zoologische Verhandelingen, 158, 1 - 55." type="journal article" year="1977">Diakonoff, 1977</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="033936B72C15DF7CFF42D015FEACF89C" blockId="0.[151,447,1843,1902]" box="[151,256,1877,1902]" pageId="0">
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<figureCitation id="9BBD2A322C15DF7CFF42D015FEACF89C" box="[151,256,1877,1902]" captionStart-0="FIGURES 1 – 2" captionStart-1="FIGURES 3 – 6" captionStart-2="FIGURE 7" captionStart-3="FIGURE 8" captionStart-4="FIGURE 9" captionStartId-0="1.[151,264,1602,1625]" captionStartId-1="2.[151,264,1813,1836]" captionStartId-2="3.[151,250,1976,1998]" captionStartId-3="6.[151,250,1771,1793]" captionStartId-4="7.[151,250,1096,1118]" captionTargetBox-0="[160,1436,917,1565]" captionTargetBox-1="[150,1436,591,1791]" captionTargetBox-2="[169,1408,1184,1955]" captionTargetBox-3="[156,1436,195,1733]" captionTargetBox-4="[357,1230,194,1063]" captionTargetId-0="figure@1.[151,1436,882,1578]" captionTargetId-1="figure@2.[147,1440,591,1791]" captionTargetId-2="figure@3.[159,1424,1175,1955]" captionTargetId-3="figure@6.[151,1436,195,1749]" captionTargetId-4="figure@7.[340,1248,193,1074]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="6" captionTargetPageId-4="7" captionText-0="FIGURES 1 – 2. Nexosa species in Vietnam. 1. N. hexaphala tamdaoana, n. subsp., holotype male (13.8 mm wingspan). 2. N. tonkinensis, n. sp., holotype female (12.3 mm wingspan)." captionText-1="FIGURES 3 – 6. Nexosa species head close-ups (lateral and dorsal views). 3 – 4. N. hexaphala tamdaoana, n. subsp., male. 5 – 6. N. tonkinensis, n. sp., female." captionText-2="FIGURE 7. Nexosa hexaphala tamdaoana, n. subsp., male genitalia, with details a) aedeagus, b) abdominal pregenital sternal plate (slide JBH 2858)." captionText-3="FIGURE 8. Nexosa tonkinensis, n. sp., female genitalia, with details a) sterigma, ostium, and basal duct of accessory bursa and ductus seminalis (slide JBH 3126)." captionText-4="FIGURE 9. Wing venation of Nexosa marmarastra (Meyrick), Java, Indonesia (redrawn from Diakonoff 1977)." httpUri-0="https://zenodo.org/record/234597/files/figure.png" httpUri-1="https://zenodo.org/record/234598/files/figure.png" httpUri-2="https://zenodo.org/record/234599/files/figure.png" httpUri-3="https://zenodo.org/record/234600/files/figure.png" httpUri-4="https://zenodo.org/record/234601/files/figure.png" pageId="0">Figs. 1–9</figureCitation>
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</paragraph>
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<paragraph id="033936B72C15DF7CFF42D0DEFB8CF846" blockId="0.[151,1056,1950,1972]" box="[151,1056,1950,1972]" pageId="0">
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Type-species:
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<taxonomicName id="C4864D342C15DF7CFEFDD0DEFD7AF846" authority="Meyrick, 1932" authorityName="Meyrick" authorityYear="1932" box="[296,726,1950,1972]" class="Insecta" family="Tortricidae" genus="Mictopsichia" kingdom="Animalia" order="Lepidoptera" pageId="0" phylum="Arthropoda" rank="species" species="marmarastra">
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<emphasis id="31F2EAA52C15DF7CFEFDD0DEFD95F846" box="[296,569,1950,1972]" italics="true" pageId="0">Mictopsichia marmarastra</emphasis>
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Meyrick, 1932
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</taxonomicName>
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, designated by
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<bibRefCitation id="67174B462C15DF7CFCA7D0DEFBB0F846" author="Diakonoff" box="[882,1052,1950,1972]" pageId="0" refString="Diakonoff, A. N. (1977) Rearrangement of certain Glyphipterygidae sensu Meyrick, 1913, with descriptions of new taxa (Lepidoptera). Zoologische Verhandelingen, 158, 1 - 55." type="journal article" year="1977">Diakonoff, 1977</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="4B9C653C2C14DF7DFF42D7D7FEB4FE21" pageId="1" type="diagnosis">
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<paragraph id="033936B72C14DF7DFF42D7D7FEB4FE21" blockId="1.[151,1437,151,827]" pageId="1">
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<emphasis id="31F2EAA52C14DF7DFF42D7D7FEA6FF42" bold="true" box="[151,266,151,176]" pageId="1">Diagnosis</emphasis>
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. Adults have three silvery lines (except for
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<taxonomicName id="C4864D342C14DF7DFD22D7D9FB96FF42" authority="Meyrick" authorityName="Meyrick" box="[759,1082,151,176]" class="Insecta" family="Tortricidae" genus="Nexosa" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="species" species="picturata">
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<emphasis id="31F2EAA52C14DF7DFD22D7D9FC14FF42" box="[759,952,153,176]" italics="true" pageId="1">Nexosa picturata</emphasis>
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(Meyrick))
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</taxonomicName>
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near the forewing apex (two in
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<taxonomicName id="C4864D342C14DF7DFF42D7FDFEEBFF26" box="[151,327,189,212]" class="Insecta" family="Tortricidae" genus="Mictocommosis" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="genus">
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<emphasis id="31F2EAA52C14DF7DFF42D7FDFEEBFF26" box="[151,327,189,212]" italics="true" pageId="1">Mictocommosis</emphasis>
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</taxonomicName>
