treatments-xml/data/03/94/35/03943535FFF64D0BE6CAE93EFA81FA5D.xml
2024-06-21 12:22:17 +02:00

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<document id="C95AD1EDBA8E3C64B6553626FE7D8E0F" ID-DOI="10.5281/zenodo.4645613" ID-ISSN="1942-1354" ID-Zenodo-Dep="4645613" ID-ZooBank="112A1F0B-1A82-4672-842B-A79A21F251D6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617040325248" checkinUser="felipe" docAuthor="Gordon, Robert D. &amp; Hanley, Guy A." docDate="2014" docId="03943535FFF64D0BE6CAE93EFA81FA5D" docLanguage="en" docName="InsectaMundi.0333.1-91.pdf" docOrigin="Insecta Mundi 2014 (333)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Glaresis zacateca Gordon and Hanley 2014, new species" docType="treatment" docVersion="3" lastPageNumber="49" masterDocId="FFAD4D4DFFC74D39E60AEF1DFF96FF86" masterDocTitle="Systematic revision of American Glaresidae (Coleoptera: Scarabaeoidea)" masterLastPageNumber="91" masterPageNumber="1" pageNumber="48" updateTime="1698895390200" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="0EA5B32ADDF524C55E0178F7FA55617A">Systematic revision of American Glaresidae (Coleoptera: Scarabaeoidea)</mods:title>
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<mods:role id="3CAD3A10849F2CF3AA4B2F2D78F4DE85">
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<mods:namePart id="0AF64DB882947D40831BBF55B7FE77F0">Gordon, Robert D.</mods:namePart>
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<mods:namePart id="23220D7E2E03049A49C49137B3926292">Hanley, Guy A.</mods:namePart>
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<mods:typeOfResource id="FB0E6E0FE01A45D223B551B4459C72C6">text</mods:typeOfResource>
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<mods:title id="1020C3DD290E41FDFF4C99C98E992C60">Insecta Mundi</mods:title>
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<mods:date id="DDD19D36B7A3284069F92FC6A2E99DED">2014</mods:date>
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<mods:number id="E7AF7F0B3E7BA5398A900BB1F927BC11">2014-01-10</mods:number>
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<mods:number id="BDC0B85598CBC1D659E40EEC2DA0F743">2014</mods:number>
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<mods:number id="CB2572194FE7C399FD68C7317A5ECC04">333</mods:number>
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<mods:identifier id="C70FD82C5A47B68006298549D2E1A5AD" type="DOI">10.5281/zenodo.4645613</mods:identifier>
<mods:identifier id="CA831AF6F2197243B65090956E314ACF" type="ISSN">1942-1354</mods:identifier>
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<treatment id="03943535FFF64D0BE6CAE93EFA81FA5D" ID-DOI="http://doi.org/10.5281/zenodo.4645782" ID-Zenodo-Dep="4645782" LSID="urn:lsid:plazi:treatment:03943535FFF64D0BE6CAE93EFA81FA5D" httpUri="http://treatment.plazi.org/id/03943535FFF64D0BE6CAE93EFA81FA5D" lastPageId="50" lastPageNumber="49" pageId="49" pageNumber="48">
<subSubSection id="C327D7A8FFF64D08E6CAE93EFCFCF9BD" box="[192,874,1571,1595]" pageId="49" pageNumber="48" type="nomenclature">
<paragraph id="8B828423FFF64D08E6CAE93EFCFCF9BD" blockId="49.[192,874,1571,1595]" box="[192,874,1571,1595]" pageId="49" pageNumber="48">
<heading id="D0CA334FFFF64D08E6CAE93EFCFCF9BD" bold="true" box="[192,874,1571,1595]" fontSize="10" level="2" pageId="49" pageNumber="48" reason="3">
<emphasis id="B9495831FFF64D08E6CAE93EFCFCF9BD" bold="true" box="[192,874,1571,1595]" pageId="49" pageNumber="48">
<taxonomicName id="4C3DFFA0FFF64D08E6CAE93EFD28F9BD" ID-CoL="3G62D" authority="Gordon and Hanley" authorityName="Gordon and Hanley" authorityYear="2014" box="[192,702,1571,1595]" class="Insecta" family="Glaresidae" genus="Glaresis" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="48" phylum="Arthropoda" rank="species" species="zacateca" status="sp. nov.">
<emphasis id="B9495831FFF64D08E6CAE93EFE38F9BC" bold="true" box="[192,430,1571,1595]" italics="true" pageId="49" pageNumber="48">Glaresis zacateca</emphasis>
Gordon and Hanley
</taxonomicName>
,
<taxonomicNameLabel id="A27AE54AFFF64D08E4C0E93EFCFCF9BD" box="[714,874,1571,1595]" pageId="49" pageNumber="48" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C327D7A8FFF64D0BE6CAE97EFD95FC3D" lastPageId="50" lastPageNumber="49" pageId="49" pageNumber="48" type="description">
<paragraph id="8B828423FFF64D0BE6CAE97EFCAAFCDD" blockId="49.[192,1441,1635,1916]" lastBlockId="50.[192,1441,195,891]" lastPageId="50" lastPageNumber="49" pageId="49" pageNumber="48">
<emphasis id="B9495831FFF64D08E6CAE97EFEF5F9FD" bold="true" box="[192,355,1635,1659]" pageId="49" pageNumber="48">Description</emphasis>
.
<emphasis id="B9495831FFF64D08E77BE97EFE22F9FD" bold="true" box="[369,436,1635,1659]" pageId="49" pageNumber="48">Male</emphasis>
. Length
<quantity id="4CC529C6FFF64D08E42AE97EFDEBF9FD" box="[544,637,1635,1659]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="49" pageNumber="48" unit="mm" value="3.2">3.2 mm</quantity>
, width
<quantity id="4CC529C6FFF64D08E4DEE97EFCA4F9FD" box="[724,818,1635,1659]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="49" pageNumber="48" unit="mm" value="1.6">1.6 mm</quantity>
; body form short, robust, widened from elytral base to apical 1/3 (
<figureCitation id="130698A6FFF64D08E76DE99EFE42F91D" box="[359,468,1667,1691]" captionStart="Figures 33" captionStartId="88.[192,287,1606,1628]" captionTargetBox="[219,1386,214,1582]" captionTargetId="figure-18@88.[192,1440,192,1604]" captionTargetPageId="88" captionText="Figures 33A-33H. Glaresis zacateca, new species. A) Habitus view. B) Clypeus and head surface. C) Elytral striae. D) Pronotal surface and elytra. E) Metasternum. F) Male genitalia. G) Metalegs and trochanter. H) Metatibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4645723" httpUri="https://zenodo.org/record/4645723/files/figure.png" pageId="49" pageNumber="48">Fig. 33A</figureCitation>
). Color dark yellowish brown. Head with clypeal surface coarsely rugose, frons finely rugose, vertex coarsely reticulate, entire head tuberculate, clypeal tubercles coarse, dense, tubercles on frons and vertex slightly smaller, sparse, setae short, barely evident. Clypeal apex truncate, with small tubercles, appearing serrate, lateral angles feebly oblique, acute (
<figureCitation id="130698A6FFF64D08E27BE9FEFB4BF97D" box="[1137,1245,1763,1787]" captionStart="Figures 33" captionStartId="88.[192,287,1606,1628]" captionTargetBox="[219,1386,214,1582]" captionTargetId="figure-18@88.[192,1440,192,1604]" captionTargetPageId="88" captionText="Figures 33A-33H. Glaresis zacateca, new species. A) Habitus view. B) Clypeus and head surface. C) Elytral striae. D) Pronotal surface and elytra. E) Metasternum. F) Male genitalia. G) Metalegs and trochanter. H) Metatibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4645723" httpUri="https://zenodo.org/record/4645723/files/figure.png" pageId="49" pageNumber="48">Fig. 33B</figureCitation>
). Mandible pair symmetrical; mesal tooth strong; lateral prominence strong, pronounced; outer margin angulate. Pronotum with deep, long, transverse fovea in anterior 1/4 extended completely across pronotum, small slightly transverse fovea on each side of middle in anterior 1/2, wide, deep fovea on each side medially near lateral margin, deep central furrow extended from base to transverse furrow; surface densely rugose, with irregular, mostly vertical, dense setae-bearing carinae, setae decumbent, 1/2 length of carinae (
<figureCitation id="130698A6FFF54D0BE322EFDEFA05FF5D" box="[1320,1427,195,219]" captionStart="Figures 33" captionStartId="88.[192,287,1606,1628]" captionTargetBox="[219,1386,214,1582]" captionTargetId="figure-18@88.[192,1440,192,1604]" captionTargetPageId="88" captionText="Figures 33A-33H. Glaresis zacateca, new species. A) Habitus view. B) Clypeus and head surface. C) Elytral striae. D) Pronotal surface and elytra. E) Metasternum. F) Male genitalia. G) Metalegs and trochanter. H) Metatibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4645723" httpUri="https://zenodo.org/record/4645723/files/figure.png" pageId="50" pageNumber="49">Fig. 33D</figureCitation>
). Anterior and lateral pronotal margins not bordered, all margins crenulate. Elytra with surface dull, densely microreticulate; all striae distinctly, strongly carinate, carinae narrowly interrupted adjacent to space between interval punctures, each carinal segment bearing an apical seta about 1/2 length of segment; intervals with deep, slightly rectangular punctures (
<figureCitation id="130698A6FFF54D0BE59EEE5EFB96FEDD" box="[916,1024,323,347]" captionStart="Figures 33" captionStartId="88.[192,287,1606,1628]" captionTargetBox="[219,1386,214,1582]" captionTargetId="figure-18@88.[192,1440,192,1604]" captionTargetPageId="88" captionText="Figures 33A-33H. Glaresis zacateca, new species. A) Habitus view. B) Clypeus and head surface. C) Elytral striae. D) Pronotal surface and elytra. E) Metasternum. F) Male genitalia. G) Metalegs and trochanter. H) Metatibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4645723" httpUri="https://zenodo.org/record/4645723/files/figure.png" pageId="50" pageNumber="49">Fig. 33C</figureCitation>
). Metasternum long, surface dull, finely, densely microreticulate, not tuberculate medially, laterally with some elongate, oblique, setaebearing ridges, median area flat, with feeble carina extended from apical keel 2/3 distance to base; metasternal groove strong, deep, both margins ridged (
<figureCitation id="130698A6FFF54D0BE541EEBEFC22FE3D" box="[843,948,419,443]" captionStart="Figures 33" captionStartId="88.[192,287,1606,1628]" captionTargetBox="[219,1386,214,1582]" captionTargetId="figure-18@88.[192,1440,192,1604]" captionTargetPageId="88" captionText="Figures 33A-33H. Glaresis zacateca, new species. A) Habitus view. B) Clypeus and head surface. C) Elytral striae. D) Pronotal surface and elytra. E) Metasternum. F) Male genitalia. G) Metalegs and trochanter. H) Metatibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4645723" httpUri="https://zenodo.org/record/4645723/files/figure.png" pageId="50" pageNumber="49">Fig. 33E</figureCitation>
). Lateral protibial teeth unevenly spaced, basal two teeth close together. Mesotibia with 4 short, widely spaced spines laterally, tibia distinctly projecting at apex, projection apically acute. Posterior metatrochanteral margin lacking teeth, smooth, posterosuperior metatrochanteral surface with 1 tooth (
<figureCitation id="130698A6FFF54D0BE577ED1EFC79FD9D" box="[893,1007,515,539]" captionStart="Figures 33" captionStartId="88.[192,287,1606,1628]" captionTargetBox="[219,1386,214,1582]" captionTargetId="figure-18@88.[192,1440,192,1604]" captionTargetPageId="88" captionText="Figures 33A-33H. Glaresis zacateca, new species. A) Habitus view. B) Clypeus and head surface. C) Elytral striae. D) Pronotal surface and elytra. E) Metasternum. F) Male genitalia. G) Metalegs and trochanter. H) Metatibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4645723" httpUri="https://zenodo.org/record/4645723/files/figure.png" pageId="50" pageNumber="49">Fig. 33G</figureCitation>
). Metafemoral surface with widely scattered, short, setae-bearing tubercles, microreticulate, dull throughout; width to length ratio 1.0:1.6, with broad flange on anterior margin; posterosuperior margin without teeth. Metatibia narrow at base, abruptly widened before apex, surface entirely, strongly microreticulate, lateral margin with strong, dentate posteromedian projection, margin anterior to projection with about 5 small, acute teeth, margin posterior to projection slightly uneven, medially with short row of about 3 small tubercles in basal 1/3, inner margin smooth, pubescent (
<figureCitation id="130698A6FFF54D0BE453EDDEFD51FD5D" box="[601,711,707,731]" captionStart="Figures 33" captionStartId="88.[192,287,1606,1628]" captionTargetBox="[219,1386,214,1582]" captionTargetId="figure-18@88.[192,1440,192,1604]" captionTargetPageId="88" captionText="Figures 33A-33H. Glaresis zacateca, new species. A) Habitus view. B) Clypeus and head surface. C) Elytral striae. D) Pronotal surface and elytra. E) Metasternum. F) Male genitalia. G) Metalegs and trochanter. H) Metatibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4645723" httpUri="https://zenodo.org/record/4645723/files/figure.png" pageId="50" pageNumber="49">Fig. 33H</figureCitation>
). Apical margin of male 5th abdominal ventrite feebly emarginate medially. Male genitalia short, basal piece much shorter than parameres, proximal end straight; median lobe slightly shorter than parameres, twice as wide at middle as paramere, slightly curved upward in apical 1/3, apex broadly rounded in ventral view; paramere tapered in lateral view, not flattened, tapered from base to nearly truncate apex (
<figureCitation id="130698A6FFF54D0BE4C6EC5EFCB9FCDD" box="[716,815,835,859]" captionStart="Figures 33" captionStartId="88.[192,287,1606,1628]" captionTargetBox="[219,1386,214,1582]" captionTargetId="figure-18@88.[192,1440,192,1604]" captionTargetPageId="88" captionText="Figures 33A-33H. Glaresis zacateca, new species. A) Habitus view. B) Clypeus and head surface. C) Elytral striae. D) Pronotal surface and elytra. E) Metasternum. F) Male genitalia. G) Metalegs and trochanter. H) Metatibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4645723" httpUri="https://zenodo.org/record/4645723/files/figure.png" pageId="50" pageNumber="49">Fig 33F</figureCitation>
).
</paragraph>
<paragraph id="8B828423FFF54D0BE6FAEC7EFE76FCFD" blockId="50.[192,1441,195,891]" box="[240,480,867,891]" pageId="50" pageNumber="49">
<emphasis id="B9495831FFF54D0BE6FAEC7EFEC5FCFD" bold="true" box="[240,339,867,891]" pageId="50" pageNumber="49">Female</emphasis>
. Unknown.
</paragraph>
<paragraph id="8B828423FFF54D0BE6CAECBEFD95FC3D" blockId="50.[192,515,931,955]" box="[192,515,931,955]" pageId="50" pageNumber="49">
<emphasis id="B9495831FFF54D0BE6CAECBEFED4FC3D" bold="true" box="[192,322,931,955]" pageId="50" pageNumber="49">Variation</emphasis>
.
<typeStatus id="54863A81FFF54D0BE758ECBEFE57FC3D" box="[338,449,931,955]" pageId="50" pageNumber="49" type="holotype">Holotype</typeStatus>
only.
</paragraph>
</subSubSection>
<subSubSection id="C327D7A8FFF54D0BE6CAECFEFD35FB9D" pageId="50" pageNumber="49" type="materials_examined">
<paragraph id="8B828423FFF54D0BE6CAECFEFD35FB9D" blockId="50.[192,1440,995,1051]" pageId="50" pageNumber="49">
<emphasis id="B9495831FFF54D0BE6CAECFEFE13FC7D" bold="true" box="[192,389,995,1019]" pageId="50" pageNumber="49">Type material</emphasis>
.
<materialsCitation id="3B558E7EFFF54D0BE792ECFEFD35FB9D" collectingDate="1962-05-10" collectionCode="CMNC" collectorName="F. D. Parker L. A. Stange Collectors" country="Mexico" location="Zac. Mex" pageId="50" pageNumber="49" specimenCount="1" stateProvince="Zacatecas" typeStatus="holotype">
<typeStatus id="54863A81FFF54D0BE792ECFEFD91FC7D" box="[408,519,995,1019]" pageId="50" pageNumber="49" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="F32AC4B3FFF54D0BE46AECFEFD2CFC7D" box="[608,698,995,1019]" name="Mexico" pageId="50" pageNumber="49">Mexico</collectingCountry>
.
