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<document id="56943B146B41CA82ECC23CD8A486BD68" ID-DOI="10.5281/zenodo.5331014" ID-ISSN="0374-1036" ID-Zenodo-Dep="5331014" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630168815873" checkinUser="diego" docAuthor="Megna, Yoandri S. &amp; Epler, John H." docDate="2012" docId="03CA7F7DFFE54C52FE3D99C94B47FDC1" docLanguage="en" docName="ActaEntMusNatPra.2012.52.2.383-410.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 52 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Copelatus insolitus Chevrolat 1863" docType="treatment" docVersion="4" lastPageNumber="398" masterDocId="FFF30705FFE84C5DFFAE99134914FFB2" masterDocTitle="A review of Copelatus from Cuba, with the description of two new species (Coleoptera: Dytiscidae: Copelatinae)" masterLastPageNumber="410" masterPageNumber="383" pageNumber="396" updateTime="1699171915230" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="7BB78C958EADB6F237AB80D8C93A08F7">A review of Copelatus from Cuba, with the description of two new species (Coleoptera: Dytiscidae: Copelatinae)</mods:title>
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<mods:namePart id="CAA0A3F1DFA9F0F92A9B6F8597A982B7">Megna, Yoandri S.</mods:namePart>
<mods:affiliation id="BA722A2DE23B148E22921676D73151B1">Departamento de Biología, Universidad de Oriente. Patricio Lumumba s / n, Santiago, Santiago de Cuba, Cuba; e-mail: ysuarez 1976 @ yahoo. es</mods:affiliation>
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<mods:namePart id="1C80E370BF1951B9E304E45711143CB2">Epler, John H.</mods:namePart>
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<treatment id="03CA7F7DFFE54C52FE3D99C94B47FDC1" ID-DOI="http://doi.org/10.5281/zenodo.5398802" ID-Zenodo-Dep="5398802" LSID="urn:lsid:plazi:treatment:03CA7F7DFFE54C52FE3D99C94B47FDC1" httpUri="http://treatment.plazi.org/id/03CA7F7DFFE54C52FE3D99C94B47FDC1" lastPageId="15" lastPageNumber="398" pageId="13" pageNumber="396">
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<paragraph id="8BDCCE6BFFE54C50FE3D99C94A0BFF41" blockId="13.[403,799,218,243]" box="[403,799,218,243]" pageId="13" pageNumber="396">
<heading id="D0947907FFE54C50FE3D99C94A0BFF41" bold="true" box="[403,799,218,243]" centered="true" fontSize="8" level="2" pageId="13" pageNumber="396" reason="3">
<taxonomicName id="4C63B5E8FFE54C50FE3D99C94A0BFF41" ID-CoL="Y39Q" authority="Chevrolat, 1863" authorityName="Chevrolat" authorityYear="1863" box="[403,799,218,243]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="396" phylum="Arthropoda" rank="species" species="insolitus">
<emphasis id="B9171279FFE54C50FE3D99C94A0BFF41" bold="true" box="[403,799,218,243]" pageId="13" pageNumber="396">
<emphasis id="B9171279FFE54C50FE3D99C94B70FF41" bold="true" box="[403,612,218,243]" italics="true" pageId="13" pageNumber="396">Copelatus insolitus</emphasis>
<bibRefCitation id="EFF2B39AFFE54C50FDC599C84A0BFF41" author="CHEVROLAT L. A. A." box="[619,799,219,243]" pageId="13" pageNumber="396" pagination="183 - 210" refId="ref12897" refString="CHEVROLAT L. A. A. 1863: Coleopteres de l'IIe de Cuba. Notes, synonymies et descriptions d'especes nouvelles. Familles des cicindeletes, carabiques, dytiscides, gyrinides et palpicornes. Annales de la Societe Entomologique de France 3: 183 - 210." type="journal article" year="1863">Chevrolat, 1863</bibRefCitation>
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<paragraph id="8BDCCE6BFFE54C50FDAB99EA4BBAFEBE" blockId="13.[113,1089,249,356]" box="[517,686,249,269]" pageId="13" pageNumber="396">
(
<figureCitation id="1358D2EEFFE54C50FDA599EA4B53FEBE" box="[523,583,249,268]" captionStart="Figs" captionStartId="6.[151,189,1533,1552]" captionTargetBox="[166,1114,432,1505]" captionTargetId="figure-11@6.[151,1126,417,1515]" captionTargetPageId="6" captionText="Figs.1–8. Habitus and color patterns of Copelatus species from Cuba.1 – C.barbouri Young, 1942; 2 – C. caelatipennis angustatus Chevrolat, 1863; 3 – C. cordovai sp. nov.; 4 – C. cubaensis Schaeffer, 1908; 5 – Copelatus darlingtoni Young, 1942; 6 – C. insolitus Chevrolat, 1863; 7 – C. montivagus Young, 1942; 8 – C. posticatus (Fabricius, 1801)." figureDoi="http://doi.org/10.5281/zenodo.5331018" httpUri="https://zenodo.org/record/5331018/files/figure.png" pageId="13" pageNumber="396">Figs. 6</figureCitation>
,
<figureCitation id="1358D2EEFFE54C50FDFF99E94B9CFEBF" box="[593,648,250,269]" captionStart="Figs" captionStartId="13.[113,152,1517,1536]" captionTargetBox="[145,1055,494,1485]" captionTargetId="figure-13@13.[133,1069,485,1499]" captionTargetPageId="13" captionText="Figs. 12–19. Male genitalia (12–15) and left protibia (16–19) of Cuban species of Copelatus. 12, 16 – C. barbouri Young, 1942; 13, 17 – C. darlingtoni Young, 1942; 14, 18 – C. insolitus Chevrolat, 1863; 15, 19 – C. montivagus Young, 1942. Parts of male genitalia: a – median lobe of the aedeagus in left lateral view, b – same in ventral lateral view, c – left paramere in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5331022" httpUri="https://zenodo.org/record/5331022/files/figure.png" pageId="13" pageNumber="396">14, 18</figureCitation>
,
<figureCitation id="1358D2EEFFE54C50FD3F99EA4BB3FEBE" box="[657,679,249,268]" captionStart="Figs" captionStartId="20.[151,190,1516,1535]" captionTargetBox="[165,1104,223,1503]" captionTargetId="figure-11@20.[161,1117,219,1504]" captionTargetPageId="20" captionText="Figs. 28–31. Known distribution of Copelatus species in Cuba. 28 – C. cubaensis Schaeffer, 1908; 29 – C. danyi sp. nov.; 30 – C. darlingtoni Young, 1942; 31 – C. insolitus Chevrolat, 1863. See Appendix 1 for additional information on the localities reported in each map." figureDoi="http://doi.org/10.5281/zenodo.5331032" httpUri="https://zenodo.org/record/5331032/files/figure.png" pageId="13" pageNumber="396">31</figureCitation>
)
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<treatmentCitation id="0AC2E87AFFE54C50FFDF980E48C7FE9D" author="CHEVROLAT L. A. A." box="[113,467,284,303]" page="201" pageId="13" pageNumber="396" year="1863">
<taxonomicName id="4C63B5E8FFE54C50FFDF980E48C7FE9D" ID-CoL="Y39Q" authority="Chevrolat, 1863: 201" authorityName="Chevrolat" authorityPageNumber="201" authorityYear="1863" box="[113,467,284,303]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="396" phylum="Arthropoda" rank="species" species="insolitus">
<emphasis id="B9171279FFE54C50FFDF980E4801FE9D" box="[113,277,285,303]" italics="true" pageId="13" pageNumber="396">Copelatus insolitus</emphasis>
<bibRefCitation id="EFF2B39AFFE54C50FEB5980F48C7FE9D" author="CHEVROLAT L. A. A." box="[283,467,284,303]" pageId="13" pageNumber="396" pagination="183 - 210" refId="ref12897" refString="CHEVROLAT L. A. A. 1863: Coleopteres de l'IIe de Cuba. Notes, synonymies et descriptions d'especes nouvelles. Familles des cicindeletes, carabiques, dytiscides, gyrinides et palpicornes. Annales de la Societe Entomologique de France 3: 183 - 210." type="journal article" year="1863">Chevrolat, 1863: 201</bibRefCitation>
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.
