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<mods:title id="EA4AC00CCB051D9B52C8039DDCA32346">A revision of the Pediacus Shuckard 1839 (Coleoptera: Cucujidae) of Asia and Australasia</mods:title>
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<treatment id="03E78793A932FFA90AEE10DACC96B4E6" ID-DOI="http://doi.org/10.5281/zenodo.6139874" ID-GBIF-Taxon="119411057" ID-Zenodo-Dep="6139874" LSID="urn:lsid:plazi:treatment:03E78793A932FFA90AEE10DACC96B4E6" httpUri="http://treatment.plazi.org/id/03E78793A932FFA90AEE10DACC96B4E6" lastPageId="7" lastPageNumber="39" pageId="5" pageNumber="37">
<subSubSection id="C354650EA932FFAB0AEE10DACF48B08B" box="[151,542,1446,1472]" pageId="5" pageNumber="37" type="nomenclature">
<paragraph id="8BF13685A932FFAB0AEE10DACF48B08B" blockId="5.[151,542,1446,1506]" box="[151,542,1446,1472]" pageId="5" pageNumber="37">
<heading id="D0B981E9A932FFAB0AEE10DACF48B08B" bold="true" box="[151,542,1446,1472]" fontSize="11" level="1" pageId="5" pageNumber="37" reason="1">
<taxonomicName id="4C4E4D06A932FFAB0AEE10DACF48B08B" ID-CoL="7692G" authority="Sen Gupta" authorityName="Sen Gupta" box="[151,542,1446,1472]" class="Insecta" family="Cucujidae" genus="Pediacus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="37" phylum="Arthropoda" rank="species" species="bhutanicus">
<emphasis id="B93AEA97A932FFAB0AEE10DACF48B08B" bold="true" box="[151,542,1446,1472]" pageId="5" pageNumber="37">
<emphasis id="B93AEA97A932FFAB0AEE10DACCC7B08B" bold="true" box="[151,401,1446,1472]" italics="true" pageId="5" pageNumber="37">Pediacus bhutanicus</emphasis>
Sen Gupta
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C354650EA932FFAB0AEE10B5CC77B0A9" box="[151,289,1481,1506]" pageId="5" pageNumber="37" type="description">
<paragraph id="8BF13685A932FFAB0AEE10B5CC77B0A9" blockId="5.[151,542,1446,1506]" box="[151,289,1481,1506]" pageId="5" pageNumber="37">
<figureCitation id="13752A00A932FFAB0AEE10B5CDB7B0A9" box="[151,225,1481,1506]" captionStart="FIGURE 3" captionStartId="8.[151,250,1807,1829]" captionTargetBox="[153,1434,201,1765]" captionTargetId="figure@8.[151,1436,195,1783]" captionTargetPageId="8" captionText="FIGURE 3. Pediacus species: (A D) Pediacus bhutanicus Sen Gupta, holotype; P. carinatus sp. nov. (E, H, I) specimen from Sumatra; (F, G) specimen from Thailand. (A, E, F) Head and pronotum; (B, I) dorsal habitus; (C, G) antenna; (D, H) aedeagus." httpUri="https://zenodo.org/record/252772/files/figure.png" pageId="5" pageNumber="37">Figs. 3</figureCitation>
AD.
</paragraph>
</subSubSection>
<subSubSection id="C354650EA932FFAB0AEE136DCF05B36C" box="[151,595,1553,1575]" pageId="5" pageNumber="37" type="reference_group">
<paragraph id="8BF13685A932FFAB0AEE136DCF05B36C" blockId="5.[151,595,1553,1575]" box="[151,595,1553,1575]" pageId="5" pageNumber="37">
<treatmentCitationGroup id="AB5E11ABA932FFAB0AEE136DCF05B36C" box="[151,595,1553,1575]" pageId="5" pageNumber="37">
<taxonomicName id="4C4E4D06A932FFAB0AEE136DCF19B36C" ID-CoL="7692G" authority="Sen Gupta, 1978: 221" authorityName="Sen Gupta" authorityPageNumber="221" authorityYear="1978" box="[151,591,1553,1575]" class="Insecta" family="Cucujidae" genus="Pediacus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="37" phylum="Arthropoda" rank="species" species="bhutanicus">
<emphasis id="B93AEA97A932FFAB0AEE136DCC3FB36C" box="[151,361,1553,1575]" italics="true" pageId="5" pageNumber="37">Pediacus bhutanicus</emphasis>
<treatmentCitation id="0AEF1094A932FFAB0B16136DCF19B36C" author="Sen" box="[367,591,1553,1575]" page="221" pageId="5" pageNumber="37" year="1978">
<bibRefCitation id="EFDF4B74A932FFAB0B16136DCF19B36C" author="Sen Gupta, T." box="[367,591,1553,1575]" firstAuthor="Sen Gupta" journalOrPublisher="Entomologica Basiliensia" page="221" pageId="5" pageNumber="37" pagination="219 - 222" part="3" refId="ref16147" refString="Sen Gupta, T. (1978) Ergebnisse der Bhutan-expedition 1972 des Naturhistorischen Museums in Basel (Coleoptera: Cucujidae). Entomologica Basiliensia, 3, 219 - 222." title="Ergebnisse der Bhutan-expedition 1972 des Naturhistorischen Museums in Basel (Coleoptera: Cucujidae)" type="journal article" year="1978">Sen Gupta, 1978: 221</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C354650EA932FFAB0AEE1328C98BB393" pageId="5" pageNumber="37" type="diagnosis">
<paragraph id="8BF13685A932FFAB0AEE1328C98BB393" blockId="5.[151,1436,1620,2005]" pageId="5" pageNumber="37">
<emphasis id="B93AEA97A932FFAB0AEE1328CC47B326" bold="true" box="[151,273,1620,1645]" pageId="5" pageNumber="37">Diagnosis.</emphasis>
Recognisable by the combination of the following characters: large size (length
<quantity id="4CB69B60A932FFAB0EB81328C848B327" box="[1217,1310,1620,1645]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="5" pageNumber="37" unit="mm" value="4.5">4.5 mm</quantity>
), head and pronotum densely and deeply punctured, pronotum with acute anterior, postero-medial and posterior denticles, elytra with sides broadly curved. This species is similar to
<taxonomicName id="4C4E4D06A932FFAB093513E0C912B3FF" box="[844,1092,1692,1716]" class="Insecta" family="Cucujidae" genus="Pediacus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="37" phylum="Arthropoda" rank="species" species="pendleburyi" status="sp. nov.">
<emphasis id="B93AEA97A932FFAB093513E0C912B3FF" box="[844,1092,1692,1716]" italics="true" pageId="5" pageNumber="37">Pediacus pendleburyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957ECA932FFAB0E3713E1C9FDB3FF" box="[1102,1195,1693,1716]" pageId="5" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
, but the sides of the pronotum are subparallel anteriorly and the anteromedial denticle is weakly produced to obsolete.
</paragraph>
</subSubSection>
<subSubSection id="C354650EA932FFA90ABE1398CDB8B456" lastPageId="7" lastPageNumber="39" pageId="5" pageNumber="37" type="description">
<paragraph id="8BF13685A932FFA90ABE1398CDB8B456" blockId="5.[151,1436,1620,2005]" lastBlockId="7.[151,1436,151,429]" lastPageId="7" lastPageNumber="39" pageId="5" pageNumber="37">
<emphasis id="B93AEA97A932FFAB0ABE1398CCEDB3B6" bold="true" box="[199,443,1764,1789]" pageId="5" pageNumber="37">Redescription. Body</emphasis>
length
<quantity id="4CB69B60A932FFAB086A1398CF3CB3B7" box="[531,618,1764,1789]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="5" pageNumber="37" unit="mm" value="4.5">4.5 mm</quantity>
(n=1), moderately elongate, 2.74× longer than wide; colour red-brown, head darker; pubescence long, pale, appressed and moderately conspicuous; surface moderately dull.
<emphasis id="B93AEA97A932FFAB0F25127BC8CDB26B" bold="true" box="[1372,1435,1799,1824]" pageId="5" pageNumber="37">Head</emphasis>
transverse, 1.91× wider than long; puncturation deeply impressed and dense medially, increasing in size and density laterally with punctures separated by less than their diameter; setae long and appressed, conspicuous; microsculpture granulate, surface dull; eyes small, moderately projecting, ocular index 0.71, finely faceted, postocular denticle absent; antennae as in
<figureCitation id="13752A00A932FFAB081412E4CFF8B2FB" box="[621,686,1944,1969]" captionStart="FIGURE 3" captionStartId="8.[151,250,1807,1829]" captionTargetBox="[153,1434,201,1765]" captionTargetId="figure@8.[151,1436,195,1783]" captionTargetPageId="8" captionText="FIGURE 3. Pediacus species: (A D) Pediacus bhutanicus Sen Gupta, holotype; P. carinatus sp. nov. (E, H, I) specimen from Sumatra; (F, G) specimen from Thailand. (A, E, F) Head and pronotum; (B, I) dorsal habitus; (C, G) antenna; (D, H) aedeagus." httpUri="https://zenodo.org/record/252772/files/figure.png" pageId="5" pageNumber="37">Fig. 3</figureCitation>
C, antennomere VII larger than VI and VIII.
<emphasis id="B93AEA97A932FFAB0EDD12E4C84AB2FA" bold="true" box="[1188,1308,1944,1969]" pageId="5" pageNumber="37">Pronotum</emphasis>
transverse, 1.30× wider than long, widest at about anterior 1/3, sides weakly rounded; lateral margins explanate, strongly reflexed with anterior, postero-medial and posterior denticles prominent and acute, antero-medial denticle weakly produced to obsolete; disc moderately impressed with a distinct pair of anterior depressions; puncturation, setation and microsculpture as for head, impunctate median strip faintly visible in basal 1/3.
<emphasis id="B93AEA97A931FFA80E2B12C4C9C8B29A" bold="true" box="[1106,1182,1976,2001]" pageId="6" pageNumber="38">Elytra</emphasis>
elongate,1.91× longer than wide, sides broadly curved; epipleura moderately explanate; puncturation moderately dense but weakly impressed basally, becoming more dense and deeply impressed laterally, puncturation faint on disc; setation moderately long, appressed and conspicuous; microsculpture granulate.
<emphasis id="B93AEA97A930FFA9098115C0C930B59E" bold="true" box="[1016,1126,188,213]" pageId="7" pageNumber="39">Genitalia</emphasis>
for male as in
<figureCitation id="13752A00A930FFA90F4E15C0C8D6B59F" box="[1335,1408,188,213]" captionStart="FIGURE 3" captionStartId="8.[151,250,1807,1829]" captionTargetBox="[153,1434,201,1765]" captionTargetId="figure@8.[151,1436,195,1783]" captionTargetPageId="8" captionText="FIGURE 3. Pediacus species: (A D) Pediacus bhutanicus Sen Gupta, holotype; P. carinatus sp. nov. (E, H, I) specimen from Sumatra; (F, G) specimen from Thailand. (A, E, F) Head and pronotum; (B, I) dorsal habitus; (C, G) antenna; (D, H) aedeagus." httpUri="https://zenodo.org/record/252772/files/figure.png" pageId="7" pageNumber="39">Fig. 3</figureCitation>
D, parameres elongate, rounded apically; internal sac with microspinules basally and lacking a sclerotized apical section.
</paragraph>
</subSubSection>
<caption id="DF31660DA931FFA80AEE1273C910B208" httpUri="https://zenodo.org/record/252771/files/figure.png" pageId="6" pageNumber="38" targetBox="[151,1405,195,1782]" targetPageId="6">
<paragraph id="8BF13685A931FFA80AEE1273C910B208" blockId="6.[151,1435,1807,1859]" pageId="6" pageNumber="38">
<emphasis id="B93AEA97A931FFA80AEE1273CC40B26E" bold="true" box="[151,278,1807,1829]" pageId="6" pageNumber="38">FIGURE 2.</emphasis>
<taxonomicName id="4C4E4D06A931FFA80B591273CC89B26E" box="[288,479,1807,1829]" class="Insecta" family="Cucujidae" genus="Pediacus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="38" phylum="Arthropoda" rank="species" species="australis" status="sp. nov.">
<emphasis id="B93AEA97A931FFA80B591273CC89B26E" box="[288,479,1807,1829]" italics="true" pageId="6" pageNumber="38">Pediacus australis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20957ECA931FFA80B901273CF6BB26E" box="[489,573,1807,1829]" pageId="6" pageNumber="38" rank="species">
<emphasis id="B93AEA97A931FFA80B901273CF6BB26E" bold="true" box="[489,573,1807,1829]" pageId="6" pageNumber="38">sp. nov.</emphasis>
</taxonomicNameLabel>
(A, D, E, H) specimen from Philippines, Mindanao; (B, C, F, G) specimen from Australia. (A, B) Head and pronotum; (C, D) antenna; (E, F) dorsal habitus; (G, H) aedeagus.
</paragraph>
</caption>
<subSubSection id="C354650EA930FFA90ABE145BCCE1B4C3" pageId="7" pageNumber="39" type="materials_examined">
<paragraph id="8BF13685A930FFA90ABE145BCCE1B4C3" blockId="7.[151,1436,151,429]" pageId="7" pageNumber="39">
<emphasis id="B93AEA97A930FFA90ABE145BCC21B40B" bold="true" box="[199,375,295,320]" pageId="7" pageNumber="39">
<typeStatus id="54F58827A930FFA90ABE145BCC54B40B" box="[199,258,295,320]" pageId="7" pageNumber="39">Type</typeStatus>
material.
</emphasis>
<materialsCitation id="3B263CD8A930FFA90B06145BCCE1B4C3" ID-GBIF-Occurrence="3923183301" collectionCode="NHMB" country="Bhutan" location="Phuntsholing" municipality="Phuntsholing" pageId="7" pageNumber="39" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
<emphasis id="B93AEA97A930FFA90B06145BCCBAB40B" bold="true" box="[383,492,295,320]" pageId="7" pageNumber="39">
<typeStatus id="54F58827A930FFA90B06145BCCBAB40B" box="[383,492,295,320]" pageId="7" pageNumber="39" type="holotype">Holotype</typeStatus>
</emphasis>
(
<specimenCount id="9D48FD0CA930FFA90B84145BCF6CB40B" box="[509,570,295,320]" pageId="7" pageNumber="39" type="male">male</specimenCount>
): “[
<collectingCountry id="F3597615A930FFA90819145BCFEEB40B" box="[608,696,295,320]" name="Bhutan" pageId="7" pageNumber="39">Bhutan</collectingCountry>
]
<locationDeviation id="9A432FA7A930FFA908B1145BCED3B40B" box="[712,901,295,320]" pageId="7" pageNumber="39">km 87 von 2215</locationDeviation>
[or 22/5]
<collectingMunicipality id="6B95ACFFA930FFA90982145BC9D9B40B" box="[1019,1167,295,320]" pageId="7" pageNumber="39">Phuntsholing</collectingMunicipality>
// Nat. - Hist. Museum Basel Bhutan Expedition 1972 // HOLOTYPE [red lettering] //
<taxonomicName id="4C4E4D06A930FFA909B11430C822B42F" authority="T. Sen Gupta" authorityName="T. Sen Gupta" box="[968,1396,332,357]" class="Insecta" family="Cucujidae" genus="Pediacus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="39" phylum="Arthropoda" rank="species" species="bhutanicus">Pediacus bhutanicus T. Sen Gupta</taxonomicName>
<specimenCount id="9D48FD0CA930FFA90FFE1431C8CAB42F" box="[1415,1436,333,356]" pageId="7" pageNumber="39" type="male"></specimenCount>
[Handwritten]” (
<collectionCode id="ED5FAE40A930FFA90B28140CCCFDB4C3" box="[337,427,368,392]" httpUri="http://grbio.org/cool/1d5a-4efs" name="Naturhistorisches Museum, Basel" pageId="7" pageNumber="39">NHMB</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C354650EA930FFA90ABE14E8CC96B4E6" box="[199,448,404,429]" pageId="7" pageNumber="39" type="distribution">
<paragraph id="8BF13685A930FFA90ABE14E8CC96B4E6" blockId="7.[151,1436,151,429]" box="[199,448,404,429]" pageId="7" pageNumber="39">
<emphasis id="B93AEA97A930FFA90ABE14E8CC36B4E6" bold="true" box="[199,352,404,429]" pageId="7" pageNumber="39">Distribution.</emphasis>
<collectingCountry id="F3597615A930FFA90B1E14E8CCEAB4E6" box="[359,444,404,429]" name="Bhutan" pageId="7" pageNumber="39">Bhutan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>