<emphasis id="B9495831FFF54D0BE4C3ECFEFCC7FC7D" bold="true" box="[713,849,995,1019]" pageId="50" pageNumber="49">
<collectingRegion id="49F94AC1FFF54D0BE4C3ECFEFCC7FC7D" box="[713,849,995,1019]" country="Mexico" name="Zacatecas" pageId="50" pageNumber="49">Zacatecas</collectingRegion>
</emphasis>
:
<locationDeviation id="9A309D01FFF54D0BE56EECFEFBD9FC7D" box="[868,1103,995,1019]" location="Fresnillo" pageId="50" pageNumber="49">
<quantity id="4CC529C6FFF54D0BE56EECFEFC22FC7D" box="[868,948,995,1019]" metricMagnitude="4" metricUnit="m" metricValue="1.609344" pageId="50" pageNumber="49" unit="mi" value="10.0">10 mi.</quantity>
N Fresnillo
</locationDeviation>
,
<location id="8EE2D2F8FFF54D0BE255ECF9FB45FC7D" LSID="urn:lsid:plazi:treatment:03943535FFF64D0BE6CAE93EFA81FA5D:8EE2D2F8FFF54D0BE255ECF9FB45FC7D" box="[1119,1235,996,1019]" country="Mexico" name="Zac. Mex" pageId="50" pageNumber="49" stateProvince="Zacatecas">Zac. Mex</location>
,
<date id="FF83A2E3FFF54D0BE2E8ECFEFAF4FC7D" box="[1250,1378,995,1019]" pageId="50" pageNumber="49" value="1962-05-10">
<collectingDate id="EFC75B0BFFF54D0BE2E8ECFEFAF4FC7D" box="[1250,1378,995,1019]" pageId="50" pageNumber="49" value="1962-05-10">V-10 1962</collectingDate>
</date>
,
<collectorName id="26C8E1F5FFF54D0BE37BECF9FDBAFB9D" pageId="50" pageNumber="49">F.D. Parker L.A. Stange Collectors</collectorName>
(
<collectionCode id="ED2C1CE6FFF54D0BE436EB1EFD01FB9D" box="[572,663,1027,1051]" pageId="50" pageNumber="49">CMNC</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C327D7A8FFF54D0BE6CAEB5EFDA5FA1D" pageId="50" pageNumber="49" type="discussion">
<paragraph id="8B828423FFF54D0BE6CAEB5EFEB6FA9D" blockId="50.[192,1440,1091,1435]" pageId="50" pageNumber="49">
<emphasis id="B9495831FFF54D0BE6CAEB5EFEADFBDD" bold="true" box="[192,315,1091,1115]" pageId="50" pageNumber="49">Remarks</emphasis>
. The unique
<typeStatus id="54863A81FFF54D0BE7E8EB5EFDDDFBDD" box="[482,587,1091,1115]" pageId="50" pageNumber="49" type="holotype">holotype</typeStatus>
is most similar to
<taxonomicName id="4C3DFFA0FFF54D0BE53FEB59FC21FBDD" authorityName="Brown. A" authorityYear="1928" box="[821,951,1091,1115]" class="Insecta" family="Glaresidae" genus="Glaresis" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="49" phylum="Arthropoda" rank="species" species="confusa">
<emphasis id="B9495831FFF54D0BE53FEB59FCDCFBDD" box="[821,842,1092,1115]" italics="true" pageId="50" pageNumber="49">G</emphasis>
.
<emphasis id="B9495831FFF54D0BE550EB5EFC21FBDD" box="[858,951,1091,1115]" italics="true" pageId="50" pageNumber="49">confusa</emphasis>
</taxonomicName>
in most characteristics, including male genitalia. It is distinguished from that species by a lack of tubercles on vertex of head; metatibia with complete row of small, median tubercles; apex of male 5th abdominal ventrite weakly emarginate; and central
<collectingCountry id="F32AC4B3FFF54D0BE716EBBEFEE4FB3D" box="[284,370,1187,1211]" name="Mexico" pageId="50" pageNumber="49">Mexico</collectingCountry>
type locality. Within Group C it is most similar to
<taxonomicName id="4C3DFFA0FFF54D0BE5C2EBB9FBDEFB3D" authorityName="Gordon and Hanley" authorityYear="2014" box="[968,1096,1187,1211]" class="Insecta" family="Glaresidae" genus="Glaresis" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="49" phylum="Arthropoda" rank="species" species="donaldi">
<emphasis id="B9495831FFF54D0BE5C2EBB9FC4BFB3D" box="[968,989,1188,1211]" italics="true" pageId="50" pageNumber="49">G</emphasis>
.
