359 lines
70 KiB
XML
359 lines
70 KiB
XML
<document id="9108E8D9E47091D42E74B2B84452126F" ID-CLB-Dataset="31822" ID-DOI="10.11646/zootaxa.4337.2.1" ID-GBIF-Dataset="bca0e227-e4fb-4f93-b99b-c8794b19de8d" ID-ISSN="1175-5326" ID-Zenodo-Dep="1018631" ID-ZooBank="A712D90D-FCD1-49C9-928F-21C95814815B" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1508314185452" checkinUser="plazi" docAuthor="Behan-Pelletier, Valerie M." docDate="2017" docId="9D02C578FF9CFFA7FF12CB4FFAC8FC8D" docLanguage="en" docName="zootaxa.4337.2.1.pdf" docOrigin="Zootaxa 4337 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Protectoribates Behan-Pelletier 2017, gen. nov." docType="treatment" docVersion="8" lastPageNumber="155" masterDocId="613BBD00FF9EFFA3FF85CC7CFF9CFFD0" masterDocTitle="Tegoribatidae of North America, with proposal of Protectoribates gen. nov., and new species (Acari, Oribatida, Tegoribatidae)" pageNumber="153" updateTime="1698469699374" updateUser="ExternalLinkService">
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<mods:titleInfo id="11B2F925E6776760B62DA4EEFE65F0DC">
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<mods:title id="64F0BF996CFA263445E6D7603A7914CD">Tegoribatidae of North America, with proposal of Protectoribates gen. nov., and new species (Acari, Oribatida, Tegoribatidae)</mods:title>
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<mods:name id="37C7AF948C3BBA9F56776FBA9C778A73" type="personal">
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<mods:namePart id="DE13F9AB54EE183C8A437003FBB2F54B">Behan-Pelletier, Valerie M.</mods:namePart>
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<mods:typeOfResource id="2D28488E39620EF6F9312A74E338BD3A">text</mods:typeOfResource>
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<mods:title id="F6CCD900F20B8C9610FC92B0B71A585F">Zootaxa</mods:title>
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<mods:part id="9836C05A2ADFBEC8902FB7B4FC822B57">
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<mods:date id="439EE43E9E6930833286591F4A5F713D">2017</mods:date>
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<mods:number id="93DA41C28C7A9C02E40B45C3039A4D1A">2017-10-18</mods:number>
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<mods:number id="93BEFCD52DA859270ED3E3AA0D746374">4337</mods:number>
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<mods:classification id="3501D2AF14887A0DAFFE140C2EEA9CC2">journal volume</mods:classification>
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<mods:identifier id="4AABE7F0711EF51A322BE8D5BBBEC77F" type="DOI">10.11646/zootaxa.4337.2.1</mods:identifier>
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<treatment id="9D02C578FF9CFFA7FF12CB4FFAC8FC8D" ID-DOI="http://doi.org/10.5281/zenodo.5664399" ID-GBIF-Taxon="135450332" ID-Zenodo-Dep="5664399" LSID="urn:lsid:plazi:treatment:9D02C578FF9CFFA7FF12CB4FFAC8FC8D" httpUri="http://treatment.plazi.org/id/9D02C578FF9CFFA7FF12CB4FFAC8FC8D" lastPageId="4" lastPageNumber="155" pageId="2" pageNumber="153">
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<subSubSection id="5DB127E5FF9CFFA1FF12CB4FFE9CF8BE" pageId="2" pageNumber="153" type="nomenclature">
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<paragraph id="1514746EFF9CFFA1FF12CB4FFE5EF89D" blockId="2.[151,450,1843,1902]" box="[151,450,1843,1869]" pageId="2" pageNumber="153">
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<heading id="4E5CC302FF9CFFA1FF12CB4FFE5EF89D" bold="true" box="[151,450,1843,1869]" fontSize="11" level="1" pageId="2" pageNumber="153" reason="1">
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<emphasis id="27DFA87CFF9CFFA1FF12CB4FFE5EF89D" bold="true" box="[151,450,1843,1869]" pageId="2" pageNumber="153">
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<taxonomicName id="D2AB0FEDFF9CFFA1FF12CB4FFED1F89D" ID-CoL="63PHT" authority="Behan-Pelletier, 2017" authorityName="Behan-Pelletier" authorityYear="2017" box="[151,333,1843,1869]" class="Arachnida" family="Tegoribatidae" genus="Protectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="153" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="27DFA87CFF9CFFA1FF12CB4FFED1F89D" bold="true" box="[151,333,1843,1869]" italics="true" pageId="2" pageNumber="153">Protectoribates</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="3CEC1507FF9CFFA1FED1CB4FFE5EF89D" box="[340,450,1843,1869]" pageId="2" pageNumber="153" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="1514746EFF9CFFA1FF12CB29FE9CF8BE" blockId="2.[151,450,1843,1902]" box="[151,256,1877,1902]" pageId="2" pageNumber="153">
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<figureCitation id="8D9068EBFF9CFFA1FF12CB29FE9CF8BE" box="[151,256,1877,1902]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 2 – 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStartId-0="6.[151,250,1985,2007]" captionStartId-1="7.[151,264,1946,1968]" captionStartId-2="8.[151,250,1974,1996]" captionStartId-3="9.[151,250,1917,1939]" captionStartId-4="10.[151,250,1882,1904]" captionStartId-5="11.[151,250,1861,1883]" captionTargetBox-0="[193,1370,231,1921]" captionTargetBox-1="[335,1280,208,1896]" captionTargetBox-2="[313,1242,203,1931]" captionTargetBox-3="[207,1364,193,1894]" captionTargetBox-4="[246,1312,207,1841]" captionTargetBox-5="[207,1359,193,1809]" captionTargetId-0="figure@6.[189,1387,195,1957]" captionTargetId-1="figure@7.[253,1333,193,1924]" captionTargetId-2="figure@8.[236,1331,193,1943]" captionTargetId-3="figure@9.[207,1364,193,1894]" captionTargetId-4="figure@10.[220,1367,193,1860]" captionTargetId-5="figure@11.[207,1359,193,1828]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionTargetPageId-4="10" captionTargetPageId-5="11" captionText-0="FIGURE 1. Protectoribates occidentalis sp. nov., adult female, dorsal aspect, legs not illustrated; dorsophragmata indicated by arrows. Scale bar = 50 µm." captionText-1="FIGURES 2 – 3. Protectoribates occidentalis sp. nov., adult female, legs not illustrated; 2, ventral aspect; 3, lateral aspect, only alveolus of seta ex illustrated, gnathosoma not illustrated. Scale bar = 50 µm." captionText-2="FIGURE 4. Protectoribates occidentalis sp. nov., adult female, A, leg I, B, leg II, C, leg IV, all antiaxial aspect. Legs I and II, trochanter not illustrated. Scale bar = 20 µm." captionText-3="FIGURE 5 A – E. Protectoribates occidentalis sp. nov., scanning electron microscope images of adult female from California; A. habitus dorsal; B, habitus ventral, with arrow to custodium; C, frontal aspect; D, dorsolateral of prodorsum, with arrow to spindle end of bothridial seta (bo), second arrow to genal incision and third arrow to carina on tibia II; E, ventrolateral of prodorsum, with arrow to ridge on genal tooth. Scale bars: A, B = 25 µm, C – E = 15 µm." captionText-4="FIGURE 6. Protectoribates occidentalis sp. nov., tritonymph, dorsal aspect. Scale bar = 50 µm." captionText-5="FIGURE 7. Protectoribates occidentalis sp. nov., differential interference contrast microscope images, A – E adults; A, bothridium and bothridial seta (5 layers), with arrow to nerve canal of seta le on lamella; insert of bothridium showing scales; B, custodium and discidium (3 layers), with arrow to broad, rounded anterior edge of custodium; C, porose organ Ah, lateral view, with arrow to surrounding cerotegument (3 layers); D, dissected specimen with Ah in direct view (2 layers); E, genu, tibia and tarsus II, with white arrow to dentate carina on tibia (2 layers) and narrow black arrow to seta pv ”; F, protonymph (2 layers), dorsal view. Scale bar: A = 25 µm; B = 10 µm; D = 7.5 µm; F = 50 µm." httpUri-0="https://zenodo.org/record/1018633/files/figure.png" httpUri-1="https://zenodo.org/record/1018635/files/figure.png" httpUri-2="https://zenodo.org/record/1018637/files/figure.png" httpUri-3="https://zenodo.org/record/1018639/files/figure.png" httpUri-4="https://zenodo.org/record/1018641/files/figure.png" httpUri-5="https://zenodo.org/record/1018645/files/figure.png" pageId="2" pageNumber="153">Figs. 1–7</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="5DB127E5FF9CFFA1FF12CBE1FD02F863" box="[151,670,1949,1971]" pageId="2" pageNumber="153" type="materials_examined">
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<paragraph id="1514746EFF9CFFA1FF12CBE1FD02F863" blockId="2.[151,670,1949,1971]" box="[151,670,1949,1971]" pageId="2" pageNumber="153">
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<emphasis id="27DFA87CFF9CFFA1FF12CBE1FE80F863" bold="true" box="[151,284,1949,1971]" pageId="2" pageNumber="153">Type species</emphasis>
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:
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<taxonomicName id="D2AB0FEDFF9CFFA1FEAFCBE1FDD4F863" authority="Behan-Pelletier, 2017" authorityName="Behan-Pelletier" authorityYear="2017" box="[298,584,1949,1971]" class="Arachnida" family="Tegoribatidae" genus="Protectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="occidentalis" status="sp. nov.">
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<emphasis id="27DFA87CFF9CFFA1FEAFCBE1FDD4F863" box="[298,584,1949,1971]" italics="true" pageId="2" pageNumber="153">Protectoribates occidentalis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="3CEC1507FF9CFFA1FDC8CBE1FD02F863" box="[589,670,1949,1971]" pageId="2" pageNumber="153" rank="species">
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<emphasis id="27DFA87CFF9CFFA1FDC8CBE1FD02F863" bold="true" box="[589,670,1949,1971]" pageId="2" pageNumber="153">sp. nov.</emphasis>
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</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="5DB127E5FF9DFFA0FF12CCEBFCD2FC8F" pageId="3" pageNumber="154" type="diagnosis">
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<paragraph id="1514746EFF9DFFA0FF12CCEBFD4EFD77" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="154">
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<emphasis id="27DFA87CFF9DFFA0FF12CCEBFE8DFF60" bold="true" box="[151,273,151,176]" pageId="3" pageNumber="154">Diagnosis.</emphasis>
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<emphasis id="27DFA87CFF9DFFA0FE98CCEBFEC5FF60" box="[285,345,151,176]" italics="true" pageId="3" pageNumber="154">Adult</emphasis>
