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<document id="94DCF4C7A30B029557EBB1A439BD07E2" ID-CLB-Dataset="8404" ID-DOI="10.11646/zootaxa.4852.2.1" ID-GBIF-Dataset="33b954d6-7939-47da-9776-6b079002355a" ID-ISSN="1175-5326" ID-Zenodo-Dep="4519043" ID-ZooBank="2A612C28-2E39-4729-9DDD-27EE6ED90F4E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609593317386" checkinUser="plazi" docAuthor="Balke, Michael, Megna, Yoandri S., Zenteno, Nilver, Figueroa, Luis &amp; Hendrich, Lars" docDate="2020" docId="03EDDF2BFFC2F219FF0CFEBE19F8F8D7" docLanguage="en" docName="zootaxa.4852.2.1.pdf" docOrigin="Zootaxa 4852 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Liodessus alpinus Balke &amp; Megna &amp; Zenteno &amp; Figueroa &amp; Hendrich 2020, sp. nov." docType="treatment" docVersion="6" lastPageNumber="155" masterDocId="FFD4A753FFC6F21DFF9BFFF4187CFF9B" masterDocTitle="New Liodessus species from the high Andes of Peru (Coleoptera: Dytiscidae Bidessini)" masterLastPageNumber="165" masterPageNumber="151" pageNumber="155" updateTime="1698850946412" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="A19625E122BDF6C3D5E2F68708DC11F1" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7ABE745296009261C09D1D3C48446FF3">
<mods:title id="4F47BE499ED976B48990B0298B6DA61F">New Liodessus species from the high Andes of Peru (Coleoptera: Dytiscidae Bidessini)</mods:title>
</mods:titleInfo>
<mods:name id="D7B76215DF085FC175523D3D604C36F8" type="personal">
<mods:role id="0C501E595F064D6E15D714F8C2845A78">
<mods:roleTerm id="83398EEBA6FA9833BF817C30DF2D6B11">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4D2C9B1DEC5448F66E3DEF83DB8B0736">Balke, Michael</mods:namePart>
<mods:nameIdentifier id="AE6D407BAFCAABDF4B0151889E0C4A5D" type="ORCID">0000-0002-3773-6586</mods:nameIdentifier>
<mods:affiliation id="4121C32D08F77E36E5641F2E4E682CB1">SNSB-Zoologische Staatssammlung München, Münchhausenstrasse 21, 81247 München, Germany.</mods:affiliation>
</mods:name>
<mods:name id="E544739BA5CE479F40390FEC5EBF0CDA" type="personal">
<mods:role id="61CEE6F9359B6743034ED9937509A3A0">
<mods:roleTerm id="C1CAB6D1905A1DC37FC129F779412546">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="477399C26C8564EDB7964ACA620920E0">Megna, Yoandri S.</mods:namePart>
<mods:nameIdentifier id="469BEDFCDEB1B6A844704E4616C6C20B" type="ORCID">0000-0002-5336-281X</mods:nameIdentifier>
<mods:affiliation id="56E49C352F91FC8BCC9F1592E36575BD">Departamento de Biología, Universidad de Oriente. Patricio Lumumba s / n, Santiago, Santiago de Cuba, Cuba. ysmegna @ gmail. com; https: // orcid. org / 0000 - 0002 - 5336 - 281 X</mods:affiliation>
<mods:nameIdentifier id="159B9021CE43C204B0E9FBFB87509725" type="email">ysmegna@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="1E3AE1A845F86B15C0B67466DE41B5BD" type="personal">
<mods:role id="F7DD305B274BA2020D9270A07EED3A16">
<mods:roleTerm id="7532A0FAC6F7667A6A22FC6970CFAAF6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6BB762EE8CB02917BC051064477898D8">Zenteno, Nilver</mods:namePart>
<mods:nameIdentifier id="B640A21C4E41263B0B354341873741E0" type="ORCID">0000-0001-8698-3797</mods:nameIdentifier>
<mods:affiliation id="2E1A4F7DDC4DC3A424F3A9CE33F854D4">Departamento de Entomología, Museo de Historía Natural, Universidad Nacional Mayor de San Marcos, Avenida Arenales 1256, Jesús María 15072, Lima, Perú. &amp; nilverjzenteno @ gmail. com; https: // orcid. org / 0000 - 0001 - 8698 - 3797</mods:affiliation>
<mods:nameIdentifier id="A558393C77480C4DBB43E8D0D0A6D5B1" type="email">nilverjzenteno@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="F409816D5C80865A257E35C2E28D0ADF" type="personal">
<mods:role id="D96BA08DFC59873F4997C5D741608636">
<mods:roleTerm id="C95AC75CB40EDA5AC100F4F619F9D137">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BD97A1C1168F06A0FF5640748EF3003D">Figueroa, Luis</mods:namePart>
<mods:nameIdentifier id="C99932CEE964C0281C4A0BC84FB1748B" type="ORCID">0000-0002-1404-6195</mods:nameIdentifier>
<mods:affiliation id="7F0AC019FB971AAEE55A078B53756172">Departamento de Entomología, Museo de Historía Natural, Universidad Nacional Mayor de San Marcos, Avenida Arenales 1256, Jesús María 15072, Lima, Perú. &amp; luis _ thecell @ hotmail. com; https: // orcid. org / 0000 - 0002 - 1404 - 6195</mods:affiliation>
<mods:nameIdentifier id="282988EEE620E254075EFD72238F3CFC" type="email">luis_thecell@hotmail.com</mods:nameIdentifier>
</mods:name>
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<mods:roleTerm id="0D224A12D70DDA10CAE31F442D6B247F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="881F10E427EFFEBDAC1F9FB23E1EA55B">Hendrich, Lars</mods:namePart>
<mods:nameIdentifier id="B15773CA392E780DEA9D44A792286E62" type="ORCID">0000-0001-8366-0749</mods:nameIdentifier>
<mods:affiliation id="E9678A6310DFD9392351119623901058">SNSB-Zoologische Staatssammlung München, Münchhausenstrasse 21, 81247 München, Germany. &amp; hendrich @ snsb. de; https: // orcid. org / 0000 - 0001 - 8366 - 0749</mods:affiliation>
<mods:nameIdentifier id="06AE9F7E59DE0E3C72653711347507F2" type="email">hendrich@snsb.de</mods:nameIdentifier>
</mods:name>
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<mods:title id="A6F6D39E249EC2A8A5CE0CCA13F4427E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="BB5FE8F7E44714CAEF114CF9CDF18969">
<mods:date id="324686D50007B369B754F35FCEDFD569">2020</mods:date>
<mods:detail id="043A3D6955DD54A4C9AB0DB7E8DF4F95" type="pubDate">
<mods:number id="E9CDAF68F7A649BC83B16BB313794063">2020-09-15</mods:number>
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<mods:number id="497A25498A83C722EF8C64B6AA28EB76">4852</mods:number>
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<mods:number id="80E59E11327252253F2B1F1FF1BB7BE1">2</mods:number>
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<mods:identifier id="D6730A284C4AEB0B8C41EF34FDE83541" type="DOI">10.11646/zootaxa.4852.2.1</mods:identifier>
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<mods:identifier id="3A957603E806E5E72CA5AFD75DEC5456" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03EDDF2BFFC2F219FF0CFEBE19F8F8D7" ID-DOI="http://doi.org/10.5281/zenodo.4519031" ID-GBIF-Taxon="177309772" ID-Zenodo-Dep="4519031" LSID="urn:lsid:plazi:treatment:03EDDF2BFFC2F219FF0CFEBE19F8F8D7" httpUri="http://treatment.plazi.org/id/03EDDF2BFFC2F219FF0CFEBE19F8F8D7" lastPageNumber="155" pageId="4" pageNumber="155">
