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<document id="75E58BE165E719DADD965DB24AF06334" ID-DOI="10.12782/specdiv.25.89" ID-GBIF-Dataset="43bd4af8-e1d9-4d5f-b5f6-73d9c339b2da" ID-ISSN="2189-7301" ID-Zenodo-Dep="3751670" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1586877905669" checkinUser="felipe" docAuthor="Karasawa, Shigenori, Kawano, Keisuke, Fukaya, Shin-ichi &amp; Tsurusaki, Nobuo" docDate="2020" docId="1C3BA22EFFE7D46DC15CFA03D8DB3100" docLanguage="en" docName="SpeciesDiversity.25.89-102.pdf.imf" docOrigin="Species Diversity 25" docStyle="DocumentStyle:27FECD2C3CD153DA405029F2D33C9514.7:SpeciesDiversity.2012-.journal_article" docStyleId="27FECD2C3CD153DA405029F2D33C9514" docStyleName="SpeciesDiversity.2012-.journal_article" docStyleVersion="7" docTitle="Eudigraphis Silvestri 1948" docType="treatment" docVersion="7" lastPageNumber="93" masterDocId="E002DA56FFE3D469C07EFFDDDB2D324B" masterDocTitle="Upgrading of Three Subspecies of Eudigraphis takakuwai to the Species Rank (Diplopoda: Penicillata: Polyxenida: Polyxenidae)" masterLastPageNumber="102" masterPageNumber="89" pageNumber="93" updateTime="1698737589781" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="21F01337773ABCE407773E8662C0E810">Upgrading of Three Subspecies of Eudigraphis takakuwai to the Species Rank (Diplopoda: Penicillata: Polyxenida: Polyxenidae)</mods:title>
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<mods:name id="0BB13A6DC04B290EC3230259E9EE4A7C" type="personal">
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<mods:roleTerm id="61891C236ABB9FF03AC30499CE361812">Author</mods:roleTerm>
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<mods:namePart id="31517DA29DF75B1CD300756418B1AA22">Karasawa, Shigenori</mods:namePart>
<mods:affiliation id="418989718E866DA723B4DAD50B43C79A">Laboratories of Biodiversity and Animal Taxonomy, Faculty of Agriculture, Tottori University, Tottori 680 - 8551, Japan E-mail: ntsuru @ tottori-u. ac. jp</mods:affiliation>
<mods:nameIdentifier id="CF0AE25838FB8C96B047C5BFD7C9E393" type="email">ntsuru@tottori-u.ac.jp</mods:nameIdentifier>
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<mods:name id="13D24F504FC0C36242C2D40405F55D39" type="personal">
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</mods:role>
<mods:namePart id="B45D01364812443D40E17B595BB4057C">Kawano, Keisuke</mods:namePart>
<mods:affiliation id="E3ECE690DEEF9F59090C5594F4575346">The Firefly Museum of Toyota Town, Nakamura 50 - 3, Toyota-cho, Shimonoseki, Yamaguchi 750 - 0441, Japan</mods:affiliation>
</mods:name>
<mods:name id="7A9EDE05DF0C318C45C5FE5EEC30BED2" type="personal">
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<mods:namePart id="15B609582E42C873093DC811E231A64E">Fukaya, Shin-ichi</mods:namePart>
<mods:affiliation id="6B2045821E692A91D92AE270B364F07C">Sugiyama-so 202, Shimo-Ochiai 4 - 18 - 8, Shinjuku Ward, Tokyo 161 - 0033, Japan</mods:affiliation>
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<mods:namePart id="937BE9DF66ADFEC4EF1BDF7BB8B87E18">Tsurusaki, Nobuo</mods:namePart>
<mods:affiliation id="034E8BFD59DACAE38BC6C28B8EFD44E7">Corresponding author</mods:affiliation>
<mods:nameIdentifier id="04261B9DF5F52AFE4A6FE6B3D347B038" type="email">ntsuru@tottori-u.ac.jp</mods:nameIdentifier>
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<mods:title id="6A96400B9C0B112AEEFB5DA0AADA808F">Species Diversity</mods:title>
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<mods:part id="816006C1EC0C304B5F81BC603A6252C4">
<mods:date id="557AB5499F17050BF30CE0BDAC5D9BF9">2020</mods:date>
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<mods:number id="29999BF42A74A8F35395C57F0FEAF01F">2020-02-15</mods:number>
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<mods:number id="A4F7728AF66CEFD39AF0218DFA876771">25</mods:number>
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<treatment id="1C3BA22EFFE7D46DC15CFA03D8DB3100" ID-DOI="http://doi.org/10.5281/zenodo.3808709" ID-GBIF-Taxon="163626650" ID-Zenodo-Dep="3808709" LSID="urn:lsid:plazi:treatment:1C3BA22EFFE7D46DC15CFA03D8DB3100" httpUri="http://treatment.plazi.org/id/1C3BA22EFFE7D46DC15CFA03D8DB3100" lastPageNumber="93" pageId="4" pageNumber="93">
<subSubSection id="DC8840B3FFE7D46DC15CFA03D95437BD" box="[290,633,1502,1526]" pageId="4" pageNumber="93" type="nomenclature">
<paragraph id="942D1338FFE7D46DC15CFA03D95437BD" blockId="4.[290,633,1502,1526]" box="[290,633,1502,1526]" pageId="4" pageNumber="93">
<heading id="CF65A454FFE7D46DC15CFA03D95437BD" box="[290,633,1502,1526]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="93" reason="2">
Genus
<taxonomicName id="539268BBFFE7D46DC117FA03D95437BD" ID-CoL="7PF3C" authority="Silvestri, 1948" authorityName="Silvestri" authorityYear="1948" box="[361,633,1502,1526]" class="Diplopoda" family="Polyxenidae" genus="Eudigraphis" kingdom="Animalia" order="Polyxenida" pageId="4" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="A6E6CF2AFFE7D46DC117FA03DACA37BD" bold="true" box="[361,487,1502,1526]" italics="true" pageId="4" pageNumber="93">Eudigraphis</emphasis>
<bibRefCitation id="F0036EC9FFE7D46DC192FA03D95437BD" author="Silvestri, F." box="[492,633,1502,1526]" pageId="4" pageNumber="93" pagination="214 - 220" refId="ref12194" refString="Silvestri, F. 1948. Tavola sinottica dei generi dei Diplopoda Penicillata. Bollettino del R. Laboratorio di entomologia agraria di Portici 8: 214 - 220." type="journal article" year="1948">Silvestri, 1948</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DC8840B3FFE7D46DC0E2F9C2DA5034D3" pageId="4" pageNumber="93" type="reference_group">
<paragraph id="942D1338FFE7D46DC0E2F9C2DA5034D3" blockId="4.[156,767,1567,2079]" pageId="4" pageNumber="93">
<treatmentCitationGroup id="B4823416FFE7D46DC0E2F9C2DA5034D3" pageId="4" pageNumber="93">
<treatmentCitation id="15333529FFE7D46DC0E2F9C2DAF9347C" author="Silvestri, F." box="[156,468,1567,1591]" page="217" pageId="4" pageNumber="93" year="1948">
<taxonomicName id="539268BBFFE7D46DC0E2F9C2DAF9347C" ID-CoL="7PF3C" authority="Silvestri, 1948: 217" authorityName="Silvestri" authorityPageNumber="217" authorityYear="1948" box="[156,468,1567,1591]" class="Diplopoda" family="Polyxenidae" genus="Eudigraphis" kingdom="Animalia" order="Polyxenida" pageId="4" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="A6E6CF2AFFE7D46DC0E2F9C2DA3F347C" box="[156,274,1567,1591]" italics="true" pageId="4" pageNumber="93">Eudigraphis</emphasis>
<bibRefCitation id="F0036EC9FFE7D46DC166F9C2DAF9347C" author="Silvestri, F." box="[280,468,1567,1591]" pageId="4" pageNumber="93" pagination="214 - 220" refId="ref12194" refString="Silvestri, F. 1948. Tavola sinottica dei generi dei Diplopoda Penicillata. Bollettino del R. Laboratorio di entomologia agraria di Portici 8: 214 - 220." type="journal article" year="1948">Silvestri, 1948: 217</bibRefCitation>
</taxonomicName>
</treatmentCitation>
[
<typeStatus id="4B29AD9AFFE7D46DC19EF9FDD9223473" box="[480,527,1568,1592]" pageId="4" pageNumber="93">type</typeStatus>
species:
<emphasis id="A6E6CF2AFFE7D46DC219F9C2DBDA341C" italics="true" pageId="4" pageNumber="93">
<taxonomicName id="539268BBFFE7D46DC219F9C2D9F0347C" ID-CoL="7PF3C" authorityName="Silvestri" authorityYear="1948" box="[615,733,1567,1591]" class="Diplopoda" family="Polyxenidae" genus="Eudigraphis" kingdom="Animalia" order="Polyxenida" pageId="4" pageNumber="93" phylum="Arthropoda" rank="genus">Eudigraphis</taxonomicName>
ja- ponica
</emphasis>
<bibRefCitation id="F0036EC9FFE7D46DC17EF99DDABC3413" author="Silvestri, F." box="[256,401,1600,1624]" pageId="4" pageNumber="93" pagination="214 - 220" refId="ref12194" refString="Silvestri, F. 1948. Tavola sinottica dei generi dei Diplopoda Penicillata. Bollettino del R. Laboratorio di entomologia agraria di Portici 8: 214 - 220." type="journal article" year="1948">Silvestri, 1948</bibRefCitation>
(=
<taxonomicName id="539268BBFFE7D46DC1C9F99DD9B23413" ID-CoL="3C24B" baseAuthorityName="Miyosi" baseAuthorityYear="1947" box="[439,671,1600,1624]" class="Diplopoda" family="Polyxenidae" genus="Eudigraphis" kingdom="Animalia" order="Polyxenida" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="takakuwai">
<emphasis id="A6E6CF2AFFE7D46DC1C9F99DD9B23413" box="[439,671,1600,1624]" italics="true" pageId="4" pageNumber="93">Eudigraphis takakuwai</emphasis>
</taxonomicName>
)]; Nguy- en
<treatmentCitation id="15333529FFE7D46DC0A4F9BDD9AA3433" box="[218,647,1632,1656]" pageId="4" pageNumber="93">
<bibRefCitation id="F0036EC9FFE7D46DC0A4F9BDD9AA3433" author="Nguyen Duy-Jacquemin, M. &amp; Geoffroy, J. - J." box="[218,647,1632,1656]" pageId="4" pageNumber="93" pagination="89 - 101" refId="ref11720" refString="Nguyen Duy-Jacquemin, M. and Geoffroy, J. - J. 2003. A revised comprehensive checklist, relational database, and taxonomic system of reference for the bristly millipedes of the world (Diplopoda, Polyxenida). African Invertebrates 44: 89 - 101." type="journal article" year="2003">Duy-Jacquemin and Geoffroy 2003: 299</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="15333529FFE7D46DC2E8F9BDDA5434D3" pageId="4" pageNumber="93">
<bibRefCitation id="F0036EC9FFE7D46DC2E8F9BDDA5434D3" author="Short, M. &amp; Vahtera, V." pageId="4" pageNumber="93" pagination="2443 - 2461" refId="ref12058" refString="Short, M., and Vahtera, V. 2017. Phylogenetic relationships of millipedes in the subclass Penicillata (Diplopoda) with a key to the genera. Journal of Natural History 51: 2443 - 2461." type="journal article" year="2017">Short and Vahtera 2017: 2456</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DC8840B3FFE7D46DC0CBF97CD98635D6" pageId="4" pageNumber="93" type="diagnosis">
<paragraph id="942D1338FFE7D46DC0CBF97CD98635D6" blockId="4.[156,767,1567,2079]" pageId="4" pageNumber="93">
<emphasis id="A6E6CF2AFFE7D46DC0CBF97CDA0834F2" bold="true" box="[181,293,1697,1721]" pageId="4" pageNumber="93">Diagnosis.</emphasis>
Head with 8 ocelli each side. Body with 10 segments, 9 pleural projections, a telson, and 13 pairs of legs in adults. Differs from
<taxonomicName id="539268BBFFE7D46DC1E8F93FD98234B1" authority="Attems, 1907" authorityName="Attems" authorityYear="1907" box="[406,687,1762,1786]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="4" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="A6E6CF2AFFE7D46DC1E8F93FD93534B1" box="[406,536,1762,1786]" italics="true" pageId="4" pageNumber="93">Monographis</emphasis>
Attems, 1907
</taxonomicName>
, in 6th antennal segments having just 3 bacilliform sensilla, respectively, and by having caudal trichomes with more numerous (range: 2–6) barbed hooks arranged in a line. Differs from
<taxonomicName id="539268BBFFE7D46DC0E2F8B8DBD73537" authorityName="Jones" authorityYear="1944" box="[156,250,1893,1916]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="4" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="A6E6CF2AFFE7D46DC0E2F8B8DBD73537" box="[156,250,1893,1916]" italics="true" pageId="4" pageNumber="93">Unixenus</emphasis>
</taxonomicName>
in the presence of a single usually continuous row of trichomes along the posterior edge of the tergites.