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). The wing venation (
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<figureCitation id="9BBD2A322C14DF7DFD95D7FCFD28FF26" box="[576,644,188,213]" captionStart="FIGURE 9" captionStartId="7.[151,250,1096,1118]" captionTargetBox="[357,1230,194,1063]" captionTargetId="figure@7.[340,1248,193,1074]" captionTargetPageId="7" captionText="FIGURE 9. Wing venation of Nexosa marmarastra (Meyrick), Java, Indonesia (redrawn from Diakonoff 1977)." httpUri="https://zenodo.org/record/234601/files/figure.png" pageId="1">Fig. 9</figureCitation>
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) is very similar to that of
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<taxonomicName id="C4864D342C14DF7DFC7DD7FDFBF4FF26" box="[936,1112,189,212]" class="Insecta" family="Tortricidae" genus="Mictocommosis" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="genus">
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<emphasis id="31F2EAA52C14DF7DFC7DD7FDFBF4FF26" box="[936,1112,189,212]" italics="true" pageId="1">Mictocommosis</emphasis>
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</taxonomicName>
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, but the hindwing discal cell is shorter in
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<taxonomicName id="C4864D342C14DF7DFEFCD7A1FED0FF0A" box="[297,380,225,248]" class="Insecta" family="Tortricidae" genus="Nexosa" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="genus">
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<emphasis id="31F2EAA52C14DF7DFEFCD7A1FED0FF0A" box="[297,380,225,248]" italics="true" pageId="1">Nexosa</emphasis>
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</taxonomicName>
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and veins R and M1 are more approximate at the discal cell, and M2 is closer to M3 than in
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<taxonomicName id="C4864D342C14DF7DFF42D648FEEBFEED" box="[151,327,264,287]" class="Insecta" family="Tortricidae" genus="Mictocommosis" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="genus">
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<emphasis id="31F2EAA52C14DF7DFF42D648FEEBFEED" box="[151,327,264,287]" italics="true" pageId="1">Mictocommosis</emphasis>
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</taxonomicName>
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. Also, the dorsal margin of the hindwing is very convex. Genital characters differ from those of
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<taxonomicName id="C4864D342C14DF7DFF42D66CFEEBFEB1" box="[151,327,300,323]" class="Insecta" family="Tortricidae" genus="Mictocommosis" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="genus">
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<emphasis id="31F2EAA52C14DF7DFF42D66CFEEBFEB1" box="[151,327,300,323]" italics="true" pageId="1">Mictocommosis</emphasis>
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</taxonomicName>
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, with the male having compact projecting apical spines on the uncus (a broadened anteroventral spine field in
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<taxonomicName id="C4864D342C14DF7DFEFBD610FE72FE95" box="[302,478,336,359]" class="Insecta" family="Tortricidae" genus="Mictocommosis" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="genus">
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<emphasis id="31F2EAA52C14DF7DFEFBD610FE72FE95" box="[302,478,336,359]" italics="true" pageId="1">Mictocommosis</emphasis>
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</taxonomicName>
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), the socius with few setae (extensive setae in
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<taxonomicName id="C4864D342C14DF7DFC30D610FB39FE95" box="[997,1173,336,359]" class="Insecta" family="Tortricidae" genus="Mictocommosis" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="genus">
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<emphasis id="31F2EAA52C14DF7DFC30D610FB39FE95" box="[997,1173,336,359]" italics="true" pageId="1">Mictocommosis</emphasis>
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</taxonomicName>
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); and the female with a large accessory bursa divergent near the ostium (or lacking in some species, but
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<taxonomicName id="C4864D342C14DF7DFBFFD634FB1FFE79" box="[1066,1203,372,395]" class="Insecta" family="Tortricidae" genus="Nexosa" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="species" species="picturata">
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<emphasis id="31F2EAA52C14DF7DFBFFD634FB1FFE79" box="[1066,1203,372,395]" italics="true" pageId="1">N. picturata</emphasis>
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</taxonomicName>
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probably belongs to another genus), and no signum in the bursa (more normal archipine-type signum in
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<taxonomicName id="C4864D342C14DF7DFB9DD6D8FB54FE5D" box="[1096,1272,408,431]" class="Insecta" family="Tortricidae" genus="Mictocommosis" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="genus">
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<emphasis id="31F2EAA52C14DF7DFB9DD6D8FB54FE5D" box="[1096,1272,408,431]" italics="true" pageId="1">Mictocommosis</emphasis>
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</taxonomicName>
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, but without a capitulum).