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<treatmentCitationGroup id="AB73E945FFE54C50FFDF982449E4FED6" pageId="13" pageNumber="396">
<treatmentCitation id="0AC2E87AFFE54C50FFDF982448F7FEFB" author="GUNDLACH J. C." box="[113,483,310,330]" page=")" pageId="13" pageNumber="396" year="1891">
<taxonomicName id="4C63B5E8FFE54C50FFDF982448F7FEFB" ID-CoL="Y39Q" authority=": GUNDLACH (1891: 37)" authorityName=": GUNDLACH" authorityPageNumber="37" authorityYear="1891" box="[113,483,310,330]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="396" phylum="Arthropoda" rank="species" species="insolitus">
<emphasis id="B9171279FFE54C50FFDF9824480CFEFB" box="[113,280,311,329]" italics="true" pageId="13" pageNumber="396">Copelatus insolitus</emphasis>
:
<bibRefCitation id="EFF2B39AFFE54C50FE8B982548F7FEFB" author="GUNDLACH J. C." box="[293,483,310,329]" pageId="13" pageNumber="396" pagination="33 - 42" refId="ref13245" refString="GUNDLACH J. C. 1891: Coleoptera. Contribucion a la entomologia Cubana. Habana. Anales de la Academia de Ciencias Medicas Fisicas y Naturales 3: 33 - 42." type="journal article" year="1891">GUNDLACH (1891: 37)</bibRefCitation>
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,
<treatmentCitation id="0AC2E87AFFE54C50FE4398244B99FEFB" author="YOUNG F. N." box="[493,653,310,330]" page="83" pageId="13" pageNumber="396" year="1942">
<bibRefCitation id="EFF2B39AFFE54C50FE4398244B99FEFB" author="YOUNG F. N." box="[493,653,310,330]" pageId="13" pageNumber="396" pagination="79 - 94" refId="ref14016" refString="YOUNG F. N. 1942: New species of Copelatus from the West Indies (Coleoptera: Dytiscidae). Proceedings of the New England Zoological Club 20: 79 - 94." type="journal article" year="1942">YOUNG (1942: 83)</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="0AC2E87AFFE54C50FD3698254A4CFEFB" author="SPANGLER P. J." box="[664,856,310,329]" page="153" pageId="13" pageNumber="396" year="1981">
<bibRefCitation id="EFF2B39AFFE54C50FD3698254A4CFEFB" author="SPANGLER P. J." box="[664,856,310,329]" pageId="13" pageNumber="396" pagination="145 - 171" refId="ref13928" refString="SPANGLER P. J. 1981: Supplement to the aquatic and semiaquatic Coleoptera of Cuba collected by the biospeleological expeditions to Cuba by the Academies of Science of Cuba and Romania. Resultats des Expeditions Biospeologiques Cubano Roumaines a Cuba 3: 145 - 171." type="journal article" year="1981">SPANGLER (1981: 153)</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="0AC2E87AFFE54C50FCCD98244D1FFEFB" author="NILSSON A. N." box="[867,1035,310,330]" page="66" pageId="13" pageNumber="396" year="2001">
<bibRefCitation id="EFF2B39AFFE54C50FCCD98244D1FFEFB" author="NILSSON A. N." box="[867,1035,310,330]" pageId="13" pageNumber="396" pagination="1 - 395" refId="ref13702" refString="NILSSON A. N. 2001: Dytiscidae (Coleoptera). World Catalogue of Insects 3: 1 - 395." type="journal article" year="2001">NILSSON (2001: 66)</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="0AC2E87AFFE54C50FBB9982449F8FED6" author="PECK S. B." page="44" pageId="13" pageNumber="396" year="2005">
<bibRefCitation id="EFF2B39AFFE54C50FBB9982449F8FED6" author="PECK S. B." pageId="13" pageNumber="396" pagination="1 - 241" refId="ref13788" refString="PECK S. B. 2005: A checklist of the beetles of Cuba with data on distribution and bionomics (Insecta: Coleoptera). Arthropods of Florida and Neighboring Land Areas 18: 1 - 241." type="journal article" year="2005">PECK (2005: 44)</bibRefCitation>
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.
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<emphasis id="B9171279FFE54C50FFDF986849FFFE3C" bold="true" box="[113,235,378,398]" pageId="13" pageNumber="396">
<typeStatus id="54D870C9FFE54C50FFDF9868498AFE3C" box="[113,158,379,398]" pageId="13" pageNumber="396">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3748EFBFFE54C50FF5E9868480BFE3C" box="[240,287,379,398]" name="Cuba" pageId="13" pageNumber="396">Cuba</collectingCountry>
,
<collectingRegion id="49A70089FFE54C50FE87986848CFFE3C" box="[297,475,379,398]" country="Cuba" name="La Habana" pageId="13" pageNumber="396">La Habana Province</collectingRegion>
. The
<typeStatus id="54D870C9FFE54C50FDA498684B3BFE3C" box="[522,559,379,398]" pageId="13" pageNumber="396">type</typeStatus>
locality of
<taxonomicName id="4C63B5E8FFE54C50FD3D986F4BE0FE3C" authorityName=": GUNDLACH" authorityYear="1891" box="[659,756,380,398]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="396" phylum="Arthropoda" rank="species" species="insolitus">
<emphasis id="B9171279FFE54C50FD3D986F4BE0FE3C" box="[659,756,380,398]" italics="true" pageId="13" pageNumber="396">C. insolitus</emphasis>
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is
<collectingRegion id="49A70089FFE54C50FCA198684A46FE3C" box="[783,850,379,398]" country="Cuba" name="La Habana" pageId="13" pageNumber="396">Havana</collectingRegion>
, but doubt exists on the occurrence of this species there due to the fact that all material examined previously from
<collectingCountry id="F3748EFBFFE54C50FCF298864A9FFE1A" box="[860,907,405,424]" name="Cuba" pageId="13" pageNumber="396">Cuba</collectingCountry>
,
<collectingCountry id="F3748EFBFFE54C50FC3A98864ACDFE1A" box="[916,985,405,424]" name="Jamaica" pageId="13" pageNumber="396">Jamaica</collectingCountry>
and Central America has been from rather high mountains (
<bibRefCitation id="EFF2B39AFFE54C50FDAB98A34B60FE71" author="YOUNG F. N." box="[517,628,432,451]" pageId="13" pageNumber="396" pagination="79 - 94" refId="ref14016" refString="YOUNG F. N. 1942: New species of Copelatus from the West Indies (Coleoptera: Dytiscidae). Proceedings of the New England Zoological Club 20: 79 - 94." type="journal article" year="1942">YOUNG 1942</bibRefCitation>
).