<emphasis id="B9495831FFF54D0BE5E3EBBEFBDEFB3D" box="[1001,1096,1187,1211]" italics="true" pageId="50" pageNumber="49">donaldi</emphasis>
</taxonomicName>
because both have a distinct posteromedian metatibial projection, but
<taxonomicName id="4C3DFFA0FFF54D0BE4A6EBD9FCA7FB5D" authorityName="Gordon and Hanley" authorityYear="2014" box="[684,817,1220,1243]" class="Insecta" family="Glaresidae" genus="Glaresis" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="49" phylum="Arthropoda" rank="species" species="zacateca">
<emphasis id="B9495831FFF54D0BE4A6EBD9FD57FB5D" box="[684,705,1220,1243]" italics="true" pageId="50" pageNumber="49">G</emphasis>
.
<emphasis id="B9495831FFF54D0BE4C4EBD9FCA7FB5D" box="[718,817,1220,1243]" italics="true" pageId="50" pageNumber="49">zacateca</emphasis>
</taxonomicName>
is distinguished from the former species by presence of tubercles on vertex of head; lack of reticulate carinae on metasternum; and metafemoral surface nearly smooth.
</paragraph>
<paragraph id="8B828423FFF54D0BE6FAEA3EFDA5FA1D" blockId="50.[192,1440,1091,1435]" pageId="50" pageNumber="49">
This is another member of Group C in a string of species extending from northern
<collectingCountry id="F32AC4B3FFF54D0BE2D5EA3EFAA3FABD" box="[1247,1333,1315,1339]" name="Mexico" pageId="50" pageNumber="49">Mexico</collectingCountry>
to
<collectingCountry id="F32AC4B3FFF54D0BE351EA3EFF6FFADD" name="Costa Rica" pageId="50" pageNumber="49">Costa Rica</collectingCountry>
. Species are now known from Coahuila, Zacatecas, Puebla,
<collectingCountry id="F32AC4B3FFF54D0BE5DBEA5EFBCDFADD" box="[977,1115,1347,1371]" name="Guatemala" pageId="50" pageNumber="49">Guatemala</collectingCountry>
, and
<collectingCountry id="F32AC4B3FFF54D0BE296EA5EFAB7FADD" box="[1180,1313,1347,1371]" name="Costa Rica" pageId="50" pageNumber="49">Costa Rica</collectingCountry>
, nearly all from unique specimens. This pattern seems to indicate the presence of other unknown southern species, particularly in central
<collectingCountry id="F32AC4B3FFF54D0BE7DEEA9EFDB8FA1D" box="[468,558,1411,1435]" name="Mexico" pageId="50" pageNumber="49">Mexico</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C327D7A8FFF54D0BE6CAEADEFA81FA5D" box="[192,1303,1475,1499]" pageId="50" pageNumber="49" type="etymology">
<paragraph id="8B828423FFF54D0BE6CAEADEFA81FA5D" blockId="50.[192,1303,1475,1499]" box="[192,1303,1475,1499]" pageId="50" pageNumber="49">
<emphasis id="B9495831FFF54D0BE6CAEADEFEC5FA5D" bold="true" box="[192,339,1475,1499]" pageId="50" pageNumber="49">Etymology</emphasis>
. This species is named for the state of
<collectingRegion id="49F94AC1FFF54D0BE52DEAD9FC0BFA5D" box="[807,925,1476,1499]" country="Mexico" name="Zacatecas" pageId="50" pageNumber="49">Zacatecas</collectingRegion>
where the
<typeStatus id="54863A81FFF54D0BE22EEADEFB1BFA5D" box="[1060,1165,1475,1499]" pageId="50" pageNumber="49" type="holotype">holotype</typeStatus>
was found.
</paragraph>
</subSubSection>
</treatment>
</document>