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</paragraph>
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</subSubSection>
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<subSubSection id="4B9C653C2C14DF7DFF12D69EFCF0FCC9" pageId="1" type="discussion">
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<paragraph id="033936B72C14DF7DFF12D69EFCF0FCC9" blockId="1.[151,1437,151,827]" pageId="1">
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<emphasis id="31F2EAA52C14DF7DFF12D69EFEE9FE05" bold="true" box="[199,325,478,503]" pageId="1">Discussion</emphasis>
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. This genus appears most related to the Asian and African genus
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<taxonomicName id="C4864D342C14DF7DFBF6D6A0FF7FFDE9" authority="Diakonoff" authorityName="Diakonoff" authorityYear="1979" class="Insecta" family="Tortricidae" genus="Mictocommosis" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="genus">
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<emphasis id="31F2EAA52C14DF7DFBF6D6A0FB7FFE05" box="[1059,1235,480,503]" italics="true" pageId="1">Mictocommosis</emphasis>
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<bibRefCitation id="67174B462C14DF7DFB0FD69EFA34FE05" author="Diakonoff" box="[1242,1432,478,503]" pageId="1" refString="Diakonoff, A. N. (1979) Descriptions of new taxa of the so-called Glyphipterigidae auctorum (Lepidoptera). Zoologische Mededelingen, 54, 291 - 312." type="journal article" year="1979">Diakonoff (1979</bibRefCitation>
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,
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<bibRefCitation id="67174B462C14DF7DFF42D542FF7FFDE9" author="Diakonoff" box="[151,211,514,539]" pageId="1" refString="Diakonoff, A. N. (1986) Glyphipterigidae auct. sensu lata (sensu Meyrick, 1913). In: Amsel, H. G., Gregor, F. & Resser, H. (Eds.), Microlepidoptera Palaearctic. Vol. 7. G. Braun, Karlsruhe, pp. 1 - 436, pls 1 - 175." type="book chapter" year="1986">1986</bibRefCitation>
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</taxonomicName>
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), although wing maculation differs considerably between the two. Besides describing the new species
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<taxonomicName id="C4864D342C14DF7DFA57D544FF41FDCD" class="Insecta" family="Tortricidae" genus="Nexosa" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="species" species="aureola">
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<emphasis id="31F2EAA52C14DF7DFA57D544FF41FDCD" italics="true" pageId="1">N. aureola</emphasis>
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</taxonomicName>
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, from
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<collectingCountry id="7B9176272C14DF7DFEE9D568FDB7FDCD" box="[316,539,550,575]" name="Papua New Guinea" pageId="1">Papua New Guinea</collectingCountry>
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,
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<bibRefCitation id="67174B462C14DF7DFDFDD566FD42FDCD" author="Diakonoff" box="[552,750,550,575]" pageId="1" refString="Diakonoff, A. N. (1977) Rearrangement of certain Glyphipterygidae sensu Meyrick, 1913, with descriptions of new taxa (Lepidoptera). Zoologische Verhandelingen, 158, 1 - 55." type="journal article" year="1977">Diakonoff (1977)</bibRefCitation>
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||
transferred to his new genus
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<taxonomicName id="C4864D342C14DF7DFB97D568FB39FDCD" box="[1090,1173,552,575]" class="Insecta" family="Tortricidae" genus="Nexosa" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="genus">
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<emphasis id="31F2EAA52C14DF7DFB97D568FB39FDCD" box="[1090,1173,552,575]" italics="true" pageId="1">Nexosa</emphasis>
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</taxonomicName>
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three other species:
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<taxonomicName id="C4864D342C14DF7DFA57D568FE5CFD91" authority="Meyrick, 1932" authorityName="Meyrick" authorityYear="1932" class="Insecta" family="Tortricidae" genus="Mictopsichia" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="species" species="marmarastra">
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<emphasis id="31F2EAA52C14DF7DFA57D568FE87FD91" italics="true" pageId="1">N. marmarastra</emphasis>
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(Meyrick, 1932)
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</taxonomicName>
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from Java (
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<collectingCountry id="7B9176272C14DF7DFDABD50AFD5DFD91" box="[638,753,586,611]" name="Indonesia" pageId="1">Indonesia</collectingCountry>
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),
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<taxonomicName id="C4864D342C14DF7DFCDDD50CFBFCFD91" authority="Meyrick 1912" authorityName="Meyrick" authorityYear="1912" box="[776,1104,586,611]" class="Insecta" family="Tortricidae" genus="Nexosa" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="species" species="picturata">
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<emphasis id="31F2EAA52C14DF7DFCDDD50CFC3FFD91" box="[776,915,588,611]" italics="true" pageId="1">N. picturata</emphasis>
|
||
(Meyrick 1912)
|
||
</taxonomicName>
|
||
from Assam (
|
||
<collectingCountry id="7B9176272C14DF7DFB2FD50AFA95FD91" box="[1274,1337,586,611]" name="India" pageId="1">India</collectingCountry>
|
||
), and
|
||
<taxonomicName id="C4864D342C14DF7DFA57D50CFE6BFD75" authority="Meyrick 1912" authorityName="Meyrick" authorityYear="1912" class="Insecta" family="Tortricidae" genus="Nexosa" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="species" species="hexaphala">
|
||
<emphasis id="31F2EAA52C14DF7DFA57D50CFEA0FD75" italics="true" pageId="1">N. hexaphala</emphasis>
|
||
(Meyrick 1912)
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="7B9176272C14DF7DFDDAD52EFD2FFD75" box="[527,643,622,647]" name="Sri Lanka" pageId="1">Sri Lanka</collectingCountry>
|
||
.