</paragraph>
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<paragraph id="8BDCCE6BFFE54C50FFDF9CFE48ADF9E2" blockId="13.[113,1089,1517,1617]" pageId="13" pageNumber="396">
Figs. 12–19. Male genitalia (12–15) and left protibia (16–19) of Cuban species of
<taxonomicName id="4C63B5E8FFE54C50FC819CFD4A92F9B2" authorityName=": GUEORGUIEV" authorityYear="1968" box="[815,902,1518,1536]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="396" phylum="Arthropoda" rank="genus">
<emphasis id="B9171279FFE54C50FC819CFD4A92F9B2" box="[815,902,1518,1536]" italics="true" pageId="13" pageNumber="396">Copelatus</emphasis>
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. 12, 16 –
<taxonomicName id="4C63B5E8FFE54C50FC729CFD49F5F9A9" authority="Young, 1942" authorityName="Young" authorityYear="1942" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="396" phylum="Arthropoda" rank="species" species="barbouri">
<emphasis id="B9171279FFE54C50FC729CFD4D55F9B2" box="[988,1089,1518,1536]" italics="true" pageId="13" pageNumber="396">C. barbouri</emphasis>
<bibRefCitation id="EFF2B39AFFE54C50FFDF9F1B49F5F9A9" author="YOUNG F. N." box="[113,225,1544,1563]" pageId="13" pageNumber="396" pagination="79 - 94" refId="ref14016" refString="YOUNG F. N. 1942: New species of Copelatus from the West Indies (Coleoptera: Dytiscidae). Proceedings of the New England Zoological Club 20: 79 - 94." type="journal article" year="1942">Young, 1942</bibRefCitation>
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; 13, 17 –
<taxonomicName id="4C63B5E8FFE54C50FE999F1A4B30F9A9" authority="Young, 1942" authorityName="Young" authorityYear="1942" box="[311,548,1544,1563]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="396" phylum="Arthropoda" rank="species" species="darlingtoni">
<emphasis id="B9171279FFE54C50FE999F1A48A4F9A9" box="[311,432,1545,1563]" italics="true" pageId="13" pageNumber="396">C. darlingtoni</emphasis>
<bibRefCitation id="EFF2B39AFFE54C50FE189F1B4B30F9A9" author="YOUNG F. N." box="[438,548,1544,1563]" pageId="13" pageNumber="396" pagination="79 - 94" refId="ref14016" refString="YOUNG F. N. 1942: New species of Copelatus from the West Indies (Coleoptera: Dytiscidae). Proceedings of the New England Zoological Club 20: 79 - 94." type="journal article" year="1942">Young, 1942</bibRefCitation>
</taxonomicName>
; 14, 18 –
<taxonomicName id="4C63B5E8FFE54C50FDD29F1A4A64F9A9" authority="Chevrolat, 1863" authorityName="Chevrolat" authorityYear="1863" box="[636,880,1544,1563]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="396" phylum="Arthropoda" rank="species" species="insolitus">
<emphasis id="B9171279FFE54C50FDD29F1A4BC9F9A9" box="[636,733,1545,1563]" italics="true" pageId="13" pageNumber="396">C. insolitus</emphasis>
<bibRefCitation id="EFF2B39AFFE54C50FD4D9F1B4A64F9A9" author="CHEVROLAT L. A. A." box="[739,880,1544,1563]" pageId="13" pageNumber="396" pagination="183 - 210" refId="ref12897" refString="CHEVROLAT L. A. A. 1863: Coleopteres de l'IIe de Cuba. Notes, synonymies et descriptions d'especes nouvelles. Familles des cicindeletes, carabiques, dytiscides, gyrinides et palpicornes. Annales de la Societe Entomologique de France 3: 183 - 210." type="journal article" year="1863">Chevrolat, 1863</bibRefCitation>
</taxonomicName>
; 15, 19 –
<taxonomicName id="4C63B5E8FFE54C50FC6B9F1A49F4F984" authority="Young, 1942" authorityName="Young" authorityYear="1942" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="396" phylum="Arthropoda" rank="species" species="montivagus">
<emphasis id="B9171279FFE54C50FC6B9F1A4D54F9A9" box="[965,1088,1545,1563]" italics="true" pageId="13" pageNumber="396">C. montivagus</emphasis>
<bibRefCitation id="EFF2B39AFFE54C50FFDF9F3049F4F984" author="YOUNG F. N." box="[113,224,1571,1590]" pageId="13" pageNumber="396" pagination="79 - 94" refId="ref14016" refString="YOUNG F. N. 1942: New species of Copelatus from the West Indies (Coleoptera: Dytiscidae). Proceedings of the New England Zoological Club 20: 79 - 94." type="journal article" year="1942">Young, 1942</bibRefCitation>
</taxonomicName>
. Parts of male genitalia: a – median lobe of the aedeagus in left lateral view, b – same in ventral lateral view, c – left paramere in lateral view.
</paragraph>
</caption>
<paragraph id="8BDCCE6BFFE64C53FF3999C84B35FE18" blockId="14.[151,1127,219,426]" pageId="14" pageNumber="397">
<emphasis id="B9171279FFE64C53FF3999C8485FFF5D" bold="true" box="[151,331,219,239]" pageId="14" pageNumber="397">Material examined.</emphasis>
<materialsCitation id="3B0BC436FFE64C53FEFD99C84D0DFF5D" box="[339,1049,219,239]" collectorName="Y. S. Megna" country="Cuba" location="Tope de Collantes" pageId="14" pageNumber="397" specimenCount="19" stateProvince="Sancti Spiritus">
<emphasis id="B9171279FFE64C53FEFD99C84B25FF5D" bold="true" box="[339,561,219,239]" pageId="14" pageNumber="397">
<collectingCountry id="F3748EFBFFE64C53FEFD99C84887FF5D" box="[339,403,219,239]" name="Cuba" pageId="14" pageNumber="397">CUBA</collectingCountry>
:
<collectingRegion id="49A70089FFE64C53FE3099C848CEFF5F" box="[414,474,219,239]" country="Cuba" name="Sancti Spiritus" pageId="14" pageNumber="397">SANCTI</collectingRegion>
SPÍRITUS:
</emphasis>
<location id="8EBC98B0FFE64C53FD9699CF4BCDFF5C" LSID="urn:lsid:plazi:treatment:03CA7F7DFFE54C52FE3D99C94B47FDC1:8EBC98B0FFE64C53FD9699CF4BCDFF5C" box="[568,729,219,239]" country="Cuba" name="Tope de Collantes" pageId="14" pageNumber="397" stateProvince="Sancti Spiritus">Tope de Collantes</location>
, 29.vi.10,
<collectorName id="2696ABBDFFE64C53FC9899CF4AB7FF5D" box="[822,931,220,239]" pageId="14" pageNumber="397">Y. S. Megna</collectorName>
leg.,
<specimenCount id="9D6505E2FFE64C53FC7999CF4D0DFF5D" box="[983,1049,220,239]" pageId="14" pageNumber="397" type="generic">19 exs.</specimenCount>
</materialsCitation>
;
<materialsCitation id="3B0BC436FFE64C53FB8D99C848F7FEB8" collectingDate="2010-07-01" collectorName="Y. S. Megna" country="Cuba" location="Codina" pageId="14" pageNumber="397" specimenCount="13" stateProvince="Sancti Spiritus">
<location id="8EBC98B0FFE64C53FB8D99C84D77FF5C" LSID="urn:lsid:plazi:treatment:03CA7F7DFFE54C52FE3D99C94B47FDC1:8EBC98B0FFE64C53FB8D99C84D77FF5C" box="[1059,1123,219,238]" country="Cuba" name="Codina" pageId="14" pageNumber="397" stateProvince="Sancti Spiritus">Codina</location>
,
<date id="FFDDE8ABFFE64C53FF3999E549EEFEBB" box="[151,250,246,265]" pageId="14" pageNumber="397" value="2010-07-01">
<collectingDate id="EF991143FFE64C53FF3999E549EEFEBB" box="[151,250,246,265]" pageId="14" pageNumber="397" value="2010-07-01">01.vii.2010</collectingDate>
</date>
,
<collectorName id="2696ABBDFFE64C53FEAA99E44864FEB8" box="[260,368,246,266]" pageId="14" pageNumber="397">Y. S. Megna</collectorName>
leg.,
<specimenCount id="9D6505E2FFE64C53FE0D99E548F7FEB8" box="[419,483,246,266]" pageId="14" pageNumber="397" type="generic">13 exs.</specimenCount>
</materialsCitation>
<materialsCitation id="3B0BC436FFE64C53FE4699E64D70FEB8" box="[488,1124,245,266]" collectingDate="2004-08-14" collectorName="La Majagua &amp; Y. S. Megna" country="Cuba" location="SANTIAGO DE CUBA" pageId="14" pageNumber="397" specimenCount="74" stateProvince="Santiago de Cuba">
<emphasis id="B9171279FFE64C53FE4699E64B80FEBB" bold="true" box="[488,660,245,265]" pageId="14" pageNumber="397">
<collectingRegion id="49A70089FFE64C53FE4699E64B9AFEBA" box="[488,654,245,265]" country="Cuba" name="Santiago de Cuba" pageId="14" pageNumber="397">SANTIAGO DE CUBA</collectingRegion>
:
</emphasis>
<collectorName id="2696ABBDFFE64C53FD3299E44A12FEBB" box="[668,774,246,266]" pageId="14" pageNumber="397">La Majagua</collectorName>
,
<date id="FFDDE8ABFFE64C53FCBE99E54A6DFEBB" box="[784,889,246,265]" pageId="14" pageNumber="397" value="2004-08-14">
<collectingDate id="EF991143FFE64C53FCBE99E54A6DFEBB" box="[784,889,246,265]" pageId="14" pageNumber="397" value="2004-08-14">14.viii.2004</collectingDate>
</date>
,
<collectorName id="2696ABBDFFE64C53FC2D99E44AFBFEB8" box="[899,1007,246,266]" pageId="14" pageNumber="397">Y. S. Megna</collectorName>
leg.,
<specimenCount id="9D6505E2FFE64C53FB8C99E54D70FEB8" box="[1058,1124,246,266]" pageId="14" pageNumber="397" type="generic">74 exs.</specimenCount>
</materialsCitation>
;
<materialsCitation id="3B0BC436FFE64C53FF3998024B23FE96" box="[151,567,273,292]" collectingDate="2004-06-12" collectorName="Y. S. Megna" country="Cuba" location="La Platica" pageId="14" pageNumber="397" specimenCount="3" stateProvince="Santiago de Cuba">
<location id="8EBC98B0FFE64C53FF39980249E5FE96" LSID="urn:lsid:plazi:treatment:03CA7F7DFFE54C52FE3D99C94B47FDC1:8EBC98B0FFE64C53FF39980249E5FE96" box="[151,241,273,292]" country="Cuba" name="La Platica" pageId="14" pageNumber="397" stateProvince="Santiago de Cuba">La Platica</location>
,
<date id="FFDDE8ABFFE64C53FF559802484CFE96" box="[251,344,273,292]" pageId="14" pageNumber="397" value="2004-06-12">
<collectingDate id="EF991143FFE64C53FF559802484CFE96" box="[251,344,273,292]" pageId="14" pageNumber="397" value="2004-06-12">12.vi.2004</collectingDate>
</date>
,
<collectorName id="2696ABBDFFE64C53FECC980248DAFE96" box="[354,462,273,292]" pageId="14" pageNumber="397">Y. S. Megna</collectorName>
leg.,
<specimenCount id="9D6505E2FFE64C53FDAF98024B23FE96" box="[513,567,273,292]" pageId="14" pageNumber="397" type="generic">3 exs.</specimenCount>
</materialsCitation>
;
<materialsCitation id="3B0BC436FFE64C53FDEF98024AEDFE96" box="[577,1017,273,292]" collectingDate="2005-12-01" collectorName="Y. S. Megna" country="Cuba" location="La Estrella" pageId="14" pageNumber="397" specimenCount="22" stateProvince="Santiago de Cuba">
<location id="8EBC98B0FFE64C53FDEF98024BB5FE96" LSID="urn:lsid:plazi:treatment:03CA7F7DFFE54C52FE3D99C94B47FDC1:8EBC98B0FFE64C53FDEF98024BB5FE96" box="[577,673,273,292]" country="Cuba" name="La Estrella" pageId="14" pageNumber="397" stateProvince="Santiago de Cuba">La Estrella</location>
,
<date id="FFDDE8ABFFE64C53FD0298024A1BFE96" box="[684,783,273,292]" pageId="14" pageNumber="397" value="2005-12-01">
<collectingDate id="EF991143FFE64C53FD0298024A1BFE96" box="[684,783,273,292]" pageId="14" pageNumber="397" value="2005-12-01">01.xii.2005</collectingDate>
</date>
,
<collectorName id="2696ABBDFFE64C53FCB798024A91FE96" box="[793,901,273,292]" pageId="14" pageNumber="397">Y. S. Megna</collectorName>
leg.,
<specimenCount id="9D6505E2FFE64C53FC1698024AEDFE96" box="[952,1017,273,292]" pageId="14" pageNumber="397" type="generic">22 exs.</specimenCount>
</materialsCitation>
;
<materialsCitation id="3B0BC436FFE64C53FBAC980248C3FE8D" collectingDate="2005-12-02" collectorName="Y. S. Megna" country="Cuba" location="La Cubana" pageId="14" pageNumber="397" specimenCount="25" stateProvince="Santiago de Cuba">
<location id="8EBC98B0FFE64C53FBAC98024D77FE96" LSID="urn:lsid:plazi:treatment:03CA7F7DFFE54C52FE3D99C94B47FDC1:8EBC98B0FFE64C53FBAC98024D77FE96" box="[1026,1123,273,292]" country="Cuba" name="La Cubana" pageId="14" pageNumber="397" stateProvince="Santiago de Cuba">La Cubana</location>
,
<date id="FFDDE8ABFFE64C53FF39983849EEFE8C" box="[151,250,299,318]" pageId="14" pageNumber="397" value="2005-12-02">
<collectingDate id="EF991143FFE64C53FF39983849EEFE8C" box="[151,250,299,318]" pageId="14" pageNumber="397" value="2005-12-02">02.xii.2005</collectingDate>
</date>
,
<collectorName id="2696ABBDFFE64C53FEAC983F487DFE8D" box="[258,361,300,319]" pageId="14" pageNumber="397">Y. S. Megna</collectorName>
leg.,
<specimenCount id="9D6505E2FFE64C53FE36983848C3FE8D" box="[408,471,299,319]" pageId="14" pageNumber="397" type="generic">25 exs.</specimenCount>
</materialsCitation>
;
<materialsCitation id="3B0BC436FFE64C53FE70983F4D70FE8D" box="[478,1124,299,319]" collectingDate="2005-11-16" collectorName="Y. S. Megna &amp; A. Deler-Hernandez" country="Cuba" location="Las Candelarias" pageId="14" pageNumber="397" specimenCount="3" stateProvince="Santiago de Cuba">
<location id="8EBC98B0FFE64C53FE70983F4B7CFE8C" LSID="urn:lsid:plazi:treatment:03CA7F7DFFE54C52FE3D99C94B47FDC1:8EBC98B0FFE64C53FE70983F4B7CFE8C" box="[478,616,299,319]" country="Cuba" name="Las Candelarias" pageId="14" pageNumber="397" stateProvince="Santiago de Cuba">Las Candelarias</location>
,
<date id="FFDDE8ABFFE64C53FDDE98384BD9FE8C" box="[624,717,299,318]" pageId="14" pageNumber="397" value="2005-11-16">
<collectingDate id="EF991143FFE64C53FDDE98384BD9FE8C" box="[624,717,299,318]" pageId="14" pageNumber="397" value="2005-11-16">16.xi.2005</collectingDate>
</date>
,
<collectorName id="2696ABBDFFE64C53FD7B983F4A28FE8D" box="[725,828,300,319]" pageId="14" pageNumber="397">Y. S. Megna</collectorName>
&amp;
<collectorName id="2696ABBDFFE64C53FCFB983F4D15FE8C" box="[853,1025,299,319]" pageId="14" pageNumber="397">A. Deler-Hernández</collectorName>
leg.,
<specimenCount id="9D6505E2FFE64C53FB81983F4D70FE8D" box="[1071,1124,300,319]" pageId="14" pageNumber="397" type="generic">3 exs.</specimenCount>
</materialsCitation>
;
<materialsCitation id="3B0BC436FFE64C53FF3998554817FEC6" collectingDate="2005-05-01" collectingDateMax="2005-08-03" collectingDateMin="2005-05-01" collectorName="Y. S. Megna &amp; Y. Alarie &amp; A. Deler-Hernandez &amp; Deler-Hernandez" country="Cuba" location="Gran Piedra" pageId="14" pageNumber="397" specimenCount="35" stateProvince="Santiago de Cuba">
<location id="8EBC98B0FFE64C53FF3998554815FEEB" LSID="urn:lsid:plazi:treatment:03CA7F7DFFE54C52FE3D99C94B47FDC1:8EBC98B0FFE64C53FF3998554815FEEB" box="[151,257,326,345]" country="Cuba" name="Gran Piedra" pageId="14" pageNumber="397" stateProvince="Santiago de Cuba">Gran Piedra</location>
,
<date id="FFDDE8ABFFE64C53FEA49855484BFEEB" box="[266,351,326,345]" pageId="14" pageNumber="397" value="2005-05-01">
<collectingDate id="EF991143FFE64C53FEA49855484BFEEB" box="[266,351,326,345]" pageId="14" pageNumber="397" value="2005-05-01">01.v.2005</collectingDate>
</date>
,
<collectorName id="2696ABBDFFE64C53FEC6985448C1FEE8" box="[360,469,326,346]" pageId="14" pageNumber="397">Y. S. Megna</collectorName>
,
<collectorName id="2696ABBDFFE64C53FE7098544B3DFEEB" box="[478,553,326,346]" pageId="14" pageNumber="397">Y. Alarie</collectorName>
&amp;
<collectorName id="2696ABBDFFE64C53FDEB98554BE6FEEB" box="[581,754,326,345]" pageId="14" pageNumber="397">A. Deler-Hernández</collectorName>
leg.,
<specimenCount id="9D6505E2FFE64C53FC8D98554A4DFEE8" box="[803,857,326,346]" pageId="14" pageNumber="397" type="generic">3 exs.</specimenCount>
,
<date id="FFDDE8ABFFE64C53FCCF98554ADFFEEB" box="[865,971,326,345]" pageId="14" pageNumber="397" value="2005-08-03">
<collectingDate id="EF991143FFE64C53FCCF98554ADFFEEB" box="[865,971,326,345]" pageId="14" pageNumber="397" value="2005-08-03">03.viii.2005</collectingDate>
</date>
,
<collectorName id="2696ABBDFFE64C53FC7A98554D72FEEB" box="[980,1126,326,345]" pageId="14" pageNumber="397">Deler-Hernández</collectorName>
leg.,
<specimenCount id="9D6505E2FFE64C53FF6C98724817FEC6" box="[194,259,353,372]" pageId="14" pageNumber="397" type="generic">32 exs.</specimenCount>
</materialsCitation>
;
<materialsCitation id="3B0BC436FFE64C53FEA398724BF5FEC6" box="[269,737,353,372]" collectingDate="2005-08-04" collectorName="Deler-Hernandez" country="Cuba" location="El Olimpo" pageId="14" pageNumber="397" specimenCount="7" stateProvince="Santiago de Cuba">
<location id="8EBC98B0FFE64C53FEA39872487DFEC6" LSID="urn:lsid:plazi:treatment:03CA7F7DFFE54C52FE3D99C94B47FDC1:8EBC98B0FFE64C53FEA39872487DFEC6" box="[269,361,353,372]" country="Cuba" name="El Olimpo" pageId="14" pageNumber="397" stateProvince="Santiago de Cuba">El Olimpo</location>
,
<date id="FFDDE8ABFFE64C53FEDD987248C9FEC6" box="[371,477,353,372]" pageId="14" pageNumber="397" value="2005-08-04">
<collectingDate id="EF991143FFE64C53FEDD987248C9FEC6" box="[371,477,353,372]" pageId="14" pageNumber="397" value="2005-08-04">04.viii.2005</collectingDate>
</date>
,
<collectorName id="2696ABBDFFE64C53FE4998724B6DFEC6" box="[487,633,353,372]" pageId="14" pageNumber="397">Deler-Hernández</collectorName>
leg.,
<specimenCount id="9D6505E2FFE64C53FD0598724BF5FEC6" box="[683,737,353,372]" pageId="14" pageNumber="397" type="generic">7 exs.</specimenCount>
</materialsCitation>
;
<materialsCitation id="3B0BC436FFE64C53FD45987249D6FE3D" collectingDate="2008-06-18" collectorName="Y. S. Megna" country="Cuba" location="Los Morones" pageId="14" pageNumber="397" specimenCount="1" stateProvince="Santiago de Cuba">
<location id="8EBC98B0FFE64C53FD4598724A4BFEC6" LSID="urn:lsid:plazi:treatment:03CA7F7DFFE54C52FE3D99C94B47FDC1:8EBC98B0FFE64C53FD4598724A4BFEC6" box="[747,863,353,372]" country="Cuba" name="Los Morones" pageId="14" pageNumber="397" stateProvince="Santiago de Cuba">Los Morones</location>
,
<date id="FFDDE8ABFFE64C53FCC798724AD2FEC6" box="[873,966,353,372]" pageId="14" pageNumber="397" value="2008-06-18">
<collectingDate id="EF991143FFE64C53FCC798724AD2FEC6" box="[873,966,353,372]" pageId="14" pageNumber="397" value="2008-06-18">18.vi.2008</collectingDate>
</date>
,
<collectorName id="2696ABBDFFE64C53FC6198724D2FFEC6" box="[975,1083,353,372]" pageId="14" pageNumber="397">Y. S. Megna</collectorName>
leg.,
<specimenCount id="9D6505E2FFE64C53FF39986F49D6FE3D" box="[151,194,380,399]" pageId="14" pageNumber="397" type="generic">1 ex.</specimenCount>
</materialsCitation>
;
<materialsCitation id="3B0BC436FFE64C53FF67986F4BB2FE3D" box="[201,678,379,399]" collectingDate="2009-06-25" collectorName="Deler-Hernandez" country="Cuba" location="Loma del Gato" pageId="14" pageNumber="397" specimenCount="8" stateProvince="Santiago de Cuba">
<location id="8EBC98B0FFE64C53FF67986F485EFE3D" LSID="urn:lsid:plazi:treatment:03CA7F7DFFE54C52FE3D99C94B47FDC1:8EBC98B0FFE64C53FF67986F485EFE3D" box="[201,330,379,399]" country="Cuba" name="Loma del Gato" pageId="14" pageNumber="397" stateProvince="Santiago de Cuba">Loma del Gato</location>
,
<date id="FFDDE8ABFFE64C53FEFE986848BAFE3C" box="[336,430,379,398]" pageId="14" pageNumber="397" value="2009-06-25">
<collectingDate id="EF991143FFE64C53FEFE986848BAFE3C" box="[336,430,379,398]" pageId="14" pageNumber="397" value="2009-06-25">25.vi.2009</collectingDate>
</date>
,
<collectorName id="2696ABBDFFE64C53FE1B98684B53FE3C" box="[437,583,379,398]" pageId="14" pageNumber="397">Deler-Hernández</collectorName>
leg.,
<specimenCount id="9D6505E2FFE64C53FDDB986F4BB2FE3D" box="[629,678,380,399]" pageId="14" pageNumber="397" type="generic">8 exs.</specimenCount>
</materialsCitation>
<materialsCitation id="3B0BC436FFE64C53FD07986849FAFE18" collectingDate="2004-02-07" collectorName="Y. S. Megna" country="Georgia" location="Monte" pageId="14" pageNumber="397" specimenCount="1" stateProvince="Guantanamo">
<emphasis id="B9171279FFE64C53FD0798684A31FE3D" bold="true" box="[681,805,379,399]" pageId="14" pageNumber="397">
<collectingRegion id="49A70089FFE64C53FD0798684A0AFE3F" box="[681,798,379,399]" country="Cuba" name="Guantanamo" pageId="14" pageNumber="397">GUANTÁNAMO</collectingRegion>
:
</emphasis>
<location id="8EBC98B0FFE64C53FC87986F4A75FE3D" LSID="urn:lsid:plazi:treatment:03CA7F7DFFE54C52FE3D99C94B47FDC1:8EBC98B0FFE64C53FC87986F4A75FE3D" box="[809,865,380,399]" country="Georgia" name="Monte" pageId="14" pageNumber="397" stateProvince="Guantanamo">Monte</location>
<collectingCountry id="F3748EFBFFE64C53FCCB98684A8DFE3C" box="[869,921,379,398]" name="Georgia" pageId="14" pageNumber="397">Iberia</collectingCountry>
,
<date id="FFDDE8ABFFE64C53FC0E98684AEDFE3C" box="[928,1017,379,398]" pageId="14" pageNumber="397" value="2004-02-07">
<collectingDate id="EF991143FFE64C53FC0E98684AEDFE3C" box="[928,1017,379,398]" pageId="14" pageNumber="397" value="2004-02-07">07.ii.2004</collectingDate>
</date>
,
<collectorName id="2696ABBDFFE64C53FBAE986F4D72FE3D" box="[1024,1126,380,399]" pageId="14" pageNumber="397">Y. S. Megna</collectorName>
leg.,
<specimenCount id="9D6505E2FFE64C53FF6F988549FAFE18" box="[193,238,406,426]" pageId="14" pageNumber="397" type="generic">1 ex.</specimenCount>
</materialsCitation>
;
<materialsCitation id="3B0BC436FFE64C53FF5898854B35FE18" box="[246,545,406,426]" collectingDate="2003-10-04" collectorName="Y. S. Megna" country="Cuba" isTailEnumeration="true" location="Monte" pageId="14" pageNumber="397" specimenCount="2" stateProvince="Guantanamo">
<date id="FFDDE8ABFFE64C53FF5898854859FE1B" box="[246,333,406,425]" pageId="14" pageNumber="397" value="2003-10-04">
<collectingDate id="EF991143FFE64C53FF5898854859FE1B" box="[246,333,406,425]" pageId="14" pageNumber="397" value="2003-10-04">04.x.2003</collectingDate>
</date>
,
<collectorName id="2696ABBDFFE64C53FEFB988448AAFE18" box="[341,446,406,426]" pageId="14" pageNumber="397">Y. S. Megna</collectorName>
leg.,
<specimenCount id="9D6505E2FFE64C53FE4098854B35FE18" box="[494,545,406,426]" pageId="14" pageNumber="397" type="generic">2 exs.</specimenCount>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3799DE0FFE64C53FF3998D24802FD8B" pageId="14" pageNumber="397" type="diagnosis">
<paragraph id="8BDCCE6BFFE64C53FF3998D24802FD8B" blockId="14.[151,1127,449,698]" pageId="14" pageNumber="397">
<emphasis id="B9171279FFE64C53FF3998D24818FE6B" bold="true" box="[151,268,449,473]" pageId="14" pageNumber="397">Diagnosis.</emphasis>
TL
<quantity id="4C9B638EFFE64C53FE9898D248ACFE6B" box="[310,440,449,473]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" metricValueMax="5.9" metricValueMin="5.3" pageId="14" pageNumber="397" unit="mm" value="5.6" valueMax="5.9" valueMin="5.3">5.3–5.9 mm</quantity>
; EW
<quantity id="4C9B638EFFE64C53FE5E98D24B66FE6B" box="[496,626,449,473]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" metricValueMax="2.9" metricValueMin="2.5" pageId="14" pageNumber="397" unit="mm" value="2.7" valueMax="2.9" valueMin="2.5">2.5–2.9 mm</quantity>
; see
<tableCitation id="C6E1FBD0FFE64C53FD0D98D14BE4FE68" box="[675,752,450,474]" captionStart="Table 1" captionStartId="16.[151,198,216,235]" captionText="Table 1. Measurements in mm (minimum, maximum, mean) of morphometric characters for Cuban species of Copelatus Erichson. HL: Head length; HW: Head width; DBE: Distance between eyes; PL: Pronotal length; PW: Pronotal width; EL: Elytral length; EW: Elytral width; TL: Total body length." pageId="14" pageNumber="397">Table 1</tableCitation>
for remaining body measurements. Female unlike male, with elytral striae undulating/anastomosing; male protibia with two ventrobasal emarginations, and median lobe in lateral aspect as in
<figureCitation id="1358D2EEFFE64C53FCF29B114AC6FDA8" box="[860,978,514,538]" captionStart="Figs" captionStartId="13.[113,152,1517,1536]" captionTargetBox="[145,1055,494,1485]" captionTargetId="figure-13@13.[133,1069,485,1499]" captionTargetPageId="13" captionText="Figs. 12–19. Male genitalia (12–15) and left protibia (16–19) of Cuban species of Copelatus. 12, 16 – C. barbouri Young, 1942; 13, 17 – C. darlingtoni Young, 1942; 14, 18 – C. insolitus Chevrolat, 1863; 15, 19 – C. montivagus Young, 1942. Parts of male genitalia: a – median lobe of the aedeagus in left lateral view, b – same in ventral lateral view, c – left paramere in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5331022" httpUri="https://zenodo.org/record/5331022/files/figure.png" pageId="14" pageNumber="397">Figure 14a</figureCitation>
. See under
<taxonomicName id="4C63B5E8FFE64C53FBE09B10481BFD88" authorityName="Young" authorityYear="1942" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="397" phylum="Arthropoda" rank="species" species="darlingtoni">
<emphasis id="B9171279FFE64C53FBE09B10481BFD88" italics="true" pageId="14" pageNumber="397">C. darlingtoni</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3799DE0FFE64C53FF399B514BD7FD08" pageId="14" pageNumber="397" type="biology_ecology">
<paragraph id="8BDCCE6BFFE64C53FF399B514BD7FD08" blockId="14.[151,1127,449,698]" pageId="14" pageNumber="397">
<emphasis id="B9171279FFE64C53FF399B5149E2FDE8" bold="true" box="[151,246,578,602]" pageId="14" pageNumber="397">Ecology.</emphasis>
All material examined previously from
<collectingCountry id="F3748EFBFFE64C53FD3A9B514BDAFDE8" box="[660,718,578,602]" name="Cuba" pageId="14" pageNumber="397">Cuba</collectingCountry>
,
<collectingCountry id="F3748EFBFFE64C53FD799B514A38FDE8" box="[727,812,578,602]" name="Jamaica" pageId="14" pageNumber="397">Jamaica</collectingCountry>
and Central America has been from rather high mountains (
<bibRefCitation id="EFF2B39AFFE64C53FE119B714B52FDC8" author="YOUNG F. N." box="[447,582,610,634]" pageId="14" pageNumber="397" pagination="79 - 94" refId="ref14016" refString="YOUNG F. N. 1942: New species of Copelatus from the West Indies (Coleoptera: Dytiscidae). Proceedings of the New England Zoological Club 20: 79 - 94." type="journal article" year="1942">YOUNG 1942</bibRefCitation>
). The species was frequently collected in backwaters of small streams with muddy-stony bottoms, abundant detritus, located in highlands with a low exposure to sun and without aquatic vegetation.
</paragraph>
</subSubSection>
<caption id="DF1C9EE3FFE64C53FF399F1B4A5EF9E2" ID-DOI="http://doi.org/10.5281/zenodo.5331026" ID-Zenodo-Dep="5331026" httpUri="https://zenodo.org/record/5331026/files/figure.png" pageId="14" pageNumber="397" startId="14.[151,190,1544,1563]" targetBox="[209,1062,729,1508]" targetPageId="14">
<paragraph id="8BDCCE6BFFE64C53FF399F1B4A5EF9E2" blockId="14.[151,1128,1544,1617]" pageId="14" pageNumber="397">
Figs. 20–23. Male genitalia of Cuban species of
<taxonomicName id="4C63B5E8FFE64C53FD999F1A4B9AF9A9" authorityName=": GUEORGUIEV" authorityYear="1968" box="[567,654,1545,1563]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="397" phylum="Arthropoda" rank="genus">
<emphasis id="B9171279FFE64C53FD999F1A4B9AF9A9" box="[567,654,1545,1563]" italics="true" pageId="14" pageNumber="397">Copelatus</emphasis>
</taxonomicName>
. 20 –
<taxonomicName id="4C63B5E8FFE64C53FD6A9F1A4AA4F9A9" authorityName=": YOUNG" authorityYear="1963" box="[708,944,1545,1563]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="397" phylum="Arthropoda" rank="subSpecies" species="caelatipennis" subSpecies="angustatus">
<emphasis id="B9171279FFE64C53FD6A9F1A4AA4F9A9" box="[708,944,1545,1563]" italics="true" pageId="14" pageNumber="397">C. caelatipennis angustatus</emphasis>
</taxonomicName>
author, year; 21 –
<taxonomicName id="4C63B5E8FFE64C53FBFD9F1A49F6F984" authority="Megna &amp; Epler, 2012" authorityName="Megna &amp; Epler" authorityYear="2012" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="397" phylum="Arthropoda" rank="species" species="cordovai" status="sp. nov.">
<emphasis id="B9171279FFE64C53FBFD9F1A49F6F984" italics="true" pageId="14" pageNumber="397">C. cordovai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A224AF02FFE64C53FF469F304839F984" box="[232,301,1571,1590]" pageId="14" pageNumber="397" rank="species">sp. nov.</taxonomicNameLabel>
; 22 –
<taxonomicName id="4C63B5E8FFE64C53FEF19F374B4EF984" authority="Schaeffer, 1908" authorityName="Schaeffer" authorityYear="1908" box="[351,602,1571,1590]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="397" phylum="Arthropoda" rank="species" species="cubaensis">
<emphasis id="B9171279FFE64C53FEF19F3748D8F984" box="[351,460,1572,1590]" italics="true" pageId="14" pageNumber="397">C. cubaensis</emphasis>
<bibRefCitation id="EFF2B39AFFE64C53FE7C9F304B4EF984" author="SCHAEFFER C." box="[466,602,1571,1590]" pageId="14" pageNumber="397" pagination="16 - 18" refId="ref13877" refString="SCHAEFFER C. 1908: On North American and some Cuban Copelatus. Journal of the New York Entomological Society 16: 16 - 18." type="journal article" year="1908">Schaeffer, 1908</bibRefCitation>
</taxonomicName>
; 23 –
<taxonomicName id="4C63B5E8FFE64C53FD209F374A8DF984" authority="(Fabricius, 1801)" baseAuthorityName="Fabricius" baseAuthorityYear="1801" box="[654,921,1571,1590]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="397" phylum="Arthropoda" rank="species" species="posticatus">
<emphasis id="B9171279FFE64C53FD209F374BEAF984" box="[654,766,1572,1590]" italics="true" pageId="14" pageNumber="397">C. posticatus</emphasis>
(
<bibRefCitation id="EFF2B39AFFE64C53FCA79F304A86F984" author="FABRICIUS J. C." box="[777,914,1571,1590]" pageId="14" pageNumber="397" refId="ref13078" refString="FABRICIUS J. C. 1801: Systema Eleutheratorum secundum ordines, genera, species; adiectis synonymis, locis, observationibus, descriptionibus. Tomus I. Bibliopolii Academici Novi, Kiliae, xxiv + 506 pp." type="book" year="1801">Fabricius, 1801</bibRefCitation>
)
</taxonomicName>
. Parts of male genitalia: a – median lobe of aedeagus in left lateral view, b – left paramere in lateral view.
</paragraph>
</caption>
<subSubSection id="C3799DE0FFE74C52FFDF99C84BD8FEA6" pageId="15" pageNumber="398" type="distribution">
<paragraph id="8BDCCE6BFFE74C52FFDF99C84BD8FEA6" blockId="15.[113,1089,219,628]" pageId="15" pageNumber="398">
<emphasis id="B9171279FFE74C52FFDF99C84810FF41" bold="true" box="[113,260,219,243]" pageId="15" pageNumber="398">Distribution.</emphasis>
This species is known from
<collectingCountry id="F3748EFBFFE74C52FD9299C84B63FF41" box="[572,631,219,243]" name="Cuba" pageId="15" pageNumber="398">Cuba</collectingCountry>
,
<collectingCountry id="F3748EFBFFE74C52FD2C99C84BCFFF41" box="[642,731,219,243]" name="Jamaica" pageId="15" pageNumber="398">Jamaica</collectingCountry>
, Hispaniola and Central America (
<bibRefCitation id="EFF2B39AFFE74C52FFD799EF4810FEA6" author="YOUNG F. N." box="[121,260,252,276]" pageId="15" pageNumber="398" pagination="79 - 94" refId="ref14016" refString="YOUNG F. N. 1942: New species of Copelatus from the West Indies (Coleoptera: Dytiscidae). Proceedings of the New England Zoological Club 20: 79 - 94." type="journal article" year="1942">YOUNG 1942</bibRefCitation>
). It is widely distributed in
<collectingCountry id="F3748EFBFFE74C52FD8699E84B74FEA1" box="[552,608,251,275]" name="Cuba" pageId="15" pageNumber="398">Cuba</collectingCountry>
(
<figureCitation id="1358D2EEFFE74C52FDC199E84BAAFEA6" box="[623,702,251,276]" captionStart="Figs" captionStartId="20.[151,190,1516,1535]" captionTargetBox="[165,1104,223,1503]" captionTargetId="figure-11@20.[161,1117,219,1504]" captionTargetPageId="20" captionText="Figs. 28–31. Known distribution of Copelatus species in Cuba. 28 – C. cubaensis Schaeffer, 1908; 29 – C. danyi sp. nov.; 30 – C. darlingtoni Young, 1942; 31 – C. insolitus Chevrolat, 1863. See Appendix 1 for additional information on the localities reported in each map." figureDoi="http://doi.org/10.5281/zenodo.5331032" httpUri="https://zenodo.org/record/5331032/files/figure.png" pageId="15" pageNumber="398">Fig. 31</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3799DE0FFE74C52FFDF980F4B47FDC1" pageId="15" pageNumber="398" type="discussion">
<paragraph id="8BDCCE6BFFE74C52FFDF980F4B47FDC1" blockId="15.[113,1089,219,628]" pageId="15" pageNumber="398">
<emphasis id="B9171279FFE74C52FFDF980F49F4FE86" bold="true" box="[113,224,284,308]" pageId="15" pageNumber="398">Remarks.</emphasis>
The
<typeStatus id="54D870C9FFE74C52FEB8980F4850FE86" box="[278,324,284,308]" pageId="15" pageNumber="398">type</typeStatus>
locality of
<taxonomicName id="4C63B5E8FFE74C52FE10980E4B2CFE86" authorityName="Chevrolat" authorityYear="1863" box="[446,568,285,308]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="398" phylum="Arthropoda" rank="species" species="insolitus">
<emphasis id="B9171279FFE74C52FE10980E4B2CFE86" box="[446,568,285,308]" italics="true" pageId="15" pageNumber="398">C. insolitus</emphasis>
</taxonomicName>
is
<collectingRegion id="49A70089FFE74C52FDF9980F4BBFFE81" box="[599,683,284,307]" country="Cuba" name="La Habana" pageId="15" pageNumber="398">Havana</collectingRegion>
, but doubt exists on the occurrence of this species there due to the fact that all material examined previously from
<collectingCountry id="F3748EFBFFE74C52FC0A98284ACBFEE1" box="[932,991,315,339]" name="Cuba" pageId="15" pageNumber="398">Cuba</collectingCountry>
,
<collectingCountry id="F3748EFBFFE74C52FC4598284D55FEE1" box="[1003,1089,315,339]" name="Jamaica" pageId="15" pageNumber="398">Jamaica</collectingCountry>
and Central America has been from rather high mountains (
<bibRefCitation id="EFF2B39AFFE74C52FD4E984F4A7EFEC6" author="YOUNG F. N." box="[736,874,348,372]" pageId="15" pageNumber="398" pagination="79 - 94" refId="ref14016" refString="YOUNG F. N. 1942: New species of Copelatus from the West Indies (Coleoptera: Dytiscidae). Proceedings of the New England Zoological Club 20: 79 - 94." type="journal article" year="1942">YOUNG 1942</bibRefCitation>
). The widespread
<taxonomicName id="4C63B5E8FFE74C52FB86984E49DFFE26" authorityName="Chevrolat" authorityYear="1863" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="398" phylum="Arthropoda" rank="species" species="insolitus">
<emphasis id="B9171279FFE74C52FB86984E49DFFE26" italics="true" pageId="15" pageNumber="398">C. insolitus</emphasis>
</taxonomicName>
is joined in
<collectingCountry id="F3748EFBFFE74C52FEE098684892FE21" box="[334,390,379,403]" name="Cuba" pageId="15" pageNumber="398">Cuba</collectingCountry>
by three other rather similar species apparently endemic to
<collectingCountry id="F3748EFBFFE74C52FBAF98684D29FE21" box="[1025,1085,379,403]" name="Cuba" pageId="15" pageNumber="398">Cuba</collectingCountry>
:
<taxonomicName id="4C63B5E8FFE74C52FFDF988E49FBFE06" authorityName="Young" authorityYear="1942" box="[113,239,413,436]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="398" phylum="Arthropoda" rank="species" species="barbouri">
<emphasis id="B9171279FFE74C52FFDF988E49FBFE06" box="[113,239,413,436]" italics="true" pageId="15" pageNumber="398">C. barbouri</emphasis>
</taxonomicName>
,
<taxonomicName id="4C63B5E8FFE74C52FF55988E4886FE06" authorityName="Young" authorityYear="1942" box="[251,402,413,436]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="398" phylum="Arthropoda" rank="species" species="darlingtoni">
<emphasis id="B9171279FFE74C52FF55988E4886FE06" box="[251,402,413,436]" italics="true" pageId="15" pageNumber="398">C. darlingtoni</emphasis>
</taxonomicName>
and
<taxonomicName id="4C63B5E8FFE74C52FE6A988E4B49FE06" authorityName="Young" authorityYear="1942" box="[452,605,413,436]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="398" phylum="Arthropoda" rank="species" species="montivagus">
<emphasis id="B9171279FFE74C52FE6A988E4B49FE06" box="[452,605,413,436]" italics="true" pageId="15" pageNumber="398">C. montivagus</emphasis>
</taxonomicName>
. These Cuban species are not known to occur together.
<bibRefCitation id="EFF2B39AFFE74C52FF7B98AF488CFE66" author="YOUNG F. N." box="[213,408,444,468]" pageId="15" pageNumber="398" pagination="79 - 94" refId="ref14016" refString="YOUNG F. N. 1942: New species of Copelatus from the West Indies (Coleoptera: Dytiscidae). Proceedings of the New England Zoological Club 20: 79 - 94." type="journal article" year="1942">YOUNG (1942: 90)</bibRefCitation>
hypothesized that these species represented “developing endemic forms on the various isolated mountain ranges.” The males all lack well defined striae and possess similar, but different, color patterns; they can be separated by the shape of their protibiae and their genitalia. Females of
<taxonomicName id="4C63B5E8FFE74C52FD899B0E4BABFD86" authorityName="Young" authorityYear="1942" box="[551,703,541,564]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="398" phylum="Arthropoda" rank="species" species="darlingtoni">
<emphasis id="B9171279FFE74C52FD899B0E4BABFD86" box="[551,703,541,564]" italics="true" pageId="15" pageNumber="398">C. darlingtoni</emphasis>
</taxonomicName>
,
<taxonomicName id="4C63B5E8FFE74C52FD639B0E4A53FD86" authorityName="Chevrolat" authorityYear="1863" box="[717,839,541,564]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="398" phylum="Arthropoda" rank="species" species="insolitus">
<emphasis id="B9171279FFE74C52FD639B0E4A53FD86" box="[717,839,541,564]" italics="true" pageId="15" pageNumber="398">C. insolitus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C63B5E8FFE74C52FCD39B0E4D0CFD86" authorityName="Young" authorityYear="1942" box="[893,1048,541,564]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="398" phylum="Arthropoda" rank="species" species="montivagus">
<emphasis id="B9171279FFE74C52FCD39B0E4D0CFD86" box="[893,1048,541,564]" italics="true" pageId="15" pageNumber="398">C. montivagus</emphasis>
</taxonomicName>
are very similar and are best separated with comparative material or by association with males; the female of
<taxonomicName id="4C63B5E8FFE74C52FEAB9B4E4897FDC6" authorityName="Young" authorityYear="1942" box="[261,387,605,628]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="398" phylum="Arthropoda" rank="species" species="barbouri">
<emphasis id="B9171279FFE74C52FEAB9B4E4897FDC6" box="[261,387,605,628]" italics="true" pageId="15" pageNumber="398">C. barbouri</emphasis>
</taxonomicName>
lacks elytral striae.
</paragraph>
</subSubSection>
</treatment>
</document>
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