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<document id="04413C00DB0AAE8DC5C31BFEAEA87956" ID-DOI="10.5070/P9361043964" ID-ISSN="0031-0298" ID-Zenodo-Dep="13750243" ID-ZooBank="3F95876E-933FF-48AF-9CF0-A840A333220B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726017767285" checkinUser="felipe" docAuthor="Cicimurri, David J. &amp; Knight, James L." docDate="2022" docId="03E787A6FE2EFF8EAAB5F9A8FE7EFDD7" docLanguage="en" docName="PaleoBios.36.43964.pdf" docOrigin="PaleoBios 36" docStyle="DocumentStyle:2D14AFDE223D5FD9EB1703D6A63DEF6C.1:PaleoBios.2018-2021.journal_article" docStyleId="2D14AFDE223D5FD9EB1703D6A63DEF6C" docStyleName="PaleoBios.2018-2021.journal_article" docStyleVersion="1" docTitle="Ginglymostomatidae GEN. ET SP. INDET." docType="treatment" docVersion="3" lastPageNumber="6" masterDocId="FFDEFFDEFE29FF86A91DFFEFFFC3FFEC" masterDocTitle="Late Eocene (Priabonian) elasmobranchs from the Dry Branch Formation (Barnwell Group) of Aiken County, South Carolina, USA" masterLastPageNumber="31" masterPageNumber="1" pageNumber="6" updateTime="1726076902459" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-SA-4.0">
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<mods:title id="0BE2E4D231AD0AD9B7B020A6CCF1C273">Late Eocene (Priabonian) elasmobranchs from the Dry Branch Formation (Barnwell Group) of Aiken County, South Carolina, USA</mods:title>
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<mods:namePart id="30126F1D5348B813424F2C2578A59E4F">Cicimurri, David J.</mods:namePart>
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<mods:namePart id="12BE27F480028D3F0246702BD8930619">Knight, James L.</mods:namePart>
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<mods:title id="6D9067B39A5C43D444C44EC9BA491715">PaleoBios</mods:title>
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<mods:date id="1B4F396C20B09EC6BC7B8E362923FAC7">2022</mods:date>
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<treatment id="03E787A6FE2EFF8EAAB5F9A8FE7EFDD7" ID-DOI="http://doi.org/10.5281/zenodo.13750210" ID-Zenodo-Dep="13750210" LSID="urn:lsid:plazi:treatment:03E787A6FE2EFF8EAAB5F9A8FE7EFDD7" httpUri="http://treatment.plazi.org/id/03E787A6FE2EFF8EAAB5F9A8FE7EFDD7" lastPageId="8" lastPageNumber="6" pageId="7" pageNumber="6">
<subSubSection id="C354653BFE2EFF81AAB5F9A8FA4AF9B3" box="[936,1417,1607,1631]" pageId="7" pageNumber="6" type="nomenclature">
<paragraph id="8BF136B0FE2EFF81AAB5F9A8FA4AF9B3" blockId="7.[936,1417,1607,1663]" box="[936,1417,1607,1631]" pageId="7" pageNumber="6">
<heading id="D0B981DCFE2EFF81AAB5F9A8FA4AF9B3" allCaps="true" box="[936,1417,1607,1631]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="6" reason="2">
<taxonomicName id="4C4E4D33FE2EFF81AAB5F9A8FA4AF9B3" ID-CoL="623HG" authority="GEN. ET SP. INDET." box="[936,1417,1607,1631]" class="Elasmobranchii" family="Ginglymostomatidae" kingdom="Animalia" order="Orectolobiformes" pageId="7" pageNumber="6" phylum="Chordata" rank="family">GINGLYMOSTOMATIDAE GEN. ET SP. INDET.</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C354653BFE2EFF81AD7BF988FC1BF933" pageId="7" pageNumber="6" type="description">
<paragraph id="8BF136B0FE2EFF81AD7BF988FB09F993" blockId="7.[936,1417,1607,1663]" box="[1126,1226,1639,1663]" pageId="7" pageNumber="6">
(
<figureCitation id="13752A35FE2EFF81AD71F988FB01F993" box="[1132,1218,1639,1663]" captionStart="Figure 2" captionStartId="6.[113,188,1698,1722]" captionTargetBox="[160,1481,195,1667]" captionTargetId="figure-259@6.[160,1481,195,1667]" captionTargetPageId="6" captionText="Figure 2. Shark teeth from the Dry Branch Formation (Priabonian) of Aiken County, South Carolina. AD. Notorynchus sp. cf. No. kempi teeth. Lower left tooth in lingual (A) and labial (B) views, SC2013.38.1. Lower right tooth in labial (C) and lingual (D) views, SC96.97.4. E. Squatina sp. cf. Sq. prima. Antero-lateral tooth in labial view, SC96.97.9. F,G. Nebrius sp. cf. Neb. thielensi. Anterolateral tooth in labial (F) and occlusal (G) views, SC2013.38.5. Labial at top in G. H. Ginglymostomatidae. Tooth in labial view, SC2013.38.7. IO. Carcharias sp. Third lower right anterior tooth in labial (I) view, SC2001.1.55. Third upper right anterior tooth in lingual (J) and labial (K) views, SC2001.1.57. First lower right anterior tooth in labial (L) view, SC2001.1.59. Second lower right anterior tooth in labial (M) view, SC96.97.44.1. Second upper right anterior tooth in lingual (N) view, SC96.97.42. Lower left lateral tooth in lingual (O) view, SC96.97.43. PS. Alopias sp. Tooth in labial (P) and lingual (Q) views, SC2001.1.65. Tooth in labial (R) and lingual (S) views, SC2001.1.66. Scale bars=1 cm in AD, IK, N, O; 5 mm in EG, LM, PS; 1 mm in H." figureDoi="http://doi.org/10.5281/zenodo.13750247" httpUri="https://zenodo.org/record/13750247/files/figure.png" pageId="7" pageNumber="6">FIG. 2H</figureCitation>
)
</paragraph>
<paragraph id="8BF136B0FE2EFF81AA7DF94DFC1BF933" blockId="7.[832,1521,1698,1934]" pageId="7" pageNumber="6">
<emphasis id="B93AEAA2FE2EFF81AA7DF94DFBA7F950" bold="true" box="[864,1124,1698,1724]" pageId="7" pageNumber="6">Referred specimen</emphasis>
—SC2013.38.7, incomplete anterior tooth.
</paragraph>
</subSubSection>
<subSubSection id="C354653BFE2EFF8EAA7DF907FE7EFDD7" lastPageId="8" lastPageNumber="7" pageId="7" pageNumber="6" type="discussion">
<paragraph id="8BF136B0FE2EFF8EAA7DF907FE7EFDD7" blockId="7.[832,1521,1698,1934]" lastBlockId="8.[112,800,194,571]" lastPageId="8" lastPageNumber="7" pageId="7" pageNumber="6">
<emphasis id="B93AEAA2FE2EFF81AA7DF907FC1BF8EE" bold="true" box="[864,984,1768,1794]" pageId="7" pageNumber="6">Remarks</emphasis>
—This specimen differs from
<taxonomicName id="4C4E4D33FE2EFF81AC5BF907FA0DF8EE" box="[1350,1486,1768,1794]" class="Elasmobranchii" family="Ginglymostomatidae" genus="Nebrius" kingdom="Animalia" order="Orectolobiformes" pageId="7" pageNumber="6" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B93AEAA2FE2EFF81AC5BF907FA66F8EE" box="[1350,1445,1768,1794]" italics="true" pageId="7" pageNumber="6">Nebrius</emphasis>
sp.
</taxonomicName>
cf.
<emphasis id="B93AEAA2FE2EFF81AA5DF8E4FC27F8C9" box="[832,996,1803,1829]" italics="true" pageId="7" pageNumber="6">Neb. thielensi</emphasis>
(see above) in having only a single pair of large lateral cusplets, and the labial crown foot is distinctively bifid but not drawn out into an elongate basal protuberance. SC2013.38.7 is similar to the anterior teeth of Cretaceous
<taxonomicName id="4C4E4D33FE21FF8EA83FFF2DFCD9FF30" authority="Case and Cappetta, 1997" authorityName="Case and Cappetta" authorityYear="1997" box="[290,794,194,220]" class="Elasmobranchii" family="Ginglymostomatidae" genus="Plicatoscyllium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orectolobiformes" pageId="8" pageNumber="7" phylum="Chordata" rank="genus">
<emphasis id="B93AEAA2FE21FF8EA83FFF2DFE18FF30" box="[290,475,194,220]" italics="true" pageId="8" pageNumber="7">Plicatoscyllium</emphasis>
<bibRefCitation id="EFDF4B41FE21FF8EA8FEFF2DFCD9FF30" author="Case, G. R. &amp; H. Cappetta" box="[483,794,194,220]" pageId="8" pageNumber="7" pagination="131 - 189" refId="ref21312" refString="Case, G. R., and H. Cappetta. 1997. A new selachian fauna from the Late Maastrichtian of Texas (Upper Cretaceous / Navarro Group; Kemp Formation). Munchner Geowissenschaftliche Abhandlungen 34: 131 - 189." type="journal article" year="1997">Case and Cappetta, 1997</bibRefCitation>
</taxonomicName>
, but there is no indication of labial crown ornamentation. The single pair of lateral cusplets also distinguishes SC2013.38.7 from the various early Paleogene species of
<taxonomicName id="4C4E4D33FE21FF8EA98EFEA1FD35FE84" authority="(Noubhani and Cappetta 1997)" baseAuthorityName="Noubhani and Cappetta" baseAuthorityYear="1997" box="[147,758,334,360]" class="Elasmobranchii" family="Ginglymostomatidae" genus="Ginglymostoma" kingdom="Animalia" order="Orectolobiformes" pageId="8" pageNumber="7" phylum="Chordata" rank="genus">
<emphasis id="B93AEAA2FE21FF8EA98EFEA1FE96FE84" box="[147,341,334,360]" italics="true" pageId="8" pageNumber="7">Ginglymostoma</emphasis>
(Noubhani and Cappetta 1997)
</taxonomicName>
. In terms of gross morphology, SC2013.38.7 is similar to
<taxonomicName id="4C4E4D33FE21FF8EA96DFE7BFD6CFE43" authority="(Casier, 1946)" baseAuthorityName="Casier" baseAuthorityYear="1946" box="[112,687,404,431]" class="Elasmobranchii" family="Ginglymostomatidae" genus="Protoginglymostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orectolobiformes" pageId="8" pageNumber="7" phylum="Chordata" rank="species" species="ypresiensis">
<emphasis id="B93AEAA2FE21FF8EA96DFE7BFE39FE42" box="[112,506,404,430]" italics="true" pageId="8" pageNumber="7">Protoginglymostoma ypresiensis</emphasis>
(
<bibRefCitation id="EFDF4B41FE21FF8EAB11FE7BFD67FE42" author="Casier, E." box="[524,676,404,430]" pageId="8" pageNumber="7" pagination="1 - 247" refId="ref21427" refString="Casier, E. 1946. La faune ichthyologique de l'Ypresien de la Belgique. Memoires du Musee Royal d'Histoire Naturelle de Belgique 104: 1 - 247." type="journal article" year="1946">Casier, 1946</bibRefCitation>
)
</taxonomicName>
from the early to middle Eocene of
<collectingCountry id="F3597620FE21FF8EA8ACFE57FDDBFE3E" box="[433,536,440,466]" name="Belgium" pageId="8" pageNumber="7">Belgium</collectingCountry>
and
<collectingCountry id="F3597620FE21FF8EAB49FE57FD03FE3E" box="[596,704,440,466]" name="Morocco" pageId="8" pageNumber="7">Morocco</collectingCountry>
(
<bibRefCitation id="EFDF4B41FE21FF8EABCFFE57FF72FE19" author="Casier, E." pageId="8" pageNumber="7" pagination="1 - 247" refId="ref21427" refString="Casier, E. 1946. La faune ichthyologique de l'Ypresien de la Belgique. Memoires du Musee Royal d'Histoire Naturelle de Belgique 104: 1 - 247." type="journal article" year="1946">Casier 1946</bibRefCitation>
,
<bibRefCitation id="EFDF4B41FE21FF8EA9DDFE33FEACFE19" author="Herman, J." box="[192,367,475,502]" pageId="8" pageNumber="7" pagination="127 - 138" refId="ref23071" refString="Herman, J. 1977. Additions to the Eocene fish fauna of Belgium. 3. Revision of the Orectolobiforms. Tertiary Research 1 (4): 127 - 138." type="journal article" year="1977">Herman 1977</bibRefCitation>
, Tabuce et al. 2005), but the poor condition of the only available tooth makes a more precise identification difficult.
</paragraph>
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