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<document id="4C4AFA5398478D856DDDF53BDD15C62D" ID-DOI="10.11646/zootaxa.3856.3.3" ID-GBIF-Dataset="d5977297-ce32-4ebb-8bee-c3c40e1e9a5f" ID-ISSN="1175-5326" ID-Zenodo-Dep="230649" ID-ZooBank="04B05A55-30E1-4A20-88F8-3974ACD8DC68" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460892518346" checkinUser="plazi" docAuthor="Ferreira, Sónia, Velo-Antón, Guillermo, Brochard, Christophe, Vieira, Cristiana, Alves, Paulo Célio, Thompson, David J., Watts, Phillip C. &amp; Brito, José Carlos" docDate="2014" docId="03DB8D13FFA99905F2BF6FF7FA4F7AB0" docLanguage="en" docName="zt03856p365.pdf" docOrigin="Zootaxa 3856 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Onychogomphus boudoti Ferreira, sp. nov." docType="treatment" docVersion="8" lastPageNumber="356" masterDocId="FFE2F56BFFAE9902F2286F29FFC67D79" masterDocTitle="A Critically Endangered new dragonfly species from Morocco: Onychogomphus boudoti sp. nov. (Odonata: Gomphidae)" masterLastPageNumber="365" masterPageNumber="349" pageNumber="356" updateTime="1698359685183" updateUser="plazi">
<mods:mods id="19EAE28D70592FA52F247319B0B5B69D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="8BC2BDF47A9F8EF439D6C2F74BEF966C">A Critically Endangered new dragonfly species from Morocco: Onychogomphus boudoti sp. nov. (Odonata: Gomphidae)</mods:title>
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<mods:namePart id="C3413D1B2158A003FE501BE849BC9591">Ferreira, Sónia</mods:namePart>
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<mods:namePart id="AA7DE0539509B436BBFC045868C07799">Velo-Antón, Guillermo</mods:namePart>
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<mods:namePart id="4956ADDEA804A7BB88C8457D24DB4E6F">Brochard, Christophe</mods:namePart>
</mods:name>
<mods:name id="5DD599C100E1F3DF87BF9F0C0D99E5C1" type="personal">
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<mods:roleTerm id="CDC9145A2744C884E5B687DBC51FE232">Author</mods:roleTerm>
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<mods:namePart id="AC581FFB3DBF537C6F1D382594686CB0">Vieira, Cristiana</mods:namePart>
</mods:name>
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<mods:role id="A9DF7B3CB3C14B63573DE2E09FC1A351">
<mods:roleTerm id="44217EA1475B8522244FDE0F6463874E">Author</mods:roleTerm>
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<mods:namePart id="4C736B191A1F4BA4573AFC03A2204A47">Alves, Paulo Célio</mods:namePart>
</mods:name>
<mods:name id="537457F99039AFF0604D7E5798674842" type="personal">
<mods:role id="D8D9C784346D1E21F9E17D15D3AD9662">
<mods:roleTerm id="0999C9DACD8CC3FD515BE22BB88F294B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DBB7C5091DC2071EA58CC006840A6217">Thompson, David J.</mods:namePart>
</mods:name>
<mods:name id="5D67FE7939148F3991BC70C1F8BE0C7D" type="personal">
<mods:role id="4A859612FF94B73E0DFE835C4460CB0F">
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<mods:namePart id="F70A2D316F46F01A0E0C9BA8EF618895">Watts, Phillip C.</mods:namePart>
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<mods:namePart id="3CE535BA1199DD8C1C95794289823B70">Brito, José Carlos</mods:namePart>
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<mods:typeOfResource id="32D61F0F1C26286BB32383F008A2B09A">text</mods:typeOfResource>
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<mods:title id="490D4DC6673C75B0F23F915C15D26A2E">Zootaxa</mods:title>
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<mods:date id="F75B8270F90EA6827C469A047C91017B">2014</mods:date>
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<mods:identifier id="4EF1A2F88F242209064209EE41987992" type="DOI">10.11646/zootaxa.3856.3.3</mods:identifier>
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<treatment id="03DB8D13FFA99905F2BF6FF7FA4F7AB0" ID-DOI="http://doi.org/10.5281/zenodo.5615890" ID-GBIF-Taxon="119587005" ID-Zenodo-Dep="5615890" LSID="urn:lsid:plazi:treatment:03DB8D13FFA99905F2BF6FF7FA4F7AB0" httpUri="http://treatment.plazi.org/id/03DB8D13FFA99905F2BF6FF7FA4F7AB0" lastPageNumber="356" pageId="7" pageNumber="356">
<subSubSection id="C3686F8EFFA99905F2BF6FF7FED47C60" pageId="7" pageNumber="356" type="nomenclature">
<paragraph id="8BCD3C05FFA99905F2BF6FF7FD5A7D81" blockId="7.[151,668,222,282]" box="[151,668,222,248]" pageId="7" pageNumber="356">
<heading id="D0858B69FFA99905F2BF6FF7FD5A7D81" bold="true" box="[151,668,222,248]" fontSize="11" level="1" pageId="7" pageNumber="356" reason="1">
<emphasis id="B906E017FFA99905F2BF6FF7FD5A7D81" bold="true" box="[151,668,222,248]" pageId="7" pageNumber="356">
<taxonomicName id="4C724786FFA99905F2BF6FF7FDF17D81" ID-CoL="9WVLM" authority="Ferreira" authorityName="Ferreira" box="[151,567,222,248]" class="Insecta" family="Gomphidae" genus="Onychogomphus" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="356" phylum="Arthropoda" rank="species" species="boudoti" status="sp. nov.">
<emphasis id="B906E017FFA99905F2BF6FF7FE007D81" bold="true" box="[151,454,222,248]" italics="true" pageId="7" pageNumber="356">Onychogomphus boudoti</emphasis>
Ferreira
</taxonomicName>
<taxonomicNameLabel id="A2355D6CFFA99905F0166FF7FD5A7D81" box="[574,668,222,248]" pageId="7" pageNumber="356" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCD3C05FFA99905F2BF6E28FED47C60" blockId="7.[151,668,222,282]" box="[151,274,257,282]" pageId="7" pageNumber="356">
(
<figureCitation id="13492080FFA99905F2B76E28FECF7C60" box="[159,265,257,282]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStart-3="FIGURE 7" captionStartId-0="7.[151,250,1589,1611]" captionStartId-1="8.[151,250,1297,1319]" captionStartId-2="9.[151,250,1849,1871]" captionStartId-3="12.[151,250,1649,1671]" captionTargetBox-0="[166,1419,486,1562]" captionTargetBox-1="[175,1415,238,1242]" captionTargetBox-2="[169,1414,229,1822]" captionTargetBox-3="[177,1413,208,1613]" captionTargetId-0="figure@7.[162,1424,481,1567]" captionTargetId-1="figure@8.[165,1421,231,1248]" captionTargetId-2="figure@9.[164,1422,221,1827]" captionTargetId-3="figure@12.[166,1420,193,1628]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="12" captionText-0="FIGURE 4. Onychogomphus boudoti; A, C F Holotype; B Paratype. (A) lateral view of holotype; (B) lateral view of paratype; (C) lateral view of cerci and epiproct; (D) dorsal view of cerci and epiproct; (E) dorsal view of thorax; (F) lateral view of secondary genitalia." captionText-1="FIGURE 5. Onychogomphus boudoti, female paratypes (both teneral). (A B) lateral view; (C) abdominal segment 8 with vulvar scale; (D) detail of head, note the presence of small tubercles behind the eyes." captionText-2="FIGURE 6. Onychogomphus boudoti, A E imagos from Morocco, 12 km east of Khenifra, 32.9262 ° N, 5.5616 ° W (photos: courtesy of Jean-Pierre Boudot). (A) fronto-dorsal view of a mature male; (B) dorsal view of a teneral female; (C) lateral view of a mature female; (D) dorsal view of a teneral male; (E) lateral view of a mature male. Note the yellow spot on the vertex, interrupted yellow mesothoracic collar and the thoracic yellow stripes disconnected dorsally (A); specific black pattern of sides of thorax (C, E); structure of the appendages in male (D, E)." captionText-3="FIGURE 7. Habitat and exuviae of Onychogomphus boudoti, 12 km east of Khenifra, 32.9262 ° N, 5.5616 ° W. (A) 27 th June 2011, type locality and surrounding area seen by the road; (B) exuvia dorsal view; (C) exuvia ventral view; (D) exuvia side view; (E) prementum ventral view; (F) dorsal view of right antenna." httpUri-0="https://zenodo.org/record/230653/files/figure.png" httpUri-1="https://zenodo.org/record/230654/files/figure.png" httpUri-2="https://zenodo.org/record/230655/files/figure.png" httpUri-3="https://zenodo.org/record/230656/files/figure.png" pageId="7" pageNumber="356">Figs. 47</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3686F8EFFA99905F2BF6E63FA4F7AB0" pageId="7" pageNumber="356" type="materials_examined">
<paragraph id="8BCD3C05FFA99905F2BF6E63FEC97CD0" blockId="7.[151,1437,329,425]" pageId="7" pageNumber="356">
<typeStatus id="54C982A7FFA99905F2BF6E63FF097C1B" box="[151,207,330,354]" pageId="7" pageNumber="356">Type</typeStatus>
material:
<collectionCode id="ED63A4C0FFA99905F3686E60FE5E7C1B" LSID="urn:lsid:biocol.org:col:34992" box="[320,408,329,354]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="7" pageNumber="356">RMNH</collectionCode>
.INS.906168
<typeStatus id="54C982A7FFA99905F0036E60FD557C1B" box="[555,659,329,354]" pageId="7" pageNumber="356" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="F3657C95FFA99905F0C86E63FC8E7C18" box="[736,840,330,353]" name="Morocco" pageId="7" pageNumber="356">Morocco</collectingCountry>
,
<quantity id="4C8A91E0FFA99905F17B6E60FC5C7C1B" box="[851,922,329,354]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="356" unit="km" value="12.0">12 km</quantity>
east of Khenifra,
<geoCoordinate id="EE465AC2FFA99905F64C6E60FB2E7C18" box="[1124,1256,329,354]" direction="north" orientation="latitude" pageId="7" pageNumber="356" precision="5" value="32.9262">32.9262° N</geoCoordinate>
,
<geoCoordinate id="EE465AC2FFA99905F6DC6E60FAA87C18" box="[1268,1390,329,354]" direction="west" orientation="longitude" pageId="7" pageNumber="356" precision="5" value="-5.5616">5.5616° W</geoCoordinate>
, alt.
<quantity id="4C8A91E0FFA99905F2BF6E47FF287CFC" box="[151,238,366,390]" metricMagnitude="3" metricUnit="m" metricValue="1.147" pageId="7" pageNumber="356" unit="m" value="1147.0">1147 m</quantity>
, collected on
<date id="FFCC1AC5FFA99905F3A46E47FDE17CFF" box="[396,551,366,390]" pageId="7" pageNumber="356" value="2012-06-12">12 June 2012</date>
, deposited at the Naturalis Biodiversity Center, Leiden,
<collectingCountry id="F3657C95FFA99905F68B6E44FA987CFF" box="[1187,1374,365,390]" name="Netherlands" pageId="7" pageNumber="356">The Netherlands</collectingCountry>
(
<figureCitation id="13492080FFA99905F7476E44FF607CD0" captionStart="FIGURE 4" captionStartId="7.[151,250,1589,1611]" captionTargetBox="[166,1419,486,1562]" captionTargetId="figure@7.[162,1424,481,1567]" captionTargetPageId="7" captionText="FIGURE 4. Onychogomphus boudoti; A, C F Holotype; B Paratype. (A) lateral view of holotype; (B) lateral view of paratype; (C) lateral view of cerci and epiproct; (D) dorsal view of cerci and epiproct; (E) dorsal view of thorax; (F) lateral view of secondary genitalia." httpUri="https://zenodo.org/record/230653/files/figure.png" pageId="7" pageNumber="356">Fig. 4</figureCitation>
A, 4CF)
</paragraph>
<caption id="DF0D6C8DFFA99905F2BF691CFEA47BF1" httpUri="https://zenodo.org/record/230653/files/figure.png" pageId="7" pageNumber="356" targetBox="[166,1419,486,1562]" targetPageId="7">
<paragraph id="8BCD3C05FFA99905F2BF691CFEA47BF1" blockId="7.[151,1436,1589,1672]" pageId="7" pageNumber="356">
<emphasis id="B906E017FFA99905F2BF691CFED47B33" bold="true" box="[151,274,1589,1611]" pageId="7" pageNumber="356">FIGURE 4.</emphasis>
<taxonomicName id="4C724786FFA99905F330691CFDD37B32" box="[280,533,1589,1611]" class="Insecta" family="Gomphidae" genus="Onychogomphus" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="356" phylum="Arthropoda" rank="species" species="boudoti">
<emphasis id="B906E017FFA99905F330691CFDD37B32" box="[280,533,1589,1611]" italics="true" pageId="7" pageNumber="356">Onychogomphus boudoti</emphasis>
</taxonomicName>
; A, CF Holotype; B Paratype. (A) lateral view of holotype; (B) lateral view of paratype; (C) lateral view of cerci and epiproct; (D) dorsal view of cerci and epiproct; (E) dorsal view of thorax; (F) lateral view of secondary genitalia.
</paragraph>
</caption>
<paragraph id="8BCD3C05FFA99905F2EF699FFE257AFB" blockId="7.[151,1437,1716,1993]" pageId="7" pageNumber="356">
<typeStatus id="54C982A7FFA99905F2EF699FFEFF7BB7" box="[199,313,1718,1742]" pageId="7" pageNumber="356" type="paratype">Paratypes</typeStatus>
:
<collectionCode id="ED63A4C0FFA99905F36B699DFE5D7BB4" LSID="urn:lsid:biocol.org:col:34992" box="[323,411,1716,1741]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="7" pageNumber="356">RMNH</collectionCode>
.INS.906169
<specimenCount id="9D74F78CFFA99905F006699CFDBF7BB4" box="[558,633,1716,1741]" pageId="7" pageNumber="356" type="male">1 male</specimenCount>
(sample code Df1030) collected on
<date id="FFCC1AC5FFA99905F63D699CFB687BB4" box="[1045,1198,1717,1741]" pageId="7" pageNumber="356" value="2011-06-27">27 June 2011</date>
,
<collectionCode id="ED63A4C0FFA99905F691699DFAD77BB4" LSID="urn:lsid:biocol.org:col:34992" box="[1209,1297,1716,1741]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="7" pageNumber="356">RMNH</collectionCode>
.INS.906170
<specimenCount id="9D74F78CFFA99905F2BF69F0FF207B8B" box="[151,230,1753,1778]" pageId="7" pageNumber="356" type="male">1 male</specimenCount>
(
<figureCitation id="13492080FFA99905F2D169F0FE867B88" box="[249,320,1753,1778]" captionStart="FIGURE 4" captionStartId="7.[151,250,1589,1611]" captionTargetBox="[166,1419,486,1562]" captionTargetId="figure@7.[162,1424,481,1567]" captionTargetPageId="7" captionText="FIGURE 4. Onychogomphus boudoti; A, C F Holotype; B Paratype. (A) lateral view of holotype; (B) lateral view of paratype; (C) lateral view of cerci and epiproct; (D) dorsal view of cerci and epiproct; (E) dorsal view of thorax; (F) lateral view of secondary genitalia." httpUri="https://zenodo.org/record/230653/files/figure.png" pageId="7" pageNumber="356">Fig. 4</figureCitation>
B) and
<collectionCode id="ED63A4C0FFA99905F3B169F0FE347B8B" LSID="urn:lsid:biocol.org:col:34992" box="[409,498,1753,1778]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="7" pageNumber="356">RMNH</collectionCode>
.INS.906171
<specimenCount id="9D74F78CFFA99905F0AF69F0FD2A7B8B" box="[647,748,1753,1778]" pageId="7" pageNumber="356" type="female">1 female</specimenCount>
(
<figureCitation id="13492080FFA99905F0D769F0FC827B8B" box="[767,836,1753,1778]" captionStart="FIGURE 5" captionStartId="8.[151,250,1297,1319]" captionTargetBox="[175,1415,238,1242]" captionTargetId="figure@8.[165,1421,231,1248]" captionTargetPageId="8" captionText="FIGURE 5. Onychogomphus boudoti, female paratypes (both teneral). (A B) lateral view; (C) abdominal segment 8 with vulvar scale; (D) detail of head, note the presence of small tubercles behind the eyes." httpUri="https://zenodo.org/record/230654/files/figure.png" pageId="7" pageNumber="356">Fig. 5</figureCitation>
A, 5C) collected on
<date id="FFCC1AC5FFA99905F61E69F0FB107B88" box="[1078,1238,1753,1777]" pageId="7" pageNumber="356" value="2012-06-12">12 June 2012</date>
, deposited at the Naturalis Biodiversity Center, Leiden,
<collectingCountry id="F3657C95FFA99905F07B69D4FCD57A6F" box="[595,787,1789,1814]" name="Netherlands" pageId="7" pageNumber="356">The Netherlands</collectingCountry>
;
<collectionCode id="ED63A4C0FFA99905F10869D4FCB17A6C" box="[800,887,1789,1813]" pageId="7" pageNumber="356">BMNH</collectionCode>
(E) 1239053
<specimenCount id="9D74F78CFFA99905F62569D4FB9C7A6F" box="[1037,1114,1789,1814]" pageId="7" pageNumber="356" type="male">1 male</specimenCount>
(sample code Df1029), and
<collectionCode id="ED63A4C0FFA99905F2BF6808FF287A40" box="[151,238,1825,1849]" pageId="7" pageNumber="356">BMNH</collectionCode>
(E) 1239054
<specimenCount id="9D74F78CFFA99905F3A96808FE257A43" box="[385,483,1825,1850]" pageId="7" pageNumber="356" type="female">1 female</specimenCount>
(
<figureCitation id="13492080FFA99905F3DC6808FDFE7A43" box="[500,568,1825,1850]" captionStart="FIGURE 5" captionStartId="8.[151,250,1297,1319]" captionTargetBox="[175,1415,238,1242]" captionTargetId="figure@8.[165,1421,231,1248]" captionTargetPageId="8" captionText="FIGURE 5. Onychogomphus boudoti, female paratypes (both teneral). (A B) lateral view; (C) abdominal segment 8 with vulvar scale; (D) detail of head, note the presence of small tubercles behind the eyes." httpUri="https://zenodo.org/record/230654/files/figure.png" pageId="7" pageNumber="356">Fig. 5</figureCitation>
B, 5D) collected on
<date id="FFCC1AC5FFA99905F1376808FC717A40" box="[799,951,1825,1849]" pageId="7" pageNumber="356" value="2012-06-12">12 June 2012</date>
deposited at the Natural History Museum, London,
<collectingCountry id="F3657C95FFA99905F2D6686FFEEC7A24" box="[254,298,1862,1885]" name="United Kingdom" pageId="7" pageNumber="356">UK</collectingCountry>
. All specimens were collected by Sónia Ferreira in
<collectingCountry id="F3657C95FFA99905F1A8686FFC2E7A24" box="[896,1000,1862,1885]" name="Morocco" pageId="7" pageNumber="356">Morocco</collectingCountry>
,
<quantity id="4C8A91E0FFA99905F1DD686CFBF97A27" box="[1013,1087,1861,1886]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="356" unit="km" value="12.0">12 km</quantity>
east of Khenifra,
<geoCoordinate id="EE465AC2FFA99905F73A686CFA5E7A24" box="[1298,1432,1861,1886]" direction="north" orientation="latitude" pageId="7" pageNumber="356" precision="5" value="32.9262">32.9262° N</geoCoordinate>
,
<geoCoordinate id="EE465AC2FFA99905F2BF6840FED77AF8" box="[151,273,1897,1922]" direction="west" orientation="longitude" pageId="7" pageNumber="356" precision="5" value="-5.5616">5.5616° W</geoCoordinate>
, alt.
<quantity id="4C8A91E0FFA99905F36E6840FE5B7AF8" box="[326,413,1897,1921]" metricMagnitude="3" metricUnit="m" metricValue="1.147" pageId="7" pageNumber="356" unit="m" value="1147.0">1147 m</quantity>
. a.s.l..
</paragraph>
<paragraph id="8BCD3C05FFA99905F2EF68A4FA4F7AB0" blockId="7.[151,1437,1716,1993]" pageId="7" pageNumber="356">
Other material: Four additional specimens were captured and released after collection of a
<collectionCode id="ED63A4C0FFA99905F6E668A4FACD7ADC" LSID="urn:lsid:biocol.org:col:15536" box="[1230,1291,1933,1957]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="7" pageNumber="356">DNA</collectionCode>
sample (one leg):
<specimenCount id="9D74F78CFFA99905F2F96898FF3B7AB0" box="[209,253,1969,1993]" pageId="7" pageNumber="356" type="generic">2 ♂</specimenCount>
,
<date id="FFCC1AC5FFA99905F3216898FE667AB0" box="[265,416,1969,1993]" pageId="7" pageNumber="356" value="2011-06-27">27 June 2011</date>
, sample codes: Df1027, Df1028; and
<specimenCount id="9D74F78CFFA99905F1606898FCB27AB0" box="[840,884,1969,1993]" pageId="7" pageNumber="356" type="generic">2 ♂</specimenCount>
,
<date id="FFCC1AC5FFA99905F1A86898FBDE7AB0" box="[896,1048,1969,1993]" pageId="7" pageNumber="356" value="2012-06-12">12 June 2012</date>
, sample codes: Df1745, Df1746.
</paragraph>
</subSubSection>
</treatment>
</document>