</paragraph>
</subSubSection>
<subSubSection id="DC8840B3FFE7D46DC0CBF87BD99535B4" pageId="4" pageNumber="93" type="reference_group">
<paragraph id="942D1338FFE7D46DC0CBF87BD99535B4" blockId="4.[156,767,1567,2079]" pageId="4" pageNumber="93">
<emphasis id="A6E6CF2AFFE7D46DC0CBF87BDA4135F6" bold="true" box="[181,364,1957,1982]" pageId="4" pageNumber="93">Included species.</emphasis>
Six species:
<emphasis id="A6E6CF2AFFE7D46DC18EF87BDA343595" italics="true" pageId="4" pageNumber="93">
<taxonomicName id="539268BBFFE7D46DC18EF87BD95B35F6" ID-CoL="3C24B" baseAuthorityName="Miyosi" baseAuthorityYear="1947" box="[496,630,1957,1981]" class="Diplopoda" family="Polyxenidae" genus="Eudigraphis" kingdom="Animalia" order="Polyxenida" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="takakuwai">E. takakuwai</taxonomicName>
,
<taxonomicName id="539268BBFFE7D46DC2FCF87BD9D135F6" ID-CoL="88SF5" authorityName="Karasawa &amp; Kawano &amp; Fukaya &amp; Tsurusaki" authorityYear="2020" baseAuthorityName="Miyosi" baseAuthorityYear="1947" box="[642,764,1958,1981]" class="Diplopoda" family="Polyxenidae" genus="Eudigraphis" kingdom="Animalia" order="Polyxenida" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="nigricans">E. nigricans</taxonomicName>
,
<taxonomicName id="539268BBFFE7D46DC0E2F81ADA343595" ID-CoL="88SF4" authorityName="Karasawa &amp; Kawano &amp; Fukaya &amp; Tsurusaki" authorityYear="2020" baseAuthorityName=": Ishii" baseAuthorityYear="1950" box="[156,281,1990,2014]" class="Diplopoda" family="Polyxenidae" genus="Eudigraphis" kingdom="Animalia" order="Polyxenida" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="kinutensis">E. kinutensis</taxonomicName>
</emphasis>
,
<taxonomicName id="539268BBFFE7D46DC15BF81AD9453595" ID-CoL="3C248" authority="Ishii and Liang, 1990" authorityName="Ishii and Liang" authorityYear="1990" box="[293,616,1990,2014]" class="Diplopoda" family="Polyxenidae" genus="Eudigraphis" kingdom="Animalia" order="Polyxenida" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="A6E6CF2AFFE7D46DC15BF81ADAA63595" box="[293,395,1991,2014]" italics="true" pageId="4" pageNumber="93">E. sinensis</emphasis>
Ishii and Liang, 1990
</taxonomicName>
,
<taxonomicName id="539268BBFFE7D46DC20FF81ADA2A35B4" ID-CoL="8J4X6" authority="Ishii, 1990" authorityName="Ishii" authorityYear="1990" class="Diplopoda" family="Polyxenidae" genus="Eudigraphis" kingdom="Animalia" order="Polyxenida" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="taiwanensis">
<emphasis id="A6E6CF2AFFE7D46DC20FF81AD9D23595" box="[625,767,1991,2014]" italics="true" pageId="4" pageNumber="93">E. taiwanensis</emphasis>
Ishii, 1990
</taxonomicName>
, and
<taxonomicName id="539268BBFFE7D46DC142F83AD99E35B4" ID-CoL="cef4ab97-7617-4a93-b4d5-c14f4d12a7a7" authority="Ishii and Yin, 2000" authorityName="Ishii and Yin" authorityYear="2000" box="[316,691,2022,2047]" class="Diplopoda" family="Polyxenidae" genus="Eudigraphis" kingdom="Animalia" order="Polyxenida" pageId="4" pageNumber="93" phylum="Arthropoda" rank="species" species="xishuangbanna">
<emphasis id="A6E6CF2AFFE7D46DC142F83ADAC335B5" box="[316,494,2022,2046]" italics="true" pageId="4" pageNumber="93">E. xishuangbanna</emphasis>
Ishii and Yin, 2000
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="DC8840B3FFE7D46DC0CBF7DAD97D3A54" box="[181,592,2055,2079]" pageId="4" pageNumber="93" type="distribution">
<paragraph id="942D1338FFE7D46DC0CBF7DAD97D3A54" blockId="4.[156,767,1567,2079]" box="[181,592,2055,2079]" pageId="4" pageNumber="93">
<emphasis id="A6E6CF2AFFE7D46DC0CBF7DADA6C3A54" bold="true" box="[181,321,2055,2079]" pageId="4" pageNumber="93">Distribution.</emphasis>
<collectingCountry id="EC8553A8FFE7D46DC139F7DADAAF3A55" box="[327,386,2055,2078]" name="Japan" pageId="4" pageNumber="93">Japan</collectingCountry>
,
<collectingCountry id="EC8553A8FFE7D46DC1F5F7DADAFB3A54" box="[395,470,2055,2079]" name="Taiwan" pageId="4" pageNumber="93">Taiwan</collectingCountry>
, and
<collectingCountry id="EC8553A8FFE7D46DC275F7DAD9613A54" box="[523,588,2055,2079]" name="China" pageId="4" pageNumber="93">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="DC8840B3FFE7D46DC330FF14D8DB3100" pageId="4" pageNumber="93" type="discussion">
<paragraph id="942D1338FFE7D46DC330FF14DF5E33C8" blockId="4.[821,1433,200,843]" pageId="4" pageNumber="93">
<emphasis id="A6E6CF2AFFE7D46DC330FF14D89F32AA" bold="true" box="[846,946,201,225]" pageId="4" pageNumber="93">Remarks.</emphasis>
<bibRefCitation id="F0036EC9FFE7D46DC3C2FF14DF6332AB" author="Miyosi, Y." box="[956,1102,200,225]" pageId="4" pageNumber="93" refId="ref11438" refString="Miyosi, Y. 1959. Uber Japanische Diplopoden. Arachnological Society of East Asia, Osaka, 223 pp., 19 pls. [In Japanese]" type="book" year="1959">Miyosi (1959)</bibRefCitation>
synonymized
<taxonomicName id="539268BBFFE7D46DC494FF14DE4D32AA" authorityName="Silvestri" authorityYear="1948" box="[1258,1376,201,225]" class="Diplopoda" family="Polyxenidae" genus="Eudigraphis" kingdom="Animalia" order="Polyxenida" pageId="4" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="A6E6CF2AFFE7D46DC494FF14DE4D32AA" box="[1258,1376,201,225]" italics="true" pageId="4" pageNumber="93">Eudigraphis</emphasis>
</taxonomicName>
with
<taxonomicName id="539268BBFFE7D46DC34BFF34D89A334A" authorityName="Attems" authorityYear="1907" box="[821,951,233,257]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="4" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="A6E6CF2AFFE7D46DC34BFF34D89A334A" box="[821,951,233,257]" italics="true" pageId="4" pageNumber="93">Monographis</emphasis>
</taxonomicName>
by expanding definition of the latter. However,
<taxonomicName id="539268BBFFE7D46DC314FED7D8CD3369" authorityName="Silvestri" authorityYear="1948" box="[874,992,266,290]" class="Diplopoda" family="Polyxenidae" genus="Eudigraphis" kingdom="Animalia" order="Polyxenida" pageId="4" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="A6E6CF2AFFE7D46DC314FED7D8CD3369" box="[874,992,266,290]" italics="true" pageId="4" pageNumber="93">Eudigraphis</emphasis>
</taxonomicName>
has been treated as a distinct genus different from
<taxonomicName id="539268BBFFE7D46DC3E8FEF7DF353309" authorityName="Attems" authorityYear="1907" box="[918,1048,298,322]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="4" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="A6E6CF2AFFE7D46DC3E8FEF7DF353309" box="[918,1048,298,322]" italics="true" pageId="4" pageNumber="93">Monographis</emphasis>
</taxonomicName>
since the revision of
<taxonomicName id="539268BBFFE7D46DC488FEF7DE553309" authorityName="Attems" authorityYear="1907" box="[1270,1400,298,322]" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="4" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="A6E6CF2AFFE7D46DC488FEF7DE553309" box="[1270,1400,298,322]" italics="true" pageId="4" pageNumber="93">Monographis</emphasis>
</taxonomicName>
by
<bibRefCitation id="F0036EC9FFE7D46DC34BFE96DFC03328" author="Nguyen Duy-Jacquemin, M. &amp; Conde, B." box="[821,1261,331,355]" pageId="4" pageNumber="93" pagination="43 - 81" refId="ref11686" refString="Nguyen Duy-Jacquemin, M. and Conde, B. 1967. Morphologie et Geonemie du genre Monographis Attems. Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut 64: 43 - 81." type="journal article" year="1967">Nguyen Duy-Jacquemin and Condé (1967)</bibRefCitation>
(also see
<bibRefCitation id="F0036EC9FFE7D46DC52AFE96D8D533C8" author="Shinohara, K. &amp; Ishii, K." pageId="4" pageNumber="93" pagination="1 - 2" refId="ref11877" refString="Shinohara, K. and Ishii, K. 1984. Review of scientific names of Japanese myriapods. VI. Generic name of Japanese Penicillata. Takakuwaia 16: 1 - 2. [In Japanese]" type="journal article" year="1984">Shinohara and Ishii 1984</bibRefCitation>
;
<bibRefCitation id="F0036EC9FFE7D46DC47FFEB6DF4533C8" author="Ishii, K." box="[1025,1128,363,387]" pageId="4" pageNumber="93" pagination="955 - 963" refId="ref10521" refString="Ishii, K. 1988. On the significance of the mandible as a diagnostic character in the taxonomy of penicillate diplopods (Diplopoda: Polyxenidae). The Canadian Entomologist 120: 955 - 963." type="journal article" year="1988">Ishii 1988</bibRefCitation>
).
</paragraph>
<paragraph id="942D1338FFE7D46DC330FE51D8DB3100" blockId="4.[821,1433,200,843]" pageId="4" pageNumber="93">
Japanese species of
<taxonomicName id="539268BBFFE7D46DC466FE51DFA333EF" authorityName="Silvestri" authorityYear="1948" box="[1048,1166,396,420]" class="Diplopoda" family="Polyxenidae" genus="Eudigraphis" kingdom="Animalia" order="Polyxenida" pageId="4" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="A6E6CF2AFFE7D46DC466FE51DFA333EF" box="[1048,1166,396,420]" italics="true" pageId="4" pageNumber="93">Eudigraphis</emphasis>
</taxonomicName>
seem to show close morphological similarity to
<taxonomicName id="539268BBFFE7D46DC449FE70DE37338F" authority="Jones, 1944" authorityName="Jones" authorityYear="1944" box="[1079,1306,428,452]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="4" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="A6E6CF2AFFE7D46DC449FE70DFB8338F" box="[1079,1173,429,452]" italics="true" pageId="4" pageNumber="93">Unixenus</emphasis>
Jones, 1944
</taxonomicName>
(
<bibRefCitation id="F0036EC9FFE7D46DC552FE71D89B33AE" author="Short, M. &amp; Huynh, C." pageId="4" pageNumber="93" pagination="75 - 90" refId="ref12021" refString="Short, M., and Huynh, C. 2013. Four new species of Unixenus Jones. 1944 (Diplopoda, Penicillata, Polyzenida) from Australia. ZooKeys 278: 75 - 90." type="journal article" year="2013">Short and Huynh 2013</bibRefCitation>
;
<bibRefCitation id="F0036EC9FFE7D46DC3BFFE10DFF133AE" author="Huynh, C. &amp; Veenstra, A. A." box="[961,1244,461,485]" pageId="4" pageNumber="93" pagination="283 - 302" refId="ref10348" refString="Huynh, C. and Veenstra, A. A. 2018 a. Two new species of penicillate millipedes (Diplopoda, Polyxenidae) from Phu Quoc Island in southern Vietnam. Zootaxa 4402: 283 - 302." type="journal article" year="2018">Huynh and Veenstra 2018a</bibRefCitation>
), especially in position and the number of bacilliform sensilla on distal segments of the antennae (
<emphasis id="A6E6CF2AFFE7D46DC44CFDD3DF64306D" box="[1074,1097,526,550]" italics="true" pageId="4" pageNumber="93">cf.</emphasis>
, fig.
<quantity id="536ABEDDFFE7D46DC4FEFDD3DF86306D" box="[1152,1195,526,550]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="4" pageNumber="93" unit="in" value="6.0">6 in</quantity>
Huynh and Veenestra 2018a) and in morphology of the gnathochilarium (
<emphasis id="A6E6CF2AFFE7D46DC52AFDF2DE46300C" box="[1364,1387,559,583]" italics="true" pageId="4" pageNumber="93">cf.</emphasis>
, fig. 4F in Huynh and Veenestra 2018a). However, Japanese
<taxonomicName id="539268BBFFE7D46DC50AFD8DD8BD30CC" authorityName="Silvestri" authorityYear="1948" class="Diplopoda" family="Polyxenidae" genus="Eudigraphis" kingdom="Animalia" order="Polyxenida" pageId="4" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="A6E6CF2AFFE7D46DC50AFD8DD8BD30CC" italics="true" pageId="4" pageNumber="93">Eudigraphis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="539268BBFFE7D46DC464FDADDF5530CC" authorityName="Jones" authorityYear="1944" box="[1050,1144,624,647]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="4" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="A6E6CF2AFFE7D46DC464FDADDF5530CC" box="[1050,1144,624,647]" italics="true" pageId="4" pageNumber="93">Unixenus</emphasis>
</taxonomicName>
in morphology of biarticuated setae on the second legs and show similarity to
<taxonomicName id="539268BBFFE7D46DC534FD4CD85F3082" authorityName="Attems" authorityYear="1907" class="Diplopoda" family="Polyxenidae" genus="Monographis" kingdom="Animalia" order="Polyxenida" pageId="4" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="A6E6CF2AFFE7D46DC534FD4CD85F3082" italics="true" pageId="4" pageNumber="93">Monographis</emphasis>
</taxonomicName>
in this character (
<emphasis id="A6E6CF2AFFE7D46DC44FFD6CDF653082" box="[1073,1096,689,713]" italics="true" pageId="4" pageNumber="93">cf.</emphasis>
, fig.
<quantity id="536ABEDDFFE7D46DC402FD6CDF9C3083" box="[1148,1201,688,713]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="4" pageNumber="93" unit="in" value="12.0">12 in</quantity>
Huynh and Veenestra 2018a). Separation of
<taxonomicName id="539268BBFFE7D46DC465FD0CDFBC30A2" authorityName="Silvestri" authorityYear="1948" box="[1051,1169,721,745]" class="Diplopoda" family="Polyxenidae" genus="Eudigraphis" kingdom="Animalia" order="Polyxenida" pageId="4" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="A6E6CF2AFFE7D46DC465FD0CDFBC30A2" box="[1051,1169,721,745]" italics="true" pageId="4" pageNumber="93">Eudigraphis</emphasis>
</taxonomicName>
from
<taxonomicName id="539268BBFFE7D46DC4ABFD0FDE1E30A2" authorityName="Jones" authorityYear="1944" box="[1237,1331,722,745]" class="Diplopoda" family="Polyxenidae" genus="Unixenus" kingdom="Animalia" order="Polyxenida" pageId="4" pageNumber="93" phylum="Arthropoda" rank="genus">
<emphasis id="A6E6CF2AFFE7D46DC4ABFD0FDE1E30A2" box="[1237,1331,722,745]" italics="true" pageId="4" pageNumber="93">Unixenus</emphasis>
</taxonomicName>
seems to be supported by a molecular phylogeny provided by
<bibRefCitation id="F0036EC9FFE7D46DC51EFD2CDF213161" author="Short, M. &amp; Vahtera, V." pageId="4" pageNumber="93" pagination="2443 - 2461" refId="ref12058" refString="Short, M., and Vahtera, V. 2017. Phylogenetic relationships of millipedes in the subclass Penicillata (Diplopoda) with a key to the genera. Journal of Natural History 51: 2443 - 2461." type="journal article" year="2017">Short and Vahtera (2017)</bibRefCitation>
. However, further detailed analyses seem to be needed.
</paragraph>
</subSubSection>
</treatment>
</document>
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