<subSubSection id="C35E3DB6FFC2F219FF0CFEBE19A8FEFF" box="[151,468,330,357]" pageId="4" pageNumber="155" type="nomenclature">
<paragraph id="8BFB6E3DFFC2F219FF0CFEBE19A8FEFF" blockId="4.[151,468,330,393]" box="[151,468,330,357]" pageId="4" pageNumber="155">
<heading id="D0B3D951FFC2F219FF0CFEBE19A8FEFF" bold="true" box="[151,468,330,357]" fontSize="11" level="1" pageId="4" pageNumber="155" reason="1">
<emphasis id="B930B22FFFC2F219FF0CFEBE19A8FEFF" bold="true" box="[151,468,330,357]" pageId="4" pageNumber="155">
<taxonomicName id="4C4415BEFFC2F219FF0CFEBE1913FEFE" ID-CoL="8LZP3" authority="Balke &amp; Megna &amp; Zenteno &amp; Figueroa &amp; Hendrich, 2020" authorityName="Balke &amp; Megna &amp; Zenteno &amp; Figueroa &amp; Hendrich" authorityYear="2020" box="[151,367,330,357]" class="Insecta" family="Dytiscidae" genus="Liodessus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="alpinus" status="sp. nov.">
<emphasis id="B930B22FFFC2F219FF0CFEBE1913FEFE" bold="true" box="[151,367,330,357]" italics="true" pageId="4" pageNumber="155">Liodessus alpinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2030F54FFC2F219FEECFEBE19A8FEFF" box="[375,468,330,356]" pageId="4" pageNumber="155" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35E3DB6FFC2F219FF0CFE9A19C1FE12" box="[151,445,366,393]" pageId="4" pageNumber="155" type="description">
<paragraph id="8BFB6E3DFFC2F219FF0CFE9A19C1FE12" blockId="4.[151,468,330,393]" box="[151,445,366,393]" pageId="4" pageNumber="155">
(
<figureCitation id="137F72B8FFC2F219FF04FE9A1961FE12" box="[159,285,366,393]" captionStart="FIGURE 2" captionStartId="7.[151,250,1936,1960]" captionTargetBox="[247,1336,202,1901]" captionTargetId="figure-17@7.[230,1357,181,1911]" captionTargetPageId="7" captionText="FIGURE 2. Dorsal habitus of Liodessus alpinus sp. nov., paratype, male (A), female (B), from Peru: Junín, Junín, Lagoon Estancia Azulmito; L. rhigos sp. nov., paratype male (C) and female (D), both from Peru: Junín, Junín, Lagoon Huacra. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4519047" httpUri="https://zenodo.org/record/4519047/files/figure.png" pageId="4" pageNumber="155">Figs 2A, B</figureCitation>
,
<figureCitation id="137F72B8FFC2F219FEB2FE9A1930FE13" box="[297,332,366,392]" captionStart="FIGURE 4" captionStartId="9.[151,250,1954,1978]" captionTargetBox="[174,1412,191,1927]" captionTargetId="figure-17@9.[172,1416,181,1930]" captionTargetPageId="9" captionText="FIGURE 4. Median lobe of aedeagus in ventral and lateral view, and paramere in lateral view of paratypes of Liodessus acollensis (A); L. andinus (B); L. alpinus sp. nov. (C), L. rhigos sp. nov. (D) and L. thespesios sp. nov. (E). Scale bar = 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.4519051" httpUri="https://zenodo.org/record/4519051/files/figure.png" pageId="4" pageNumber="155">4C</figureCitation>
,
<figureCitation id="137F72B8FFC2F219FECCFE9A19E5FE12" box="[343,409,366,393]" captionStart="FIGURE 5" captionStartId="10.[151,250,1812,1836]" captionTargetBox="[155,1432,273,1787]" captionTargetId="figure-54@10.[155,1432,273,1789]" captionTargetPageId="10" captionText="FIGURE 5. Habitat of: Liodessus acollensis (A), Junín, Juaja, Lagoon Tragadero, 3,459 m, (PER_YSM_2018_03); (B) Junín, La Oroya, Nueva Ciudad Morococha, 4,244 m, (PER_YSM_2018_04); L. alpinus sp. nov. (C), Junín, Junín, Shalacancha, 4,320 m, (PER_YSM_2018_10); L. acollensis and L. rhigos sp. nov. (D), Junín, Junín, Lagoon Huacra, 4,461 m, (PER_YSM_2018_ 13); L. acollensis and L. alpinus sp. nov. (E), Junín, Junín, Lagoon Estancia Azulmito, 4,452 m, (PER_YSM_2018_14); L. acollensis and L. rhigos sp. nov. (F), Junín, Junín, Lagoon Yanamyacu, 4,327 m, (PER_YSM_2018_16)." figureDoi="http://doi.org/10.5281/zenodo.4519053" httpUri="https://zenodo.org/record/4519053/files/figure.png" pageId="4" pageNumber="155">5C, E</figureCitation>
,
<figureCitation id="137F72B8FFC2F219FE3EFE9B19C8FE12" box="[421,436,367,393]" captionStart="FIGURE 7" captionStartId="12.[151,250,1110,1134]" captionTargetBox="[151,1435,181,1086]" captionTargetId="figure-22@12.[151,1435,181,1087]" captionTargetPageId="12" captionText="FIGURE 7. Distribution map of Liodessus species treated in this publication." figureDoi="http://doi.org/10.5281/zenodo.4519057" httpUri="https://zenodo.org/record/4519057/files/figure.png" pageId="4" pageNumber="155">7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35E3DB6FFC2F219FF0CFE42193BFCC7" pageId="4" pageNumber="155" type="materials_examined">
<paragraph id="8BFB6E3DFFC2F219FF0CFE421CFBFE4A" blockId="4.[151,1437,438,1868]" box="[151,1159,438,465]" pageId="4" pageNumber="155">
<emphasis id="B930B22FFFC2F219FF0CFE42194AFE4B" bold="true" box="[151,310,438,464]" pageId="4" pageNumber="155">Type locality.</emphasis>
<materialsCitation id="3B2C6460FFC2F219FEA7FE431BD6FE4A" ID-GBIF-Occurrence="3032722317" box="[316,938,438,465]" country="Peru" elevation="4452" location="Lagoon Estancia Azulmito" pageId="4" pageNumber="155" specimenCount="1" stateProvince="Junin">
<collectingCountry id="F3532EADFFC2F219FEA7FE431908FE4A" box="[316,372,439,465]" name="Peru" pageId="4" pageNumber="155">Peru</collectingCountry>
:
<collectingRegion id="4980A0DFFFC2F219FEE5FE4219C1FE4B" box="[382,445,438,464]" country="Peru" name="Junin" pageId="4" pageNumber="155">Junín</collectingRegion>
,
<collectingRegion id="4980A0DFFFC2F219FE52FE421A74FE4B" box="[457,520,438,464]" country="Peru" name="Junin" pageId="4" pageNumber="155">Junín</collectingRegion>
,
<location id="8E9B38E6FFC2F219FD8FFE431B3EFE4B" LSID="urn:lsid:plazi:treatment:03EDDF2BFFC2F219FF0CFEBE19F8F8D7:8E9B38E6FFC2F219FD8FFE431B3EFE4B" box="[532,834,438,465]" country="Peru" name="Lagoon Estancia Azulmito" pageId="4" pageNumber="155" stateProvince="Junin">Lagoon Estancia Azulmito</location>
,
<quantity id="4CBCC3D8FFC2F219FCD7FE421BD6FE4A" box="[844,938,438,465]" metricMagnitude="3" metricUnit="m" metricValue="4.452" pageId="4" pageNumber="155" unit="m" value="4452.0">
<elevation id="0069890EFFC2F219FCD7FE421BD6FE4A" box="[844,938,438,465]" metricMagnitude="3" metricUnit="m" metricValue="4.452" pageId="4" pageNumber="155" unit="m" value="4452.0">4,452 m</elevation>
</quantity>
</materialsCitation>
, -11.303° -75.989°.
</paragraph>
<paragraph id="8BFB6E3DFFC2F219FF5CFE2E1ACEFDA6" blockId="4.[151,1437,438,1868]" pageId="4" pageNumber="155">
<emphasis id="B930B22FFFC2F219FF5CFE2E190EFE6F" bold="true" box="[199,370,474,500]" pageId="4" pageNumber="155">Type material.</emphasis>
<materialsCitation id="3B2C6460FFC2F219FEECFE2E1ACEFDA6" ID-GBIF-Occurrence="3032722334" collectingDate="2018-08-14" collectionCode="MUSM, PER" collectorName="Y. S. Megna &amp; N. Zenteno" country="Peru" elevation="4452" location="Male" pageId="4" pageNumber="155" specimenCount="2" stateProvince="Junin" typeStatus="holotype">
<emphasis id="B930B22FFFC2F219FEECFE2E1991FE6F" bold="true" box="[375,493,474,500]" pageId="4" pageNumber="155">
<typeStatus id="54FFD09FFFC2F219FEECFE2E1994FE6F" box="[375,488,474,500]" pageId="4" pageNumber="155" type="holotype">Holotype</typeStatus>
:
</emphasis>
<location id="8E9B38E6FFC2F219FE6AFE2E1A57FE6F" LSID="urn:lsid:plazi:treatment:03EDDF2BFFC2F219FF0CFEBE19F8F8D7:8E9B38E6FFC2F219FE6AFE2E1A57FE6F" box="[497,555,474,500]" country="Peru" name="Male" pageId="4" pageNumber="155" stateProvince="Junin">Male</location>
(
<collectionCode id="ED55F6F8FFC2F219FDACFE2E1AE8FE6F" box="[567,660,474,500]" pageId="4" pageNumber="155">MUSM</collectionCode>
): “
<collectingCountry id="F3532EADFFC2F219FD35FE2E1A94FE6F" box="[686,744,474,500]" name="Peru" pageId="4" pageNumber="155">Peru</collectingCountry>
:
<collectingRegion id="4980A0DFFFC2F219FD74FE2E1B52FE6F" box="[751,814,474,500]" country="Peru" name="Junin" pageId="4" pageNumber="155">Junín</collectingRegion>
,
<collectingRegion id="4980A0DFFFC2F219FCA3FE2E1B0BFE6F" box="[824,887,474,500]" country="Peru" name="Junin" pageId="4" pageNumber="155">Junín</collectingRegion>
, Lagoon Estancia Azulmito,
<quantity id="4CBCC3D8FFC2F219FB2BFE2E1D79FE6E" box="[1200,1285,474,501]" metricMagnitude="3" metricUnit="m" metricValue="4.452" pageId="4" pageNumber="155" unit="m" value="4452.0">
<elevation id="0069890EFFC2F219FB2BFE2E1D79FE6E" box="[1200,1285,474,501]" metricMagnitude="3" metricUnit="m" metricValue="4.452" pageId="4" pageNumber="155" unit="m" value="4452.0">4452 m</elevation>
</quantity>
,
<date id="FFFA48FDFFC2F219FA95FE2E1DE4FE6F" box="[1294,1432,474,500]" pageId="4" pageNumber="155" value="2018-08-14">
<collectingDate id="EFBEB115FFC2F219FA95FE2E1DE4FE6F" box="[1294,1432,474,500]" pageId="4" pageNumber="155" value="2018-08-14">14.viii.2018</collectingDate>
</date>
, -11.303° -75.989°,
<collectorName id="26B10BEBFFC2F219FEF1FE0B198EFD82" box="[362,498,510,537]" pageId="4" pageNumber="155">Y. S. Megna</collectorName>
&amp;
<collectorName id="26B10BEBFFC2F219FD88FE0B1AECFD82" box="[531,656,511,537]" pageId="4" pageNumber="155">N. Zenteno</collectorName>
(
<collectionCode id="ED55F6F8FFC2F219FD06FE0A1AA8FD83" box="[669,724,510,536]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:12821" name="City Museum" pageId="4" pageNumber="155" type="Herbarium">PER</collectionCode>
_YSM_2018_14)”; “
<typeStatus id="54FFD09FFFC2F219FC22FE0A1C58FD83" box="[953,1060,510,536]" pageId="4" pageNumber="155" type="holotype">Holotype</typeStatus>
<taxonomicName id="4C4415BEFFC2F219FBB1FE0A1C8DFD83" authority="Balke &amp; Megna &amp; Zenteno &amp; Figueroa &amp; Hendrich, 2020" authorityName="Balke &amp; Megna &amp; Zenteno &amp; Figueroa &amp; Hendrich" authorityYear="2020" box="[1066,1265,510,536]" class="Insecta" family="Dytiscidae" genus="Liodessus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="alpinus" status="sp. nov.">Liodessus alpinus</taxonomicName>
<emphasis id="B930B22FFFC2F219FB6DFE0A1D32FD83" bold="true" box="[1270,1358,510,536]" pageId="4" pageNumber="155">
<taxonomicNameLabel id="A2030F54FFC2F219FB6DFE0A1D32FD83" box="[1270,1358,510,536]" pageId="4" pageNumber="155" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Balke, Megna, Hendrich des. 2020” [red printed label].
</materialsCitation>
</paragraph>
<paragraph id="8BFB6E3DFFC2F219FF5CFDB2193BFCC7" blockId="4.[151,1437,438,1868]" pageId="4" pageNumber="155">
<materialsCitation id="3B2C6460FFC2F219FF5CFDB21A63FDFB" ID-GBIF-Occurrence="3032722318" box="[199,543,582,608]" collectionCode="MUSM, ZSM" pageId="4" pageNumber="155" specimenCount="85" typeStatus="paratype">
<emphasis id="B930B22FFFC2F219FF5CFDB2191FFDFB" bold="true" box="[199,355,582,608]" pageId="4" pageNumber="155">
<specimenCount id="9D42A5B4FFC2F219FF5CFDB2191FFDFB" box="[199,355,582,608]" pageId="4" pageNumber="155" type="generic" typeStatus="Paratypes">85 Paratypes</specimenCount>
</emphasis>
(
<collectionCode id="ED55F6F8FFC2F219FEE9FDB219B0FDFB" box="[370,460,582,608]" pageId="4" pageNumber="155">MUSM</collectionCode>
,
<collectionCode id="ED55F6F8FFC2F219FE43FDB21A6BFDFB" box="[472,535,582,608]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="4" pageNumber="155" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
:
<materialsCitation id="3B2C6460FFC2F219FDB1FDB21B04FDFB" ID-GBIF-Occurrence="3032722308" box="[554,888,582,609]" collectingDate="2018-08-14" collectionCode="MUSM, PER" collectorName="Y. S. Megna &amp; N. Zenteno" country="Peru" elevation="4452" location="Male" pageId="4" pageNumber="155" specimenCount="5" stateProvince="Junin" typeStatus="paratype">
<specimenCount id="9D42A5B4FFC2F219FDB1FDB21A0CFDFA" box="[554,624,582,609]" pageId="4" pageNumber="155" type="generic">5 exs.</specimenCount>
: same data as holotype
</materialsCitation>
;
<materialsCitation id="3B2C6460FFC2F219FC1FFDB21C1DFD1F" ID-GBIF-Occurrence="3032722335" collectingDate="2018-08-11" collectionCode="PER" collectorName="Y. S. Megna &amp; N. Zenteno" country="Peru" elevation="4600" location="Shalacancha" pageId="4" pageNumber="155" specimenCount="55" stateProvince="Junin" typeStatus="paratype">
<specimenCount id="9D42A5B4FFC2F219FC1FFDB21BA4FDFA" box="[900,984,582,609]" pageId="4" pageNumber="155" type="generic">55 exs.</specimenCount>
: “
<collectingCountry id="F3532EADFFC2F219FC75FDB21C54FDFB" box="[1006,1064,582,608]" name="Peru" pageId="4" pageNumber="155">Peru</collectingCountry>
:
<collectingRegion id="4980A0DFFFC2F219FBA8FDB21C0EFDFB" box="[1075,1138,582,608]" country="Peru" name="Junin" pageId="4" pageNumber="155">Junín</collectingRegion>
,
<collectingRegion id="4980A0DFFFC2F219FBE5FDB21CC1FDFB" box="[1150,1213,582,608]" country="Peru" name="Junin" pageId="4" pageNumber="155">Junín</collectingRegion>
,
<location id="8E9B38E6FFC2F219FB52FDB21D24FDFB" LSID="urn:lsid:plazi:treatment:03EDDF2BFFC2F219FF0CFEBE19F8F8D7:8E9B38E6FFC2F219FB52FDB21D24FDFB" box="[1225,1368,582,608]" country="Peru" name="Shalacancha" pageId="4" pageNumber="155" stateProvince="Junin">Shalacancha</location>
,
<quantity id="4CBCC3D8FFC2F219FAFFFDB218D3FD1E" metricMagnitude="3" metricUnit="m" metricValue="4.6" pageId="4" pageNumber="155" unit="m" value="4600.0">
<elevation id="0069890EFFC2F219FAFFFDB218D3FD1E" metricMagnitude="3" metricUnit="m" metricValue="4.6" pageId="4" pageNumber="155" unit="m" value="4600.0">4600 m</elevation>
</quantity>
,
<date id="FFFA48FDFFC2F219FF27FD9E1938FD1F" box="[188,324,618,644]" pageId="4" pageNumber="155" value="2018-08-11">
<collectingDate id="EFBEB115FFC2F219FF27FD9E1938FD1F" box="[188,324,618,644]" pageId="4" pageNumber="155" value="2018-08-11">11.viii.2018</collectingDate>
</date>
, -11.09° -75.879°,
<collectorName id="26B10BEBFFC2F219FD81FD9F1AD9FD1E" box="[538,677,618,645]" pageId="4" pageNumber="155">Y. S. Megna</collectorName>
&amp;
<collectorName id="26B10BEBFFC2F219FD51FD9F1B35FD1E" box="[714,841,619,645]" pageId="4" pageNumber="155">N. Zenteno</collectorName>
(
<collectionCode id="ED55F6F8FFC2F219FCC3FD9E1BF3FD1F" box="[856,911,618,644]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:12821" name="City Museum" pageId="4" pageNumber="155" type="Herbarium">PER</collectionCode>
_YSM_2018_09)”
</materialsCitation>
;
<materialsCitation id="3B2C6460FFC2F219FBF6FD9E1D0DFD33" ID-GBIF-Occurrence="3032722305" collectingDate="2018-08-11" collectionCode="PER" collectorName="Y. S. Megna &amp; N. Zenteno" country="Peru" elevation="4320" location="Shalacancha" pageId="4" pageNumber="155" specimenCount="2" stateProvince="Junin" typeStatus="paratype">
<specimenCount id="9D42A5B4FFC2F219FBF6FD9E1CCEFD1E" box="[1133,1202,618,645]" pageId="4" pageNumber="155" type="generic">2 exs.</specimenCount>
: “
<collectingCountry id="F3532EADFFC2F219FB53FD9E1D7EFD1F" box="[1224,1282,618,644]" name="Peru" pageId="4" pageNumber="155">Peru</collectingCountry>
:
<collectingRegion id="4980A0DFFFC2F219FA96FD9E1D30FD1F" box="[1293,1356,618,644]" country="Peru" name="Junin" pageId="4" pageNumber="155">Junín</collectingRegion>
,
<collectingRegion id="4980A0DFFFC2F219FAC3FD9E1DEBFD1F" box="[1368,1431,618,644]" country="Peru" name="Junin" pageId="4" pageNumber="155">Junín</collectingRegion>
,
<location id="8E9B38E6FFC2F219FF0CFD7A195AFD33" LSID="urn:lsid:plazi:treatment:03EDDF2BFFC2F219FF0CFEBE19F8F8D7:8E9B38E6FFC2F219FF0CFD7A195AFD33" box="[151,294,654,680]" country="Peru" name="Shalacancha" pageId="4" pageNumber="155" stateProvince="Junin">Shalacancha</location>
,
<quantity id="4CBCC3D8FFC2F219FEAEFD7A19ECFD32" box="[309,400,654,681]" metricMagnitude="3" metricUnit="m" metricValue="4.32" pageId="4" pageNumber="155" unit="m" value="4320.0">
<elevation id="0069890EFFC2F219FEAEFD7A19ECFD32" box="[309,400,654,681]" metricMagnitude="3" metricUnit="m" metricValue="4.32" pageId="4" pageNumber="155" unit="m" value="4320.0">4320 m</elevation>
</quantity>
,
<date id="FFFA48FDFFC2F219FE3BFD7A1A54FD33" box="[416,552,654,680]" pageId="4" pageNumber="155" value="2018-08-11">
<collectingDate id="EFBEB115FFC2F219FE3BFD7A1A54FD33" box="[416,552,654,680]" pageId="4" pageNumber="155" value="2018-08-11">11.viii.2018</collectingDate>
</date>
, -11.075° -75.863°,
<collectorName id="26B10BEBFFC2F219FC8DFD7B1BD4FD32" box="[790,936,654,681]" pageId="4" pageNumber="155">Y. S. Megna</collectorName>
&amp;
<collectorName id="26B10BEBFFC2F219FC48FD7B1C2AFD32" box="[979,1110,655,681]" pageId="4" pageNumber="155">N. Zenteno</collectorName>
(
<collectionCode id="ED55F6F8FFC2F219FBF3FD7A1CE3FD33" box="[1128,1183,654,680]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:12821" name="City Museum" pageId="4" pageNumber="155" type="Herbarium">PER</collectionCode>
_YSM_2018_10)”
</materialsCitation>
;
<materialsCitation id="3B2C6460FFC2F219FA1BFD7A196DFD6B" ID-GBIF-Occurrence="3032722348" collectingDate="2018-08-12" collectionCode="PER" collectorName="Y. S. Megna &amp; N. Zenteno" country="Peru" elevation="4099" location="Junin" pageId="4" pageNumber="155" specimenCount="15" stateProvince="Junin" typeStatus="paratype">
<specimenCount id="9D42A5B4FFC2F219FA1BFD7A18BBFD56" pageId="4" pageNumber="155" type="generic">15 exs.</specimenCount>
: “
<collectingCountry id="F3532EADFFC2F219FF46FD46196BFD57" box="[221,279,690,716]" name="Peru" pageId="4" pageNumber="155">Peru</collectingCountry>
:
<collectingRegion id="4980A0DFFFC2F219FEBAFD46191CFD57" box="[289,352,690,716]" country="Peru" name="Junin" pageId="4" pageNumber="155">Junín</collectingRegion>
,
<collectingRegion id="4980A0DFFFC2F219FEF7FD4619D7FD57" box="[364,427,690,716]" country="Peru" name="Junin" pageId="4" pageNumber="155">Junín</collectingRegion>
,
<collectingRegion id="4980A0DFFFC2F219FE2CFD46198AFD57" box="[439,502,690,716]" country="Peru" name="Junin" pageId="4" pageNumber="155">Junín</collectingRegion>
,
<quantity id="4CBCC3D8FFC2F219FD98FD461A26FD56" box="[515,602,690,717]" metricMagnitude="3" metricUnit="m" metricValue="4.098999999999999" pageId="4" pageNumber="155" unit="m" value="4099.0">
<elevation id="0069890EFFC2F219FD98FD461A26FD56" box="[515,602,690,717]" metricMagnitude="3" metricUnit="m" metricValue="4.098999999999999" pageId="4" pageNumber="155" unit="m" value="4099.0">4099 m</elevation>
</quantity>
,
<date id="FFFA48FDFFC2F219FDFDFD461A8CFD57" box="[614,752,690,716]" pageId="4" pageNumber="155" value="2018-08-12">
<collectingDate id="EFBEB115FFC2F219FDFDFD461A8CFD57" box="[614,752,690,716]" pageId="4" pageNumber="155" value="2018-08-12">12.viii.2018</collectingDate>
</date>
, -11.164° -76.008°,
<collectorName id="26B10BEBFFC2F219FC49FD471C21FD56" box="[978,1117,690,717]" pageId="4" pageNumber="155">Y. S. Megna</collectorName>
&amp;
<collectorName id="26B10BEBFFC2F219FB1AFD471D7CFD56" box="[1153,1280,691,717]" pageId="4" pageNumber="155">N. Zenteno</collectorName>
(
<collectionCode id="ED55F6F8FFC2F219FA95FD461D3FFD57" box="[1294,1347,690,716]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:12821" name="City Museum" pageId="4" pageNumber="155" type="Herbarium">PER</collectionCode>
_YSM_ 2018_11)”
</materialsCitation>
;
<materialsCitation id="3B2C6460FFC2F219FE87FD2219B2FC8F" ID-GBIF-Occurrence="3032722332" collectingDate="2018-08-15" collectionCode="PER" collectorName="Y. S. Megna" country="Peru" elevation="4324" location="Shoqui" pageId="4" pageNumber="155" specimenCount="18" stateProvince="Junin" typeStatus="paratype">
<specimenCount id="9D42A5B4FFC2F219FE87FD221912FD6A" box="[284,366,726,753]" pageId="4" pageNumber="155" type="generic">18 exs.</specimenCount>
: “
<collectingCountry id="F3532EADFFC2F219FE18FD2219C1FD6B" box="[387,445,726,752]" name="Peru" pageId="4" pageNumber="155">Peru</collectingCountry>
:
<collectingRegion id="4980A0DFFFC2F219FE5DFD221A79FD6B" box="[454,517,726,752]" country="Peru" name="Junin" pageId="4" pageNumber="155">Junín</collectingRegion>
,
<collectingRegion id="4980A0DFFFC2F219FD8BFD221A33FD6B" box="[528,591,726,752]" country="Peru" name="Junin" pageId="4" pageNumber="155">Junín</collectingRegion>
,
<location id="8E9B38E6FFC2F219FDC1FD221AD5FD6B" LSID="urn:lsid:plazi:treatment:03EDDF2BFFC2F219FF0CFEBE19F8F8D7:8E9B38E6FFC2F219FDC1FD221AD5FD6B" box="[602,681,726,752]" country="Peru" name="Shoqui" pageId="4" pageNumber="155" stateProvince="Junin">Shoqui</location>
,
<quantity id="4CBCC3D8FFC2F219FD26FD221B6FFD6A" box="[701,787,726,753]" metricMagnitude="3" metricUnit="m" metricValue="4.324" pageId="4" pageNumber="155" unit="m" value="4324.0">
<elevation id="0069890EFFC2F219FD26FD221B6FFD6A" box="[701,787,726,753]" metricMagnitude="3" metricUnit="m" metricValue="4.324" pageId="4" pageNumber="155" unit="m" value="4324.0">4324 m</elevation>
</quantity>
,
<date id="FFFA48FDFFC2F219FC85FD221BDBFD6A" box="[798,935,726,753]" pageId="4" pageNumber="155" value="2018-08-15">
<collectingDate id="EFBEB115FFC2F219FC85FD221BDBFD6A" box="[798,935,726,753]" pageId="4" pageNumber="155" value="2018-08-15">15.viii.2018</collectingDate>
</date>
, -11.046° -75.897°,
<collectorName id="26B10BEBFFC2F219FB1CFD231D6CFD6A" box="[1159,1296,726,753]" pageId="4" pageNumber="155">Y. S. Megna</collectorName>
&amp; N. Zenteno (
<collectionCode id="ED55F6F8FFC2F219FF5EFD0E1880FC8F" box="[197,252,762,788]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:12821" name="City Museum" pageId="4" pageNumber="155" type="Herbarium">PER</collectionCode>
_YSM_2018_18)”
</materialsCitation>
;
<materialsCitation id="3B2C6460FFC2F219FE46FD0F1938FCC7" ID-GBIF-Occurrence="3032722310" collectingDate="2018-08-25" collectionCode="PER" collectorName="Y. S. Megna &amp; N. Zenteno" country="Peru" county="Canchis" elevation="4906" location="Cayena" municipality="Checacupe" pageId="4" pageNumber="155" specimenCount="7" stateProvince="Cusco" typeStatus="paratype">
<specimenCount id="9D42A5B4FFC2F219FE46FD0F1A59FC8E" box="[477,549,763,789]" pageId="4" pageNumber="155" type="generic">7 exs.</specimenCount>
: “
<collectingCountry id="F3532EADFFC2F219FDA6FD0E1A0BFC8F" box="[573,631,762,788]" name="Peru" pageId="4" pageNumber="155">Peru</collectingCountry>
:
<collectingRegion id="4980A0DFFFC2F219FD1EFD0E1AB2FC8F" box="[645,718,762,788]" country="Peru" name="Cusco" pageId="4" pageNumber="155">Cusco</collectingRegion>
,
<collectingCounty id="629A16B1FFC2F219FD47FD0E1B45FC8F" box="[732,825,762,788]" pageId="4" pageNumber="155">Canchis</collectingCounty>
,
<collectingMunicipality id="6B9FF447FFC2F219FCD3FD0E1BBAFC8F" box="[840,966,762,788]" pageId="4" pageNumber="155">Checacupe</collectingMunicipality>
,
<location id="8E9B38E6FFC2F219FC4FFD0E1C57FC8F" LSID="urn:lsid:plazi:treatment:03EDDF2BFFC2F219FF0CFEBE19F8F8D7:8E9B38E6FFC2F219FC4FFD0E1C57FC8F" box="[980,1067,762,788]" country="Peru" county="Canchis" municipality="Checacupe" name="Cayena" pageId="4" pageNumber="155" stateProvince="Cusco">Cayena</location>
,
<quantity id="4CBCC3D8FFC2F219FBA2FD0E1CEFFC8E" box="[1081,1171,762,789]" metricMagnitude="3" metricUnit="m" metricValue="4.906" pageId="4" pageNumber="155" unit="m" value="4906.0">
<elevation id="0069890EFFC2F219FBA2FD0E1CEFFC8E" box="[1081,1171,762,789]" metricMagnitude="3" metricUnit="m" metricValue="4.906" pageId="4" pageNumber="155" unit="m" value="4906.0">4906 m</elevation>
</quantity>
,
<date id="FFFA48FDFFC2F219FB39FD0E1D50FC8E" box="[1186,1324,762,789]" pageId="4" pageNumber="155" value="2018-08-25">
<collectingDate id="EFBEB115FFC2F219FB39FD0E1D50FC8E" box="[1186,1324,762,789]" pageId="4" pageNumber="155" value="2018-08-25">25.viii.2018</collectingDate>
</date>
, -14.009° -71.067°,
<collectorName id="26B10BEBFFC2F219FE91FCEB19E0FCA2" box="[266,412,798,825]" pageId="4" pageNumber="155">Y. S. Megna</collectorName>
&amp;
<collectorName id="26B10BEBFFC2F219FE5CFCEB1A35FCA2" box="[455,585,799,825]" pageId="4" pageNumber="155">N. Zenteno</collectorName>
(
<collectionCode id="ED55F6F8FFC2F219FDC0FCEA1AEEFCA3" box="[603,658,798,824]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:12821" name="City Museum" pageId="4" pageNumber="155" type="Herbarium">PER</collectionCode>
_YSM_2018_28)”. All
<typeStatus id="54FFD09FFFC2F219FC3BFCEB1C77FCA2" box="[928,1035,799,825]" pageId="4" pageNumber="155" type="paratype">paratypes</typeStatus>
are provided with our red printed
<typeStatus id="54FFD09FFFC2F219FF0CFCB7188BFCC6" box="[151,247,835,861]" pageId="4" pageNumber="155" type="paratype">paratype</typeStatus>
labels
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35E3DB6FFC2F219FF5CFC92188CF903" pageId="4" pageNumber="155" type="description">
<paragraph id="8BFB6E3DFFC2F219FF5CFC92191BFC52" blockId="4.[151,1437,438,1868]" pageId="4" pageNumber="155">
<emphasis id="B930B22FFFC2F219FF5CFC92199BFC1B" bold="true" box="[199,487,870,896]" pageId="4" pageNumber="155">
Description of
<typeStatus id="54FFD09FFFC2F219FEE1FC92199FFC1B" box="[378,483,870,896]" pageId="4" pageNumber="155" type="holotype">holotype</typeStatus>
.
</emphasis>
Habitus with distinct discontinuity between pronotum and elytra; broadest at about 1/3 of elytral length and body therefore appearing slightly pear shaped (as in
<figureCitation id="137F72B8FFC2F219FB9BFC7E1C02FC3E" box="[1024,1150,906,933]" captionStart="FIGURE 2" captionStartId="7.[151,250,1936,1960]" captionTargetBox="[247,1336,202,1901]" captionTargetId="figure-17@7.[230,1357,181,1911]" captionTargetPageId="7" captionText="FIGURE 2. Dorsal habitus of Liodessus alpinus sp. nov., paratype, male (A), female (B), from Peru: Junín, Junín, Lagoon Estancia Azulmito; L. rhigos sp. nov., paratype male (C) and female (D), both from Peru: Junín, Junín, Lagoon Huacra. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4519047" httpUri="https://zenodo.org/record/4519047/files/figure.png" pageId="4" pageNumber="155">Figs 2A, B</figureCitation>
). TL: 2.0 mm; TL-H:
<quantity id="4CBCC3D8FFC2F219FAE2FC7E18BAFC52" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="4" pageNumber="155" unit="mm" value="1.7">1.7 mm</quantity>
; TW:
<quantity id="4CBCC3D8FFC2F219FE91FC5A191FFC52" box="[266,355,942,969]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="4" pageNumber="155" unit="mm" value="0.9">0.9 mm</quantity>
.
</paragraph>
<paragraph id="8BFB6E3DFFC2F219FF5CFC261B96FC76" blockId="4.[151,1437,438,1868]" box="[199,1002,978,1005]" pageId="4" pageNumber="155">
<emphasis id="B930B22FFFC2F219FF5CFC261923FC77" bold="true" box="[199,351,978,1004]" pageId="4" pageNumber="155">Colouration.</emphasis>
Mostly blackish dorsally and ventrally (as in
<figureCitation id="137F72B8FFC2F219FCFBFC261BA1FC76" box="[864,989,978,1005]" captionStart="FIGURE 2" captionStartId="7.[151,250,1936,1960]" captionTargetBox="[247,1336,202,1901]" captionTargetId="figure-17@7.[230,1357,181,1911]" captionTargetPageId="7" captionText="FIGURE 2. Dorsal habitus of Liodessus alpinus sp. nov., paratype, male (A), female (B), from Peru: Junín, Junín, Lagoon Estancia Azulmito; L. rhigos sp. nov., paratype male (C) and female (D), both from Peru: Junín, Junín, Lagoon Huacra. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4519047" httpUri="https://zenodo.org/record/4519047/files/figure.png" pageId="4" pageNumber="155">Figs 2A, B</figureCitation>
).
</paragraph>
<paragraph id="8BFB6E3DFFC2F219FF5CFC021BE3FBC3" blockId="4.[151,1437,438,1868]" pageId="4" pageNumber="155">
<emphasis id="B930B22FFFC2F219FF5CFC0219DDFB8B" bold="true" box="[199,417,1014,1040]" pageId="4" pageNumber="155">Surface sculpture.</emphasis>
Head with faint microreticulation at base and along hind angles of eyes, frons and clypeus more or less smooth and with few fine setiferous punctures; pronotum shiny with with sparser and fine setiferous punctation; elytra shiny with dense and coarser setiferous punctation.
</paragraph>
<paragraph id="8BFB6E3DFFC2F219FF5CFB961D3FFB5F" blockId="4.[151,1437,438,1868]" pageId="4" pageNumber="155">
<emphasis id="B930B22FFFC2F219FF5CFB961930FBE7" bold="true" box="[199,332,1122,1148]" pageId="4" pageNumber="155">Structures.</emphasis>
Antenna stout. Head without cervical line but few serial punctures instead. Pronotum on each lateral side (or margin) with distinct bead and with distinct and deep basal plicae. Elytron with short and faint basal plicae only visible using diffuse lateral lighting. Metathoracic wings short, about half the length of elytron.
</paragraph>
<paragraph id="8BFB6E3DFFC2F219FF5CFB3A19A9FAAB" blockId="4.[151,1437,438,1868]" pageId="4" pageNumber="155">
<emphasis id="B930B22FFFC2F219FF5CFB3A1941FB73" bold="true" box="[199,317,1230,1256]" pageId="4" pageNumber="155">Genitalia.</emphasis>
Median lobe of aedeagus thick, bulky in lateral view, tip with short nose which is also prominent in ventral view, dorsally on both sides at about mid-length with shallow lateral keels; lateral lobes (parameres) with distal part longish (
<figureCitation id="137F72B8FFC2F219FEF4FAE219B4FAAB" box="[367,456,1302,1328]" captionStart="FIGURE 4" captionStartId="9.[151,250,1954,1978]" captionTargetBox="[174,1412,191,1927]" captionTargetId="figure-17@9.[172,1416,181,1930]" captionTargetPageId="9" captionText="FIGURE 4. Median lobe of aedeagus in ventral and lateral view, and paramere in lateral view of paratypes of Liodessus acollensis (A); L. andinus (B); L. alpinus sp. nov. (C), L. rhigos sp. nov. (D) and L. thespesios sp. nov. (E). Scale bar = 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.4519051" httpUri="https://zenodo.org/record/4519051/files/figure.png" pageId="4" pageNumber="155">Fig. 4C</figureCitation>
).
</paragraph>
<paragraph id="8BFB6E3DFFC2F219FF5CFACE1A48FA5B" blockId="4.[151,1437,438,1868]" pageId="4" pageNumber="155">
<emphasis id="B930B22FFFC2F219FF5CFACE1943FACF" bold="true" box="[199,319,1338,1364]" pageId="4" pageNumber="155">Variation.</emphasis>
TL: 2.0–
<quantity id="4CBCC3D8FFC2F219FE30FACE1A79FACE" box="[427,517,1338,1365]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="4" pageNumber="155" unit="mm" value="2.2">2.2 mm</quantity>
; TL-H:
<quantity id="4CBCC3D8FFC2F219FDF9FACE1A91FACE" box="[610,749,1338,1365]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="4" pageNumber="155" unit="mm" value="1.75" valueMax="1.8" valueMin="1.7">1.7–1.8 mm</quantity>
; TW: 0.9–1.0 mm. Few specimens show an extremely faint, interrupted cervical line. We assessed the length of the metathoracic wings in
<specimenCount id="9D42A5B4FFC2F219FC63FAAA1CEFFAE3" box="[1016,1171,1374,1400]" pageId="4" pageNumber="155" type="generic">10 specimens</specimenCount>
, which all had the same short wing length. While these specimens are certainly not able to fly actively, there might be few fully winged individuals that are capable of flight.
</paragraph>
<paragraph id="8BFB6E3DFFC2F219FF5CFA3E19C0F9B7" blockId="4.[151,1437,438,1868]" pageId="4" pageNumber="155">
<emphasis id="B930B22FFFC2F219FF5CFA3E1959FA7F" bold="true" box="[199,293,1482,1508]" pageId="4" pageNumber="155">Female.</emphasis>
Dorsal surface as in male, few specimens with more or less extensive microreticulation on clypeus and along eyes, pronotal disc and laterally as well as on elytron apically only or from about basal 1/3 up to apex (dullest form depicted in
<figureCitation id="137F72B8FFC2F219FECCF9E619D2F9B7" box="[343,430,1554,1580]" captionStart="FIGURE 2" captionStartId="7.[151,250,1936,1960]" captionTargetBox="[247,1336,202,1901]" captionTargetId="figure-17@7.[230,1357,181,1911]" captionTargetPageId="7" captionText="FIGURE 2. Dorsal habitus of Liodessus alpinus sp. nov., paratype, male (A), female (B), from Peru: Junín, Junín, Lagoon Estancia Azulmito; L. rhigos sp. nov., paratype male (C) and female (D), both from Peru: Junín, Junín, Lagoon Huacra. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4519047" httpUri="https://zenodo.org/record/4519047/files/figure.png" pageId="4" pageNumber="155">Fig. 2B</figureCitation>
).
</paragraph>
<paragraph id="8BFB6E3DFFC2F219FF5CF9C2188CF903" blockId="4.[151,1437,438,1868]" pageId="4" pageNumber="155">
<emphasis id="B930B22FFFC2F219FF5CF9C219ECF9CB" bold="true" box="[199,400,1590,1616]" pageId="4" pageNumber="155">BOLD platform.</emphasis>
We provided 17 entries in the “COLLI” project, all retrieved in one cluster. Assignment to that cluster was unambiguous, meaning all specimens were correctly assigned to this morphologically delineated species.
</paragraph>
</subSubSection>
<subSubSection id="C35E3DB6FFC2F219FF5CF9561C24F926" box="[199,1112,1698,1725]" pageId="4" pageNumber="155" type="etymology">
<paragraph id="8BFB6E3DFFC2F219FF5CF9561C24F926" blockId="4.[151,1437,438,1868]" box="[199,1112,1698,1725]" pageId="4" pageNumber="155">
<emphasis id="B930B22FFFC2F219FF5CF9561932F927" bold="true" box="[199,334,1698,1724]" pageId="4" pageNumber="155">Etymology.</emphasis>
The specific name is the Latin adjective alpinus (of high mountains).
</paragraph>
</subSubSection>
<subSubSection id="C35E3DB6FFC2F219FF5CF93219F8F8D7" pageId="4" pageNumber="155" type="distribution">
<paragraph id="8BFB6E3DFFC2F219FF5CF9321CE7F97A" blockId="4.[151,1437,438,1868]" box="[199,1179,1734,1761]" pageId="4" pageNumber="155">
<emphasis id="B930B22FFFC2F219FF5CF932191DF97B" bold="true" box="[199,353,1734,1760]" pageId="4" pageNumber="155">Distribution.</emphasis>
High Andes of central and southeastern
<collectingCountry id="F3532EADFFC2F219FCBCF9331B26F97A" box="[807,858,1735,1761]" name="Peru" pageId="4" pageNumber="155">Peru</collectingCountry>
(
<figureCitation id="137F72B8FFC2F219FCF1F9321BD2F97A" box="[874,942,1734,1761]" captionStart="FIGURE 7" captionStartId="12.[151,250,1110,1134]" captionTargetBox="[151,1435,181,1086]" captionTargetId="figure-22@12.[151,1435,181,1087]" captionTargetPageId="12" captionText="FIGURE 7. Distribution map of Liodessus species treated in this publication." figureDoi="http://doi.org/10.5281/zenodo.4519057" httpUri="https://zenodo.org/record/4519057/files/figure.png" pageId="4" pageNumber="155">Fig. 7</figureCitation>
), ca.
<quantity id="4CBCC3D8FFC2F219FC71F9321CEAF97A" box="[1002,1174,1734,1761]" metricMagnitude="3" metricUnit="m" metricValue="4.6" metricValueMax="4.9" metricValueMin="4.3" pageId="4" pageNumber="155" unit="m" value="4600.0" valueMax="4900.0" valueMin="4300.0">
<geoCoordinate id="EE7008FAFFC2F219FC71F9321C55F97B" box="[1002,1065,1734,1760]" degrees="4,300" direction="north" orientation="latitude" pageId="4" pageNumber="155" precision="55" value="4.3">4,300</geoCoordinate>
<geoCoordinate id="EE7008FAFFC2F219FBB2F9321C0BF97B" box="[1065,1143,1734,1760]" degrees="4,900" direction="west" orientation="longitude" pageId="4" pageNumber="155" precision="55" value="-4.9">–4,900</geoCoordinate>
m
</quantity>
.
</paragraph>
<paragraph id="8BFB6E3DFFC2F219FF5CF91E19F8F8D7" blockId="4.[151,1437,438,1868]" pageId="4" pageNumber="155">
<emphasis id="B930B22FFFC2F219FF5CF91E1956F89F" bold="true" box="[199,298,1770,1796]" pageId="4" pageNumber="155">Habitat.</emphasis>
Shallow and exposed puddles, collected with strainer out of mats of vegetation (
<figureCitation id="137F72B8FFC2F219FB48F91E1D3AF89E" box="[1235,1350,1770,1797]" captionStart="FIGURE 5" captionStartId="10.[151,250,1812,1836]" captionTargetBox="[155,1432,273,1787]" captionTargetId="figure-54@10.[155,1432,273,1789]" captionTargetPageId="10" captionText="FIGURE 5. Habitat of: Liodessus acollensis (A), Junín, Juaja, Lagoon Tragadero, 3,459 m, (PER_YSM_2018_03); (B) Junín, La Oroya, Nueva Ciudad Morococha, 4,244 m, (PER_YSM_2018_04); L. alpinus sp. nov. (C), Junín, Junín, Shalacancha, 4,320 m, (PER_YSM_2018_10); L. acollensis and L. rhigos sp. nov. (D), Junín, Junín, Lagoon Huacra, 4,461 m, (PER_YSM_2018_ 13); L. acollensis and L. alpinus sp. nov. (E), Junín, Junín, Lagoon Estancia Azulmito, 4,452 m, (PER_YSM_2018_14); L. acollensis and L. rhigos sp. nov. (F), Junín, Junín, Lagoon Yanamyacu, 4,327 m, (PER_YSM_2018_16)." figureDoi="http://doi.org/10.5281/zenodo.4519053" httpUri="https://zenodo.org/record/4519053/files/figure.png" pageId="4" pageNumber="155">Fig.5C, E</figureCitation>
). Other
<taxonomicName id="4C4415BEFFC2F219FF0CF8FA1973F8B3" box="[151,271,1806,1832]" class="Insecta" family="Dytiscidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="family">Dytiscidae</taxonomicName>
associated with this species were species of
<taxonomicName id="4C4415BEFFC2F219FC99F8FA1B86F8B3" authority="Dejean, 1833" authorityName="Dejean" authorityYear="1833" box="[770,1018,1806,1832]" class="Insecta" family="Dytiscidae" genus="Rhantus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="B930B22FFFC2F219FC99F8FA1B22F8B3" box="[770,862,1806,1832]" italics="true" pageId="4" pageNumber="155">Rhantus</emphasis>
Dejean, 1833
</taxonomicName>
and
<taxonomicName id="4C4415BEFFC2F219FBABF8FB1D59F8B3" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" box="[1072,1317,1806,1832]" class="Insecta" family="Dytiscidae" genus="Lancetes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="B930B22FFFC2F219FBABF8FB1CE8F8B3" box="[1072,1172,1807,1832]" italics="true" pageId="4" pageNumber="155">Lancetes</emphasis>
Sharp, 1882
</taxonomicName>
, as well as
<emphasis id="B930B22FFFC2F219FF0CF8C619F8F8D7" box="[151,388,1842,1868]" italics="true" pageId="4" pageNumber="155">
<taxonomicName id="4C4415BEFFC2F219FF0CF8C619FDF8D7" authorityName="Guignot" authorityYear="1955" box="[151,385,1842,1868]" class="Insecta" family="Dytiscidae" genus="Liodessus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="acollensis">Liodessus acollensis</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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