|
||
<taxonomicName id="C4864D342C14DF7DFD5BD530FCFCFD75" box="[654,848,624,647]" class="Insecta" family="Tortricidae" genus="Nexosa" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="species" species="picturata">
|
||
<emphasis id="31F2EAA52C14DF7DFD5BD530FCFCFD75" box="[654,848,624,647]" italics="true" pageId="1">Nexosa picturata</emphasis>
|
||
</taxonomicName>
|
||
likely belongs in a different genus, as the genitalia differ markedly from other
|
||
<taxonomicName id="C4864D342C14DF7DFE0CD5D4FD80FD59" box="[473,556,660,683]" class="Insecta" family="Tortricidae" genus="Nexosa" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="31F2EAA52C14DF7DFE0CD5D4FD80FD59" box="[473,556,660,683]" italics="true" pageId="1">Nexosa</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="C4864D342C14DF7DFDBCD5D4FCB5FD59" box="[617,793,660,683]" class="Insecta" family="Tortricidae" genus="Mictocommosis" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="31F2EAA52C14DF7DFDBCD5D4FCB5FD59" box="[617,793,660,683]" italics="true" pageId="1">Mictocommosis</emphasis>
|
||
</taxonomicName>
|
||
species in both the male and female.
|
||
<taxonomicName id="C4864D342C14DF7DFB0ED5D4FA81FD59" box="[1243,1325,660,683]" class="Insecta" family="Tortricidae" genus="Nexosa" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="31F2EAA52C14DF7DFB0ED5D4FA81FD59" box="[1243,1325,660,683]" italics="true" pageId="1">Nexosa</emphasis>
|
||
</taxonomicName>
|
||
was first reported from northern
|
||
<collectingCountry id="7B9176272C14DF7DFE48D5F6FE50FD3D" box="[413,508,694,719]" name="Vietnam" pageId="1">Vietnam</collectingCountry>
|
||
by
|
||
<bibRefCitation id="67174B462C14DF7DFDF1D5F6FD4BFD3D" author="Razowski" box="[548,743,694,719]" pageId="1" refString="Razowski, J. (2008) Tortricidae (Lepidoptera) from Vietnam in the collection of the Berlin Museum. 2. Chlidanotinae and descriptions of one species of Tortricini (Lepidoptera: Tortricidae). Polskie Pismo Entomologicne, 77, 199 - 210." type="journal article" year="2008">Razowski (2008)</bibRefCitation>
|
||
, based on a single male of
|
||
<taxonomicName id="C4864D342C14DF7DFBC4D5F8FB0AFD3D" box="[1041,1190,694,719]" class="Insecta" family="Tortricidae" genus="Nexosa" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="species" species="hexaphala">
|
||
<emphasis id="31F2EAA52C14DF7DFBC4D5F8FB0AFD3D" box="[1041,1190,694,719]" italics="true" pageId="1">N. hexaphala</emphasis>
|
||
</taxonomicName>
|
||
captured at Tam Dao, Vinh Phuc Province; but we herein treat that specimen as a new subspecies. The related genus
|
||
<taxonomicName id="C4864D342C14DF7DFB39D59CFA30FD01" box="[1260,1436,732,755]" class="Insecta" family="Tortricidae" genus="Mictocommosis" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="31F2EAA52C14DF7DFB39D59CFA30FD01" box="[1260,1436,732,755]" italics="true" pageId="1">Mictocommosis</emphasis>
|
||
</taxonomicName>
|
||
includes four species: two Asian species (one in
|
||
<collectingCountry id="7B9176272C14DF7DFD67D5BEFCB3FCE5" box="[690,799,766,791]" name="Indonesia" pageId="1">Indonesia</collectingCountry>
|
||
and one from
|
||
<collectingCountry id="7B9176272C14DF7DFC15D440FBACFCE5" box="[960,1024,768,791]" name="Japan" pageId="1">Japan</collectingCountry>
|
||
to
|
||
<collectingCountry id="7B9176272C14DF7DFBF7D5BEFB2FFCE5" box="[1058,1155,766,791]" name="Vietnam" pageId="1">Vietnam</collectingCountry>
|
||
) and two African species (
|
||
<bibRefCitation id="67174B462C14DF7DFF4BD463FEEFFCC9" author="Heppner" box="[158,323,803,827]" pageId="1" refString="Heppner, J. B. (1977) The status of the Glyphipterigidae and a reassessment of relationships in yponomeutoid families and ditrysian superfamilies. Journal of the Lepidopterists' Society, 31, 124 - 134." type="journal article" year="1977">Heppner 1977</bibRefCitation>
|
||
), although the African species need more study.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="57F9663F2C14DF7DFF42D102FD73F985" httpUri="https://zenodo.org/record/234597/files/figure.png" pageId="1" targetBox="[160,1436,917,1565]" targetPageId="1">
|
||
<paragraph id="033936B72C14DF7DFF42D102FD73F985" blockId="1.[151,1437,1602,1655]" pageId="1">
|
||
<emphasis id="31F2EAA52C14DF7DFF42D102FE97F9AA" bold="true" box="[151,315,1602,1625]" pageId="1">FIGURES 1–2.</emphasis>
|
||
<taxonomicName id="C4864D342C14DF7DFE97D104FE20F9AB" box="[322,396,1604,1625]" class="Insecta" family="Tortricidae" genus="Nexosa" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="31F2EAA52C14DF7DFE97D104FE20F9AB" box="[322,396,1604,1625]" italics="true" pageId="1">Nexosa</emphasis>
|
||
</taxonomicName>
|
||
species in Vietnam. 1.
|
||
<taxonomicName id="C4864D342C14DF7DFDA9D104FCD1F9AB" box="[636,893,1603,1625]" class="Insecta" family="Tortricidae" genus="Nexosa" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="subSpecies" species="hexaphala" status="subsp. nov." subSpecies="tamdaoana">
|
||
<emphasis id="31F2EAA52C14DF7DFDA9D104FCD1F9AB" box="[636,893,1603,1625]" italics="true" pageId="1">N. hexaphala tamdaoana</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="2AC157DE2C14DF7DFC5FD104FC4BF9AB" box="[906,999,1603,1625]" pageId="1" rank="subSpecies">n. subsp.</taxonomicNameLabel>
|
||
, holotype male (13.8 mm wingspan). 2.
|
||
<taxonomicName id="C4864D342C14DF7DFA51D104FEA5F985" class="Insecta" family="Tortricidae" genus="Nexosa" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="species" species="tonkinensis" status="sp. nov.">
|
||
<emphasis id="31F2EAA52C14DF7DFA51D104FEA5F985" italics="true" pageId="1">N. tonkinensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="2AC157DE2C14DF7DFEC0D121FEE3F984" box="[277,335,1633,1654]" pageId="1" rank="species">
|
||
<emphasis id="31F2EAA52C14DF7DFEC0D121FEE3F984" bold="true" box="[277,335,1633,1654]" pageId="1">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype female (12.3 mm wingspan).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="4B9C653C2C14DF7EFF12D1DEFD8BFDEA" lastPageId="2" pageId="1" type="reference_group">
|
||
<paragraph id="033936B72C14DF7DFF12D1DEFA31F825" blockId="1.[151,1437,1694,2008]" pageId="1">
|
||
Whereas
|
||
<taxonomicName id="C4864D342C14DF7DFEE4D1E0FE4DF945" ID-CoL="92FRW" box="[305,481,1696,1719]" class="Insecta" family="Tortricidae" genus="Mictocommosis" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="31F2EAA52C14DF7DFEE4D1E0FE4DF945" box="[305,481,1696,1719]" italics="true" pageId="1">Mictocommosis</emphasis>
|
||
</taxonomicName>
|
||
species have a more typical archipine female signum,
|
||
<taxonomicName id="C4864D342C14DF7DFB80D1E0FB04F945" ID-CoL="92GNV" box="[1109,1192,1696,1719]" class="Insecta" family="Tortricidae" genus="Nexosa" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="31F2EAA52C14DF7DFB80D1E0FB04F945" box="[1109,1192,1696,1719]" italics="true" pageId="1">Nexosa</emphasis>
|
||
</taxonomicName>
|
||
species lack this, yet have virtually the same wing venation and similar male genitalia as
|
||
<taxonomicName id="C4864D342C14DF7DFC10D184FBD9F929" ID-CoL="92FRW" box="[965,1141,1732,1755]" class="Insecta" family="Tortricidae" genus="Mictocommosis" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="31F2EAA52C14DF7DFC10D184FBD9F929" box="[965,1141,1732,1755]" italics="true" pageId="1">Mictocommosis</emphasis>
|
||
</taxonomicName>
|
||
, as well as similar wing maculation and head morphology.
|
||
<taxonomicName id="C4864D342C14DF7DFDC9D1A8FDC3F90D" ID-CoL="92GNV" box="[540,623,1768,1791]" class="Insecta" family="Tortricidae" genus="Nexosa" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="31F2EAA52C14DF7DFDC9D1A8FDC3F90D" box="[540,623,1768,1791]" italics="true" pageId="1">Nexosa</emphasis>
|
||
</taxonomicName>
|
||
appears to be a primitive relative of
|
||
<taxonomicName id="C4864D342C14DF7DFBDAD1A8FB13F90D" ID-CoL="92FRW" box="[1039,1215,1768,1791]" class="Insecta" family="Tortricidae" genus="Mictocommosis" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="31F2EAA52C14DF7DFBDAD1A8FB13F90D" box="[1039,1215,1768,1791]" italics="true" pageId="1">Mictocommosis</emphasis>
|
||
</taxonomicName>
|
||
, and best conforms to characters for the tribe
|
||
<taxonomicName id="C4864D342C14DF7DFE6DD04AFD8FF8D1" ID-CoL="B6NRN" box="[440,547,1802,1827]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="tribe" tribe="Archipini">Archipini</taxonomicName>
|
||
rather than
|
||
<taxonomicName id="C4864D342C14DF7DFD7FD04AFCFEF8D1" ID-CoL="B6NT8" box="[682,850,1802,1827]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="tribe" tribe="Hilarographini">Hilarographini</taxonomicName>
|
||
, based on the tribal characteristics (
|
||
<bibRefCitation id="67174B462C14DF7DFB34D04AFA34F8D1" author="Diakonoff" box="[1249,1432,1802,1827]" pageId="1" refString="Diakonoff, A. N. (1977) Rearrangement of certain Glyphipterygidae sensu Meyrick, 1913, with descriptions of new taxa (Lepidoptera). Zoologische Verhandelingen, 158, 1 - 55." type="journal article" year="1977">Diakonoff 1977</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="67174B462C14DF7DFF42D06EFF7FF8B5" author="Diakonoff" box="[151,211,1838,1863]" pageId="1" refString="Diakonoff, A. N. (1986) Glyphipterigidae auct. sensu lata (sensu Meyrick, 1913). In: Amsel, H. G., Gregor, F. & Resser, H. (Eds.), Microlepidoptera Palaearctic. Vol. 7. G. Braun, Karlsruhe, pp. 1 - 436, pls 1 - 175." type="book chapter" year="1986">1986</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="67174B462C14DF7DFF0BD06EFECAF8B5" author="Horak" box="[222,358,1838,1863]" pageId="1" refString="Horak, M. (1984) Assessment of taxonomically significant structures in Tortricinae (Lep., Tortricidae). Mitteilungen der Schweizerische Entomologische Gesellschaft, 57, 3 - 64." type="journal article" year="1984">Horak 1984</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="67174B462C14DF7DFEA4D06FFE01F8B5" author="Horak" box="[369,429,1839,1863]" pageId="1" refString="Horak, M. (1998) The Tortricoidea. In: Kristensen, N. P. (Ed.), Lepidoptera, Moths and Butterflies. Vol. 1. Evolution, Systematics, and Biogeography. Handbuch der Zoologie. Band IV. Arthropoda: Insecta. Teilband 35. W. De Gruyter, Berlin, pp. 199 - 215." type="journal article" year="1998">1998</bibRefCitation>
|
||
). Archipine features of
|
||
<taxonomicName id="C4864D342C14DF7DFD60D070FCABF8B5" ID-CoL="92GNV" box="[693,775,1840,1863]" class="Insecta" family="Tortricidae" genus="Nexosa" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="31F2EAA52C14DF7DFD60D070FCABF8B5" box="[693,775,1840,1863]" italics="true" pageId="1">Nexosa</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="C4864D342C14DF7DFCEAD070FC43F8B5" ID-CoL="92FRW" box="[831,1007,1840,1863]" class="Insecta" family="Tortricidae" genus="Mictocommosis" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="31F2EAA52C14DF7DFCEAD070FC43F8B5" box="[831,1007,1840,1863]" italics="true" pageId="1">Mictocommosis</emphasis>
|
||
</taxonomicName>
|
||
include the toothed knob at the valval costal base, which is a putative pulvinus that lacks setae (a pulvinus is absent in a few other
|
||
<taxonomicName id="C4864D342C14DF7DFB1DD012FA94F899" ID-CoL="B6NRN" box="[1224,1336,1874,1899]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="tribe" tribe="Archipini">Archipini</taxonomicName>
|
||
), but the genera also have a complete and flattened or slender transtilla (split in some
|
||
<taxonomicName id="C4864D342C14DF7DFBC6D036FB28F87D" ID-CoL="B6NRN" box="[1043,1156,1910,1935]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="tribe" tribe="Archipini">Archipini</taxonomicName>
|
||
); and the socius is welldeveloped although elongated and with only a few setae in
|
||