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. With character states of
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<taxonomicName id="D2AB0FEDFF9DFFA0FD16CCEBFCB0FF60" authorityName="Grandjean" authorityYear="1954" box="[659,812,151,176]" class="Arachnida" family="Tegoribatidae" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="154" phylum="Arthropoda" rank="family">Tegoribatidae</taxonomicName>
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(
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<bibRefCitation id="713A099FFF9DFFA0FCC5CCEBFB9AFF60" author="Grandjean, F." box="[832,1030,151,176]" journalOrPublisher="Bulletin du Museum national d´Histoire naturelle, 2 e Serie" pageId="3" pageNumber="154" pagination="286 - 293" part="25" refString="Grandjean, F. (1953) Observations sur les Oribates (26 e serie). Bulletin du Museum national d´Histoire naturelle, 2 e Serie, 25 (3), 286 - 293." title="Observations sur les Oribates (26 e serie)" type="journal article" year="1953">Grandjean (1953</bibRefCitation>
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,
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<bibRefCitation id="713A099FFF9DFFA0FB9DCCEBFBC8FF60" author="Grandjean, F." box="[1048,1108,151,176]" journalOrPublisher="Bulletin de la Societe Zoologique de France" pageId="3" pageNumber="154" pagination="421 - 446" part="78" refString="Grandjean, F. (1954) Essai de classification des Oribates (Acariens). Bulletin de la Societe Zoologique de France, 78, 421 - 446." title="Essai de classification des Oribates (Acariens)" type="journal article" year="1954">1954</bibRefCitation>
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),
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<bibRefCitation id="713A099FFF9DFFA0FBE8CCEBFABEFF60" author="Woolley, T. A." box="[1133,1314,151,176]" journalOrPublisher="Transaction of the American Microscopical Society" pageId="3" pageNumber="154" pagination="409 – 412" part="84" publicationUrl="https://doi.org/10.2307/3224729" refString="Woolley, T. A. (1965) A New Species of Tegoribates from Colorado with Notes on the Genus (Acarina: Oribatei, Tegoribatidae). Transaction of the American Microscopical Society, 84, 409 - 412. https: // doi. org / 10.2307 / 3224729" title="A New Species of Tegoribates from Colorado with Notes on the Genus (Acarina: Oribatei, Tegoribatidae)" type="journal article" year="1965">Woolley (1965)</bibRefCitation>
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<bibRefCitation id="713A099FFF9DFFA0FAAACCEBFECBFF05" author="Fredes, N. A. & Martinez, P. A." journalOrPublisher="Zootaxa" pageId="3" pageNumber="154" pagination="191 - 200" part="4098" publicationUrl="https://doi.org/10.11646/zootaxa.4098.1.10" refString="Fredes, N. A. & Martinez, P. A. (2016) Revision of Physobates (Acari, Oribatida, Tegoribatidae) and redescription of Physobates spinipes Hammer, 1962 with comments on tegoribatid genera. Zootaxa, 4098 (1), 191 - 200. https: // doi. org / 10.11646 / zootaxa. 4098.1.10" title="Revision of Physobates (Acari, Oribatida, Tegoribatidae) and redescription of Physobates spinipes Hammer, 1962 with comments on tegoribatid genera" type="journal article" year="2016">Fredes & Martínez (2016)</bibRefCitation>
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, and see below). Species comprising this genus are unique among poronotic
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<taxonomicName id="D2AB0FEDFF9DFFA0FA80CCC0FEC3FF28" authority="(Grandjean 1954)" authorityName="Hull" authorityYear="1918" baseAuthorityName="Grandjean" baseAuthorityYear="1954" class="Arachnida" infraOrder="Brachypylina" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="154" phylum="Arthropoda" rank="subTribe" subTribe="Brachypylina">
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Brachypylina (
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<bibRefCitation id="713A099FFF9DFFA0FF1BCCA3FECBFF28" author="Grandjean, F." box="[158,343,223,248]" journalOrPublisher="Bulletin de la Societe Zoologique de France" pageId="3" pageNumber="154" pagination="421 - 446" part="78" refString="Grandjean, F. (1954) Essai de classification des Oribates (Acariens). Bulletin de la Societe Zoologique de France, 78, 421 - 446." title="Essai de classification des Oribates (Acariens)" type="journal article" year="1954">Grandjean 1954</bibRefCitation>
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)
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</taxonomicName>
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by having the following combination of character states. Rostrum with medial keel extending posteriorly; rostrum rounded anteriorly, without medial incision. Lamella long, fused, with V-shaped indentation anteriorly, covering all of prodorsum in dorsal aspect, without longitudinal line medially. Tutorium narrow lamelliform, cusp triangular. Genal tooth well-developed, pointed distally, with ridge along length. Bothridium with lateral and medial scales. Porose areas
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<emphasis id="27DFA87CFF9DFFA0FD17CD13FD2DFE58" box="[658,689,367,392]" italics="true" pageId="3" pageNumber="154">Ad</emphasis>
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and
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<emphasis id="27DFA87CFF9DFFA0FD6ECD0CFC8CFE57" box="[747,784,368,391]" italics="true" pageId="3" pageNumber="154">Am</emphasis>
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present,
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<emphasis id="27DFA87CFF9DFFA0FCFFCD13FC0FFE58" box="[890,915,367,392]" italics="true" pageId="3" pageNumber="154">Al</emphasis>
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absent,
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<emphasis id="27DFA87CFF9DFFA0FC71CD13FB88FE58" box="[1012,1044,367,392]" italics="true" pageId="3" pageNumber="154">Ah</emphasis>
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expressed as deep, round saccule. Dorsophragmata medially separated. Octotaxic system as 4 pairs of porose areas. Notogaster with non-overlapping posterior tectum. Pteromorph well-developed, without hinge; lenticulus absent; hexagonal pattern absent anteriorly on notogaster. Notogaster with 10 pairs of setae; setae not apobasic. Epimere I without necklace of tubercles. Epimeral setation 3-1-3-3. Custodium broadly rounded distally, or not. Sejugal apodeme and apodeme 3 welldeveloped. Genital setation 6 pairs, with 3 setae along anterior margin of each plate. Band of darkened integument posterior to genital plates. Postanal porose area absent. Subcapitulum diarthric, without tectum on mentum. Axillary saccule present. Legs monodactylous. Tibia II with large, dentate carina anterodorsally. Tibia IV with solenidion. Femur I without bend at level of seta
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<emphasis id="27DFA87CFF9DFFA0FD38CEF3FD57FD78" box="[701,715,655,680]" italics="true" pageId="3" pageNumber="154">d</emphasis>
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.
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</paragraph>
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<paragraph id="1514746EFF9DFFA0FF42CEC8FCD2FC8F" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="154">
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<emphasis id="27DFA87CFF9DFFA0FF42CEC8FEE0FD1C" box="[199,380,692,717]" italics="true" pageId="3" pageNumber="154">Juvenile instars</emphasis>
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Apheredermous; line of dehiscense incomplete; integument plicate, without hysterosomal macrosclerites or excentrosclerites, cerotegument as tightly packed platelets,
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<quantity id="D253D98BFF9DFFA0FB8ECEA4FBA4FD20" box="[1035,1080,727,752]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="154" unit="in" value="1.0">1 in</quantity>
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diameter, covering all of body and leg segments. Prodorsal porose region present. Gastronotal setation unideficient; nymphs with 15 pairs of setae (
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<emphasis id="27DFA87CFF9DFFA0FF25CF5CFF2CFCEC" box="[160,176,800,828]" italics="true" pageId="3" pageNumber="154">f1</emphasis>
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absent). Gastronotal setae of
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<emphasis id="27DFA87CFF9DFFA0FE78CF5DFDA0FCE8" box="[509,572,799,824]" italics="true" pageId="3" pageNumber="154">c, d, l</emphasis>
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and
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<emphasis id="27DFA87CFF9DFFA0FDF6CF5DFD1DFCE8" box="[627,641,801,824]" italics="true" pageId="3" pageNumber="154">p</emphasis>
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series monomorphic, short, without barbs, acuminate (
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<figureCitation id="8D9068EBFF9DFFA0FB6FCF63FAB3FCE8" box="[1258,1327,799,824]" captionStart="FIGURE 6" captionStartId="10.[151,250,1882,1904]" captionTargetBox="[246,1312,207,1841]" captionTargetId="figure@10.[220,1367,193,1860]" captionTargetPageId="10" captionText="FIGURE 6. Protectoribates occidentalis sp. nov., tritonymph, dorsal aspect. Scale bar = 50 µm." httpUri="https://zenodo.org/record/1018641/files/figure.png" pageId="3" pageNumber="154">Fig. 6</figureCitation>
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). Rostral, lamellar, and interlamellar setae subequal to gastronotal setae.
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</paragraph>
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</subSubSection>
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<subSubSection id="5DB127E5FF9DFFA0FF42CF16FCC6F936" pageId="3" pageNumber="154" type="description">
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<paragraph id="1514746EFF9DFFA0FF42CF16FD97FC1B" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="154">
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<emphasis id="27DFA87CFF9DFFA0FF42CF16FECDFC53" bold="true" box="[199,337,874,899]" pageId="3" pageNumber="154">Description</emphasis>
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.
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<emphasis id="27DFA87CFF9DFFA0FEE7CF16FE32FC53" bold="true" box="[354,430,874,899]" pageId="3" pageNumber="154">Adult.</emphasis>
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<emphasis id="27DFA87CFF9DFFA0FE32CF10FDA9FC53" box="[439,565,876,899]" italics="true" pageId="3" pageNumber="154">Integument</emphasis>
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. Cerotegument present laterally on podosoma, finely and densely granulate (
|
||
<figureCitation id="8D9068EBFF9DFFA0FF1ACFF2FF67FC77" box="[159,251,910,935]" captionStart="FIGURE 5" captionStartId="9.[151,250,1917,1939]" captionTargetBox="[207,1364,193,1894]" captionTargetId="figure@9.[207,1364,193,1894]" captionTargetPageId="9" captionText="FIGURE 5 A – E. Protectoribates occidentalis sp. nov., scanning electron microscope images of adult female from California; A. habitus dorsal; B, habitus ventral, with arrow to custodium; C, frontal aspect; D, dorsolateral of prodorsum, with arrow to spindle end of bothridial seta (bo), second arrow to genal incision and third arrow to carina on tibia II; E, ventrolateral of prodorsum, with arrow to ridge on genal tooth. Scale bars: A, B = 25 µm, C – E = 15 µm." httpUri="https://zenodo.org/record/1018639/files/figure.png" pageId="3" pageNumber="154">Fig. 5E</figureCitation>
|
||
). Integument microtuberculate throughout; U-shaped depression extending from coxisternal region to posterior of anal plates (
|
||
<figureCitation id="8D9068EBFF9DFFA0FE23CFCEFE62FC1B" box="[422,510,946,971]" captionStart="FIGURE 5" captionStartId="9.[151,250,1917,1939]" captionTargetBox="[207,1364,193,1894]" captionTargetId="figure@9.[207,1364,193,1894]" captionTargetPageId="9" captionText="FIGURE 5 A – E. Protectoribates occidentalis sp. nov., scanning electron microscope images of adult female from California; A. habitus dorsal; B, habitus ventral, with arrow to custodium; C, frontal aspect; D, dorsolateral of prodorsum, with arrow to spindle end of bothridial seta (bo), second arrow to genal incision and third arrow to carina on tibia II; E, ventrolateral of prodorsum, with arrow to ridge on genal tooth. Scale bars: A, B = 25 µm, C – E = 15 µm." httpUri="https://zenodo.org/record/1018639/files/figure.png" pageId="3" pageNumber="154">Fig. 5B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="1514746EFF9DFFA0FF42CFAAFD8AFB3B" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="154">
|
||
<emphasis id="27DFA87CFF9DFFA0FF42CFAAFED8FC3F" box="[199,324,982,1007]" italics="true" pageId="3" pageNumber="154">Prodorsum</emphasis>
|
||
. Rostrum with smooth margin, without medial incision (
|
||
<figureCitation id="8D9068EBFF9DFFA0FC42CFAAFBBCFC3F" box="[967,1056,982,1007]" captionStart="FIGURE 5" captionStartId="9.[151,250,1917,1939]" captionTargetBox="[207,1364,193,1894]" captionTargetId="figure@9.[207,1364,193,1894]" captionTargetPageId="9" captionText="FIGURE 5 A – E. Protectoribates occidentalis sp. nov., scanning electron microscope images of adult female from California; A. habitus dorsal; B, habitus ventral, with arrow to custodium; C, frontal aspect; D, dorsolateral of prodorsum, with arrow to spindle end of bothridial seta (bo), second arrow to genal incision and third arrow to carina on tibia II; E, ventrolateral of prodorsum, with arrow to ridge on genal tooth. Scale bars: A, B = 25 µm, C – E = 15 µm." httpUri="https://zenodo.org/record/1018639/files/figure.png" pageId="3" pageNumber="154">Fig. 5C</figureCitation>
|
||
); broadly rounded keel extending from margin posteriorly to underneath lamella (
|
||
<figureCitation id="8D9068EBFF9DFFA0FD51CF86FCFBFBC3" box="[724,871,1018,1043]" captionStart="FIGURE 5" captionStartId="9.[151,250,1917,1939]" captionTargetBox="[207,1364,193,1894]" captionTargetId="figure@9.[207,1364,193,1894]" captionTargetPageId="9" captionText="FIGURE 5 A – E. Protectoribates occidentalis sp. nov., scanning electron microscope images of adult female from California; A. habitus dorsal; B, habitus ventral, with arrow to custodium; C, frontal aspect; D, dorsolateral of prodorsum, with arrow to spindle end of bothridial seta (bo), second arrow to genal incision and third arrow to carina on tibia II; E, ventrolateral of prodorsum, with arrow to ridge on genal tooth. Scale bars: A, B = 25 µm, C – E = 15 µm." httpUri="https://zenodo.org/record/1018639/files/figure.png" pageId="3" pageNumber="154">Figs. 5C, D</figureCitation>
|
||
). Rostral seta heavily barbed laterally, directed anteromedially (
|
||
<figureCitation id="8D9068EBFF9DFFA0FECBC862FE0FFBE7" box="[334,403,1054,1079]" captionStart="FIGURES 2 – 3" captionStartId="7.[151,264,1946,1968]" captionTargetBox="[335,1280,208,1896]" captionTargetId="figure@7.[253,1333,193,1924]" captionTargetPageId="7" captionText="FIGURES 2 – 3. Protectoribates occidentalis sp. nov., adult female, legs not illustrated; 2, ventral aspect; 3, lateral aspect, only alveolus of seta ex illustrated, gnathosoma not illustrated. Scale bar = 50 µm." httpUri="https://zenodo.org/record/1018635/files/figure.png" pageId="3" pageNumber="154">Fig. 3</figureCitation>
|
||
). Lamella long, pair fused, covering all of prodorsum in dorsal aspect (
|
||
<figureCitation id="8D9068EBFF9DFFA0FB34C862FB63FBE7" box="[1201,1279,1054,1079]" captionStart="FIGURE 1" captionStartId="6.[151,250,1985,2007]" captionTargetBox="[193,1370,231,1921]" captionTargetId="figure@6.[189,1387,195,1957]" captionTargetPageId="6" captionText="FIGURE 1. Protectoribates occidentalis sp. nov., adult female, dorsal aspect, legs not illustrated; dorsophragmata indicated by arrows. Scale bar = 50 µm." httpUri="https://zenodo.org/record/1018633/files/figure.png" pageId="3" pageNumber="154">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="8D9068EBFF9DFFA0FA88C862FAAEFBE7" box="[1293,1330,1054,1079]" captionStart="FIGURE 5" captionStartId="9.[151,250,1917,1939]" captionTargetBox="[207,1364,193,1894]" captionTargetId="figure@9.[207,1364,193,1894]" captionTargetPageId="9" captionText="FIGURE 5 A – E. Protectoribates occidentalis sp. nov., scanning electron microscope images of adult female from California; A. habitus dorsal; B, habitus ventral, with arrow to custodium; C, frontal aspect; D, dorsolateral of prodorsum, with arrow to spindle end of bothridial seta (bo), second arrow to genal incision and third arrow to carina on tibia II; E, ventrolateral of prodorsum, with arrow to ridge on genal tooth. Scale bars: A, B = 25 µm, C – E = 15 µm." httpUri="https://zenodo.org/record/1018639/files/figure.png" pageId="3" pageNumber="154">5A</figureCitation>
|
||
), without longitudinal line medially; nerve canal for seta
|
||
<emphasis id="27DFA87CFF9DFFA0FD21C83EFD24FB8B" box="[676,696,1090,1115]" italics="true" pageId="3" pageNumber="154">le</emphasis>
|
||
evident (
|
||
<figureCitation id="8D9068EBFF9DFFA0FCA5C83EFCE5FB8B" box="[800,889,1090,1115]" captionStart="FIGURE 7" captionStartId="11.[151,250,1861,1883]" captionTargetBox="[207,1359,193,1809]" captionTargetId="figure@11.[207,1359,193,1828]" captionTargetPageId="11" captionText="FIGURE 7. Protectoribates occidentalis sp. nov., differential interference contrast microscope images, A – E adults; A, bothridium and bothridial seta (5 layers), with arrow to nerve canal of seta le on lamella; insert of bothridium showing scales; B, custodium and discidium (3 layers), with arrow to broad, rounded anterior edge of custodium; C, porose organ Ah, lateral view, with arrow to surrounding cerotegument (3 layers); D, dissected specimen with Ah in direct view (2 layers); E, genu, tibia and tarsus II, with white arrow to dentate carina on tibia (2 layers) and narrow black arrow to seta pv ”; F, protonymph (2 layers), dorsal view. Scale bar: A = 25 µm; B = 10 µm; D = 7.5 µm; F = 50 µm." httpUri="https://zenodo.org/record/1018645/files/figure.png" pageId="3" pageNumber="154">Fig. 7A</figureCitation>
|
||
). Seta
|
||
<emphasis id="27DFA87CFF9DFFA0FC46C83EFC4BFB8B" box="[963,983,1090,1115]" italics="true" pageId="3" pageNumber="154">le</emphasis>
|
||
arising anterodorsally on lamella. Fused lamella with short, V-shaped indentation anteriorly; concave posterolaterally, accommodating bothridium (
|
||
<figureCitation id="8D9068EBFF9DFFA0FAC2C81AFA09FBAF" box="[1351,1429,1126,1151]" captionStart="FIGURE 1" captionStartId="6.[151,250,1985,2007]" captionTargetBox="[193,1370,231,1921]" captionTargetId="figure@6.[189,1387,195,1957]" captionTargetPageId="6" captionText="FIGURE 1. Protectoribates occidentalis sp. nov., adult female, dorsal aspect, legs not illustrated; dorsophragmata indicated by arrows. Scale bar = 50 µm." httpUri="https://zenodo.org/record/1018633/files/figure.png" pageId="3" pageNumber="154">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="8D9068EBFF9DFFA0FF12C8F7FF20FB73" box="[151,188,1163,1187]" captionStart="FIGURE 7" captionStartId="11.[151,250,1861,1883]" captionTargetBox="[207,1359,193,1809]" captionTargetId="figure@11.[207,1359,193,1828]" captionTargetPageId="11" captionText="FIGURE 7. Protectoribates occidentalis sp. nov., differential interference contrast microscope images, A – E adults; A, bothridium and bothridial seta (5 layers), with arrow to nerve canal of seta le on lamella; insert of bothridium showing scales; B, custodium and discidium (3 layers), with arrow to broad, rounded anterior edge of custodium; C, porose organ Ah, lateral view, with arrow to surrounding cerotegument (3 layers); D, dissected specimen with Ah in direct view (2 layers); E, genu, tibia and tarsus II, with white arrow to dentate carina on tibia (2 layers) and narrow black arrow to seta pv ”; F, protonymph (2 layers), dorsal view. Scale bar: A = 25 µm; B = 10 µm; D = 7.5 µm; F = 50 µm." httpUri="https://zenodo.org/record/1018645/files/figure.png" pageId="3" pageNumber="154">7A</figureCitation>
|
||
, and insert). Translamella absent. Exobothridial seta present (alveolus only indicated on
|
||
<figureCitation id="8D9068EBFF9DFFA0FB1EC8F6FB42FB73" box="[1179,1246,1162,1187]" captionStart="FIGURES 2 – 3" captionStartId="7.[151,264,1946,1968]" captionTargetBox="[335,1280,208,1896]" captionTargetId="figure@7.[253,1333,193,1924]" captionTargetPageId="7" captionText="FIGURES 2 – 3. Protectoribates occidentalis sp. nov., adult female, legs not illustrated; 2, ventral aspect; 3, lateral aspect, only alveolus of seta ex illustrated, gnathosoma not illustrated. Scale bar = 50 µm." httpUri="https://zenodo.org/record/1018635/files/figure.png" pageId="3" pageNumber="154">Fig. 3</figureCitation>
|
||
). Bothridial wall expanded medially and laterally, with scales, without lateral indentation (
|
||
<figureCitation id="8D9068EBFF9DFFA0FC51C8D2FBB3FB17" box="[980,1071,1198,1223]" captionStart="FIGURE 7" captionStartId="11.[151,250,1861,1883]" captionTargetBox="[207,1359,193,1809]" captionTargetId="figure@11.[207,1359,193,1828]" captionTargetPageId="11" captionText="FIGURE 7. Protectoribates occidentalis sp. nov., differential interference contrast microscope images, A – E adults; A, bothridium and bothridial seta (5 layers), with arrow to nerve canal of seta le on lamella; insert of bothridium showing scales; B, custodium and discidium (3 layers), with arrow to broad, rounded anterior edge of custodium; C, porose organ Ah, lateral view, with arrow to surrounding cerotegument (3 layers); D, dissected specimen with Ah in direct view (2 layers); E, genu, tibia and tarsus II, with white arrow to dentate carina on tibia (2 layers) and narrow black arrow to seta pv ”; F, protonymph (2 layers), dorsal view. Scale bar: A = 25 µm; B = 10 µm; D = 7.5 µm; F = 50 µm." httpUri="https://zenodo.org/record/1018645/files/figure.png" pageId="3" pageNumber="154">Fig. 7A</figureCitation>
|
||
, insert). Porose area
|
||
<emphasis id="27DFA87CFF9DFFA0FA98C8D2FAA0FB17" box="[1309,1340,1198,1223]" italics="true" pageId="3" pageNumber="154">Ad</emphasis>
|
||
present. Dorsophragmata separate (
|
||
<figureCitation id="8D9068EBFF9DFFA0FE41C8AEFD99FB3B" box="[452,517,1234,1259]" captionStart="FIGURE 1" captionStartId="6.[151,250,1985,2007]" captionTargetBox="[193,1370,231,1921]" captionTargetId="figure@6.[189,1387,195,1957]" captionTargetPageId="6" captionText="FIGURE 1. Protectoribates occidentalis sp. nov., adult female, dorsal aspect, legs not illustrated; dorsophragmata indicated by arrows. Scale bar = 50 µm." httpUri="https://zenodo.org/record/1018633/files/figure.png" pageId="3" pageNumber="154">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="1514746EFF9DFFA0FF42C884FAABFAAA" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="154">
|
||
<emphasis id="27DFA87CFF9DFFA0FF42C884FEDEFADF" box="[199,322,1272,1295]" italics="true" pageId="3" pageNumber="154">Notogaster</emphasis>
|
||
. Longer than wide; with immovable pteromorphs having smooth margins, without dens anteroventrally (
|
||
<figureCitation id="8D9068EBFF9DFFA0FED5C966FE35FAE3" box="[336,425,1306,1331]" captionStart="FIGURE 5" captionStartId="9.[151,250,1917,1939]" captionTargetBox="[207,1364,193,1894]" captionTargetId="figure@9.[207,1364,193,1894]" captionTargetPageId="9" captionText="FIGURE 5 A – E. Protectoribates occidentalis sp. nov., scanning electron microscope images of adult female from California; A. habitus dorsal; B, habitus ventral, with arrow to custodium; C, frontal aspect; D, dorsolateral of prodorsum, with arrow to spindle end of bothridial seta (bo), second arrow to genal incision and third arrow to carina on tibia II; E, ventrolateral of prodorsum, with arrow to ridge on genal tooth. Scale bars: A, B = 25 µm, C – E = 15 µm." httpUri="https://zenodo.org/record/1018639/files/figure.png" pageId="3" pageNumber="154">Fig. 5B</figureCitation>
|
||
). Anterior margin straight, transverse, without ridges in region lateral to bothridium (
|
||
<figureCitation id="8D9068EBFF9DFFA0FAEFC966FF39FA87" captionStart="FIGURE 1" captionStartId="6.[151,250,1985,2007]" captionTargetBox="[193,1370,231,1921]" captionTargetId="figure@6.[189,1387,195,1957]" captionTargetPageId="6" captionText="FIGURE 1. Protectoribates occidentalis sp. nov., adult female, dorsal aspect, legs not illustrated; dorsophragmata indicated by arrows. Scale bar = 50 µm." httpUri="https://zenodo.org/record/1018633/files/figure.png" pageId="3" pageNumber="154">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="8D9068EBFF9DFFA0FF36C942FF45FA87" box="[179,217,1342,1367]" captionStart="FIGURE 5" captionStartId="9.[151,250,1917,1939]" captionTargetBox="[207,1364,193,1894]" captionTargetId="figure@9.[207,1364,193,1894]" captionTargetPageId="9" captionText="FIGURE 5 A – E. Protectoribates occidentalis sp. nov., scanning electron microscope images of adult female from California; A. habitus dorsal; B, habitus ventral, with arrow to custodium; C, frontal aspect; D, dorsolateral of prodorsum, with arrow to spindle end of bothridial seta (bo), second arrow to genal incision and third arrow to carina on tibia II; E, ventrolateral of prodorsum, with arrow to ridge on genal tooth. Scale bars: A, B = 25 µm, C – E = 15 µm." httpUri="https://zenodo.org/record/1018639/files/figure.png" pageId="3" pageNumber="154">5A</figureCitation>
|
||
). Without subsurface anteromedial hexagonal pattern. Octotaxic system expressed as 4 pairs of porose areas (
|
||
<figureCitation id="8D9068EBFF9DFFA0FF1AC91EFF7DFAAB" box="[159,225,1378,1403]" captionStart="FIGURE 1" captionStartId="6.[151,250,1985,2007]" captionTargetBox="[193,1370,231,1921]" captionTargetId="figure@6.[189,1387,195,1957]" captionTargetPageId="6" captionText="FIGURE 1. Protectoribates occidentalis sp. nov., adult female, dorsal aspect, legs not illustrated; dorsophragmata indicated by arrows. Scale bar = 50 µm." httpUri="https://zenodo.org/record/1018633/files/figure.png" pageId="3" pageNumber="154">Fig. 1</figureCitation>
|
||
); without sexual dimorphism. With 10 pairs of short setae; distance
|
||
<emphasis id="27DFA87CFF9DFFA0FC5AC91EFB88FAAF" box="[991,1044,1378,1407]" italics="true" pageId="3" pageNumber="154">h1-h1</emphasis>
|
||
wider than distance
|
||
<emphasis id="27DFA87CFF9DFFA0FB79C918FAACFAAF" box="[1276,1328,1380,1407]" italics="true" pageId="3" pageNumber="154">p1-p1</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="1514746EFF9DFFA0FF42C9F5FC02F986" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="154">
|
||
<emphasis id="27DFA87CFF9DFFA0FF42C9F5FD92FA72" box="[199,526,1417,1442]" italics="true" pageId="3" pageNumber="154">Lateral Region of Podosoma</emphasis>
|
||
. Genal tooth, pointed triangular, with longitudinal ridge (
|
||
<figureCitation id="8D9068EBFF9DFFA0FB18C9F5FB72FA72" box="[1181,1262,1417,1442]" captionStart="FIGURES 2 – 3" captionStartId="7.[151,264,1946,1968]" captionTargetBox="[335,1280,208,1896]" captionTargetId="figure@7.[253,1333,193,1924]" captionTargetPageId="7" captionText="FIGURES 2 – 3. Protectoribates occidentalis sp. nov., adult female, legs not illustrated; 2, ventral aspect; 3, lateral aspect, only alveolus of seta ex illustrated, gnathosoma not illustrated. Scale bar = 50 µm." httpUri="https://zenodo.org/record/1018635/files/figure.png" pageId="3" pageNumber="154">Figs. 3</figureCitation>
|
||
,
|
||
<figureCitation id="8D9068EBFF9DFFA0FB7EC9F5FA82FA72" box="[1275,1310,1417,1442]" captionStart="FIGURE 5" captionStartId="9.[151,250,1917,1939]" captionTargetBox="[207,1364,193,1894]" captionTargetId="figure@9.[207,1364,193,1894]" captionTargetPageId="9" captionText="FIGURE 5 A – E. Protectoribates occidentalis sp. nov., scanning electron microscope images of adult female from California; A. habitus dorsal; B, habitus ventral, with arrow to custodium; C, frontal aspect; D, dorsolateral of prodorsum, with arrow to spindle end of bothridial seta (bo), second arrow to genal incision and third arrow to carina on tibia II; E, ventrolateral of prodorsum, with arrow to ridge on genal tooth. Scale bars: A, B = 25 µm, C – E = 15 µm." httpUri="https://zenodo.org/record/1018639/files/figure.png" pageId="3" pageNumber="154">5E</figureCitation>
|
||
). Tutorium narrow, lamelliform, lying parallel to dorsal contour of prodorsum in lateral aspect, seta
|
||
<emphasis id="27DFA87CFF9DFFA0FB26C9D2FB25FA15" box="[1187,1209,1454,1477]" italics="true" pageId="3" pageNumber="154">ro</emphasis>
|
||
inserted at base of tutorial cusp (
|
||
<figureCitation id="8D9068EBFF9DFFA0FEB9C9ADFE0CFA3A" box="[316,400,1489,1514]" captionStart="FIGURES 2 – 3" captionStartId="7.[151,264,1946,1968]" captionTargetBox="[335,1280,208,1896]" captionTargetId="figure@7.[253,1333,193,1924]" captionTargetPageId="7" captionText="FIGURES 2 – 3. Protectoribates occidentalis sp. nov., adult female, legs not illustrated; 2, ventral aspect; 3, lateral aspect, only alveolus of seta ex illustrated, gnathosoma not illustrated. Scale bar = 50 µm." httpUri="https://zenodo.org/record/1018635/files/figure.png" pageId="3" pageNumber="154">Figs. 3</figureCitation>
|
||
,
|
||
<figureCitation id="8D9068EBFF9DFFA0FE24C9ADFE5AFA3A" box="[417,454,1489,1514]" captionStart="FIGURE 5" captionStartId="9.[151,250,1917,1939]" captionTargetBox="[207,1364,193,1894]" captionTargetId="figure@9.[207,1364,193,1894]" captionTargetPageId="9" captionText="FIGURE 5 A – E. Protectoribates occidentalis sp. nov., scanning electron microscope images of adult female from California; A. habitus dorsal; B, habitus ventral, with arrow to custodium; C, frontal aspect; D, dorsolateral of prodorsum, with arrow to spindle end of bothridial seta (bo), second arrow to genal incision and third arrow to carina on tibia II; E, ventrolateral of prodorsum, with arrow to ridge on genal tooth. Scale bars: A, B = 25 µm, C – E = 15 µm." httpUri="https://zenodo.org/record/1018639/files/figure.png" pageId="3" pageNumber="154">5D</figureCitation>
|
||
). Pedotectum I convex dorsally, without ventral depression (
|
||
<figureCitation id="8D9068EBFF9DFFA0FB10C9ADFB43FA3A" box="[1173,1247,1489,1514]" captionStart="FIGURES 2 – 3" captionStartId="7.[151,264,1946,1968]" captionTargetBox="[335,1280,208,1896]" captionTargetId="figure@7.[253,1333,193,1924]" captionTargetPageId="7" captionText="FIGURES 2 – 3. Protectoribates occidentalis sp. nov., adult female, legs not illustrated; 2, ventral aspect; 3, lateral aspect, only alveolus of seta ex illustrated, gnathosoma not illustrated. Scale bar = 50 µm." httpUri="https://zenodo.org/record/1018635/files/figure.png" pageId="3" pageNumber="154">Fig. 3</figureCitation>
|
||
). Pedotectum II present, without tubercle close to body wall. Custodium present, with long, free distal region, broadly rounded or not (
|
||
<figureCitation id="8D9068EBFF9DFFA0FF49CA65FE81F9E2" box="[204,285,1561,1586]" captionStart="FIGURES 2 – 3" captionStartId="7.[151,264,1946,1968]" captionTargetBox="[335,1280,208,1896]" captionTargetId="figure@7.[253,1333,193,1924]" captionTargetPageId="7" captionText="FIGURES 2 – 3. Protectoribates occidentalis sp. nov., adult female, legs not illustrated; 2, ventral aspect; 3, lateral aspect, only alveolus of seta ex illustrated, gnathosoma not illustrated. Scale bar = 50 µm." httpUri="https://zenodo.org/record/1018635/files/figure.png" pageId="3" pageNumber="154">Figs. 3</figureCitation>
|
||
,
|
||
<figureCitation id="8D9068EBFF9DFFA0FEA9CA66FECCF9E2" box="[300,336,1562,1586]" captionStart="FIGURE 7" captionStartId="11.[151,250,1861,1883]" captionTargetBox="[207,1359,193,1809]" captionTargetId="figure@11.[207,1359,193,1828]" captionTargetPageId="11" captionText="FIGURE 7. Protectoribates occidentalis sp. nov., differential interference contrast microscope images, A – E adults; A, bothridium and bothridial seta (5 layers), with arrow to nerve canal of seta le on lamella; insert of bothridium showing scales; B, custodium and discidium (3 layers), with arrow to broad, rounded anterior edge of custodium; C, porose organ Ah, lateral view, with arrow to surrounding cerotegument (3 layers); D, dissected specimen with Ah in direct view (2 layers); E, genu, tibia and tarsus II, with white arrow to dentate carina on tibia (2 layers) and narrow black arrow to seta pv ”; F, protonymph (2 layers), dorsal view. Scale bar: A = 25 µm; B = 10 µm; D = 7.5 µm; F = 50 µm." httpUri="https://zenodo.org/record/1018645/files/figure.png" pageId="3" pageNumber="154">7B</figureCitation>
|
||
). Discidium strongly curved lamina (
|
||
<figureCitation id="8D9068EBFF9DFFA0FD79CA65FCD2F9E2" box="[764,846,1561,1586]" captionStart="FIGURES 2 – 3" captionStartId="7.[151,264,1946,1968]" captionTargetBox="[335,1280,208,1896]" captionTargetId="figure@7.[253,1333,193,1924]" captionTargetPageId="7" captionText="FIGURES 2 – 3. Protectoribates occidentalis sp. nov., adult female, legs not illustrated; 2, ventral aspect; 3, lateral aspect, only alveolus of seta ex illustrated, gnathosoma not illustrated. Scale bar = 50 µm." httpUri="https://zenodo.org/record/1018635/files/figure.png" pageId="3" pageNumber="154">Figs. 3</figureCitation>
|
||
,
|
||
<figureCitation id="8D9068EBFF9DFFA0FCD9CA66FC1CF9E2" box="[860,896,1562,1586]" captionStart="FIGURE 7" captionStartId="11.[151,250,1861,1883]" captionTargetBox="[207,1359,193,1809]" captionTargetId="figure@11.[207,1359,193,1828]" captionTargetPageId="11" captionText="FIGURE 7. Protectoribates occidentalis sp. nov., differential interference contrast microscope images, A – E adults; A, bothridium and bothridial seta (5 layers), with arrow to nerve canal of seta le on lamella; insert of bothridium showing scales; B, custodium and discidium (3 layers), with arrow to broad, rounded anterior edge of custodium; C, porose organ Ah, lateral view, with arrow to surrounding cerotegument (3 layers); D, dissected specimen with Ah in direct view (2 layers); E, genu, tibia and tarsus II, with white arrow to dentate carina on tibia (2 layers) and narrow black arrow to seta pv ”; F, protonymph (2 layers), dorsal view. Scale bar: A = 25 µm; B = 10 µm; D = 7.5 µm; F = 50 µm." httpUri="https://zenodo.org/record/1018645/files/figure.png" pageId="3" pageNumber="154">7B</figureCitation>
|
||
). Porose area
|
||
<emphasis id="27DFA87CFF9DFFA0FBA6CA65FBA0F9E2" box="[1059,1084,1561,1586]" italics="true" pageId="3" pageNumber="154">Al</emphasis>
|
||
absent. Humerosejugal porose organs
|
||
<emphasis id="27DFA87CFF9DFFA0FF6DCA42FE91F985" box="[232,269,1598,1621]" italics="true" pageId="3" pageNumber="154">Am</emphasis>
|
||
and
|
||
<emphasis id="27DFA87CFF9DFFA0FEC6CA41FEFFF986" box="[323,355,1597,1622]" italics="true" pageId="3" pageNumber="154">Ah</emphasis>
|
||
present,
|
||
<emphasis id="27DFA87CFF9DFFA0FE4CCA42FE72F985" box="[457,494,1598,1621]" italics="true" pageId="3" pageNumber="154">Am</emphasis>
|
||
porose area,
|
||
<emphasis id="27DFA87CFF9DFFA0FD06CA41FD3EF986" box="[643,674,1597,1622]" italics="true" pageId="3" pageNumber="154">Ah</emphasis>
|
||
saccule (
|
||
<figureCitation id="8D9068EBFF9DFFA0FC8ECA41FC0DF986" box="[779,913,1597,1622]" captionStart="FIGURE 7" captionStartId="11.[151,250,1861,1883]" captionTargetBox="[207,1359,193,1809]" captionTargetId="figure@11.[207,1359,193,1828]" captionTargetPageId="11" captionText="FIGURE 7. Protectoribates occidentalis sp. nov., differential interference contrast microscope images, A – E adults; A, bothridium and bothridial seta (5 layers), with arrow to nerve canal of seta le on lamella; insert of bothridium showing scales; B, custodium and discidium (3 layers), with arrow to broad, rounded anterior edge of custodium; C, porose organ Ah, lateral view, with arrow to surrounding cerotegument (3 layers); D, dissected specimen with Ah in direct view (2 layers); E, genu, tibia and tarsus II, with white arrow to dentate carina on tibia (2 layers) and narrow black arrow to seta pv ”; F, protonymph (2 layers), dorsal view. Scale bar: A = 25 µm; B = 10 µm; D = 7.5 µm; F = 50 µm." httpUri="https://zenodo.org/record/1018645/files/figure.png" pageId="3" pageNumber="154">Figs. 7C, D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="1514746EFF9DFFA0FF42CA1DFCC6F936" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="154">
|
||
<emphasis id="27DFA87CFF9DFFA0FF42CA1DFE89F9AA" box="[199,277,1633,1658]" italics="true" pageId="3" pageNumber="154">Ventral</emphasis>
|
||
. Epimere I without anterior necklace of small tubercles. Epimeral setal formula 3-1-3-3. Genital plates with 6 setae, 3 on anterior margin; 1 pair aggenital setae; 3 pairs adanal setae and 2 pairs anal setae (
|
||
<figureCitation id="8D9068EBFF9DFFA0FAC3CAF9FA13F94E" box="[1350,1423,1669,1694]" captionStart="FIGURES 2 – 3" captionStartId="7.[151,264,1946,1968]" captionTargetBox="[335,1280,208,1896]" captionTargetId="figure@7.[253,1333,193,1924]" captionTargetPageId="7" captionText="FIGURES 2 – 3. Protectoribates occidentalis sp. nov., adult female, legs not illustrated; 2, ventral aspect; 3, lateral aspect, only alveolus of seta ex illustrated, gnathosoma not illustrated. Scale bar = 50 µm." httpUri="https://zenodo.org/record/1018635/files/figure.png" pageId="3" pageNumber="154">Fig. 2</figureCitation>
|
||
). Lyrifissure
|
||
<emphasis id="27DFA87CFF9DFFA0FE9BCAD5FEDEF912" box="[286,322,1705,1730]" italics="true" pageId="3" pageNumber="154">iad</emphasis>
|
||
at anterolateral edge of anal plate. Postanal porose area absent. Band of darker integument extending transversely between genital and anal plates (
|
||
<figureCitation id="8D9068EBFF9DFFA0FC8DCAB1FCD0F936" box="[776,844,1741,1766]" captionStart="FIGURES 2 – 3" captionStartId="7.[151,264,1946,1968]" captionTargetBox="[335,1280,208,1896]" captionTargetId="figure@7.[253,1333,193,1924]" captionTargetPageId="7" captionText="FIGURES 2 – 3. Protectoribates occidentalis sp. nov., adult female, legs not illustrated; 2, ventral aspect; 3, lateral aspect, only alveolus of seta ex illustrated, gnathosoma not illustrated. Scale bar = 50 µm." httpUri="https://zenodo.org/record/1018635/files/figure.png" pageId="3" pageNumber="154">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5DB127E5FF9DFFA0FF42CA8DFA00F832" pageId="3" pageNumber="154" type="reference_group">
|
||
<paragraph id="1514746EFF9DFFA0FF42CA8DFAFDF882" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="154">
|
||
<taxonomicName id="D2AB0FEDFF9DFFA0FF42CA8DFECAF8DA" box="[199,342,1777,1802]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="3" pageNumber="154" phylum="Nematoda" rank="genus">
|
||
<emphasis id="27DFA87CFF9DFFA0FF42CA8DFECAF8DA" box="[199,342,1777,1802]" italics="true" pageId="3" pageNumber="154">Gnathosoma</emphasis>
|
||
</taxonomicName>
|
||
. Chelicera chelate-dentate. Mentum without tectum, without recurved ridge distally (
|
||
<figureCitation id="8D9068EBFF9DFFA0FAB0CA8DFA13F8DA" box="[1333,1423,1777,1802]" captionStart="FIGURE 5" captionStartId="9.[151,250,1917,1939]" captionTargetBox="[207,1364,193,1894]" captionTargetId="figure@9.[207,1364,193,1894]" captionTargetPageId="9" captionText="FIGURE 5 A – E. Protectoribates occidentalis sp. nov., scanning electron microscope images of adult female from California; A. habitus dorsal; B, habitus ventral, with arrow to custodium; C, frontal aspect; D, dorsolateral of prodorsum, with arrow to spindle end of bothridial seta (bo), second arrow to genal incision and third arrow to carina on tibia II; E, ventrolateral of prodorsum, with arrow to ridge on genal tooth. Scale bars: A, B = 25 µm, C – E = 15 µm." httpUri="https://zenodo.org/record/1018639/files/figure.png" pageId="3" pageNumber="154">Fig. 5E</figureCitation>
|
||
). Palp setal formula 0-2-1-3-9(1); eupathidium
|
||
<emphasis id="27DFA87CFF9DFFA0FD20CB6AFD48F8FD" box="[677,724,1814,1837]" italics="true" pageId="3" pageNumber="154">acm</emphasis>
|
||
shorter than solenidion, forming double horn with solenidion along length, but partially separated basally (as in
|
||
<taxonomicName id="D2AB0FEDFF9DFFA0FD4ECB45FB25F882" authority="Behan-Pelletier & Walter 2013" authorityName="Behan-Pelletier & Walter" authorityYear="2013" box="[715,1209,1849,1874]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="154" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="27DFA87CFF9DFFA0FD4ECB45FCC8F882" box="[715,852,1849,1874]" italics="true" pageId="3" pageNumber="154">Tectoribates</emphasis>
|
||
<bibRefCitation id="713A099FFF9DFFA0FCDECB45FB25F882" author="Behan-Pelletier, V. M. & Walter, D. E." box="[859,1209,1849,1874]" journalOrPublisher="Zootaxa" pageId="3" pageNumber="154" pagination="459 - 489" part="3741" publicationUrl="https://doi.org/10.11646/zootaxa.3741.4.2" refString="Behan-Pelletier, V. M. & Walter, D. E. (2013) Phylogenetic relationships of Tectoribates: nymphal characters of new North American species place the genus in Tegoribatidae (Acari, Oribatida). Zootaxa, 3741 (4), 459 - 489. https: // doi. org / 10.11646 / zootaxa. 3741.4.2" title="Phylogenetic relationships of Tectoribates: nymphal characters of new North American species place the genus in Tegoribatidae (Acari, Oribatida)" type="journal article" year="2013">Behan-Pelletier & Walter 2013</bibRefCitation>
|
||
</taxonomicName>
|
||
, their
|
||
<figureCitation id="8D9068EBFF9DFFA0FB7BCB45FAC8F882" box="[1278,1364,1849,1874]" captionStart="FIGURE 7" captionStartId="11.[151,250,1861,1883]" captionTargetBox="[207,1359,193,1809]" captionTargetId="figure@11.[207,1359,193,1828]" captionTargetPageId="11" captionText="FIGURE 7. Protectoribates occidentalis sp. nov., differential interference contrast microscope images, A – E adults; A, bothridium and bothridial seta (5 layers), with arrow to nerve canal of seta le on lamella; insert of bothridium showing scales; B, custodium and discidium (3 layers), with arrow to broad, rounded anterior edge of custodium; C, porose organ Ah, lateral view, with arrow to surrounding cerotegument (3 layers); D, dissected specimen with Ah in direct view (2 layers); E, genu, tibia and tarsus II, with white arrow to dentate carina on tibia (2 layers) and narrow black arrow to seta pv ”; F, protonymph (2 layers), dorsal view. Scale bar: A = 25 µm; B = 10 µm; D = 7.5 µm; F = 50 µm." httpUri="https://zenodo.org/record/1018645/files/figure.png" pageId="3" pageNumber="154">Fig. 7E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="1514746EFF9DFFA0FF42CB23FA00F832" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="154">
|
||
<emphasis id="27DFA87CFF9DFFA0FF42CB23FF60F8A6" box="[199,252,1887,1910]" italics="true" pageId="3" pageNumber="154">Legs</emphasis>
|
||
. Monodactylous (
|
||
<figureCitation id="8D9068EBFF9DFFA0FE43CB21FDD0F8A6" box="[454,588,1885,1910]" captionStart="FIGURE 4" captionStartId="8.[151,250,1974,1996]" captionTargetBox="[313,1242,203,1931]" captionTargetId="figure@8.[236,1331,193,1943]" captionTargetPageId="8" captionText="FIGURE 4. Protectoribates occidentalis sp. nov., adult female, A, leg I, B, leg II, C, leg IV, all antiaxial aspect. Legs I and II, trochanter not illustrated. Scale bar = 20 µm." httpUri="https://zenodo.org/record/1018637/files/figure.png" pageId="3" pageNumber="154">Figs. 4A–C</figureCitation>
|
||
). Leg setal formula I–IV, solenidia in parentheses: I: 1-5-3(1)-4(2)-20(2); II: 1-5-3(1)-4(1)-15(2); III: 2-2-1(1)-3(1)- 15; IV: 1-2-2-3(1)-12. Genua I and II without tooth ventrally (
|
||
<figureCitation id="8D9068EBFF9DFFA0FA8CCBFDFA12F84A" box="[1289,1422,1921,1946]" captionStart="FIGURE 4" captionStartId="8.[151,250,1974,1996]" captionTargetBox="[313,1242,203,1931]" captionTargetId="figure@8.[236,1331,193,1943]" captionTargetPageId="8" captionText="FIGURE 4. Protectoribates occidentalis sp. nov., adult female, A, leg I, B, leg II, C, leg IV, all antiaxial aspect. Legs I and II, trochanter not illustrated. Scale bar = 20 µm." httpUri="https://zenodo.org/record/1018637/files/figure.png" pageId="3" pageNumber="154">Figs. 4A, B</figureCitation>
|
||
). Porose areas on femora I to IV and trochanters III and IV. Femur I without bend medially at level of seta
|
||
<emphasis id="27DFA87CFF9DFFA0FAA9CBD9FAA6F86E" box="[1324,1338,1957,1982]" italics="true" pageId="3" pageNumber="154">d</emphasis>
|
||
. Tibia II with anterodorsal dentate carina (
|
||
<figureCitation id="8D9068EBFF9DFFA0FD93CBB5FDE6F832" box="[534,634,1993,2018]" captionStart="FIGURE 4" captionStartId="8.[151,250,1974,1996]" captionTargetBox="[313,1242,203,1931]" captionTargetId="figure@8.[236,1331,193,1943]" captionTargetPageId="8" captionText="FIGURE 4. Protectoribates occidentalis sp. nov., adult female, A, leg I, B, leg II, C, leg IV, all antiaxial aspect. Legs I and II, trochanter not illustrated. Scale bar = 20 µm." httpUri="https://zenodo.org/record/1018637/files/figure.png" pageId="3" pageNumber="154">Figs. 4B</figureCitation>
|
||
,
|
||
<figureCitation id="8D9068EBFF9DFFA0FD0DCBB5FD31F832" box="[648,685,1993,2018]" captionStart="FIGURE 5" captionStartId="9.[151,250,1917,1939]" captionTargetBox="[207,1364,193,1894]" captionTargetId="figure@9.[207,1364,193,1894]" captionTargetPageId="9" captionText="FIGURE 5 A – E. Protectoribates occidentalis sp. nov., scanning electron microscope images of adult female from California; A. habitus dorsal; B, habitus ventral, with arrow to custodium; C, frontal aspect; D, dorsolateral of prodorsum, with arrow to spindle end of bothridial seta (bo), second arrow to genal incision and third arrow to carina on tibia II; E, ventrolateral of prodorsum, with arrow to ridge on genal tooth. Scale bars: A, B = 25 µm, C – E = 15 µm." httpUri="https://zenodo.org/record/1018639/files/figure.png" pageId="3" pageNumber="154">5D</figureCitation>
|
||
,
|
||
<figureCitation id="8D9068EBFF9DFFA0FD3ECBB6FD41F832" box="[699,733,1994,2018]" captionStart="FIGURE 7" captionStartId="11.[151,250,1861,1883]" captionTargetBox="[207,1359,193,1809]" captionTargetId="figure@11.[207,1359,193,1828]" captionTargetPageId="11" captionText="FIGURE 7. Protectoribates occidentalis sp. nov., differential interference contrast microscope images, A – E adults; A, bothridium and bothridial seta (5 layers), with arrow to nerve canal of seta le on lamella; insert of bothridium showing scales; B, custodium and discidium (3 layers), with arrow to broad, rounded anterior edge of custodium; C, porose organ Ah, lateral view, with arrow to surrounding cerotegument (3 layers); D, dissected specimen with Ah in direct view (2 layers); E, genu, tibia and tarsus II, with white arrow to dentate carina on tibia (2 layers) and narrow black arrow to seta pv ”; F, protonymph (2 layers), dorsal view. Scale bar: A = 25 µm; B = 10 µm; D = 7.5 µm; F = 50 µm." httpUri="https://zenodo.org/record/1018645/files/figure.png" pageId="3" pageNumber="154">7E</figureCitation>
|
||
). Seta
|
||
<emphasis id="27DFA87CFF9DFFA0FCA8CBB7FCA4F832" box="[813,824,1995,2018]" italics="true" pageId="3" pageNumber="154">s</emphasis>
|
||
on tarsus I eupathidial, positioned distal to setae (
|
||
<emphasis id="27DFA87CFF9DFFA0FAF8CBB7FA17F832" box="[1405,1419,1995,2018]" italics="true" pageId="3" pageNumber="154">a</emphasis>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5DB127E5FF9AFFA7FF12CCEBFE25FECC" pageId="4" pageNumber="155" type="description">
|
||
<paragraph id="1514746EFF9AFFA7FF12CCEBFE25FECC" blockId="4.[151,1437,151,861]" pageId="4" pageNumber="155">
|
||
Solenidia on tarsus II shorter than setae on segment (
|
||
<figureCitation id="8D9068EBFF9AFFA7FD66CCEBFCA6FF60" box="[739,826,151,176]" captionStart="FIGURE 7" captionStartId="11.[151,250,1861,1883]" captionTargetBox="[207,1359,193,1809]" captionTargetId="figure@11.[207,1359,193,1828]" captionTargetPageId="11" captionText="FIGURE 7. Protectoribates occidentalis sp. nov., differential interference contrast microscope images, A – E adults; A, bothridium and bothridial seta (5 layers), with arrow to nerve canal of seta le on lamella; insert of bothridium showing scales; B, custodium and discidium (3 layers), with arrow to broad, rounded anterior edge of custodium; C, porose organ Ah, lateral view, with arrow to surrounding cerotegument (3 layers); D, dissected specimen with Ah in direct view (2 layers); E, genu, tibia and tarsus II, with white arrow to dentate carina on tibia (2 layers) and narrow black arrow to seta pv ”; F, protonymph (2 layers), dorsal view. Scale bar: A = 25 µm; B = 10 µm; D = 7.5 µm; F = 50 µm." httpUri="https://zenodo.org/record/1018645/files/figure.png" pageId="4" pageNumber="155">Fig. 7E</figureCitation>
|
||
);
|
||
<emphasis id="27DFA87CFF9AFFA7FCC8CCE4FCF3FF60" box="[845,879,152,176]" italics="true" pageId="4" pageNumber="155">ω1</emphasis>
|
||
not most proximal setiform structure on segment. Femur III with seta
|
||
<emphasis id="27DFA87CFF9AFFA7FE07CCC0FE0FFF05" box="[386,403,188,213]" italics="true" pageId="4" pageNumber="155">l’</emphasis>
|
||
absent (2 setae); seta
|
||
<emphasis id="27DFA87CFF9AFFA7FD1FCCC1FD31FF04" box="[666,685,189,212]" italics="true" pageId="4" pageNumber="155">v'</emphasis>
|
||
of genua I and II present. On genua and tibiae I and II setae
|
||
<emphasis id="27DFA87CFF9AFFA7FA01CCC0FA00FF05" box="[1412,1436,188,213]" italics="true" pageId="4" pageNumber="155">l”</emphasis>
|
||
subequal in shape to
|
||
<emphasis id="27DFA87CFF9AFFA7FE02CCA3FE04FF28" box="[391,408,223,248]" italics="true" pageId="4" pageNumber="155">l’</emphasis>
|
||
, setiform (
|
||
<figureCitation id="8D9068EBFF9AFFA7FD93CCA3FD01FF28" box="[534,669,223,248]" captionStart="FIGURE 4" captionStartId="8.[151,250,1974,1996]" captionTargetBox="[313,1242,203,1931]" captionTargetId="figure@8.[236,1331,193,1943]" captionTargetPageId="8" captionText="FIGURE 4. Protectoribates occidentalis sp. nov., adult female, A, leg I, B, leg II, C, leg IV, all antiaxial aspect. Legs I and II, trochanter not illustrated. Scale bar = 20 µm." httpUri="https://zenodo.org/record/1018637/files/figure.png" pageId="4" pageNumber="155">Figs. 4A, B</figureCitation>
|
||
). Seta
|
||
<emphasis id="27DFA87CFF9AFFA7FD69CC9CFC8AFF27" box="[748,790,224,247]" italics="true" pageId="4" pageNumber="155">pv”</emphasis>
|
||
on tarsus II, comb-like (
|
||
<figureCitation id="8D9068EBFF9AFFA7FBB4CCA3FB16FF28" box="[1073,1162,223,248]" captionStart="FIGURE 7" captionStartId="11.[151,250,1861,1883]" captionTargetBox="[207,1359,193,1809]" captionTargetId="figure@11.[207,1359,193,1828]" captionTargetPageId="11" captionText="FIGURE 7. Protectoribates occidentalis sp. nov., differential interference contrast microscope images, A – E adults; A, bothridium and bothridial seta (5 layers), with arrow to nerve canal of seta le on lamella; insert of bothridium showing scales; B, custodium and discidium (3 layers), with arrow to broad, rounded anterior edge of custodium; C, porose organ Ah, lateral view, with arrow to surrounding cerotegument (3 layers); D, dissected specimen with Ah in direct view (2 layers); E, genu, tibia and tarsus II, with white arrow to dentate carina on tibia (2 layers) and narrow black arrow to seta pv ”; F, protonymph (2 layers), dorsal view. Scale bar: A = 25 µm; B = 10 µm; D = 7.5 µm; F = 50 µm." httpUri="https://zenodo.org/record/1018645/files/figure.png" pageId="4" pageNumber="155">Fig. 7E</figureCitation>
|
||
). Anterodorsal knobs or spines absent from tibia I.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5DB127E5FF9AFFA7FF42CD5BFE2CFE57" pageId="4" pageNumber="155" type="etymology">
|
||
<paragraph id="1514746EFF9AFFA7FF42CD5BFE2CFE57" blockId="4.[151,1437,151,861]" pageId="4" pageNumber="155">
|
||
<emphasis id="27DFA87CFF9AFFA7FF42CD5BFEDAFE90" bold="true" box="[199,326,295,320]" pageId="4" pageNumber="155">Etymology</emphasis>
|
||
. The derivation of
|
||
<taxonomicName id="D2AB0FEDFF9AFFA7FDC5CD5BFD71FE90" authority="Behan-Pelletier, 2017" authorityName="Behan-Pelletier" authorityYear="2017" box="[576,749,295,320]" class="Arachnida" family="Tegoribatidae" genus="Protectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="194" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="27DFA87CFF9AFFA7FDC5CD5BFD71FE90" box="[576,749,295,320]" italics="true" pageId="4" pageNumber="155">Protectoribates</emphasis>
|
||
</taxonomicName>
|
||
is from the Latin ‘protector’, meaning "protector", and ‘oribates’ from Greek, a common ending for generic names in
|
||
<taxonomicName id="D2AB0FEDFF9AFFA7FCDFCD30FC6BFEB5" authorityName="Grandjean" authorityYear="1954" box="[858,1015,332,357]" class="Arachnida" family="Tegoribatidae" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="155" phylum="Arthropoda" rank="family">Tegoribatidae</taxonomicName>
|
||
. It refers to a possible function of the dentate carina on tibia II.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5DB127E5FF9AFFA7FF42CDE8FAC8FC8D" pageId="4" pageNumber="155" type="discussion">
|
||
<paragraph id="1514746EFF9AFFA7FF42CDE8FEB8FDED" blockId="4.[151,1437,151,861]" pageId="4" pageNumber="155">
|
||
<emphasis id="27DFA87CFF9AFFA7FF42CDE8FEA7FE7D" bold="true" box="[199,315,404,429]" pageId="4" pageNumber="155">Remarks.</emphasis>
|
||
Adults of
|
||
<taxonomicName id="D2AB0FEDFF9AFFA7FE4BCDE8FDE7FE7D" authority="Behan-Pelletier, 2017" authorityName="Behan-Pelletier" authorityYear="2017" box="[462,635,404,429]" class="Arachnida" family="Tegoribatidae" genus="Protectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="194" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="27DFA87CFF9AFFA7FE4BCDE8FDE7FE7D" box="[462,635,404,429]" italics="true" pageId="4" pageNumber="155">Protectoribates</emphasis>
|
||
</taxonomicName>
|
||
are distinguished from those of all other genera by having the combination of: dentate carina on tibia II,
|
||
<taxonomicName id="D2AB0FEDFF9AFFA7FDF7CDCBFD65FE00" box="[626,761,439,464]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="4" pageNumber="190" phylum="Nematoda" rank="genus">gnathosoma</taxonomicName>
|
||
without tectum on mentum, pteromorphs without hinge,
|
||
<emphasis id="27DFA87CFF9AFFA7FAF8CDCBFA00FE00" box="[1405,1436,439,464]" italics="true" pageId="4" pageNumber="155">Ah</emphasis>
|
||
developed as saccule; band of darkened integument posterior to genital plates, postanal porose area absent (
|
||
<tableCitation id="582941D5FF9AFFA7FAD9CDA0FF34FDC8" captionStart="TABLE 4" captionStartId="38.[120,196,495,515]" captionText="TABLE 4. Adult character states of genera of Tegoribatidae; boldface indicates potential apomorphies. Abbreviations: NGTLP = posterior notogastral tectum present; w / = with; w / o = without; octotaxic system as A = porose areas, S = saccules, T = tubules." pageId="4" pageNumber="155">Table 4</tableCitation>
|
||
). Among these traits, the dentate carina on tibia II and the form of humerosejugal organ
|
||
<emphasis id="27DFA87CFF9AFFA7FB4ECD83FB77FDC8" box="[1227,1259,511,536]" italics="true" pageId="4" pageNumber="155">Ah</emphasis>
|
||
are considered apomorphic.
|
||
</paragraph>
|
||
<paragraph id="1514746EFF9AFFA7FF42CE34FAC8FC8D" blockId="4.[151,1437,151,861]" pageId="4" pageNumber="155">
|
||
Immatures of
|
||
<taxonomicName id="D2AB0FEDFF9AFFA7FEE0CE3BFD8EFDB0" authority="Behan-Pelletier, 2017" authorityName="Behan-Pelletier" authorityYear="2017" box="[357,530,583,608]" class="Arachnida" family="Tegoribatidae" genus="Protectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="194" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="27DFA87CFF9AFFA7FEE0CE3BFD8EFDB0" box="[357,530,583,608]" italics="true" pageId="4" pageNumber="155">Protectoribates</emphasis>
|
||
</taxonomicName>
|
||
are similar to those of
|
||
<taxonomicName id="D2AB0FEDFF9AFFA7FC9DCE3BFC3EFDB0" authorityName="Behan-Pelletier & Walter" authorityYear="2013" box="[792,930,583,608]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="27DFA87CFF9AFFA7FC9DCE3BFC3EFDB0" box="[792,930,583,608]" italics="true" pageId="4" pageNumber="155">Tectoribates</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="D2AB0FEDFF9AFFA7FC37CE35FF53FD54" authority="Behan-Pelletier & Walter, 2013" authorityName="Behan-Pelletier & Walter" authorityYear="2013" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="alcicampestris">
|
||
<emphasis id="27DFA87CFF9AFFA7FC37CE35FBEDFDB0" box="[946,1137,583,608]" italics="true" pageId="4" pageNumber="155">T. alcicampestris</emphasis>
|
||
<bibRefCitation id="713A099FFF9AFFA7FBFCCE3BFF53FD54" author="Behan-Pelletier, V. M. & Walter, D. E." journalOrPublisher="Zootaxa" pageId="4" pageNumber="155" pagination="459 - 489" part="3741" publicationUrl="https://doi.org/10.11646/zootaxa.3741.4.2" refString="Behan-Pelletier, V. M. & Walter, D. E. (2013) Phylogenetic relationships of Tectoribates: nymphal characters of new North American species place the genus in Tegoribatidae (Acari, Oribatida). Zootaxa, 3741 (4), 459 - 489. https: // doi. org / 10.11646 / zootaxa. 3741.4.2" title="Phylogenetic relationships of Tectoribates: nymphal characters of new North American species place the genus in Tegoribatidae (Acari, Oribatida)" type="journal article" year="2013">Behan-Pelletier & Walter, 2013</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="D2AB0FEDFF9AFFA7FE90CE11FC80FD54" authority="Behan-Pelletier & Walter, 2013" authorityName="Behan-Pelletier & Walter" authorityYear="2013" box="[277,796,620,645]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="borealis">
|
||
<emphasis id="27DFA87CFF9AFFA7FE90CE11FE0FFD55" box="[277,403,620,645]" italics="true" pageId="4" pageNumber="155">T. borealis</emphasis>
|
||
<bibRefCitation id="713A099FFF9AFFA7FE27CE10FC80FD54" author="Behan-Pelletier, V. M. & Walter, D. E." box="[418,796,620,645]" journalOrPublisher="Zootaxa" pageId="4" pageNumber="155" pagination="459 - 489" part="3741" publicationUrl="https://doi.org/10.11646/zootaxa.3741.4.2" refString="Behan-Pelletier, V. M. & Walter, D. E. (2013) Phylogenetic relationships of Tectoribates: nymphal characters of new North American species place the genus in Tegoribatidae (Acari, Oribatida). Zootaxa, 3741 (4), 459 - 489. https: // doi. org / 10.11646 / zootaxa. 3741.4.2" title="Phylogenetic relationships of Tectoribates: nymphal characters of new North American species place the genus in Tegoribatidae (Acari, Oribatida)" type="journal article" year="2013">Behan-Pelletier & Walter, 2013</bibRefCitation>
|
||
</taxonomicName>
|
||
) in being plicate, apheredermous with monomorphic hysterosomal setae of series
|
||
<emphasis id="27DFA87CFF9AFFA7FE5ECEEDFDBEFD78" box="[475,546,655,680]" italics="true" pageId="4" pageNumber="155">c, d, l,</emphasis>
|
||
and
|
||
<emphasis id="27DFA87CFF9AFFA7FDDECEF3FDF5FD78" box="[603,617,655,680]" italics="true" pageId="4" pageNumber="155">h</emphasis>
|
||
. Hysterosomal setae of immature
|
||
<taxonomicName id="D2AB0FEDFF9AFFA7FC6BCEEDFAEEFD78" authority="Aoki, 1964" authorityName="Aoki" authorityYear="1964" box="[1006,1394,655,680]" class="Arachnida" family="Austrachipteriidae" genus="Hypozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="laysanensis">
|
||
<emphasis id="27DFA87CFF9AFFA7FC6BCEEDFB77FD78" box="[1006,1259,655,680]" italics="true" pageId="4" pageNumber="155">Hypozetes laysanensis</emphasis>
|
||
<bibRefCitation id="713A099FFF9AFFA7FB76CEF3FAEEFD78" author="Aoki, J." box="[1267,1394,655,680]" journalOrPublisher="Pacific Insects" pageId="4" pageNumber="155" pagination="649 - 664" part="6" refString="Aoki, J. (1964) Some oribatid mites (Acarina) from Laysan Island. Pacific Insects, 6 (4), 649 - 664." title="Some oribatid mites (Acarina) from Laysan Island" type="journal article" year="1964">Aoki, 1964</bibRefCitation>
|
||
</taxonomicName>
|
||
are also monomorphic, but larval seta
|
||
<emphasis id="27DFA87CFF9AFFA7FDA3CEC8FDDEFD1D" box="[550,578,692,717]" italics="true" pageId="4" pageNumber="155">dp</emphasis>
|
||
are twice the length of other hysterosomal setae and are borne on distinct tubercles (
|
||
<bibRefCitation id="713A099FFF9AFFA7FE94CEABFD90FD20" author="Behan-Pelletier, V. M." box="[273,524,727,752]" editor="Halliday, R. B." journalOrPublisher="CSIRO Publishing, Melbourne" pageId="4" pageNumber="155" pagination="50 - 57" refString="Behan-Pelletier, V. M. (2001) Phylogenetic relationships of Hypozetes (Acari: Tegoribatidae). In: Halliday, R. B., Walter, D. E., Proctor, H. C., Norton, R. A. & Colloff, M. J. (Eds.), Acarology: Proceedings of the 10 th International Congress. CSIRO Publishing, Melbourne, pp. 50 - 57." title="Phylogenetic relationships of Hypozetes (Acari: Tegoribatidae)" type="book chapter" volumeTitle="Acarology: Proceedings of the 10 th International Congress" year="2001">Behan-Pelletier 2001</bibRefCitation>
|
||
). Similarly, in larva and nymphs of
|
||
<taxonomicName id="D2AB0FEDFF9AFFA7FC4BCEABFA00FD20" authority="Popp, 1960" authorityName="Popp" authorityYear="1960" box="[974,1436,727,752]" class="Arachnida" family="Achipteriidae" genus="Plakoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="multicuspidus">
|
||
<emphasis id="27DFA87CFF9AFFA7FC4BCEABFA94FD20" box="[974,1288,727,752]" italics="true" pageId="4" pageNumber="155">Plakoribates multicuspidus</emphasis>
|
||
<bibRefCitation id="713A099FFF9AFFA7FA90CEA4FA00FD20" author="Popp, E." box="[1301,1436,727,752]" journalOrPublisher="Bulletin de la Societe Entomologique d'Egypte" pageId="4" pageNumber="155" pagination="203 - 221" part="54" refString="Popp, E. (1960) Neue Oribatiden aus Agypten (Acarina). Bulletin de la Societe Entomologique d'Egypte, 54, 203 - 221." title="Neue Oribatiden aus Agypten (Acarina)" type="journal article" year="1960">Popp, 1960</bibRefCitation>
|
||
</taxonomicName>
|
||
hysterosomal setae are monomorphic in shape, but seta
|
||
<emphasis id="27DFA87CFF9AFFA7FC8DCE80FCB8FCC5" box="[776,804,764,789]" italics="true" pageId="4" pageNumber="155">dp</emphasis>
|
||
are twice the length of other hysterosomal setae and are also borne on distinct tubercles (
|
||
<bibRefCitation id="713A099FFF9AFFA7FD9ECF63FD26FCE8" author="Shereef, G. M." box="[539,698,799,824]" journalOrPublisher="Acarologia" pageId="4" pageNumber="155" pagination="748 - 753" part="18" refString="Shereef, G. M. (1977) Biological studies and description of developmental stages of Plakoribates multicuspidatus Popp. and Xylobates souchnaiensis Abdel - Hamid (Acarina, Oribatei) in Egypt. Acarologia, 18 (4), 748 - 753." title="Biological studies and description of developmental stages of Plakoribates multicuspidatus Popp. and Xylobates souchnaiensis Abdel - Hamid (Acarina, Oribatei) in Egypt" type="journal article" year="1977">Shereef 1977</bibRefCitation>
|
||
). In contrast, known immatures of
|
||
<taxonomicName id="D2AB0FEDFF9AFFA7FBDACF63FB7EFCE8" authorityName="Ewing" authorityYear="1917" box="[1119,1250,799,824]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="27DFA87CFF9AFFA7FBDACF63FB7EFCE8" box="[1119,1250,799,824]" italics="true" pageId="4" pageNumber="155">Tegoribates</emphasis>
|
||
</taxonomicName>
|
||
are plicate and eupheredermous, integridorsal (retain centrodorsal setae) and hysterosomal setae are dimorphic (see below).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |