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<document id="BB4BB006F50018BA8F6DDD375F57F7D3" ID-CLB-Dataset="8651" ID-DOI="10.11646/zootaxa.4920.1.5" ID-GBIF-Dataset="6e8d5acc-6318-4e76-a10e-b13dba83c98b" ID-ISSN="1175-5326" ID-Zenodo-Dep="4471693" ID-ZooBank="9E967AD2-CF5E-4C7E-8ACA-DEEC544D0773" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1611748427741" checkinUser="plazi" docAuthor="Li, Qi-Qi &amp; Yin, Zi-Wei" docDate="2021" docId="03E787BF062DFFA4FF45FF3FFB86F9AA" docLanguage="en" docName="zootaxa.4920.1.5.pdf" docOrigin="Zootaxa 4920 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tribasodites tubericeps Li &amp; Yin 2021, sp. nov." docType="treatment" docVersion="5" lastPageNumber="121" masterDocId="FFDEFFC7062BFFA3FFD2FFE6FFD5FFA3" masterDocTitle="Four new species of Scydmaeninae and Pselaphinae (Coleoptera: Staphylinidae) from Yunnan, China" masterLastPageNumber="122" masterPageNumber="114" pageNumber="120" updateTime="1698925668827" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="E583BF5860BE3AC24DCE4E22E49A421C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F8B69E30FCC67B9F1D9A4939D814849D">
<mods:title id="D9690452E782A323BD4EC28E8B1EB671">Four new species of Scydmaeninae and Pselaphinae (Coleoptera: Staphylinidae) from Yunnan, China</mods:title>
</mods:titleInfo>
<mods:name id="DDB20217E37A0591D5DFD4E37D3CAFA4" type="personal">
<mods:role id="6B62F390732794AC0144EA09A519BB75">
<mods:roleTerm id="0DC8D15DA121EA5D2873527E8C6108D4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E17200D485031F4671A2C3E5BA765E95">Li, Qi-Qi</mods:namePart>
<mods:nameIdentifier id="F368B32386A677DA6B0BC836AF43F4F8" type="ORCID">0000-0003-0651-8401</mods:nameIdentifier>
<mods:affiliation id="98A8597E3A81F2BCB990C1D6DC931A8C">Laboratory of Systematic Entomology, College of Life Sciences, Shanghai Normal University, 100 Guilin Road, Xuhui District, Shanghai 200234, China. &amp; 409429668 @ qq. com; https: // orcid. org / 0000 - 0003 - 0651 - 8401</mods:affiliation>
<mods:nameIdentifier id="D1FFB29073508B8955EACAEBA1C05692" type="email">409429668@qq.com</mods:nameIdentifier>
</mods:name>
<mods:name id="876D7BC1CE5F95564E5560954A9B0F58" type="personal">
<mods:role id="C5BEDEFA9107E87115DA7CEE77DB5738">
<mods:roleTerm id="30054C042744BF3FA5A87CEE247B6736">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="86178AB3F4740645181FF1A0B0D57A6A">Yin, Zi-Wei</mods:namePart>
<mods:nameIdentifier id="7847795F70A393AAF07F4635F96031A2" type="ORCID">0000-0001-6659-9448</mods:nameIdentifier>
<mods:affiliation id="FFEC197CBDF132CFF55CF6F6845C62FC">Laboratory of Systematic Entomology, College of Life Sciences, Shanghai Normal University, 100 Guilin Road, Xuhui District, Shanghai 200234, China. &amp; pselaphinae @ gmail. com; https: // orcid. org / 0000 - 0001 - 6659 - 9448</mods:affiliation>
<mods:nameIdentifier id="1B85423EFAFAB0BF5F4CEF173ED43D5B" type="email">pselaphinae@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="1C758F9D984E1E9B206C85E116CAF3F5">text</mods:typeOfResource>
<mods:relatedItem id="ABAFEA2DE05C4F4534B37DD3FD65BA67" type="host">
<mods:titleInfo id="49D6529BB2672DE53E325A94817FB5FC">
<mods:title id="569E71C02A58EB172BB05CDB3FE9AD25">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="9A6FCC9B909626D24B5375B8DC46AA45">
<mods:date id="E4A261A7762D2D5DF6308CD1124F7B56">2021</mods:date>
<mods:detail id="8FD8E35E52F015E5F4F6877FB4D9F402" type="pubDate">
<mods:number id="A537B9DD54581182D92598ED1D081C36">2021-01-27</mods:number>
</mods:detail>
<mods:detail id="E69873EF4D5A81BFC9658597A16B0157" type="volume">
<mods:number id="3EF45606E06E7DF6AB04AEC92A01B48E">4920</mods:number>
</mods:detail>
<mods:detail id="F78E43E30E74471B25ABED09DB551CD8" type="issue">
<mods:number id="97B257E64E52C9B0B9C27A882038A7D5">1</mods:number>
</mods:detail>
<mods:extent id="840A19523255B616052A70F80ECC9082" unit="page">
<mods:start id="49DA690469BFE7715D3AA87042D9674D">114</mods:start>
<mods:end id="01A1EA609E18C9B9181E1740CCAEC340">122</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="5BF8E46D451A2FD43B102CD4C0FB8826">journal article</mods:classification>
<mods:identifier id="BAAF5577014992BFC61204034B69F9ED" type="CLB-Dataset">8651</mods:identifier>
<mods:identifier id="2F74399D94A9031A8104B377E2E8E9A6" type="DOI">10.11646/zootaxa.4920.1.5</mods:identifier>
<mods:identifier id="588FD324BA142EAE0156750D63E9A4E2" type="GBIF-Dataset">6e8d5acc-6318-4e76-a10e-b13dba83c98b</mods:identifier>
<mods:identifier id="DB8319887313BBBC9AB5D7446200F457" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="AC8B528D795AE4251058AE2BF7A96300" type="Zenodo-Dep">4471693</mods:identifier>
<mods:identifier id="69553ECF8C35AAC127E3036EB82B3B30" type="ZooBank">9E967AD2-CF5E-4C7E-8ACA-DEEC544D0773</mods:identifier>
</mods:mods>
<treatment id="03E787BF062DFFA4FF45FF3FFB86F9AA" ID-DOI="http://doi.org/10.5281/zenodo.4471709" ID-GBIF-Taxon="177131988" ID-Zenodo-Dep="4471709" LSID="urn:lsid:plazi:treatment:03E787BF062DFFA4FF45FF3FFB86F9AA" httpUri="http://treatment.plazi.org/id/03E787BF062DFFA4FF45FF3FFB86F9AA" lastPageId="7" lastPageNumber="121" pageId="6" pageNumber="120">
<subSubSection id="C3546522062DFFA5FF45FF3FFDC1FF57" box="[151,532,217,244]" pageId="6" pageNumber="120" type="nomenclature">
<paragraph id="8BF136A9062DFFA5FF45FF3FFDC1FF57" blockId="6.[151,532,217,280]" box="[151,532,217,244]" pageId="6" pageNumber="120">
<heading id="D0B981C5062DFFA5FF45FF3FFDC1FF57" bold="true" box="[151,532,217,244]" fontSize="11" level="1" pageId="6" pageNumber="120" reason="1">
<emphasis id="B93AEABB062DFFA5FF45FF3FFDC1FF57" bold="true" box="[151,532,217,244]" pageId="6" pageNumber="120">
<taxonomicName id="4C4E4D2A062DFFA5FF45FF3FFE7BFF57" ID-CoL="8YBY5" authority="Li &amp; Yin, 2021" authorityName="Li &amp; Yin" authorityYear="2021" box="[151,430,217,244]" class="Insecta" family="Staphylinidae" genus="Tribasodites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="120" phylum="Arthropoda" rank="species" species="tubericeps" status="sp. nov.">
<emphasis id="B93AEABB062DFFA5FF45FF3FFE7BFF57" bold="true" box="[151,430,217,244]" italics="true" pageId="6" pageNumber="120">Tribasodites tubericeps</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957C0062DFFA5FE67FF3CFDC1FF57" box="[437,532,218,244]" pageId="6" pageNumber="120" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3546522062DFFA5FF45FF18FF3EFEBB" box="[151,235,254,280]" pageId="6" pageNumber="120" type="description">
<paragraph id="8BF136A9062DFFA5FF45FF18FF3EFEBB" blockId="6.[151,532,217,280]" box="[151,235,254,280]" pageId="6" pageNumber="120">
(
<figureCitation id="13752A2C062DFFA5FF72FF18FF36FEBB" box="[160,227,254,280]" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1965]" captionTargetBox="[163,1428,442,1908]" captionTargetId="figure-63@6.[160,1428,442,1917]" captionTargetPageId="6" captionText="FIGURE 4. Tribasodites tubericeps sp. nov. (A–G. Male. H–J. Female). A. Habitus. B. Head and pronotum. C. Head, in dorsolateral view. D. Mesotibia. E. Metatrochanter. F–G. Aedeagus, lateral (F) and ventral (G). H. Tergite 5 (VIII). I. Sternite 6 (VIII). J. Genital complex. Scale bars: 0.5 mm in A; 0.2 mm in B, C; 0.1 mm in D–J." figureDoi="http://doi.org/10.5281/zenodo.4471705" httpUri="https://zenodo.org/record/4471705/files/figure.png" pageId="6" pageNumber="120">Fig. 4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3546522062DFFA5FF45FEA7FD69FE00" pageId="6" pageNumber="120" type="materials_examined">
<paragraph id="8BF136A9062DFFA5FF45FEA7FD69FE00" blockId="6.[151,1437,320,419]" pageId="6" pageNumber="120">
<emphasis id="B93AEABB062DFFA5FF45FEA7FEEAFEF9" bold="true" box="[151,319,320,347]" pageId="6" pageNumber="120">Type material</emphasis>
(15 exx.).
<materialsCitation id="3B263CF4062DFFA5FE6EFEA7FB89FEDC" ID-GBIF-Occurrence="3026410307" collectingDate="2020-02-25" collectionCode="SNUC" collectorName="Ji-Shen Wang" country="China" elevation="1758" latitude="24.021666" location="Mt. Tangli" longLatPrecision="21" longitude="99.238884" municipality="Lincang" pageId="6" pageNumber="120" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B93AEABB062DFFA5FE6EFEA7FD74FEF9" bold="true" box="[444,673,320,347]" pageId="6" pageNumber="120">
<typeStatus id="54F5880B062DFFA5FE6EFEA7FDF8FEF8" box="[444,557,321,347]" pageId="6" pageNumber="120" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F3597639062DFFA5FDE8FEA6FD49FEF9" box="[570,668,320,346]" name="China" pageId="6" pageNumber="120">CHINA</collectingCountry>
:
</emphasis>
Ƌ, labeled ‘
<collectingRegion id="498AF84B062DFFA5FCF8FEA7FC52FEF8" box="[810,903,321,347]" country="China" name="Yunnan" pageId="6" pageNumber="120">Yunnan</collectingRegion>
,
<collectingMunicipality id="6B95ACD3062DFFA5FC46FEA6FC26FEF9" box="[916,1011,320,346]" pageId="6" pageNumber="120">Lincang</collectingMunicipality>
,
<location id="8E916072062DFFA5FC2DFEA7FBADFEF9" LSID="urn:lsid:plazi:treatment:03E787BF062DFFA4FF45FF3FFB86F9AA:8E916072062DFFA5FC2DFEA7FBADFEF9" box="[1023,1144,320,347]" country="China" latitude="24.021666" longLatPrecision="21" longitude="99.238884" municipality="Lincang" name="Mt. Tangli" pageId="6" pageNumber="120" stateProvince="Yunnan">Mt. Tangli</location>
,
<geoCoordinate id="EE7A506E062DFFA5FB56FEA7FADCFEF8" box="[1156,1289,321,347]" degrees="24" direction="north" minutes="01" orientation="latitude" pageId="6" pageNumber="120" precision="15" seconds="18" value="24.021666">24°01'18''N</geoCoordinate>
,
<geoCoordinate id="EE7A506E062DFFA5FAC4FEA7FA4DFEF8" box="[1302,1432,321,347]" degrees="99" direction="east" minutes="14" orientation="longitude" pageId="6" pageNumber="120" precision="15" seconds="20" value="99.238884">99°14'20''E</geoCoordinate>
,
<quantity id="4CB69B4C062DFFA5FF45FE82FF25FEDC" box="[151,240,356,383]" metricMagnitude="3" metricUnit="m" metricValue="1.758" pageId="6" pageNumber="120" unit="m" value="1758.0">
<elevation id="0063D19A062DFFA5FF45FE82FF25FEDC" box="[151,240,356,383]" metricMagnitude="3" metricUnit="m" metricValue="1.758" pageId="6" pageNumber="120" unit="m" value="1758.0">1758 m</elevation>
</quantity>
,
<date id="FFF01069062DFFA5FF2CFE82FEA7FEDC" box="[254,370,356,383]" pageId="6" pageNumber="120" value="2020-02-25">
<collectingDate id="EFB4E981062DFFA5FF2CFE82FEA7FEDC" box="[254,370,356,383]" pageId="6" pageNumber="120" value="2020-02-25">2020.ii.25</collectingDate>
</date>
, with
<taxonomicName id="4C4E4D2A062DFFA5FE69FE82FD8FFEDD" authorityName="Holmgren" authorityYear="1910" box="[443,602,356,382]" class="Insecta" family="Termitidae" genus="Odontotermes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="120" phylum="Arthropoda" rank="genus">
<emphasis id="B93AEABB062DFFA5FE69FE82FD8FFEDD" box="[443,602,356,382]" italics="true" pageId="6" pageNumber="120">Odontotermes</emphasis>
</taxonomicName>
termites, sifted,
<collectorName id="26BB537F062DFFA5FCC9FE82FC6CFEDC" box="[795,953,356,383]" pageId="6" pageNumber="120">Ji-Shen Wang</collectorName>
leg.’ (
<collectionCode id="ED5FAE6C062DFFA5FBD1FE83FB81FEDC" box="[1027,1108,357,383]" pageId="6" pageNumber="120">SNUC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B263CF4062DFFA5FBBAFE82FD6DFE00" ID-GBIF-Occurrence="3026410304" collectingDate="2020-02-25" collectionCode="SNUC" collectorName="Ji-Shen Wang" country="China" elevation="1758" latitude="24.021666" location="Mt. Tangli" longLatPrecision="21" longitude="99.238884" municipality="Lincang" pageId="6" pageNumber="120" specimenCount="8" specimenCount-female="8" stateProvince="Yunnan" typeStatus="paratype">
<emphasis id="B93AEABB062DFFA5FBBAFE82FA50FEDD" bold="true" box="[1128,1413,356,382]" pageId="6" pageNumber="120">
<typeStatus id="54F5880B062DFFA5FBBAFE82FAC5FEDD" box="[1128,1296,356,382]" pageId="6" pageNumber="120" type="paratype">PARATYPES</typeStatus>
:
<collectingCountry id="F3597639062DFFA5FACCFE82FA55FEDD" box="[1310,1408,356,382]" name="China" pageId="6" pageNumber="120">CHINA</collectingCountry>
:
</emphasis>
6 ƋƋ,
<specimenCount id="9D48FD20062DFFA5FF1DFE6FFEDAFE00" box="[207,271,393,419]" pageId="6" pageNumber="120" type="female">8 ♀♀</specimenCount>
, same label data as holotype (
<collectionCode id="ED5FAE6C062DFFA5FD8DFE6FFD65FE00" box="[607,688,393,419]" pageId="6" pageNumber="120">SNUC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF316621062DFFA5FF45F873FC22F857" ID-DOI="http://doi.org/10.5281/zenodo.4471705" ID-Zenodo-Dep="4471705" httpUri="https://zenodo.org/record/4471705/files/figure.png" pageId="6" pageNumber="120" startId="6.[151,250,1941,1965]" targetBox="[163,1428,442,1908]" targetPageId="6">
<paragraph id="8BF136A9062DFFA5FF45F873FC22F857" blockId="6.[151,1437,1941,2037]" pageId="6" pageNumber="120">
<emphasis id="B93AEABB062DFFA5FF45F873FEC6F80E" bold="true" box="[151,275,1941,1965]" pageId="6" pageNumber="120">FIGURE 4.</emphasis>
<taxonomicName id="4C4E4D2A062DFFA5FEC8F873FDDDF80E" authority="Li &amp; Yin, 2021" authorityName="Li &amp; Yin" authorityYear="2021" box="[282,520,1941,1965]" class="Insecta" family="Staphylinidae" genus="Tribasodites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="120" phylum="Arthropoda" rank="species" species="tubericeps" status="sp. nov.">
<emphasis id="B93AEABB062DFFA5FEC8F873FDDDF80E" box="[282,520,1941,1965]" italics="true" pageId="6" pageNumber="120">Tribasodites tubericeps</emphasis>
</taxonomicName>
<emphasis id="B93AEABB062DFFA5FDDDF873FDB5F80E" bold="true" box="[527,608,1941,1965]" pageId="6" pageNumber="120">
<taxonomicNameLabel id="A20957C0062DFFA5FDDDF873FDB5F80E" box="[527,608,1941,1965]" pageId="6" pageNumber="120" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A–G. Male. H–J. Female).
<emphasis id="B93AEABB062DFFA5FC57F873FC4BF80E" bold="true" box="[901,926,1941,1965]" pageId="6" pageNumber="120">A.</emphasis>
Habitus.
<emphasis id="B93AEABB062DFFA5FBD3F873FBCDF80E" bold="true" box="[1025,1048,1941,1965]" pageId="6" pageNumber="120">B.</emphasis>
Head and pronotum.
<emphasis id="B93AEABB062DFFA5FB24F873FADBF80E" bold="true" box="[1270,1294,1941,1965]" pageId="6" pageNumber="120">C.</emphasis>
Head, in dorsolateral view.
<emphasis id="B93AEABB062DFFA5FEE0F85FFE9EF872" bold="true" box="[306,331,1977,2001]" pageId="6" pageNumber="120">D.</emphasis>
Mesotibia.
<emphasis id="B93AEABB062DFFA5FE17F85FFE08F872" bold="true" box="[453,477,1977,2001]" pageId="6" pageNumber="120">E.</emphasis>
Metatrochanter.
<emphasis id="B93AEABB062DFFA5FD59F85FFD17F872" bold="true" box="[651,706,1977,2001]" pageId="6" pageNumber="120">F–G.</emphasis>
Aedeagus, lateral (F) and ventral (G).
<emphasis id="B93AEABB062DFFA5FB81F85FFBB8F872" bold="true" box="[1107,1133,1977,2001]" pageId="6" pageNumber="120">H.</emphasis>
Tergite 5 (VIII).
<emphasis id="B93AEABB062DFFA5FAF3F85FFAE4F872" bold="true" box="[1313,1329,1977,2001]" pageId="6" pageNumber="120">I.</emphasis>
Sternite 6 (VIII).
<emphasis id="B93AEABB062DFFA5FF32F83BFF26F856" bold="true" box="[224,243,2013,2037]" pageId="6" pageNumber="120">J.</emphasis>
Genital complex. Scale bars: 0.5 mm in A; 0.2 mm in B, C; 0.1 mm in D–J.
</paragraph>
</caption>
<subSubSection id="C3546522062CFFA4FF15FF70FACEFEBE" pageId="7" pageNumber="121" type="diagnosis">
<paragraph id="8BF136A9062CFFA4FF15FF70FACEFEBE" blockId="7.[151,1437,150,1545]" pageId="7" pageNumber="121">
<emphasis id="B93AEABB062CFFA4FF15FF70FE97FF13" bold="true" box="[199,322,150,176]" pageId="7" pageNumber="121">Diagnosis.</emphasis>
<emphasis id="B93AEABB062CFFA4FE9BFF71FE56FF12" box="[329,387,151,177]" italics="true" pageId="7" pageNumber="121">Male</emphasis>
. Body length
<quantity id="4CB69B4C062CFFA4FDF7FF71FD19FF12" box="[549,716,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="2.41" metricValueMax="2.44" metricValueMin="2.38" pageId="7" pageNumber="121" unit="mm" value="2.41" valueMax="2.44" valueMin="2.38">2.38–2.44 mm</quantity>
; with simple antennae, vertex with greatly raised sub-triangular projection in deep central cavity, pronotum with three spines on each discal carina, aedeagus sub-triangular, median lobe with greatly constricted basal capsule, ventral lobe broad, dorsal lobe strongly curved downwards, with a pointed apex.
<emphasis id="B93AEABB062CFFA4FEE7FEE5FE5CFEBE" box="[309,393,259,285]" italics="true" pageId="7" pageNumber="121">Female</emphasis>
. Body length
<quantity id="4CB69B4C062CFFA4FDFAFEE5FD1BFEBE" box="[552,718,259,285]" metricMagnitude="-3" metricUnit="m" metricValue="2.5549999999999997" metricValueMax="2.63" metricValueMin="2.48" pageId="7" pageNumber="121" unit="mm" value="2.5549999999999997" valueMax="2.63" valueMin="2.48">2.48–2.63 mm</quantity>
; genital complex transverse, with short lateral lobes.
</paragraph>
</subSubSection>
<subSubSection id="C3546522062CFFA4FF15FEC0FBEEFA62" pageId="7" pageNumber="121" type="description">
<paragraph id="8BF136A9062CFFA4FF15FEC0FCC7FC6A" blockId="7.[151,1437,150,1545]" pageId="7" pageNumber="121">
<emphasis id="B93AEABB062CFFA4FF15FEC0FE8CFEE3" bold="true" box="[199,345,294,320]" pageId="7" pageNumber="121">Description.</emphasis>
<emphasis id="B93AEABB062CFFA4FE8CFEC1FE4DFEE2" box="[350,408,295,321]" italics="true" pageId="7" pageNumber="121">Male</emphasis>
. Body (
<figureCitation id="13752A2C062CFFA4FE22FEC1FD93FEE2" box="[496,582,295,321]" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1965]" captionTargetBox="[163,1428,442,1908]" captionTargetId="figure-63@6.[160,1428,442,1917]" captionTargetPageId="6" captionText="FIGURE 4. Tribasodites tubericeps sp. nov. (A–G. Male. H–J. Female). A. Habitus. B. Head and pronotum. C. Head, in dorsolateral view. D. Mesotibia. E. Metatrochanter. F–G. Aedeagus, lateral (F) and ventral (G). H. Tergite 5 (VIII). I. Sternite 6 (VIII). J. Genital complex. Scale bars: 0.5 mm in A; 0.2 mm in B, C; 0.1 mm in D–J." figureDoi="http://doi.org/10.5281/zenodo.4471705" httpUri="https://zenodo.org/record/4471705/files/figure.png" pageId="7" pageNumber="121">Fig. 4A</figureCitation>
) length (combined length of head, pronotum, elytra and abdomen)
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. Coloration red-brown, mouthparts and tarsi paler. Head (
<figureCitation id="13752A2C062CFFA4FCB5FEADFC31FEC6" box="[871,996,331,357]" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1965]" captionTargetBox="[163,1428,442,1908]" captionTargetId="figure-63@6.[160,1428,442,1917]" captionTargetPageId="6" captionText="FIGURE 4. Tribasodites tubericeps sp. nov. (A–G. Male. H–J. Female). A. Habitus. B. Head and pronotum. C. Head, in dorsolateral view. D. Mesotibia. E. Metatrochanter. F–G. Aedeagus, lateral (F) and ventral (G). H. Tergite 5 (VIII). I. Sternite 6 (VIII). J. Genital complex. Scale bars: 0.5 mm in A; 0.2 mm in B, C; 0.1 mm in D–J." figureDoi="http://doi.org/10.5281/zenodo.4471705" httpUri="https://zenodo.org/record/4471705/files/figure.png" pageId="7" pageNumber="121">Fig. 4B, C</figureCitation>
) slightly wider than long, subtriangular, length from anterior margin of clypeus to head base
<quantity id="4CB69B4C062CFFA4FCCEFE88FC16FE2A" box="[796,963,366,393]" metricMagnitude="-4" metricUnit="m" metricValue="4.85" metricValueMax="5.0" metricValueMin="4.699999999999999" pageId="7" pageNumber="121" unit="mm" value="0.485" valueMax="0.5" valueMin="0.47">0.47–0.50 mm</quantity>
, width across eyes
<quantity id="4CB69B4C062CFFA4FB70FE88FA9CFE2A" box="[1186,1353,366,393]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="5.3" metricValueMin="4.699999999999999" pageId="7" pageNumber="121" unit="mm" value="0.5" valueMax="0.53" valueMin="0.47">0.47–0.53 mm</quantity>
; vertex with large sub-triangular projection in deep central cavity, anterior margin of cavity also greatly raised to same level as projection; frons depressed between moderately raised antennal tubercles. Eyes fully developed, large and prominent, each composed of approximately 55 facets. Gular area slightly depressed; with thin median longitudinal carina. Antenna long and slender, unmodified, length
<quantity id="4CB69B4C062CFFA4FCD4FE19FC7AFDBA" box="[774,943,511,537]" metricMagnitude="-3" metricUnit="m" metricValue="1.23" metricValueMax="1.24" metricValueMin="1.22" pageId="7" pageNumber="121" unit="mm" value="1.23" valueMax="1.24" valueMin="1.22">1.22–1.24 mm</quantity>
; antennomeres 9–11 forming distinct club. Pronotum (
<figureCitation id="13752A2C062CFFA4FEC6FDC5FEBFFD9E" box="[276,362,547,573]" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1965]" captionTargetBox="[163,1428,442,1908]" captionTargetId="figure-63@6.[160,1428,442,1917]" captionTargetPageId="6" captionText="FIGURE 4. Tribasodites tubericeps sp. nov. (A–G. Male. H–J. Female). A. Habitus. B. Head and pronotum. C. Head, in dorsolateral view. D. Mesotibia. E. Metatrochanter. F–G. Aedeagus, lateral (F) and ventral (G). H. Tergite 5 (VIII). I. Sternite 6 (VIII). J. Genital complex. Scale bars: 0.5 mm in A; 0.2 mm in B, C; 0.1 mm in D–J." figureDoi="http://doi.org/10.5281/zenodo.4471705" httpUri="https://zenodo.org/record/4471705/files/figure.png" pageId="7" pageNumber="121">Fig. 4B</figureCitation>
) as long as wide, length along midline and maximum width
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; lateral sides spinose, each with one large spine and few smaller denticles; with broad median longitudinal impression followed by short median carina at base, with pair of sinuate lateral sulci and pair of discal carinae, with three discal spines on each carina. Elytra transverse, length along suture
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, maximum width
<quantity id="4CB69B4C062CFFA4FC24FD69FB4FFD0A" box="[1014,1178,655,681]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" metricValueMax="8.9" metricValueMin="8.7" pageId="7" pageNumber="121" unit="mm" value="0.88" valueMax="0.89" valueMin="0.87">0.87–0.89 mm</quantity>
, with truncate base and straight posterior margin; discal stria approximately half length of elytra; sutural stria complete. Metathoracic wings fully developed. Legs elongate; mesotrochanter with long spine at ventral margin, mesotibia (
<figureCitation id="13752A2C062CFFA4FB6FFD31FAC0FD52" box="[1213,1301,727,753]" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1965]" captionTargetBox="[163,1428,442,1908]" captionTargetId="figure-63@6.[160,1428,442,1917]" captionTargetPageId="6" captionText="FIGURE 4. Tribasodites tubericeps sp. nov. (A–G. Male. H–J. Female). A. Habitus. B. Head and pronotum. C. Head, in dorsolateral view. D. Mesotibia. E. Metatrochanter. F–G. Aedeagus, lateral (F) and ventral (G). H. Tergite 5 (VIII). I. Sternite 6 (VIII). J. Genital complex. Scale bars: 0.5 mm in A; 0.2 mm in B, C; 0.1 mm in D–J." figureDoi="http://doi.org/10.5281/zenodo.4471705" httpUri="https://zenodo.org/record/4471705/files/figure.png" pageId="7" pageNumber="121">Fig. 4D</figureCitation>
) with small protuberance at apex; metatrochanter (
<figureCitation id="13752A2C062CFFA4FD95FD1DFD4EFCB6" box="[583,667,763,789]" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1965]" captionTargetBox="[163,1428,442,1908]" captionTargetId="figure-63@6.[160,1428,442,1917]" captionTargetPageId="6" captionText="FIGURE 4. Tribasodites tubericeps sp. nov. (A–G. Male. H–J. Female). A. Habitus. B. Head and pronotum. C. Head, in dorsolateral view. D. Mesotibia. E. Metatrochanter. F–G. Aedeagus, lateral (F) and ventral (G). H. Tergite 5 (VIII). I. Sternite 6 (VIII). J. Genital complex. Scale bars: 0.5 mm in A; 0.2 mm in B, C; 0.1 mm in D–J." figureDoi="http://doi.org/10.5281/zenodo.4471705" httpUri="https://zenodo.org/record/4471705/files/figure.png" pageId="7" pageNumber="121">Fig. 4E</figureCitation>
) with long, apically curved projection ventrally. Abdomen relatively short in dorsal view, length along midline
<quantity id="4CB69B4C062CFFA4FDBDFCF9FCC1FC9A" box="[623,788,799,825]" metricMagnitude="-4" metricUnit="m" metricValue="7.6" metricValueMax="7.8" metricValueMin="7.4" pageId="7" pageNumber="121" unit="mm" value="0.76" valueMax="0.78" valueMin="0.74">0.74–0.78 mm</quantity>
, maximum width
<quantity id="4CB69B4C062CFFA4FC0DFCF9FB50FC9A" box="[991,1157,799,825]" metricMagnitude="-4" metricUnit="m" metricValue="8.45" metricValueMax="8.6" metricValueMin="8.299999999999999" pageId="7" pageNumber="121" unit="mm" value="0.845" valueMax="0.86" valueMin="0.83">0.83–0.86 mm</quantity>
; tergite 1 (IV) with basal sulcus interrupted by short discal carinae, with oblique inner and complete outer marginal carinae; sternite 2 (IV) along middle as long as sternites 3–5 (V–VII) combined. Aedeagus (
<figureCitation id="13752A2C062CFFA4FC7FFC81FBF1FC22" box="[941,1060,871,897]" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1965]" captionTargetBox="[163,1428,442,1908]" captionTargetId="figure-63@6.[160,1428,442,1917]" captionTargetPageId="6" captionText="FIGURE 4. Tribasodites tubericeps sp. nov. (A–G. Male. H–J. Female). A. Habitus. B. Head and pronotum. C. Head, in dorsolateral view. D. Mesotibia. E. Metatrochanter. F–G. Aedeagus, lateral (F) and ventral (G). H. Tergite 5 (VIII). I. Sternite 6 (VIII). J. Genital complex. Scale bars: 0.5 mm in A; 0.2 mm in B, C; 0.1 mm in D–J." figureDoi="http://doi.org/10.5281/zenodo.4471705" httpUri="https://zenodo.org/record/4471705/files/figure.png" pageId="7" pageNumber="121">Fig. 4F, G</figureCitation>
)
<quantity id="4CB69B4C062CFFA4FBE4FC81FB49FC22" box="[1078,1180,871,897]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="7" pageNumber="121" unit="mm" value="0.38">0.38 mm</quantity>
long, triangular, basal capsule strongly constricted, with large foramen, ventral lobe broad, dorsal lobe strongly curved downwards and pointed at apex; parameres fused to form ventral lamina.
</paragraph>
<paragraph id="8BF136A9062CFFA4FF15FC35FAD6FBDE" blockId="7.[151,1437,150,1545]" pageId="7" pageNumber="121">
<emphasis id="B93AEABB062CFFA4FF15FC35FECEFC4E" box="[199,283,979,1005]" italics="true" pageId="7" pageNumber="121">Female</emphasis>
. External morphology similar to that of male, except head lacking modifications, antennae slightly shorter, eyes smaller, and legs simple. Each eye composed of about 58 facets. Measurements (as for male): body length
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, length/width of head 0.47–0.52/
<quantity id="4CB69B4C062CFFA4FCD5FBFCFC78FB96" box="[775,941,1050,1077]" metricMagnitude="-4" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="7" pageNumber="121" unit="mm" value="0.525" valueMax="0.55" valueMin="0.5">0.50–0.55 mm</quantity>
, pronotum 0.52–0.57/
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, elytra 0.75–0.77/
<quantity id="4CB69B4C062CFFA4FEDDFBD8FE61FBFA" box="[271,436,1086,1113]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" metricValueMax="8.9" metricValueMin="8.7" pageId="7" pageNumber="121" unit="mm" value="0.88" valueMax="0.89" valueMin="0.87">0.87–0.89 mm</quantity>
, abdomen 0.74–0.77/
<quantity id="4CB69B4C062CFFA4FD75FBD8FC98FBFA" box="[679,845,1086,1113]" metricMagnitude="-4" metricUnit="m" metricValue="8.35" metricValueMax="8.5" metricValueMin="8.2" pageId="7" pageNumber="121" unit="mm" value="0.835" valueMax="0.85" valueMin="0.82">0.82–0.85 mm</quantity>
. Tergite 5 (VIII) (
<figureCitation id="13752A2C062CFFA4FBC7FBD9FBB9FBFA" box="[1045,1132,1087,1113]" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1965]" captionTargetBox="[163,1428,442,1908]" captionTargetId="figure-63@6.[160,1428,442,1917]" captionTargetPageId="6" captionText="FIGURE 4. Tribasodites tubericeps sp. nov. (A–G. Male. H–J. Female). A. Habitus. B. Head and pronotum. C. Head, in dorsolateral view. D. Mesotibia. E. Metatrochanter. F–G. Aedeagus, lateral (F) and ventral (G). H. Tergite 5 (VIII). I. Sternite 6 (VIII). J. Genital complex. Scale bars: 0.5 mm in A; 0.2 mm in B, C; 0.1 mm in D–J." figureDoi="http://doi.org/10.5281/zenodo.4471705" httpUri="https://zenodo.org/record/4471705/files/figure.png" pageId="7" pageNumber="121">Fig. 4H</figureCitation>
) and sternite 6 (VIII) (
<figureCitation id="13752A2C062CFFA4FABDFBD9FF65FBDE" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1965]" captionTargetBox="[163,1428,442,1908]" captionTargetId="figure-63@6.[160,1428,442,1917]" captionTargetPageId="6" captionText="FIGURE 4. Tribasodites tubericeps sp. nov. (A–G. Male. H–J. Female). A. Habitus. B. Head and pronotum. C. Head, in dorsolateral view. D. Mesotibia. E. Metatrochanter. F–G. Aedeagus, lateral (F) and ventral (G). H. Tergite 5 (VIII). I. Sternite 6 (VIII). J. Genital complex. Scale bars: 0.5 mm in A; 0.2 mm in B, C; 0.1 mm in D–J." figureDoi="http://doi.org/10.5281/zenodo.4471705" httpUri="https://zenodo.org/record/4471705/files/figure.png" pageId="7" pageNumber="121">Fig. 4I</figureCitation>
) each transverse; genital complex (
<figureCitation id="13752A2C062CFFA4FDEEFB85FD5FFBDE" box="[572,650,1123,1149]" captionStart="FIGURE 4" captionStartId="6.[151,250,1941,1965]" captionTargetBox="[163,1428,442,1908]" captionTargetId="figure-63@6.[160,1428,442,1917]" captionTargetPageId="6" captionText="FIGURE 4. Tribasodites tubericeps sp. nov. (A–G. Male. H–J. Female). A. Habitus. B. Head and pronotum. C. Head, in dorsolateral view. D. Mesotibia. E. Metatrochanter. F–G. Aedeagus, lateral (F) and ventral (G). H. Tergite 5 (VIII). I. Sternite 6 (VIII). J. Genital complex. Scale bars: 0.5 mm in A; 0.2 mm in B, C; 0.1 mm in D–J." figureDoi="http://doi.org/10.5281/zenodo.4471705" httpUri="https://zenodo.org/record/4471705/files/figure.png" pageId="7" pageNumber="121">Fig. 4J</figureCitation>
) strongly transverse, lateral lobes short, width
<quantity id="4CB69B4C062CFFA4FB4AFB85FB2AFBDE" box="[1176,1279,1123,1149]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="7" pageNumber="121" unit="mm" value="0.33">0.33 mm</quantity>
.
</paragraph>
<paragraph id="8BF136A9062CFFA4FF15FB60FBEEFA62" blockId="7.[151,1437,150,1545]" pageId="7" pageNumber="121">
<emphasis id="B93AEABB062CFFA4FF15FB60FE7AFB02" bold="true" box="[199,431,1158,1185]" pageId="7" pageNumber="121">Comparative notes.</emphasis>
A total of 16 Chinese species of this genus have been described (
<bibRefCitation id="EFDF4B58062CFFA4FBAEFB61FAC1FB02" author="Li, L. - Z. &amp; Hu, J. - Y. &amp; Peng, Z. &amp; Tang, L. &amp; Yin, Z. - W. &amp; Zhao, M. - J." box="[1148,1300,1159,1185]" pageId="7" pageNumber="121" refId="ref4942" refString="Li, L. - Z., Hu, J. - Y., Peng, Z., Tang, L., Yin, Z. - W. &amp; Zhao, M. - J. (2019) Catalogue of Chinese Coleoptera. Vol. 3. Staphylinidae. Science Press, Beijing, xix + 720 pp." type="book" year="2019">
Li
<emphasis id="B93AEABB062CFFA4FB4EFB61FB19FB02" box="[1180,1228,1159,1185]" italics="true" pageId="7" pageNumber="121">et al</emphasis>
. 2019
</bibRefCitation>
), with a majority of them (13 spp.) restricted to cave environments in southern and southwestern areas of the country (Yin et al. 2015;
<bibRefCitation id="EFDF4B58062CFFA4FF0EFB29FE70FB4A" author="Yin, Z. - W. &amp; Zhou, G. - C." box="[220,421,1231,1257]" pageId="7" pageNumber="121" refId="ref5398" refString="Yin, Z. - W. &amp; Zhou, G. - C. (2018) Two new cavernicolous Pselaphinae (Coleoptera: Staphylinidae)" type="book" year="2018">Yin &amp; Zhou 2018</bibRefCitation>
). The male of
<taxonomicName id="4C4E4D2A062CFFA4FD91FB29FC9DFB4A" authority="Li &amp; Yin, 2021" authorityName="Li &amp; Yin" authorityYear="2021" box="[579,840,1231,1257]" class="Insecta" family="Staphylinidae" genus="Tribasodites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="121" phylum="Arthropoda" rank="species" species="tubericeps" status="sp. nov.">
<emphasis id="B93AEABB062CFFA4FD91FB29FC9DFB4A" box="[579,840,1231,1257]" italics="true" pageId="7" pageNumber="121">Tribasodites tubericeps</emphasis>
</taxonomicName>
<emphasis id="B93AEABB062CFFA4FC9CFB29FC7CFB4A" bold="true" box="[846,937,1231,1257]" pageId="7" pageNumber="121">
<taxonomicNameLabel id="A20957C0062CFFA4FC9CFB29FC7CFB4A" box="[846,937,1231,1257]" pageId="7" pageNumber="121" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is highly distinctive in the strongly modified vertex (cephalic modification lacking in the three free-living Chinese species), and the median lobe of the aedeagus has a greatly constricted basal capsule. The new species can be further separated from
<taxonomicName id="4C4E4D2A062CFFA4FB86FAF1FAABFA91" authority="(Raffray)" baseAuthorityName="Raffray" baseAuthorityYear="1912" box="[1108,1406,1303,1330]" class="Insecta" family="Staphylinidae" genus="Tribasodites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="121" phylum="Arthropoda" rank="species" species="semipunctatus">
<emphasis id="B93AEABB062CFFA4FB86FAF1FBB1FA93" box="[1108,1124,1303,1328]" italics="true" pageId="7" pageNumber="121">T</emphasis>
.
<emphasis id="B93AEABB062CFFA4FBA3FAF1FAC4FA93" box="[1137,1297,1303,1328]" italics="true" pageId="7" pageNumber="121">semipunctatus</emphasis>
(Raffray)
</taxonomicName>
of
<collectingCountry id="F3597639062CFFA4FF45FADDFF3CFAF6" box="[151,233,1339,1365]" name="Taiwan" pageId="7" pageNumber="121">Taiwan</collectingCountry>
by the more elongate antennomeres and larger lateral spines of the pronotum; from
<taxonomicName id="4C4E4D2A062CFFA4FB4CFADDFF38FADA" authority="Yin, Li &amp; Zhao" authorityName="Yin, Li &amp; Zhao" class="Insecta" family="Staphylinidae" genus="Tribasodites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="121" phylum="Arthropoda" rank="species" species="spinacaritus">
<emphasis id="B93AEABB062CFFA4FB4CFADDFB7BFAF7" box="[1182,1198,1339,1364]" italics="true" pageId="7" pageNumber="121">T</emphasis>
.
<emphasis id="B93AEABB062CFFA4FB6EFADDFA93FAF7" box="[1212,1350,1339,1364]" italics="true" pageId="7" pageNumber="121">spinacaritus</emphasis>
Yin, Li &amp; Zhao
</taxonomicName>
of Zhejiang (
<bibRefCitation id="EFDF4B58062CFFA4FE50FAB9FD93FADA" author="Zhao, M. - J. &amp; Yin, Z. - W. &amp; Li, L. - Z." box="[386,582,1375,1401]" pageId="7" pageNumber="121" pagination="25 - 31" refId="ref5571" refString="Zhao, M. - J., Yin, Z. - W. &amp; Li, L. - Z. (2010 a) Description of a new species of the genus Tribasodites Jeannel (Coleoptera, Staphylinidae, Pselaphinae) from East China with a key to world species. ZooKeys, 64, 25 - 31. https: // doi. org / 10.3897 / zookeys. 64.454" type="journal article" year="2010">Zhao et al. 2010a</bibRefCitation>
) by the unmodified antennal clubs of the male and much shorter apical spine of the mesotibia; and from
<taxonomicName id="4C4E4D2A062CFFA4FE1BFA65FCF4FA3E" authority="Yin, Zhao &amp; Li" authorityName="Yin, Zhao &amp; Li" box="[457,801,1411,1437]" class="Insecta" family="Staphylinidae" genus="Tribasodites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="121" phylum="Arthropoda" rank="species" species="tianmuensis">
<emphasis id="B93AEABB062CFFA4FE1BFA65FE0CFA3F" box="[457,473,1411,1436]" italics="true" pageId="7" pageNumber="121">T</emphasis>
.
<emphasis id="B93AEABB062CFFA4FE35FA65FDB8FA3F" box="[487,621,1411,1436]" italics="true" pageId="7" pageNumber="121">tianmuensis</emphasis>
Yin, Zhao &amp; Li
</taxonomicName>
of Zhejiang (
<bibRefCitation id="EFDF4B58062CFFA4FC6BFA65FB57FA3D" author="Zhao, M. - J. &amp; Yin, Z. - W. &amp; Li, L. - Z." box="[953,1154,1411,1438]" pageId="7" pageNumber="121" pagination="527 - 535" refId="ref5645" refString="Zhao, M. - J., Yin, Z. - W. &amp; Li, L. - Z. (2010 b) Two new species of the subtribe Batrisina (Coleoptera, Staphylinidae, Pselaphinae) associated with Pachycondyla luteipes (Formicidae) in China. Sociobiology, 56 (2), 527 - 535." type="journal article" year="2010">Zhao et al. 2010b</bibRefCitation>
) by the relatively thicker mesotibiae, and much larger elytra in relation to the size of the head and pronotum.
</paragraph>
</subSubSection>
<subSubSection id="C3546522062CFFA4FF15FA2CFD54FA46" box="[199,641,1482,1509]" pageId="7" pageNumber="121" type="distribution">
<paragraph id="8BF136A9062CFFA4FF15FA2CFD54FA46" blockId="7.[151,1437,150,1545]" box="[199,641,1482,1509]" pageId="7" pageNumber="121">
<emphasis id="B93AEABB062CFFA4FF15FA2CFEB4FA47" bold="true" box="[199,353,1482,1508]" pageId="7" pageNumber="121">Distribution.</emphasis>
Southern
<collectingCountry id="F3597639062CFFA4FE06FA2DFDCEFA46" box="[468,539,1483,1509]" name="China" pageId="7" pageNumber="121">China</collectingCountry>
:
<collectingRegion id="498AF84B062CFFA4FDF6FA2DFDA8FA46" box="[548,637,1483,1509]" country="China" name="Yunnan" pageId="7" pageNumber="121">Yunnan</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C3546522062CFFA4FF15FA09FB86F9AA" box="[199,1107,1519,1545]" pageId="7" pageNumber="121" type="etymology">
<paragraph id="8BF136A9062CFFA4FF15FA09FB86F9AA" blockId="7.[151,1437,150,1545]" box="[199,1107,1519,1545]" pageId="7" pageNumber="121">
<emphasis id="B93AEABB062CFFA4FF15FA09FE9BF9AA" bold="true" box="[199,334,1519,1545]" pageId="7" pageNumber="121">Etymology.</emphasis>
The new specific epithet refers to the modification of the male head.
</paragraph>
</subSubSection>
</treatment>
</document>
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