<taxonomicName id="C4864D342C14DF7DFCE1D0DCFC2AF841" ID-CoL="92GNV" box="[820,902,1948,1971]" class="Insecta" family="Tortricidae" genus="Nexosa" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="31F2EAA52C14DF7DFCE1D0DCFC2AF841" box="[820,902,1948,1971]" italics="true" pageId="1">Nexosa</emphasis>
|
||
</taxonomicName>
|
||
. The socius in
|
||
<taxonomicName id="C4864D342C14DF7DFBE7D0DCFB28F841" ID-CoL="92GNV" box="[1074,1156,1948,1971]" class="Insecta" family="Tortricidae" genus="Nexosa" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="31F2EAA52C14DF7DFBE7D0DCFB28F841" box="[1074,1156,1948,1971]" italics="true" pageId="1">Nexosa</emphasis>
|
||
</taxonomicName>
|
||
was erroneously labeled as hami by
|
||
<bibRefCitation id="67174B462C14DF7DFEC9D0FEFE48F825" author="Diakonoff" box="[284,484,1982,2007]" pageId="1" refString="Diakonoff, A. N. (1977) Rearrangement of certain Glyphipterygidae sensu Meyrick, 1913, with descriptions of new taxa (Lepidoptera). Zoologische Verhandelingen, 158, 1 - 55." type="journal article" year="1977">Diakonoff (1977)</bibRefCitation>
|
||
, thereby a further reason the genus may have been described in the
|
||
<taxonomicName id="C4864D342C14DF7DFB25D0FEFA35F825" ID-CoL="B6NT8" box="[1264,1433,1982,2007]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="1" phylum="Arthropoda" rank="tribe" tribe="Hilarographini">Hilarographini</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="033936B72C17DF7EFF42D7D7FD0CFE97" blockId="2.[151,1437,151,536]" pageId="2">
|
||
Razowski (1987) corrected the matter of the socius in
|
||
<taxonomicName id="C4864D342C17DF7EFD2BD7D9FCFDFF42" ID-CoL="92GNV" box="[766,849,153,176]" class="Insecta" family="Tortricidae" genus="Nexosa" kingdom="Animalia" order="Lepidoptera" pageId="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="31F2EAA52C17DF7EFD2BD7D9FCFDFF42" box="[766,849,153,176]" italics="true" pageId="2">Nexosa</emphasis>
|
||
</taxonomicName>
|
||
, but retained the genus in
|
||
<taxonomicName id="C4864D342C17DF7EFBABD7D7FA8BFF42" ID-CoL="B6NT8" box="[1150,1319,151,176]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="2" phylum="Arthropoda" rank="tribe" tribe="Hilarographini">Hilarographini</taxonomicName>
|
||
. The male valva is strongly sclerotized both on the dorsal and ventral margins in
|
||
<taxonomicName id="C4864D342C17DF7EFC05D7FDFB8EFF26" ID-CoL="92GNV" box="[976,1058,189,212]" class="Insecta" family="Tortricidae" genus="Nexosa" kingdom="Animalia" order="Lepidoptera" pageId="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="31F2EAA52C17DF7EFC05D7FDFB8EFF26" box="[976,1058,189,212]" italics="true" pageId="2">Nexosa</emphasis>
|
||
</taxonomicName>
|
||
, but mostly membranous on the margins in
|
||
<taxonomicName id="C4864D342C17DF7EFEC1D7A1FE68FF0A" ID-CoL="92FRW" box="[276,452,225,248]" class="Insecta" family="Tortricidae" genus="Mictocommosis" kingdom="Animalia" order="Lepidoptera" pageId="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="31F2EAA52C17DF7EFEC1D7A1FE68FF0A" box="[276,452,225,248]" italics="true" pageId="2">Mictocommosis</emphasis>
|
||
</taxonomicName>
|
||
(usually sclerotized only on the ventral margin in most
|
||
<taxonomicName id="C4864D342C17DF7EFBE1D79FFB09FF0A" ID-CoL="B6NRN" box="[1076,1189,223,248]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="2" phylum="Arthropoda" rank="tribe" tribe="Archipini">Archipini</taxonomicName>
|
||
). The female genitalia in
|
||
<taxonomicName id="C4864D342C17DF7EFF62D645FEA6FEEE" ID-CoL="92GNV" box="[183,266,261,284]" class="Insecta" family="Tortricidae" genus="Nexosa" kingdom="Animalia" order="Lepidoptera" pageId="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="31F2EAA52C17DF7EFF62D645FEA6FEEE" box="[183,266,261,284]" italics="true" pageId="2">Nexosa</emphasis>
|
||
</taxonomicName>
|
||
have a large accessory bursa in some species and lacking in others, but which varies in presence or absence in many groups; and lacking in
|
||
<taxonomicName id="C4864D342C17DF7EFD81D669FCA8FEB2" ID-CoL="92FRW" box="[596,772,297,320]" class="Insecta" family="Tortricidae" genus="Mictocommosis" kingdom="Animalia" order="Lepidoptera" pageId="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="31F2EAA52C17DF7EFD81D669FCA8FEB2" box="[596,772,297,320]" italics="true" pageId="2">Mictocommosis</emphasis>
|
||
</taxonomicName>
|
||
. No signum is present in
|
||
<taxonomicName id="C4864D342C17DF7EFBCAD669FBDEFEB2" ID-CoL="92GNV" box="[1055,1138,297,320]" class="Insecta" family="Tortricidae" genus="Nexosa" kingdom="Animalia" order="Lepidoptera" pageId="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="31F2EAA52C17DF7EFBCAD669FBDEFEB2" box="[1055,1138,297,320]" italics="true" pageId="2">Nexosa</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="9BBD2A322C17DF7EFB55D667FB6EFEB2" box="[1152,1218,295,320]" captionStart="FIGURE 8" captionStartId="6.[151,250,1771,1793]" captionTargetBox="[156,1436,195,1733]" captionTargetId="figure@6.[151,1436,195,1749]" captionTargetPageId="6" captionText="FIGURE 8. Nexosa tonkinensis, n. sp., female genitalia, with details a) sterigma, ostium, and basal duct of accessory bursa and ductus seminalis (slide JBH 3126)." httpUri="https://zenodo.org/record/234600/files/figure.png" pageId="2">Fig. 8</figureCitation>
|
||
) (archipine signum in
|
||
<taxonomicName id="C4864D342C17DF7EFF61D60DFEC8FE96" ID-CoL="92FRW" box="[180,356,333,356]" class="Insecta" family="Tortricidae" genus="Mictocommosis" kingdom="Animalia" order="Lepidoptera" pageId="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="31F2EAA52C17DF7EFF61D60DFEC8FE96" box="[180,356,333,356]" italics="true" pageId="2">Mictocommosis</emphasis>
|
||
</taxonomicName>
|
||
, but lacking the capitulum).
|
||
</paragraph>
|
||
<paragraph id="033936B72C17DF7EFF12D62FFD8BFDEA" blockId="2.[151,1437,151,536]" pageId="2">
|
||
The correct placement within the
|
||
<taxonomicName id="C4864D342C17DF7EFD92D62FFD1FFE7A" ID-CoL="B6NRN" box="[583,691,367,392]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="2" phylum="Arthropoda" rank="tribe" tribe="Archipini">Archipini</taxonomicName>
|
||
of both
|
||
<taxonomicName id="C4864D342C17DF7EFCC1D631FCCBFE7A" ID-CoL="92GNV" box="[788,871,369,392]" class="Insecta" family="Tortricidae" genus="Nexosa" kingdom="Animalia" order="Lepidoptera" pageId="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="31F2EAA52C17DF7EFCC1D631FCCBFE7A" box="[788,871,369,392]" italics="true" pageId="2">Nexosa</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="C4864D342C17DF7EFC75D631FBFCFE7A" ID-CoL="92FRW" box="[928,1104,369,392]" class="Insecta" family="Tortricidae" genus="Mictocommosis" kingdom="Animalia" order="Lepidoptera" pageId="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="31F2EAA52C17DF7EFC75D631FBFCFE7A" box="[928,1104,369,392]" italics="true" pageId="2">Mictocommosis</emphasis>
|
||
</taxonomicName>
|
||
, and possibly related genera, requires further study. Unfortunately, even recent studies, including those using new DNA techniques (
|
||
<bibRefCitation id="67174B462C17DF7EFAC5D6D4FEF6FE22" author="Dombroskie" pageId="2" refString="Dombroskie, J. J. & Sperling, F. A. H. (2013) Phylogeny of the tribe Archipini (Lepidoptera: Tortricidae: Tortricinae) and the evolutionary correlates of novel secondary sexual structures. Zootaxa, 3729 (1), 1 - 62. http: // dx. doi. org / 10.11646 / zootaxa. 3729.1.1" type="journal article" year="2013">Dombroskie & Sperling 2013</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="67174B462C17DF7EFEB2D6F7FD90FE22" author="Regier" box="[359,572,439,464]" pageId="2" refString="Regier, J. C., Brown, J. W., Mitter, C., Baixeras, J., Cho, S. - W., Cummings, M. P. & Zwick, A. (2012) A molecular phylogeny for the leaf-roller moths (Lepidoptera: Tortricidae) and its implications for classification and life history evolution. PLoS One, 7 (e 35574), 1 - 14." type="journal article" year="2012">
|
||
Regier
|
||
<emphasis id="31F2EAA52C17DF7EFE69D6F9FE43FE22" box="[444,495,439,464]" italics="true" pageId="2">et al</emphasis>
|
||
. 2012
|
||
</bibRefCitation>
|
||
), have not included these genera because they have remained misplaced in
|
||
<taxonomicName id="C4864D342C17DF7EFF42D69CFE5EFE07" ID-CoL="B6NT8" authority="(Chlidaontinae)" baseAuthorityName="Chlidaontinae" box="[151,498,476,501]" class="Insecta" family="Tortricidae" kingdom="Animalia" order="Lepidoptera" pageId="2" phylum="Arthropoda" rank="tribe" tribe="Hilarographini">Hilarographini (Chlidaontinae)</taxonomicName>
|
||
and not specifically studied even as hilarographine genera, or among various exotic genera not included in such studies.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |