treatments-xml/data/03/BF/5F/03BF5F1AFFEFFFD3CBB4FE0EBD3FCECD.xml

<document id="BF2EA3A530D1B9F285A9DAD8238831EC" ID-DOI="10.5281/zenodo.5305424" ID-ISSN="0374-1036" ID-Zenodo-Dep="5305424" ID-ZooBank="34AECE17-DCFC-4373-A0E1-3D48B5BFE915" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630111498801" checkinUser="marcus" docAuthor="Janák, Jiří &amp; Makranczy, György" docDate="2016" docId="03BF5F1AFFEFFFD3CBB4FE0EBD3FCECD" docLanguage="en" docName="ActaEntMusNatPra.2016.56.1.203-210.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 56 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Thinobius crypticola Janák &amp; Makranczy 2016, sp. nov." docType="treatment" docVersion="3" lastPageNumber="210" masterDocId="FF862762FFEDFFD4CA6BFFF5BF78CF3E" masterDocTitle="Description of a blind and flightless species of Thinobius from South Africa (Coleoptera: Staphylinidae: Oxytelinae)" masterLastPageNumber="210" masterPageNumber="203" pageNumber="205" updateTime="1699155369259" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="DA0E893BA1C0DACA7E89A4F5CC53A237">Description of a blind and flightless species of Thinobius from South Africa (Coleoptera: Staphylinidae: Oxytelinae)</mods:title>
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<mods:namePart id="45F515211CCC44528B3471C89736242D">Janák, Jiří</mods:namePart>
<mods:affiliation id="DDA1361BF53BCC6BC86263845CC9AF57">CZ- 417 62 Rtyně nad Bílinou, č. 4, Czech Republic; e-mail: jiri. janak @ heineken. com</mods:affiliation>
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<mods:namePart id="65B82F538317E5FB8970352AC9028FEF">Makranczy, György</mods:namePart>
<mods:affiliation id="5EB173CF968C7CB91E3AAA6F330474D7">Department of Zoology, Hungarian Natural History Museum, H- 1088 Budapest, Baross utca 13, Hungary; e-mail: makranczy @ zoo. nhmus. hu</mods:affiliation>
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<mods:title id="B21271A62AD2373BBBBBDCA2A143E2E0">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="BFEE015A4F85BC7338E117E1B683B871">2016</mods:date>
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<treatment id="03BF5F1AFFEFFFD3CBB4FE0EBD3FCECD" LSID="urn:lsid:plazi:treatment:03BF5F1AFFEFFFD3CBB4FE0EBD3FCECD" httpUri="http://treatment.plazi.org/id/03BF5F1AFFEFFFD3CBB4FE0EBD3FCECD" lastPageId="7" lastPageNumber="210" pageId="2" pageNumber="205">
<subSubSection id="C30CBD87FFEFFFD6CBB4FE0EBC67CD2C" box="[479,799,507,530]" pageId="2" pageNumber="205" type="nomenclature">
<paragraph id="8BA9EE0CFFEFFFD6CBB4FE0EBC67CD2C" blockId="2.[479,799,507,530]" box="[479,799,507,530]" pageId="2" pageNumber="205">
<heading id="D0E15960FFEFFFD6CBB4FE0EBC67CD2C" bold="true" box="[479,799,507,530]" centered="true" fontSize="8" level="2" pageId="2" pageNumber="205" reason="3">
<emphasis id="B962321EFFEFFFD6CBB4FE0EBC67CD2C" bold="true" box="[479,799,507,530]" pageId="2" pageNumber="205">
<taxonomicName id="4C16958FFFEFFFD6CBB4FE0EBDBACD2C" ID-CoL="8YBR5" authority="Janák &amp; Makranczy, 2016" authorityName="Janák &amp; Makranczy" authorityYear="2016" box="[479,706,507,530]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="205" phylum="Arthropoda" rank="species" species="crypticola" status="sp. nov.">
<emphasis id="B962321EFFEFFFD6CBB4FE0EBDBACD2C" bold="true" box="[479,706,507,530]" italics="true" pageId="2" pageNumber="205">Thinobius crypticola</emphasis>
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<taxonomicNameLabel id="A2518F65FFEFFFD6C8A2FE0EBC67CD2C" box="[713,799,507,530]" pageId="2" pageNumber="205" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C30CBD87FFEFFFD6C853FDECBDBDCD12" box="[568,709,537,556]" pageId="2" pageNumber="205" type="description">
<paragraph id="8BA9EE0CFFEFFFD6C853FDECBDBDCD12" blockId="2.[568,709,537,556]" box="[568,709,537,556]" pageId="2" pageNumber="205">
(
<figureCitation id="132DF289FFEFFFD6C855FDECBDF3CD12" box="[574,651,537,556]" captionStart="Figs 1–2" captionStartId="2.[841,879,1480,1499]" captionTargetBox="[151,830,690,1602]" captionTargetId="figure-190@2.[151,830,687,1605]" captionTargetPageId="2" captionText="Figs 1–2. Thinobius crypticola sp. nov.: 1 – male holotype, dorsal habitus; 2 – suspected larva (maximum width of head capsule 0.16 mm)." figureDoi="http://doi.org/10.5281/zenodo.5305426" httpUri="https://zenodo.org/record/5305426/files/figure.png" pageId="2" pageNumber="205">Figs 1–2</figureCitation>
, 
<figureCitation id="132DF289FFEFFFD6C8FFFDECBDB8CD12" box="[660,704,537,556]" captionStart="Figs 5–13" captionStartId="6.[151,188,1514,1533]" captionTargetBox="[152,1099,222,1451]" captionTargetId="figure-9@6.[151,1103,219,1466]" captionTargetPageId="6" captionText="Figs 5–13. Thinobius crypticola sp. nov. 5 – antenna; 6 – maxillary palp; 7 – scutellum; 8 – male sternite VIII; 9 – male tergite IX; 10 – male sternite IX; 11 – aedeagus, frontal; 12 – female genital appendage; 13 – spermatheca. Scale bars: 0.075 mm for 11, 13; 0.09 mm for 5–7, 9–10, 12; 0.1 mm for 8." figureDoi="http://doi.org/10.5281/zenodo.5305430" httpUri="https://zenodo.org/record/5305430/files/figure.png" pageId="2" pageNumber="205">5–13</figureCitation>
)
</paragraph>
</subSubSection>
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<paragraph id="8BA9EE0CFFEFFFD6CAFCFDB6BEBCCD4F" blockId="2.[151,1127,579,678]" pageId="2" pageNumber="205">
<emphasis id="B962321EFFEFFFD6CAFCFDB6BE68CD68" bold="true" box="[151,272,579,598]" pageId="2" pageNumber="205">
<typeStatus id="54AD50AEFFEFFFD6CAFCFDB6BFBCCD68" box="[151,196,579,598]" pageId="2" pageNumber="205">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F301AE9CFFEFFFD6CB7FFDB6BEFBCD68" box="[276,387,579,598]" name="South Africa" pageId="2" pageNumber="205">South Africa</collectingCountry>
, 
<collectingRegion id="49D220EEFFEFFFD6CBE1FDB6BD2BCD68" box="[394,595,579,598]" country="South Africa" name="Western Cape" pageId="2" pageNumber="205">Western Cape Province</collectingRegion>
, Ysternek Nature Reserve, Dal van Varings Trail, off Highway R339, 
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, 
<geoCoordinate id="EE2288CBFFEFFFD6CB45FDABBE01CD4F" box="[302,377,606,625]" degrees="23" direction="east" minutes="8.7" orientation="longitude" pageId="2" pageNumber="205" precision="92" value="23.145">23°8.7′E</geoCoordinate>
(
<figureCitation id="132DF289FFEFFFD6CBEFFDABBEC0CD4F" box="[388,440,606,625]" captionStart="Figs 3–4" captionStartId="3.[113,150,1560,1579]" captionTargetBox="[113,1088,219,1548]" captionTargetId="figure-41@3.[113,1089,216,1551]" captionTargetPageId="3" captionText="Figs 3–4. Habitats of Thinobius crypticola sp. nov. 3 – Knysna, Dal van Varings Trail (type locality); 4 – Knysna, Diepwalle State Forest (photos by J. Janák)." figureDoi="http://doi.org/10.5281/zenodo.5305428" httpUri="https://zenodo.org/record/5305428/files/figure.png" pageId="2" pageNumber="205">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8BA9EE0CFFEFFFD0CAFCFD8DBDC5CDA4" blockId="2.[151,1127,579,678]" lastBlockId="4.[151,1128,220,666]" lastPageId="4" lastPageNumber="207" pageId="2" pageNumber="205">
<emphasis id="B962321EFFEFFFD6CAFCFD8DBE67CDB5" bold="true" box="[151,287,632,651]" pageId="2" pageNumber="205">Type material.</emphasis>
<materialsCitation id="3B7EE451FFEFFFD6CB4CFD8DBB23CD98" collectingDate="2014-02-07" collectorName="Ysternek N. R. &amp; Dal van Varings Tr. &amp; Hwy R &amp; J. Janak" country="South Africa" latitude="-33.913334" location="Hwy" longLatPrecision="119" longitude="23.145" pageId="2" pageNumber="205" specimenCount="1" stateProvince="Western Cape" typeStatus="holotype">
<typeStatus id="54AD50AEFFEFFFD6CB4CFD8DBE02CDB4" box="[295,378,632,651]" pageId="2" pageNumber="205" type="holotype">HOLOTYPE</typeStatus>
: J, ‘ 
<collectingCountry id="F301AE9CFFEFFFD6CBC2FD8DBD66CDB5" box="[425,542,632,651]" name="South Africa" pageId="2" pageNumber="205">South Africa</collectingCountry>
, 
<collectingRegion id="49D220EEFFEFFFD6C842FD8DBDDDCDB5" box="[553,677,632,651]" country="South Africa" name="Western Cape" pageId="2" pageNumber="205">Western Cape</collectingRegion>
, 
<collectorName id="26E38BDAFFEFFFD6C8C4FD8DBC56CDB5" box="[687,814,632,651]" pageId="2" pageNumber="205">
Ysternek 
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</collectorName>
[Nature Reserve], 
<collectorName id="26E38BDAFFEFFFD6C9B3FD8DBFCBCD98" pageId="2" pageNumber="205">Dal van Varings Tr.</collectorName>
[ail], off 
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<location id="8EC9B8D7FFEFFFD6CA97FD66BE5DCD98" LSID="urn:lsid:plazi:treatment:03BF5F1AFFEFFFD3CBB4FE0EBD3FCECD:8EC9B8D7FFEFFFD6CA97FD66BE5DCD98" box="[252,293,659,678]" country="South Africa" latitude="-33.913334" longLatPrecision="119" longitude="23.145" name="Hwy" pageId="2" pageNumber="205" stateProvince="Western Cape">Hwy</location>
R
</collectorName>
339; 
<geoCoordinate id="EE2288CBFFEFFFD6CB0DFD66BEC4CD98" box="[358,444,659,678]" degrees="33" direction="south" minutes="54.8" orientation="latitude" pageId="2" pageNumber="205" precision="92" value="-33.913334">33°54.8’S</geoCoordinate>
<geoCoordinate id="EE2288CBFFEFFFD6CBA8FD66BD6ACD98" box="[451,530,659,678]" degrees="23" direction="east" minutes="8.7" orientation="longitude" pageId="2" pageNumber="205" precision="92" value="23.145">23°8.7’E</geoCoordinate>
; 
<date id="FFA8C8CCFFEFFFD6C877FD66BD12CD98" box="[540,618,659,678]" pageId="2" pageNumber="205" value="2014-02-07">
<collectingDate id="EFEC3124FFEFFFD6C877FD66BD12CD98" box="[540,618,659,678]" pageId="2" pageNumber="205" value="2014-02-07">7.ii.2014</collectingDate>
</date>
, 
<collectorName id="26E38BDAFFEFFFD6C818FD66BDC0CD98" box="[627,696,659,678]" pageId="2" pageNumber="205">J. Janák</collectorName>
lgt. \ Berlese extraction, leaf &amp; log litter, sifting
</materialsCitation>
\ 
<typeStatus id="54AD50AEFFE9FFD0CAFCFF29BF88CFD1" box="[151,240,220,239]" pageId="4" pageNumber="207" type="holotype">Holotypus</typeStatus>
<taxonomicName id="4C16958FFFE9FFD0CA9EFF29BEDECFD1" authority="Janák &amp; Makranczy, 2016" authorityName="Janák &amp; Makranczy" authorityYear="2016" box="[245,422,220,239]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="207" phylum="Arthropoda" rank="species" species="crypticola" status="sp. nov.">
<emphasis id="B962321EFFE9FFD0CA9EFF29BEDECFD1" box="[245,422,220,239]" italics="true" pageId="4" pageNumber="207">Thinobius crypticola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2518F65FFE9FFD0CBC1FF29BE96CFD1" box="[426,494,220,239]" pageId="4" pageNumber="207" rank="species">sp. nov.</taxonomicNameLabel>
, J. Janák &amp; Gy. Makranczy det. 2015’ (
<collectionCode id="ED0776C9FFE9FFD0C95DFF29BC0ECFD1" box="[822,886,220,239]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="4" pageNumber="207">TMSA</collectionCode>
). 
<materialsCitation id="3B7EE451FFE9FFD0C9EFFF29BECCCE1A" collectingDate="2014-02-07" collectionCode="TMSA, NMPC, NHMW, HNHM, JJRC" collectorName="Ysternek N. R. &amp; Dal van Varings Tr. &amp; Hwy R &amp; J. Janak" country="South Africa" latitude="-33.913334" location="Hwy" longLatPrecision="119" longitude="23.145" pageId="4" pageNumber="207" specimenCount="106" specimenCount-female="30" stateProvince="Western Cape" typeStatus="paratype">
<typeStatus id="54AD50AEFFE9FFD0C9EFFF29BCA3CFD3" box="[900,987,220,239]" pageId="4" pageNumber="207" type="paratype">PARATYPES</typeStatus>
(
<specimenCount id="9D102585FFE9FFD0C98DFF29BB26CFD1" box="[998,1118,220,239]" pageId="4" pageNumber="207" type="generic">76 specimens</specimenCount>
): 19 JJ 
<specimenCount id="9D102585FFE9FFD0CABDFF02BE77CE37" box="[214,271,240,266]" pageId="4" pageNumber="207" type="female">15 ♀♀</specimenCount>
: same data as holotype (1 J 
<specimenCount id="9D102585FFE9FFD0C86DFF02BD5DCE37" box="[518,549,240,266]" pageId="4" pageNumber="207" type="female">1 ♀</specimenCount>
in 
<collectionCode id="ED0776C9FFE9FFD0C82BFF02BD05CE34" box="[576,637,247,266]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="4" pageNumber="207">TMSA</collectionCode>
, 1 J 
<specimenCount id="9D102585FFE9FFD0C8C1FF02BDB1CE37" box="[682,713,240,266]" pageId="4" pageNumber="207" type="female">1 ♀</specimenCount>
in 
<collectionCode id="ED0776C9FFE9FFD0C88FFF02BC5BCE34" box="[740,803,247,266]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="4" pageNumber="207">NMPC</collectionCode>
, 1 J 
<specimenCount id="9D102585FFE9FFD0C924FF02BC16CE37" box="[847,878,240,266]" pageId="4" pageNumber="207" type="female">1 ♀</specimenCount>
<collectionCode id="ED0776C9FFE9FFD0C918FF02BCC1CE34" box="[883,953,247,266]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="4" pageNumber="207" type="Museum">NHMW</collectionCode>
, 1 J 
<specimenCount id="9D102585FFE9FFD0C98DFF02BB7DCE37" box="[998,1029,240,266]" pageId="4" pageNumber="207" type="female">1 ♀</specimenCount>
in 
<collectionCode id="ED0776C9FFE9FFD0CE4BFF02BB1BCE34" box="[1056,1123,247,266]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="4" pageNumber="207">HNHM</collectionCode>
, 15 JJ 
<specimenCount id="9D102585FFE9FFD0CABCFEE4BE77CE1A" box="[215,271,267,292]" pageId="4" pageNumber="207" type="female">11 ♀♀</specimenCount>
in 
<collectionCode id="ED0776C9FFE9FFD0CB40FEE4BE21CE1A" box="[299,345,273,292]" pageId="4" pageNumber="207">JJRC</collectionCode>
) {
<quantity id="4CEE43E9FFE9FFD0CB00FEE4BECCCE1A" box="[363,436,273,292]" metricMagnitude="1" metricUnit="kg" metricValue="1.94" pageId="4" pageNumber="207" unit="kg" value="19.4">19.4 kg</quantity>
</materialsCitation>
}; 
<materialsCitation id="3B7EE451FFE9FFD0CBAFFEE4BE6BCE01" collectorName="Diepwalle S. F." country="South Africa" location="State Forest" pageId="4" pageNumber="207" specimenCount="2" specimenCount-female="2" stateProvince="Western Cape" typeStatus="paratype">
2 JJ 
<specimenCount id="9D102585FFE9FFD0CB92FEE4BD50CE1A" box="[505,552,267,292]" pageId="4" pageNumber="207" type="female">2 ♀♀</specimenCount>
: ‘ 
<collectingCountry id="F301AE9CFFE9FFD0C85CFEE4BDD2CE1A" box="[567,682,273,292]" name="South Africa" pageId="4" pageNumber="207">South Africa</collectingCountry>
, 
<collectingRegion id="49D220EEFFE9FFD0C8DAFEE4BC52CE1A" box="[689,810,273,292]" country="South Africa" name="Western Cape" pageId="4" pageNumber="207">Western Cape</collectingRegion>
, 
<collectorName id="26E38BDAFFE9FFD0C958FEE4BCD7CE1A" box="[819,943,273,292]" pageId="4" pageNumber="207">Diepwalle S.F.</collectorName>
[
<location id="8EC9B8D7FFE9FFD0C9D8FEE4BB58CE1A" LSID="urn:lsid:plazi:treatment:03BF5F1AFFEFFFD3CBB4FE0EBD3FCECD:8EC9B8D7FFE9FFD0C9D8FEE4BB58CE1A" box="[947,1056,273,292]" country="South Africa" name="State Forest" pageId="4" pageNumber="207" stateProvince="Western Cape">State Forest</location>
] (
<locationDeviation id="9A1BF72EFFE9FFD0CE5BFEE4BE75CE01" location="Knysna" pageId="4" pageNumber="207">
<quantity id="4CEE43E9FFE9FFD0CE5BFEE4BB1ECE1A" box="[1072,1126,273,292]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="4" pageNumber="207" unit="km" value="14.0">14 km</quantity>
ENE Knysna
</locationDeviation>
)
</materialsCitation>
; 
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<geoCoordinate id="EE2288CBFFE9FFD0CB17FED9BEB4CE01" box="[380,460,300,319]" degrees="23" direction="east" minutes="9.5" orientation="longitude" pageId="4" pageNumber="207" precision="92" value="23.158333">23°9.5´E</geoCoordinate>
; 
<materialsCitation id="3B7EE451FFE9FFD0CBBDFED9BE59CE64" collectingDate="2014-02-07" collectionCode="JJRC" country="South Africa" location="Berlese" municipality="Janak" pageId="4" pageNumber="207" specimenCount="1" stateProvince="Western Cape" typeStatus="paratype">
<date id="FFA8C8CCFFE9FFD0CBBDFED9BD5DCE01" box="[470,549,300,319]" pageId="4" pageNumber="207" value="2014-02-07">
<collectingDate id="EFEC3124FFE9FFD0CBBDFED9BD5DCE01" box="[470,549,300,319]" pageId="4" pageNumber="207" value="2014-02-07">7.ii.2014</collectingDate>
</date>
, J. 
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lgt., 
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extraction, leaf &amp; log litter, sifting’ (
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) {
<quantity id="4CEE43E9FFE9FFD0CAF5FEB2BF9BCE64" box="[158,227,327,346]" metricMagnitude="1" metricUnit="kg" metricValue="1.67" pageId="4" pageNumber="207" unit="kg" value="16.7">16.7 kg</quantity>
, 
<figureCitation id="132DF289FFE9FFD0CA80FEB2BE59CE64" box="[235,289,327,346]" captionStart="Figs 3–4" captionStartId="3.[113,150,1560,1579]" captionTargetBox="[113,1088,219,1548]" captionTargetId="figure-41@3.[113,1089,216,1551]" captionTargetPageId="3" captionText="Figs 3–4. Habitats of Thinobius crypticola sp. nov. 3 – Knysna, Dal van Varings Trail (type locality); 4 – Knysna, Diepwalle State Forest (photos by J. Janák)." figureDoi="http://doi.org/10.5281/zenodo.5305428" httpUri="https://zenodo.org/record/5305428/files/figure.png" pageId="4" pageNumber="207">Fig. 4</figureCitation>
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6 JJ 
<specimenCount id="9D102585FFE9FFD0CB0EFEB2BEEACE67" box="[357,402,320,346]" pageId="4" pageNumber="207" type="female">4 ♀♀</specimenCount>
: ‘ 
<collectingCountry id="F301AE9CFFE9FFD0CBCAFEB2BD6CCE64" box="[417,532,327,346]" name="South Africa" pageId="4" pageNumber="207">South Africa</collectingCountry>
, 
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, 
<collectorName id="26E38BDAFFE9FFD0C8F0FEB2BC58CE64" box="[667,800,327,346]" pageId="4" pageNumber="207">Diepwalle S. F.</collectorName>
, 
<location id="8EC9B8D7FFE9FFD0C94CFEB2BCEBCE64" LSID="urn:lsid:plazi:treatment:03BF5F1AFFEFFFD3CBB4FE0EBD3FCECD:8EC9B8D7FFE9FFD0C94CFEB2BCEBCE64" box="[807,915,327,346]" country="South Africa" name="Kom Se Pad" pageId="4" pageNumber="207" stateProvince="Western Cape">Kom Se Pad</location>
, 
<collectorName id="26E38BDAFFE9FFD0C9F0FEB2BB55CE64" box="[923,1069,327,346]" pageId="4" pageNumber="207">Red Elephant Tr.</collectorName>
[Trail]
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; 
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; 
<materialsCitation id="3B7EE451FFE9FFD0CB27FE94BE5CCEB1" collectingDate="2014-02-09" collectionCode="TMSA, JJRC" country="South Africa" location="Berlese" municipality="Janak" pageId="4" pageNumber="207" specimenCount="4" specimenCount-female="4" stateProvince="Western Cape" typeStatus="paratype">
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</date>
, J. 
<collectingMunicipality id="6BCD7476FFE9FFD0CBDCFE94BE90CE4A" box="[439,488,353,372]" pageId="4" pageNumber="207">Janák</collectingMunicipality>
lgt., 
<location id="8EC9B8D7FFE9FFD0C87FFE94BD2CCE4A" LSID="urn:lsid:plazi:treatment:03BF5F1AFFEFFFD3CBB4FE0EBD3FCECD:8EC9B8D7FFE9FFD0C87FFE94BD2CCE4A" box="[532,596,353,372]" country="South Africa" municipality="Janak" name="Berlese" pageId="4" pageNumber="207" stateProvince="Western Cape">Berlese</location>
extraction, leaf &amp; log litter, sifting’ (1 J 
<collectionCode id="ED0776C9FFE9FFD0C9D1FE94BC8FCE4A" box="[954,1015,353,372]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="4" pageNumber="207">TMSA</collectionCode>
, 5 JJ 
<specimenCount id="9D102585FFE9FFD0CE5CFE94BB1FCE4A" box="[1079,1127,347,372]" pageId="4" pageNumber="207" type="female">4 ♀♀</specimenCount>
<collectionCode id="ED0776C9FFE9FFD0CAFCFE89BFBDCEB1" box="[151,197,380,399]" pageId="4" pageNumber="207">JJRC</collectionCode>
) {
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<materialsCitation id="3B7EE451FFE9FFD0CB5EFE89BE16CE94" collectorName="Diepwalle S. F." country="South Africa" location="Kom Se Pad" pageId="4" pageNumber="207" specimenCount="5" specimenCount-female="5" stateProvince="Western Cape" typeStatus="paratype">
6 JJ 
<specimenCount id="9D102585FFE9FFD0CB06FE89BEE5CEB0" box="[365,413,373,399]" pageId="4" pageNumber="207" type="female">5 ♀♀</specimenCount>
: ‘ 
<collectingCountry id="F301AE9CFFE9FFD0CBC6FE89BD5ACEB1" box="[429,546,380,399]" name="South Africa" pageId="4" pageNumber="207">South Africa</collectingCountry>
, 
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, 
<collectorName id="26E38BDAFFE9FFD0C8C4FE89BC55CEB1" box="[687,813,380,399]" pageId="4" pageNumber="207">Diepwalle S.F.</collectorName>
[State Forest], 
<location id="8EC9B8D7FFE9FFD0C9DBFE89BB67CEB1" LSID="urn:lsid:plazi:treatment:03BF5F1AFFEFFFD3CBB4FE0EBD3FCECD:8EC9B8D7FFE9FFD0C9DBFE89BB67CEB1" box="[944,1055,380,399]" country="South Africa" name="Kom Se Pad" pageId="4" pageNumber="207" stateProvince="Western Cape">Kom Se Pad</location>
, 
<locationDeviation id="9A1BF72EFFE9FFD0CE41FE89BFA4CE94" location="Hwy" pageId="4" pageNumber="207">
<quantity id="4CEE43E9FFE9FFD0CE41FE89BB1ECEB1" box="[1066,1126,380,399]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="4" pageNumber="207" unit="km" value="1.7">1.7 km</quantity>
W Hwy
</locationDeviation>
[Highway] R339
</materialsCitation>
; 
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<geoCoordinate id="EE2288CBFFE9FFD0CBB8FE62BD5ACE94" box="[467,546,407,426]" degrees="23" direction="east" minutes="8.0" orientation="longitude" pageId="4" pageNumber="207" precision="92" value="23.133333">23°8.0´E</geoCoordinate>
; 
<materialsCitation id="3B7EE451FFE9FFD0C840FE62BD63CEFA" collectingDate="2014-02-09" collectionCode="TMSA, JJRC" country="South Africa" location="Berlese" municipality="Janak" pageId="4" pageNumber="207" specimenCount="5" specimenCount-female="5" stateProvince="Western Cape" typeStatus="paratype">
<date id="FFA8C8CCFFE9FFD0C840FE62BD02CE94" box="[555,634,407,426]" pageId="4" pageNumber="207" value="2014-02-09">
<collectingDate id="EFEC3124FFE9FFD0C840FE62BD02CE94" box="[555,634,407,426]" pageId="4" pageNumber="207" value="2014-02-09">9.ii.2014</collectingDate>
</date>
, J. 
<collectingMunicipality id="6BCD7476FFE9FFD0C8FDFE62BDBFCE94" box="[662,711,407,426]" pageId="4" pageNumber="207">Janák</collectingMunicipality>
lgt., 
<location id="8EC9B8D7FFE9FFD0C89FFE62BC4CCE94" LSID="urn:lsid:plazi:treatment:03BF5F1AFFEFFFD3CBB4FE0EBD3FCECD:8EC9B8D7FFE9FFD0C89FFE62BC4CCE94" box="[756,820,407,426]" country="South Africa" municipality="Janak" name="Berlese" pageId="4" pageNumber="207" stateProvince="Western Cape">Berlese</location>
extraction, leaf &amp; log litter, sifting’ (1 J 
<specimenCount id="9D102585FFE9FFD0CAAFFE44BF9DCEFA" box="[196,229,427,452]" pageId="4" pageNumber="207" type="female">1 ♀</specimenCount>
<collectionCode id="ED0776C9FFE9FFD0CA81FE44BE5FCEFA" box="[234,295,433,452]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="4" pageNumber="207">TMSA</collectionCode>
, 5 JJ 
<specimenCount id="9D102585FFE9FFD0CB0EFE44BEEDCEFA" box="[357,405,427,452]" pageId="4" pageNumber="207" type="female">4 ♀♀</specimenCount>
<collectionCode id="ED0776C9FFE9FFD0CBF0FE44BEB1CEFA" box="[411,457,433,452]" pageId="4" pageNumber="207">JJRC</collectionCode>
) {
<quantity id="4CEE43E9FFE9FFD0CBB0FE44BD63CEFA" box="[475,539,433,452]" metricMagnitude="0" metricUnit="kg" metricValue="5.6" pageId="4" pageNumber="207" unit="kg" value="5.6">5.6 kg</quantity>
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}; 
<materialsCitation id="3B7EE451FFE9FFD0C840FE44BB1BCEE1" collectingDate="2004-01-30" collectorName="Ysternek N. R. &amp; Dal &amp; Varings Tr. &amp; Hwy R" country="South Africa" elevation="339650" latitude="-33.913334" location="Afromontane forest" longLatPrecision="119" longitude="23.145" pageId="4" pageNumber="207" specimenCount="4" specimenCount-female="4" stateProvince="Western Cape" typeStatus="paratype">
4 JJ 
<specimenCount id="9D102585FFE9FFD0C80BFE44BDE8CEFA" box="[608,656,427,452]" pageId="4" pageNumber="207" type="female">4 ♀♀</specimenCount>
: ‘ 
<collectingCountry id="F301AE9CFFE9FFD0C8F4FE44BC6BCEFA" box="[671,787,433,452]" name="South Africa" pageId="4" pageNumber="207">South Africa</collectingCountry>
: 
<collectingRegion id="49D220EEFFE9FFD0C970FE44BCEDCEFA" box="[795,917,433,452]" country="South Africa" name="Western Cape" pageId="4" pageNumber="207">Western Cape</collectingRegion>
: 
<collectorName id="26E38BDAFFE9FFD0C9F6FE44BB63CEFA" box="[925,1051,433,452]" pageId="4" pageNumber="207">
Ysternek 
<taxonomicNameLabel id="A2518F65FFE9FFD0C99BFE44BB63CEFA" box="[1008,1051,433,452]" pageId="4" pageNumber="207" rank="subSpecies">N.R.</taxonomicNameLabel>
</collectorName>
, 
<collectorName id="26E38BDAFFE9FFD0CE48FE44BB3ACEFA" box="[1059,1090,433,452]" pageId="4" pageNumber="207">Dal</collectorName>
van 
<collectorName id="26E38BDAFFE9FFD0CAFCFE39BF80CEE1" box="[151,248,460,479]" pageId="4" pageNumber="207">Varings Tr.</collectorName>
, off 
<collectorName id="26E38BDAFFE9FFD0CB76FE39BE22CEE1" box="[285,346,460,479]" pageId="4" pageNumber="207">Hwy R</collectorName>
<quantity id="4CEE43E9FFE9FFD0CB31FE39BECFCEE1" box="[346,439,460,479]" metricMagnitude="5" metricUnit="m" metricValue="3.3964999999999996" pageId="4" pageNumber="207" unit="m" value="339650.0">
<elevation id="003B093FFFE9FFD0CB31FE39BECFCEE1" box="[346,439,460,479]" metricMagnitude="5" metricUnit="m" metricValue="3.3964999999999996" pageId="4" pageNumber="207" unit="m" value="339650.0">339, 650m</elevation>
</quantity>
, 
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, 
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, 
<date id="FFA8C8CCFFE9FFD0C8EAFE39BDACCEE1" box="[641,724,460,479]" pageId="4" pageNumber="207" value="2004-01-30">
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, tall 
<location id="8EC9B8D7FFE9FFD0C897FE39BCDBCEE1" LSID="urn:lsid:plazi:treatment:03BF5F1AFFEFFFD3CBB4FE0EBD3FCECD:8EC9B8D7FFE9FFD0C897FE39BCDBCEE1" box="[764,931,460,479]" country="South Africa" latitude="-33.913334" longLatPrecision="119" longitude="23.145" name="Afromontane forest" pageId="4" pageNumber="207" stateProvince="Western Cape">Afromontane forest</location>
w/abundant tree ferns
</materialsCitation>
; 
<materialsCitation id="3B7EE451FFE9FFD0CAFCFE12BE2ECD2A" collectionCode="FMNH" collectorName="Newton &amp; Thayer" country="South Africa" location="Field Museum Nat. Hist." pageId="4" pageNumber="207" specimenCount="1" stateProvince="Western Cape" typeStatus="paratype">
FMHD #2004-022, berl., leaf &amp; log litter, 
<collectorName id="26E38BDAFFE9FFD0C867FE12BD2ACEC4" box="[524,594,487,506]" pageId="4" pageNumber="207">Newton</collectorName>
, 
<collectorName id="26E38BDAFFE9FFD0C837FE12BDE0CEC4" box="[604,664,487,506]" pageId="4" pageNumber="207">Thayer</collectorName>
et al. 1080 
<location id="8EC9B8D7FFE9FFD0C968FE12BCA7CEC4" LSID="urn:lsid:plazi:treatment:03BF5F1AFFEFFFD3CBB4FE0EBD3FCECD:8EC9B8D7FFE9FFD0C968FE12BCA7CEC4" box="[771,991,487,506]" country="South Africa" name="Field Museum Nat. Hist." pageId="4" pageNumber="207" stateProvince="Western Cape">Field Museum Nat. Hist.</location>
’ (all 
<collectionCode id="ED0776C9FFE9FFD0CE67FE12BB34CEC4" box="[1036,1100,487,506]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="4" pageNumber="207" type="Museum">FMNH</collectionCode>
, in 70% ethanol) {
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}; 
<materialsCitation id="3B7EE451FFE9FFD0CB0DFDF4BC53CD11" collectingDate="2004-01-30" collectorName="Diepwalle S. F. &amp; Kom Se Pad" country="South Africa" elevation="339420" latitude="-33.945" location="Afromontane forest" longLatPrecision="772" longitude="23.133333" pageId="4" pageNumber="207" specimenCount="2" specimenCount-female="1" specimenCount-larva="1" stateProvince="Western Cape" typeStatus="paratype">
5 JJ 
<specimenCount id="9D102585FFE9FFD0CBF0FDF4BEC5CD2A" box="[411,445,507,532]" pageId="4" pageNumber="207" type="female">1 ♀</specimenCount>
(+ 
<specimenCount id="9D102585FFE9FFD0CBB1FDF4BD62CD2A" box="[474,538,513,532]" pageId="4" pageNumber="207" type="larva">1 larva</specimenCount>
): ‘ 
<collectingCountry id="F301AE9CFFE9FFD0C847FDF4BDD8CD2A" box="[556,672,513,532]" name="South Africa" pageId="4" pageNumber="207">South Africa</collectingCountry>
: 
<collectingRegion id="49D220EEFFE9FFD0C8C3FDF4BC5ACD2A" box="[680,802,513,532]" country="South Africa" name="Western Cape" pageId="4" pageNumber="207">Western Cape</collectingRegion>
: 
<collectorName id="26E38BDAFFE9FFD0C940FDF4BCD3CD2A" box="[811,939,513,532]" pageId="4" pageNumber="207">Diepwalle S.F.</collectorName>
, 
<collectorName id="26E38BDAFFE9FFD0C9DFFDF4BB5ACD2A" box="[948,1058,513,532]" pageId="4" pageNumber="207">Kom Se Pad</collectorName>
, 
<locationDeviation id="9A1BF72EFFE9FFD0CE40FDF4BFA1CD11" location="Hwy" pageId="4" pageNumber="207">
<quantity id="4CEE43E9FFE9FFD0CE40FDF4BB1ECD2A" box="[1067,1126,513,532]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="4" pageNumber="207" unit="km" value="1.7">1.7 km</quantity>
W Hwy
</locationDeviation>
R 
<quantity id="4CEE43E9FFE9FFD0CA85FDE9BE33CD11" box="[238,331,540,559]" metricMagnitude="5" metricUnit="m" metricValue="3.394199999999999" pageId="4" pageNumber="207" unit="m" value="339420.0">
<elevation id="003B093FFFE9FFD0CA85FDE9BE33CD11" box="[238,331,540,559]" metricMagnitude="5" metricUnit="m" metricValue="3.394199999999999" pageId="4" pageNumber="207" unit="m" value="339420.0">339, 420m</elevation>
</quantity>
, 
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, 
<geoCoordinate id="EE2288CBFFE9FFD0CBDFFDE9BE85CD11" box="[436,509,540,559]" degrees="23" direction="east" minutes="08" orientation="longitude" pageId="4" pageNumber="207" precision="925" value="23.133333">23°08’E</geoCoordinate>
, 
<date id="FFA8C8CCFFE9FFD0C86DFDE9BD21CD11" box="[518,601,540,559]" pageId="4" pageNumber="207" value="2004-01-30">
<collectingDate id="EFEC3124FFE9FFD0C86DFDE9BD21CD11" box="[518,601,540,559]" pageId="4" pageNumber="207" value="2004-01-30">30.i.2004</collectingDate>
</date>
, tall 
<location id="8EC9B8D7FFE9FFD0C8EAFDE9BC53CD11" LSID="urn:lsid:plazi:treatment:03BF5F1AFFEFFFD3CBB4FE0EBD3FCECD:8EC9B8D7FFE9FFD0C8EAFDE9BC53CD11" box="[641,811,540,559]" country="South Africa" latitude="-33.945" longLatPrecision="772" longitude="23.133333" name="Afromontane forest" pageId="4" pageNumber="207" stateProvince="Western Cape">Afromontane forest</location>
</materialsCitation>
; 
<materialsCitation id="3B7EE451FFE9FFD0C95FFDE9BB5DCD74" collectionCode="FMNH" collectorName="Newton &amp; Solodovnikov" country="South Africa" location="Field Museum Nat. Hist." pageId="4" pageNumber="207" specimenCount="1" stateProvince="Western Cape" typeStatus="paratype">
FMHD #2004-024, berl., leaf &amp; log litter, 
<collectorName id="26E38BDAFFE9FFD0CAA1FDC2BE68CD74" box="[202,272,567,586]" pageId="4" pageNumber="207">Newton</collectorName>
, 
<collectorName id="26E38BDAFFE9FFD0CB72FDC2BEE9CD74" box="[281,401,567,586]" pageId="4" pageNumber="207">Solodovnikov</collectorName>
et al. 1081 
<location id="8EC9B8D7FFE9FFD0CB93FDC2BDA9CD74" LSID="urn:lsid:plazi:treatment:03BF5F1AFFEFFFD3CBB4FE0EBD3FCECD:8EC9B8D7FFE9FFD0CB93FDC2BDA9CD74" box="[504,721,567,586]" country="South Africa" name="Field Museum Nat. Hist." pageId="4" pageNumber="207" stateProvince="Western Cape">Field Museum Nat. Hist.</location>
’ (all 
<collectionCode id="ED0776C9FFE9FFD0C890FDC2BC43CD74" box="[763,827,567,586]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="4" pageNumber="207" type="Museum">FMNH</collectionCode>
, in 70% ethanol) {10.0 kg
</materialsCitation>
}; 
<materialsCitation id="3B7EE451FFE9FFD0CE5EFDC2BDFBCD41" collectingDate="2004-01-31" collectorName="Gouna S. F. &amp; Kom Se Pad" country="South Africa" elevation="339440" latitude="-33.945" location="Afromontane forest" longLatPrecision="119" longitude="23.093334" pageId="4" pageNumber="207" specimenCount="2" specimenCount-female="2" stateProvince="Western Cape" typeStatus="paratype">
1 J 
<specimenCount id="9D102585FFE9FFD0CE37FDC2BFCECD5A" pageId="4" pageNumber="207" type="female">2 ♀♀</specimenCount>
: ‘ 
<collectingCountry id="F301AE9CFFE9FFD0CAADFDA4BE42CD5A" box="[198,314,593,612]" name="South Africa" pageId="4" pageNumber="207">South Africa</collectingCountry>
: 
<collectingRegion id="49D220EEFFE9FFD0CB29FDA4BEC5CD5A" box="[322,445,593,612]" country="South Africa" name="Western Cape" pageId="4" pageNumber="207">Western Cape</collectingRegion>
: 
<collectorName id="26E38BDAFFE9FFD0CBAEFDA4BD5FCD5A" box="[453,551,593,612]" pageId="4" pageNumber="207">Gouna S.F.</collectorName>
, 
<collectorName id="26E38BDAFFE9FFD0C85BFDA4BDE6CD5A" box="[560,670,593,612]" pageId="4" pageNumber="207">Kom Se Pad</collectorName>
, 
<locationDeviation id="9A1BF72EFFE9FFD0C8C3FDA4BC54CD5A" box="[680,812,593,612]" location="Hwy" pageId="4" pageNumber="207">
<quantity id="4CEE43E9FFE9FFD0C8C3FDA4BD9BCD5A" box="[680,739,593,612]" metricMagnitude="3" metricUnit="m" metricValue="6.2" pageId="4" pageNumber="207" unit="km" value="6.2">6.2 km</quantity>
W Hwy
</locationDeviation>
R 
<quantity id="4CEE43E9FFE9FFD0C929FDA4BCE7CD5A" box="[834,927,593,612]" metricMagnitude="5" metricUnit="m" metricValue="3.3944" pageId="4" pageNumber="207" unit="m" value="339440.0">
<elevation id="003B093FFFE9FFD0C929FDA4BCE7CD5A" box="[834,927,593,612]" metricMagnitude="5" metricUnit="m" metricValue="3.3944" pageId="4" pageNumber="207" unit="m" value="339440.0">339, 440m</elevation>
</quantity>
, 
<geoCoordinate id="EE2288CBFFE9FFD0C9C3FDA4BB79CD5A" box="[936,1025,593,612]" degrees="33" direction="south" minutes="56.7" orientation="latitude" pageId="4" pageNumber="207" precision="92" value="-33.945">33°56.7’S</geoCoordinate>
, 
<geoCoordinate id="EE2288CBFFE9FFD0CE61FDA4BB1BCD5A" box="[1034,1123,593,612]" degrees="23" direction="east" minutes="05.6" orientation="longitude" pageId="4" pageNumber="207" precision="92" value="23.093334">23°05.6’E</geoCoordinate>
, 
<date id="FFA8C8CCFFE9FFD0CAFCFD99BF91CD41" box="[151,233,620,639]" pageId="4" pageNumber="207" value="2004-01-31">
<collectingDate id="EFEC3124FFE9FFD0CAFCFD99BF91CD41" box="[151,233,620,639]" pageId="4" pageNumber="207" value="2004-01-31">31.i.2004</collectingDate>
</date>
, 
<location id="8EC9B8D7FFE9FFD0CA9AFD99BEE1CD41" LSID="urn:lsid:plazi:treatment:03BF5F1AFFEFFFD3CBB4FE0EBD3FCECD:8EC9B8D7FFE9FFD0CA9AFD99BEE1CD41" box="[241,409,620,639]" country="South Africa" latitude="-33.945" longLatPrecision="119" longitude="23.093334" name="Afromontane forest" pageId="4" pageNumber="207" stateProvince="Western Cape">Afromontane forest</location>
w/ground ferns, few lianas
</materialsCitation>
; 
<materialsCitation id="3B7EE451FFE9FFD0C8E7FD99BDCBCDA4" collectionCode="FMNH" collectorName="Newton" country="South Africa" location="Field Museum Nat. Hist." pageId="4" pageNumber="207" specimenCount="1" stateProvince="Western Cape" typeStatus="paratype">
FMHD #2004-030, berl., leaf &amp; log litter, 
<collectorName id="26E38BDAFFE9FFD0C998FD99BB4FCD41" box="[1011,1079,620,639]" pageId="4" pageNumber="207">Newton</collectorName>
1083 
<location id="8EC9B8D7FFE9FFD0CAFCFD72BE16CDA4" LSID="urn:lsid:plazi:treatment:03BF5F1AFFEFFFD3CBB4FE0EBD3FCECD:8EC9B8D7FFE9FFD0CAFCFD72BE16CDA4" box="[151,366,647,666]" country="South Africa" name="Field Museum Nat. Hist." pageId="4" pageNumber="207" stateProvince="Western Cape">Field Museum Nat. Hist.</location>
’ (all 
<collectionCode id="ED0776C9FFE9FFD0CBFCFD72BEAFCDA4" box="[407,471,647,666]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="4" pageNumber="207" type="Museum">FMNH</collectionCode>
, in 70% ethanol) {3.0 kg
</materialsCitation>
}.
</paragraph>
</subSubSection>
<caption id="DF69BE84FFEFFFD6C922FA3DBCDBC978" ID-DOI="http://doi.org/10.5281/zenodo.5305426" ID-Zenodo-Dep="5305426" httpUri="https://zenodo.org/record/5305426/files/figure.png" pageId="2" pageNumber="205" startId="2.[841,879,1480,1499]" targetBox="[151,830,690,1602]" targetPageId="2">
<paragraph id="8BA9EE0CFFEFFFD6C922FA3DBCDBC978" blockId="2.[841,1126,1480,1606]" pageId="2" pageNumber="205">
Figs 1–2. 
<taxonomicName id="4C16958FFFEFFFD6C9C3FA3DBB1ECAE5" authority="Janák &amp; Makranczy, 2016" authorityName="Janák &amp; Makranczy" authorityYear="2016" box="[936,1126,1480,1499]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="205" phylum="Arthropoda" rank="species" species="crypticola" status="sp. nov.">
<emphasis id="B962321EFFEFFFD6C9C3FA3DBB1ECAE5" box="[936,1126,1480,1499]" italics="true" pageId="2" pageNumber="205">Thinobius crypticola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2518F65FFEFFFD6C922FA16BCE8CAC8" box="[841,912,1507,1526]" pageId="2" pageNumber="205" rank="species">sp. nov.</taxonomicNameLabel>
: 1 – male holotype, dorsal habitus; 2 – suspected larva (maximum width of head capsule 0.16 mm).
</paragraph>
</caption>
<caption id="DF69BE84FFEEFFD7CA1AF9EDBE92C97B" ID-DOI="http://doi.org/10.5281/zenodo.5305428" ID-Zenodo-Dep="5305428" httpUri="https://zenodo.org/record/5305428/files/figure.png" pageId="3" pageNumber="206" startId="3.[113,150,1560,1579]" targetBox="[113,1088,219,1548]" targetPageId="3">
<paragraph id="8BA9EE0CFFEEFFD7CA1AF9EDBE92C97B" blockId="3.[113,1089,1560,1605]" pageId="3" pageNumber="206">
Figs 3–4. Habitats of 
<taxonomicName id="4C16958FFFEEFFD7CB40F9EDBEA5C915" authority="Janák &amp; Makranczy, 2016" authorityName="Janák &amp; Makranczy" authorityYear="2016" box="[299,477,1560,1579]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="206" phylum="Arthropoda" rank="species" species="crypticola" status="sp. nov.">
<emphasis id="B962321EFFEEFFD7CB40F9EDBEA5C915" box="[299,477,1560,1579]" italics="true" pageId="3" pageNumber="206">Thinobius crypticola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2518F65FFEEFFD7CB88F9EDBD5EC915" box="[483,550,1560,1579]" pageId="3" pageNumber="206" rank="species">sp. nov.</taxonomicNameLabel>
3 – Knysna, Dal van Varings Trail (type locality); 4 – Knysna, Diepwalle State Forest (photos by J. Janák).
</paragraph>
</caption>
<subSubSection id="C30CBD87FFE9FFD1CAFCFD47BD93CD0D" lastPageId="5" lastPageNumber="208" pageId="4" pageNumber="207" type="description">
<paragraph id="8BA9EE0CFFE9FFD0CAFCFD47BB1FCCF7" blockId="4.[151,1127,690,1609]" pageId="4" pageNumber="207">
<emphasis id="B962321EFFE9FFD0CAFCFD47BE59CDF7" bold="true" box="[151,289,690,713]" pageId="4" pageNumber="207">Description.</emphasis>
<emphasis id="B962321EFFE9FFD0CB4EFD47BEC6CDF7" box="[293,446,690,713]" italics="true" pageId="4" pageNumber="207">Measurements</emphasis>
(in mm, n = 10): HW = 0.175 (
<geoCoordinate id="EE2288CBFFE9FFD0C890FD47BC41CDF7" box="[763,825,690,713]" degrees="0.165" direction="north" orientation="latitude" pageId="4" pageNumber="207" precision="55" value="0.165">0.165</geoCoordinate>
<geoCoordinate id="EE2288CBFFE9FFD0C952FD47BCFDCDF7" box="[825,901,690,713]" degrees="0.185" direction="west" orientation="longitude" pageId="4" pageNumber="207" precision="55" value="-0.185">–0.185</geoCoordinate>
); PW = 0.180 (
<geoCoordinate id="EE2288CBFFE9FFD0CE77FD47BB21CDF7" box="[1052,1113,690,713]" degrees="0.170" direction="north" orientation="latitude" pageId="4" pageNumber="207" precision="55" value="0.17">0.170</geoCoordinate>
<geoCoordinate id="EE2288CBFFE9FFD0CE32FD47BFA0CDD7" degrees="0.185" direction="west" orientation="longitude" pageId="4" pageNumber="207" precision="55" value="-0.185">– 0.185</geoCoordinate>
); EW = 0.170 (
<geoCoordinate id="EE2288CBFFE9FFD0CB11FD27BEC1CDD7" box="[378,441,722,745]" degrees="0.160" direction="north" orientation="latitude" pageId="4" pageNumber="207" precision="55" value="0.16">0.160</geoCoordinate>
<geoCoordinate id="EE2288CBFFE9FFD0CBD2FD27BD7ECDD7" box="[441,518,722,745]" degrees="0.180" direction="west" orientation="longitude" pageId="4" pageNumber="207" precision="55" value="-0.18">–0.180</geoCoordinate>
); AW = 0.210 (
<geoCoordinate id="EE2288CBFFE9FFD0C8C3FD27BD9FCDD7" box="[680,743,722,745]" degrees="0.190" direction="north" orientation="latitude" pageId="4" pageNumber="207" precision="55" value="0.19">0.190</geoCoordinate>
<geoCoordinate id="EE2288CBFFE9FFD0C88DFD27BC4CCDD7" box="[742,820,722,745]" degrees="0.230" direction="west" orientation="longitude" pageId="4" pageNumber="207" precision="55" value="-0.23">–0.230</geoCoordinate>
); HL = 0.160 (
<geoCoordinate id="EE2288CBFFE9FFD0C9A4FD27BB76CDD7" box="[975,1038,722,745]" degrees="0.150" direction="north" orientation="latitude" pageId="4" pageNumber="207" precision="55" value="0.15">0.150</geoCoordinate>
<geoCoordinate id="EE2288CBFFE9FFD0CE65FD27BB23CDD7" box="[1038,1115,722,745]" degrees="0.170" direction="west" orientation="longitude" pageId="4" pageNumber="207" precision="55" value="-0.17">–0.170</geoCoordinate>
); PL = 0.145 (
<geoCoordinate id="EE2288CBFFE9FFD0CB73FD07BE2ECC37" box="[280,342,754,777]" degrees="0.135" direction="north" orientation="latitude" pageId="4" pageNumber="207" precision="55" value="0.135">0.135</geoCoordinate>
<geoCoordinate id="EE2288CBFFE9FFD0CB3DFD07BEDBCC37" box="[342,419,754,777]" degrees="0.150" direction="west" orientation="longitude" pageId="4" pageNumber="207" precision="55" value="-0.15">–0.150</geoCoordinate>
); SL = 0.130 (
<geoCoordinate id="EE2288CBFFE9FFD0C85EFD07BD0CCC37" box="[565,628,754,777]" degrees="0.120" direction="north" orientation="latitude" pageId="4" pageNumber="207" precision="55" value="0.12">0.120</geoCoordinate>
<geoCoordinate id="EE2288CBFFE9FFD0C81FFD07BDB9CC37" box="[628,705,754,777]" degrees="0.135" direction="west" orientation="longitude" pageId="4" pageNumber="207" precision="55" value="-0.135">–0.135</geoCoordinate>
); SC = 0.115 (
<geoCoordinate id="EE2288CBFFE9FFD0C93FFD07BCEACC37" box="[852,914,754,777]" degrees="0.105" direction="north" orientation="latitude" pageId="4" pageNumber="207" precision="55" value="0.105">0.105</geoCoordinate>
<geoCoordinate id="EE2288CBFFE9FFD0C9F9FD07BCA7CC37" box="[914,991,754,777]" degrees="0.120" direction="west" orientation="longitude" pageId="4" pageNumber="207" precision="55" value="-0.12">–0.120</geoCoordinate>
); FB = 0.440 (
<geoCoordinate id="EE2288CBFFE9FFD0CAF4FCE7BFA6CC17" box="[159,222,786,809]" degrees="0.420" direction="north" orientation="latitude" pageId="4" pageNumber="207" precision="55" value="0.42">0.420</geoCoordinate>
<geoCoordinate id="EE2288CBFFE9FFD0CAB6FCE7BE53CC17" box="[221,299,786,809]" degrees="0.460" direction="west" orientation="longitude" pageId="4" pageNumber="207" precision="55" value="-0.46">–0.460</geoCoordinate>
); BL = 1.110 (
<geoCoordinate id="EE2288CBFFE9FFD0CBAEFCE7BD7DCC17" box="[453,517,786,809]" degrees="0.955" direction="north" orientation="latitude" pageId="4" pageNumber="207" precision="55" value="0.955">0.955</geoCoordinate>
<geoCoordinate id="EE2288CBFFE9FFD0C86FFCE7BD28CC17" box="[516,592,786,809]" degrees="1.235" direction="west" orientation="longitude" pageId="4" pageNumber="207" precision="55" value="-1.235">–1.235</geoCoordinate>
). 
<emphasis id="B962321EFFE9FFD0C809FCE7BDE1CC17" box="[610,665,786,809]" italics="true" pageId="4" pageNumber="207">Body</emphasis>
(
<figureCitation id="132DF289FFE9FFD0C8C3FCE7BD9ECC17" box="[680,742,786,809]" captionStart="Figs 1–2" captionStartId="2.[841,879,1480,1499]" captionTargetBox="[151,830,690,1602]" captionTargetId="figure-190@2.[151,830,687,1605]" captionTargetPageId="2" captionText="Figs 1–2. Thinobius crypticola sp. nov.: 1 – male holotype, dorsal habitus; 2 – suspected larva (maximum width of head capsule 0.16 mm)." figureDoi="http://doi.org/10.5281/zenodo.5305426" httpUri="https://zenodo.org/record/5305426/files/figure.png" pageId="4" pageNumber="207">Fig. 1</figureCitation>
) straw yellow, moderately lustrous, although microsculpture weak (inconspicuous on forebody). Forebody sparsely punctate, punctures moderately small, interspaces on average 4–5× puncture diameters. Body rather translucent, area between supraantennal tubercles (around epistomal suture) darker, pronotal marginal bead laterally and posteriorly marked as thin darker line. A few thicker setae dark brown, finer setae match overall body colour.Apex of antennomere II with slightly darker rim.
</paragraph>
<paragraph id="8BA9EE0CFFE9FFD0CADEFC27BDACCA17" blockId="4.[151,1127,690,1609]" pageId="4" pageNumber="207">
<emphasis id="B962321EFFE9FFD0CADEFC27BF97CCD7" box="[181,239,978,1001]" italics="true" pageId="4" pageNumber="207">Head</emphasis>
slightly transverse, eyes absent, sides of head gently curved. Neck not separated by groove, although sides of head constricted before it and narrowly rounded there. Disc of head gently convex, with feeble coriaceous microsculpture (with isodiametric cells of varying sizes), at obscure demarcation line of neck microsculpture turning transverse; supraantennal tubercles slightly elevated. Maxillary palps (
<figureCitation id="132DF289FFE9FFD0C866FBA7BD35CB57" box="[525,589,1106,1129]" captionStart="Figs 5–13" captionStartId="6.[151,188,1514,1533]" captionTargetBox="[152,1099,222,1451]" captionTargetId="figure-9@6.[151,1103,219,1466]" captionTargetPageId="6" captionText="Figs 5–13. Thinobius crypticola sp. nov. 5 – antenna; 6 – maxillary palp; 7 – scutellum; 8 – male sternite VIII; 9 – male tergite IX; 10 – male sternite IX; 11 – aedeagus, frontal; 12 – female genital appendage; 13 – spermatheca. Scale bars: 0.075 mm for 11, 13; 0.09 mm for 5–7, 9–10, 12; 0.1 mm for 8." figureDoi="http://doi.org/10.5281/zenodo.5305430" httpUri="https://zenodo.org/record/5305430/files/figure.png" pageId="4" pageNumber="207">Fig. 6</figureCitation>
) with penultimate segment fusiform, last segment not acicular but asymmetrical, bulbous at base. 
<emphasis id="B962321EFFE9FFD0C8E6FB87BD8ACBB7" box="[653,754,1138,1161]" italics="true" pageId="4" pageNumber="207">Antennae</emphasis>
(
<figureCitation id="132DF289FFE9FFD0C894FB87BC46CBB7" box="[767,830,1138,1161]" captionStart="Figs 5–13" captionStartId="6.[151,188,1514,1533]" captionTargetBox="[152,1099,222,1451]" captionTargetId="figure-9@6.[151,1103,219,1466]" captionTargetPageId="6" captionText="Figs 5–13. Thinobius crypticola sp. nov. 5 – antenna; 6 – maxillary palp; 7 – scutellum; 8 – male sternite VIII; 9 – male tergite IX; 10 – male sternite IX; 11 – aedeagus, frontal; 12 – female genital appendage; 13 – spermatheca. Scale bars: 0.075 mm for 11, 13; 0.09 mm for 5–7, 9–10, 12; 0.1 mm for 8." figureDoi="http://doi.org/10.5281/zenodo.5305430" httpUri="https://zenodo.org/record/5305430/files/figure.png" pageId="4" pageNumber="207">Fig. 5</figureCitation>
) rather short, antennomeres I, II and XI quite stout, mid antennomeres strongly transverse; antennomere V significantly smaller than neighbouring antennomeres, latter may appear somewhat asymmetrical from a certain viewpoint. Antennomeres IX–XI with modified setae (remarkably thicker, less point- ed), similar to those in other endogean staphylinids like leptotyphlines, antennomere VII also occasionally with few small and short modified setae.
</paragraph>
<paragraph id="8BA9EE0CFFE9FFD1CADEFAC7BE9ECE2D" blockId="4.[151,1127,690,1609]" lastBlockId="5.[113,1091,220,1587]" lastPageId="5" lastPageNumber="208" pageId="4" pageNumber="207">
<emphasis id="B962321EFFE9FFD0CADEFAC7BE67CA77" box="[181,287,1330,1353]" italics="true" pageId="4" pageNumber="207">Pronotum</emphasis>
transverse, insignificantly wider than head, corners rounded, posteriorly more broadly than anteriorly. Arched sides somewhat constricted towards base, although widest point still around or little behind middle. Pronotal marginal bead very thin, anteriorly mostly missing. Disc with similar microsculpture as vertex, in middle with two parallel longitudinal impressions (leaving free anterior fourth). Scutellum as in 
<figureCitation id="132DF289FFE9FFD0C964FA47BC28CAF7" box="[783,848,1458,1481]" captionStart="Figs 5–13" captionStartId="6.[151,188,1514,1533]" captionTargetBox="[152,1099,222,1451]" captionTargetId="figure-9@6.[151,1103,219,1466]" captionTargetPageId="6" captionText="Figs 5–13. Thinobius crypticola sp. nov. 5 – antenna; 6 – maxillary palp; 7 – scutellum; 8 – male sternite VIII; 9 – male tergite IX; 10 – male sternite IX; 11 – aedeagus, frontal; 12 – female genital appendage; 13 – spermatheca. Scale bars: 0.075 mm for 11, 13; 0.09 mm for 5–7, 9–10, 12; 0.1 mm for 8." figureDoi="http://doi.org/10.5281/zenodo.5305430" httpUri="https://zenodo.org/record/5305430/files/figure.png" pageId="4" pageNumber="207">Fig. 7</figureCitation>
, sculpture rather reduced, short setae only in postero-lateral and lateral parts. 
<emphasis id="B962321EFFE9FFD0C8A0FA27BC76CAD7" box="[715,782,1490,1513]" italics="true" pageId="4" pageNumber="207">Elytra</emphasis>
combined significantly broader than long, dilated towards apex, together less wide than pronotum, anterior margin at sutural corners slightly concave. Humeri poorly developed, posterior elytral corners broadly round- ed, disc mostly convex, along suture with marginal bead. Hind wings absent. 
<emphasis id="B962321EFFE9FFD0C9A7F9C7BC86C977" box="[972,1022,1586,1609]" italics="true" pageId="4" pageNumber="207">Legs</emphasis>
short and rather stout, tibiae spindle-shaped. Tarsi two segmented as in other 
<taxonomicName id="4C16958FFFE8FFD1C92CFF29BCCACFCD" authorityName="Kiesenwetter" authorityYear="1844" box="[839,946,220,243]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="208" phylum="Arthropoda" rank="genus">
<emphasis id="B962321EFFE8FFD1C92CFF29BCCACFCD" box="[839,946,220,243]" italics="true" pageId="5" pageNumber="208">Thinobius</emphasis>
</taxonomicName>
, basal article with setae modified as tarsal lobes.
</paragraph>
<paragraph id="8BA9EE0CFFE8FFD1CAFBFEE9BD93CD0D" blockId="5.[113,1091,220,1587]" pageId="5" pageNumber="208">
<emphasis id="B962321EFFE8FFD1CAFBFEE9BF8DCE0D" box="[144,245,284,307]" italics="true" pageId="5" pageNumber="208">Abdomen</emphasis>
weakly fusiform, sides more or less straight and parallel, widest around middle and slightly wider than elytra. Tergites with narrow laterosclerites, basal ridges and basolateral ridges separated and running parallel in the middle, basolateral ridges marked in dark brown colour medially. Surface with strongly transverse coriaceous microsculpture, apical margin of tergite VII with thin palisade fringe consisting of short and somewhat sparse setae (not tightly arranged). 
<emphasis id="B962321EFFE8FFD1CB5AFE49BD89CEED" box="[305,753,444,467]" italics="true" pageId="5" pageNumber="208">Primary and secondary sexual characters</emphasis>
. Sexes do not differ in external characters except terminalia. Male sternite VIII as in 
<figureCitation id="132DF289FFE8FFD1C8C1FE29BD92CECD" box="[682,746,476,499]" captionStart="Figs 5–13" captionStartId="6.[151,188,1514,1533]" captionTargetBox="[152,1099,222,1451]" captionTargetId="figure-9@6.[151,1103,219,1466]" captionTargetPageId="6" captionText="Figs 5–13. Thinobius crypticola sp. nov. 5 – antenna; 6 – maxillary palp; 7 – scutellum; 8 – male sternite VIII; 9 – male tergite IX; 10 – male sternite IX; 11 – aedeagus, frontal; 12 – female genital appendage; 13 – spermatheca. Scale bars: 0.075 mm for 11, 13; 0.09 mm for 5–7, 9–10, 12; 0.1 mm for 8." figureDoi="http://doi.org/10.5281/zenodo.5305430" httpUri="https://zenodo.org/record/5305430/files/figure.png" pageId="5" pageNumber="208">Fig. 8</figureCitation>
, tergites IX as in 
<figureCitation id="132DF289FFE8FFD1C9CDFE29BC9ECECD" box="[934,998,476,499]" captionStart="Figs 5–13" captionStartId="6.[151,188,1514,1533]" captionTargetBox="[152,1099,222,1451]" captionTargetId="figure-9@6.[151,1103,219,1466]" captionTargetPageId="6" captionText="Figs 5–13. Thinobius crypticola sp. nov. 5 – antenna; 6 – maxillary palp; 7 – scutellum; 8 – male sternite VIII; 9 – male tergite IX; 10 – male sternite IX; 11 – aedeagus, frontal; 12 – female genital appendage; 13 – spermatheca. Scale bars: 0.075 mm for 11, 13; 0.09 mm for 5–7, 9–10, 12; 0.1 mm for 8." figureDoi="http://doi.org/10.5281/zenodo.5305430" httpUri="https://zenodo.org/record/5305430/files/figure.png" pageId="5" pageNumber="208">Fig. 9</figureCitation>
, sternite IX as in 
<figureCitation id="132DF289FFE8FFD1CABAFE09BE59CD2D" box="[209,289,508,531]" captionStart="Figs 5–13" captionStartId="6.[151,188,1514,1533]" captionTargetBox="[152,1099,222,1451]" captionTargetId="figure-9@6.[151,1103,219,1466]" captionTargetPageId="6" captionText="Figs 5–13. Thinobius crypticola sp. nov. 5 – antenna; 6 – maxillary palp; 7 – scutellum; 8 – male sternite VIII; 9 – male tergite IX; 10 – male sternite IX; 11 – aedeagus, frontal; 12 – female genital appendage; 13 – spermatheca. Scale bars: 0.075 mm for 11, 13; 0.09 mm for 5–7, 9–10, 12; 0.1 mm for 8." figureDoi="http://doi.org/10.5281/zenodo.5305430" httpUri="https://zenodo.org/record/5305430/files/figure.png" pageId="5" pageNumber="208">Fig. 10</figureCitation>
, tergite X similar in both sexes. Aedeagus as in 
<figureCitation id="132DF289FFE8FFD1C95BFE09BC06CD2D" box="[816,894,508,531]" captionStart="Figs 5–13" captionStartId="6.[151,188,1514,1533]" captionTargetBox="[152,1099,222,1451]" captionTargetId="figure-9@6.[151,1103,219,1466]" captionTargetPageId="6" captionText="Figs 5–13. Thinobius crypticola sp. nov. 5 – antenna; 6 – maxillary palp; 7 – scutellum; 8 – male sternite VIII; 9 – male tergite IX; 10 – male sternite IX; 11 – aedeagus, frontal; 12 – female genital appendage; 13 – spermatheca. Scale bars: 0.075 mm for 11, 13; 0.09 mm for 5–7, 9–10, 12; 0.1 mm for 8." figureDoi="http://doi.org/10.5281/zenodo.5305430" httpUri="https://zenodo.org/record/5305430/files/figure.png" pageId="5" pageNumber="208">Fig. 11</figureCitation>
. Female modified genital appendages as in 
<figureCitation id="132DF289FFE8FFD1CB17FDE9BEB2CD0D" box="[380,458,540,563]" captionStart="Figs 5–13" captionStartId="6.[151,188,1514,1533]" captionTargetBox="[152,1099,222,1451]" captionTargetId="figure-9@6.[151,1103,219,1466]" captionTargetPageId="6" captionText="Figs 5–13. Thinobius crypticola sp. nov. 5 – antenna; 6 – maxillary palp; 7 – scutellum; 8 – male sternite VIII; 9 – male tergite IX; 10 – male sternite IX; 11 – aedeagus, frontal; 12 – female genital appendage; 13 – spermatheca. Scale bars: 0.075 mm for 11, 13; 0.09 mm for 5–7, 9–10, 12; 0.1 mm for 8." figureDoi="http://doi.org/10.5281/zenodo.5305430" httpUri="https://zenodo.org/record/5305430/files/figure.png" pageId="5" pageNumber="208">Fig. 12</figureCitation>
, spermatheca as in 
<figureCitation id="132DF289FFE8FFD1C8F2FDE9BD9FCD0D" box="[665,743,540,563]" captionStart="Figs 5–13" captionStartId="6.[151,188,1514,1533]" captionTargetBox="[152,1099,222,1451]" captionTargetId="figure-9@6.[151,1103,219,1466]" captionTargetPageId="6" captionText="Figs 5–13. Thinobius crypticola sp. nov. 5 – antenna; 6 – maxillary palp; 7 – scutellum; 8 – male sternite VIII; 9 – male tergite IX; 10 – male sternite IX; 11 – aedeagus, frontal; 12 – female genital appendage; 13 – spermatheca. Scale bars: 0.075 mm for 11, 13; 0.09 mm for 5–7, 9–10, 12; 0.1 mm for 8." figureDoi="http://doi.org/10.5281/zenodo.5305430" httpUri="https://zenodo.org/record/5305430/files/figure.png" pageId="5" pageNumber="208">Fig. 13</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30CBD87FFE8FFD1CA1AFDC9BD83CCAD" pageId="5" pageNumber="208" type="diagnosis">
<paragraph id="8BA9EE0CFFE8FFD1CA1AFDC9BD83CCAD" blockId="5.[113,1091,220,1587]" pageId="5" pageNumber="208">
<emphasis id="B962321EFFE8FFD1CA1AFDC9BE14CD6D" bold="true" box="[113,364,572,595]" pageId="5" pageNumber="208">Differential diagnosis.</emphasis>
The new species bears similarity only to 
<taxonomicName id="4C16958FFFE8FFD1C946FDC9BE70CD4D" authority="Lobl &amp; Rychlik, 1994" authorityName="Lobl &amp; Rychlik" authorityYear="1994" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="208" phylum="Arthropoda" rank="species" species="korbeli">
<emphasis id="B962321EFFE8FFD1C946FDC9BC93CD6D" box="[813,1003,572,595]" italics="true" pageId="5" pageNumber="208">Thinobius korbeli</emphasis>
<bibRefCitation id="EF8793FDFFE8FFD1C999FDC9BE70CD4D" author="LOBL I. &amp; RYCHLIK I." pageId="5" pageNumber="208" pagination="25 - 32" refId="ref3747" refString="LOBL I. &amp; RYCHLIK I. 1994: Thinobius korbeli sp. nov., an anophthalmous oxyteline (Coleoptera: Staphylinidae) from Slovakia. Entomological Problems 25: 25 - 32." type="journal article" year="1994">Löbl &amp; Rychlík, 1994</bibRefCitation>
</taxonomicName>
from 
<collectingCountry id="F301AE9CFFE8FFD1CB27FDA9BED1CD4D" box="[332,425,604,627]" name="Slovakia" pageId="5" pageNumber="208">Slovakia</collectingCountry>
and 
<collectingCountry id="F301AE9CFFE8FFD1CB8BFDA9BD45CD4D" box="[480,573,604,627]" name="Hungary" pageId="5" pageNumber="208">Hungary</collectingCountry>
(
<bibRefCitation id="EF8793FDFFE8FFD1C826FDA9BC39CD4D" author="LOBL I. &amp; RYCHLIK I." box="[589,833,604,627]" pageId="5" pageNumber="208" pagination="25 - 32" refId="ref3747" refString="LOBL I. &amp; RYCHLIK I. 1994: Thinobius korbeli sp. nov., an anophthalmous oxyteline (Coleoptera: Staphylinidae) from Slovakia. Entomological Problems 25: 25 - 32." type="journal article" year="1994">LÖBL &amp; RYCHLÍK 1994</bibRefCitation>
, 
<bibRefCitation id="EF8793FDFFE8FFD1C926FDA9BB73CD4D" author="MAKRANCZY GY" box="[845,1035,604,627]" pageId="5" pageNumber="208" pagination="173 - 177" refId="ref3817" refString="MAKRANCZY GY. 2008: Thinobius korbeli found in the Szigetkoz, Hungary (Coleoptera: Staphylinidae: Oxytelinae). Folia Entomologica Hungarica 69: 173 - 177." type="journal article" year="2008">MAKRANCZY 2008</bibRefCitation>
) and 
<taxonomicName id="4C16958FFFE8FFD1CA1AFD89BEE5CDAD" authority="(Sawada, 1971)" baseAuthorityName="Sawada" baseAuthorityYear="1971" box="[113,413,636,659]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="208" phylum="Arthropoda" rank="species" species="kuroshio">
<emphasis id="B962321EFFE8FFD1CA1AFD89BF92CDAD" box="[113,234,636,659]" italics="true" pageId="5" pageNumber="208">T. kuroshio</emphasis>
(
<bibRefCitation id="EF8793FDFFE8FFD1CA91FD89BEECCDAD" author="SAWADA K." box="[250,404,636,659]" pageId="5" pageNumber="208" pagination="327 - 330" refId="ref3967" refString="SAWADA K. 1971: On the new genus and species of Leptotyphlinae of Japan (Coleoptera, Staphylinidae). Revue d'Ecologie et de Biologie du Sol 8: 327 - 330." type="journal article" year="1971">Sawada, 1971</bibRefCitation>
)
</taxonomicName>
from 
<collectingCountry id="F301AE9CFFE8FFD1CB89FD89BD67CDAD" box="[482,543,636,659]" name="Japan" pageId="5" pageNumber="208">Japan</collectingCountry>
(
<bibRefCitation id="EF8793FDFFE8FFD1C844FD89BC0ECDAD" author="GUSAROV V. I. &amp; MAKRANCZY GY" box="[559,886,636,659]" pageId="5" pageNumber="208" pagination="1 - 4" refId="ref3567" refString="GUSAROV V. I. &amp; MAKRANCZY GY. 2004: Yosiityphlus Sawada, 1971, a new synonym of Thinobius Kiesenwetter, 1844 (Coleoptera: Staphylinidae). Zootaxa 748: 1 - 4." type="journal article" year="2004">GUSAROV &amp; MAKRANCZY 2004</bibRefCitation>
) in the absence of eyes and functional wings. The male genitalia of 
<taxonomicName id="4C16958FFFE8FFD1C8ECFD69BC76CD8D" authority="Janák &amp; Makranczy, 2016" authorityName="Janák &amp; Makranczy" authorityYear="2016" box="[647,782,668,691]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="208" phylum="Arthropoda" rank="species" species="crypticola" status="sp. nov.">
<emphasis id="B962321EFFE8FFD1C8ECFD69BC76CD8D" box="[647,782,668,691]" italics="true" pageId="5" pageNumber="208">T. crypticola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2518F65FFE8FFD1C97DFD69BC12CD8D" box="[790,874,668,691]" pageId="5" pageNumber="208" rank="species">sp. nov.</taxonomicNameLabel>
are similar to those of many other 
<taxonomicName id="4C16958FFFE8FFD1CB7BFD49BEAFCDED" box="[272,471,700,723]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="208" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B962321EFFE8FFD1CB7BFD49BE03CDED" box="[272,379,700,723]" italics="true" pageId="5" pageNumber="208">Thinobius</emphasis>
species.
</taxonomicName>
<taxonomicName id="4C16958FFFE8FFD1CBB6FD49BDD3CDED" baseAuthorityName="Sawada" baseAuthorityYear="1971" box="[477,683,700,723]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="208" phylum="Arthropoda" rank="species" species="kuroshio">
<emphasis id="B962321EFFE8FFD1CBB6FD49BDD3CDED" box="[477,683,700,723]" italics="true" pageId="5" pageNumber="208">Thinobius kuroshio</emphasis>
</taxonomicName>
has much longer antennae with more elongate antennomeres, while 
<taxonomicName id="4C16958FFFE8FFD1CBD3FD29BD66CDCD" authorityName="Lobl &amp; Rychlik" authorityYear="1994" box="[440,542,732,755]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="208" phylum="Arthropoda" rank="species" species="korbeli">
<emphasis id="B962321EFFE8FFD1CBD3FD29BD66CDCD" box="[440,542,732,755]" italics="true" pageId="5" pageNumber="208">T. korbeli</emphasis>
</taxonomicName>
also has less transverse mid-antennal segments. In the new species these are conspicuously transverse (approximately twice as broad as long) and maxillary palps are remarkably different from both other blind species: the normally acicular last palpomere is swollen and curved in the middle and the penultimate palpomere is more slender. The two species of the Northern Hemisphere seem to be united by a more swollen penultimate palpomere and an acicular last segment.
</paragraph>
</subSubSection>
<subSubSection id="C30CBD87FFE8FFD1CA1AFC69BE12CCED" pageId="5" pageNumber="208" type="etymology">
<paragraph id="8BA9EE0CFFE8FFD1CA1AFC69BE12CCED" blockId="5.[113,1091,220,1587]" pageId="5" pageNumber="208">
<emphasis id="B962321EFFE8FFD1CA1AFC69BF8ACC8D" bold="true" box="[113,242,924,947]" pageId="5" pageNumber="208">Etymology.</emphasis>
The species epithet refers to its hidden way of life, the Latin word ‘ 
<taxonomicName id="4C16958FFFE8FFD1C9A5FC69BB42CC8D" authorityName="Janák &amp; Makranczy" authorityYear="2016" box="[974,1082,924,947]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="208" phylum="Arthropoda" rank="species" species="crypticola">
<emphasis id="B962321EFFE8FFD1C9A5FC69BB42CC8D" box="[974,1082,924,947]" italics="true" pageId="5" pageNumber="208">crypticola</emphasis>
</taxonomicName>
’ is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C30CBD87FFE8FFD3CA1AFC29BD3FCECD" lastPageId="7" lastPageNumber="210" pageId="5" pageNumber="208" type="distribution">
<paragraph id="8BA9EE0CFFE8FFD1CA1AFC29BDF6CA2D" blockId="5.[113,1091,220,1587]" pageId="5" pageNumber="208">
<emphasis id="B962321EFFE8FFD1CA1AFC29BEE8CCCD" bold="true" box="[113,400,988,1011]" pageId="5" pageNumber="208">Distribution and habitat.</emphasis>
Only known from 
<collectingCountry id="F301AE9CFFE8FFD1C80AFC29BD96CCCD" box="[609,750,988,1011]" name="South Africa" pageId="5" pageNumber="208">South Africa</collectingCountry>
. The 
<typeStatus id="54AD50AEFFE8FFD1C942FC29BC2FCCCD" box="[809,855,988,1011]" pageId="5" pageNumber="208">type</typeStatus>
locality is situated in the Ysternek Nature Reserve (currently part of the Knysna Section of Garden Route National Park) at an elevation of about 
<quantity id="4CEE43E9FFE8FFD1CBC9FBE9BE9CCB0D" box="[418,484,1052,1075]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="5" pageNumber="208" unit="m" value="650.0">650 m</quantity>
a.s.l. It is an Afromontane indigenous forest with abundant fern trees. All the other collecting sites are within a 
<quantity id="4CEE43E9FFE8FFD1C8C8FBC9BDA2CB6D" box="[675,730,1084,1107]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="208" unit="km" value="6.0">6 km</quantity>
sided triangle, in a rather similar Afromontane forest (with less frequent fern trees); the FMNH localities, in the sequence listed, are roughly from wetter to drier sites with elevations 
<quantity id="4CEE43E9FFE8FFD1C8F0FB89BC6BCBAD" box="[667,787,1148,1171]" metricMagnitude="2" metricUnit="m" metricValue="5.35" metricValueMax="6.5" metricValueMin="4.2" pageId="5" pageNumber="208" unit="m" value="535.0" valueMax="650.0" valueMin="420.0">420–650 m</quantity>
a.s.l. The highest abundance of 
<taxonomicName id="4C16958FFFE8FFD1CAE6FB69BE6ACB8D" authority="Janák &amp; Makranczy, 2016" authorityName="Janák &amp; Makranczy" authorityYear="2016" box="[141,274,1180,1203]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="208" phylum="Arthropoda" rank="species" species="crypticola" status="sp. nov.">
<emphasis id="B962321EFFE8FFD1CAE6FB69BE6ACB8D" box="[141,274,1180,1203]" italics="true" pageId="5" pageNumber="208">T. crypticola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2518F65FFE8FFD1CB7CFB69BE11CB8D" box="[279,361,1180,1203]" pageId="5" pageNumber="208" rank="species">sp. nov.</taxonomicNameLabel>
in sifted material was about 
<specimenCount id="9D102585FFE8FFD1C8FFFB69BC49CB8D" box="[660,817,1180,1203]" pageId="5" pageNumber="208" type="generic">two specimens</specimenCount>
per kg in Dal van Varings Trail (2014, Janák lgt.) and Kom Se Pad, 
<quantity id="4CEE43E9FFE8FFD1C845FB49BD00CBED" box="[558,632,1212,1235]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="5" pageNumber="208" unit="km" value="1.7">1.7 km</quantity>
W Hwy R339 (2014, Janák lgt.). As far as habitats are concerned, 
<taxonomicName id="4C16958FFFE8FFD1CB00FB29BEA8CBCD" authorityName="Lobl &amp; Rychlik" authorityYear="1994" box="[363,464,1244,1267]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="208" phylum="Arthropoda" rank="species" species="korbeli">
<emphasis id="B962321EFFE8FFD1CB00FB29BEA8CBCD" box="[363,464,1244,1267]" italics="true" pageId="5" pageNumber="208">T. korbeli</emphasis>
</taxonomicName>
is more similar to the new species described here, as it was collected 
<quantity id="4CEE43E9FFE8FFD1CAB3FB09BE3BCA2D" box="[216,323,1276,1299]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="5" pageNumber="208" unit="cm" value="25.0" valueMax="40.0" valueMin="10.0">10–40 cm</quantity>
deep in soil in riverine forests.
</paragraph>
<paragraph id="8BA9EE0CFFE8FFD3CA1AFAE9BCD9CE0D" blockId="5.[113,1091,220,1587]" lastBlockId="7.[113,1089,220,499]" lastPageId="7" lastPageNumber="210" pageId="5" pageNumber="208">
<emphasis id="B962321EFFE8FFD1CA1AFAE9BFC6CA0D" bold="true" box="[113,190,1308,1331]" pageId="5" pageNumber="208">Larva.</emphasis>
In one of the three FMNH samples (collected in 2004) one oxyteline larva was found that almost certainly is conspecific with 
<taxonomicName id="4C16958FFFE8FFD1C843FAC9BDD7CA6D" authority="Janák &amp; Makranczy, 2016" authorityName="Janák &amp; Makranczy" authorityYear="2016" box="[552,687,1340,1363]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="208" phylum="Arthropoda" rank="species" species="crypticola" status="sp. nov.">
<emphasis id="B962321EFFE8FFD1C843FAC9BDD7CA6D" box="[552,687,1340,1363]" italics="true" pageId="5" pageNumber="208">T. crypticola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2518F65FFE8FFD1C8DCFAC9BC74CA6D" box="[695,780,1340,1363]" pageId="5" pageNumber="208" rank="species">sp. nov.</taxonomicNameLabel>
This larva was identified by A. Newton as 
<taxonomicName id="4C16958FFFE8FFD1CB78FAA9BEF0CA4D" authorityName="Janák &amp; Makranczy" authorityYear="2016" box="[275,392,1372,1395]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="208" phylum="Arthropoda" rank="subFamily" subFamily="Oxytelinae">Oxytelinae</taxonomicName>
based on agreement with published characterizations of larvae of this subfamily by 
<bibRefCitation id="EF8793FDFFE8FFD1CB33FA89BE8FCAAD" author="KASULE F. K." box="[344,503,1404,1427]" pageId="5" pageNumber="208" pagination="115 - 138" refId="ref3644" refString="KASULE F. K. 1968: The larval characters of some subfamilies of British Staphylinidae (Coleoptera) with keys to the known genera. Transactions of the Royal Entomological Society of London 120: 115 - 138." type="journal article" year="1968">KASULE (1968)</bibRefCitation>
and 
<bibRefCitation id="EF8793FDFFE8FFD1C85BFA89BDAFCAAD" author="NEWTON A. F. JR." box="[560,727,1404,1427]" pageId="5" pageNumber="208" pagination="1 - 24" refId="ref3920" refString="NEWTON A. F. JR. 1982: A new genus and species of Oxytelinae from Australia, with a description of its larva, systematic position, and phylogenetic relationships (Coleoptera, Staphylinidae). American Museum Novitates 2744: 1 - 24." type="journal article" year="1982">NEWTON (1982)</bibRefCitation>
and general knowledge of larval characteristics of this group, including the presence of two subapical teeth on each mandible giving a ‘trifid’ appearance, a penultimate maxillary palp segment that is shorter than the preceding and ultimate segments, a long looped gut with visible but unidentifiable organic matter, and short pigmented urogomphi that appear one-segmented. It was further attributed to this 
<taxonomicName id="4C16958FFFE8FFD1CADDF9E9BE58C90D" authorityName="Kiesenwetter" authorityYear="1844" box="[182,288,1564,1587]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="208" phylum="Arthropoda" rank="genus">
<emphasis id="B962321EFFE8FFD1CADDF9E9BE58C90D" box="[182,288,1564,1587]" italics="true" pageId="5" pageNumber="208">Thinobius</emphasis>
</taxonomicName>
by association with adults in the same microhabitat, the absence of stemmata (unusual in oxyteline larvae, and in agreement with the lack of eyes in adults), and agreement with the only described larva of 
<taxonomicName id="4C16958FFFEAFFD3CBA6FF09BD40CE2D" authorityName="Kiesenwetter" authorityYear="1844" box="[461,568,252,275]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="genus">
<emphasis id="B962321EFFEAFFD3CBA6FF09BD40CE2D" box="[461,568,252,275]" italics="true" pageId="7" pageNumber="210">Thinobius</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C16958FFFEAFFD3C82DFF09BC42CE2D" authority="Hatch, 1957" authorityName="Hatch" authorityYear="1957" box="[582,826,252,275]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="species" species="frizzelli">
<emphasis id="B962321EFFEAFFD3C82DFF09BDC8CE2D" box="[582,688,252,275]" italics="true" pageId="7" pageNumber="210">T. frizzelli</emphasis>
Hatch, 1957
</taxonomicName>
(a partial description by 
<bibRefCitation id="EF8793FDFFEAFFD3CA1AFEE9BE7ECE0D" author="KINCAID T." box="[113,262,284,307]" pageId="7" pageNumber="210" pagination="1 - 3" refId="ref3684" refString="KINCAID T. 1961: The ecology and morphology of Thinobius frizzelli Hatch, an intertidal beetle. The Calliostoma Co., Seattle, Washington, 12 or 15 pp. + 6 pls [There are two editions: the title page, pp. 1 - 3 and plates are identical, the rest of the text differs]." type="book chapter" year="1961">KINCAID 1961</bibRefCitation>
). In addition, there were no other 
<taxonomicName id="4C16958FFFEAFFD3C804FEE9BD9CCE0D" authorityName="Janák &amp; Makranczy" authorityYear="2016" box="[623,740,284,307]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="subFamily" subFamily="Oxytelinae">Oxytelinae</taxonomicName>
in these samples.
</paragraph>
<caption id="DF69BE84FFEBFFD2CAFCFA1FBC61C90D" ID-DOI="http://doi.org/10.5281/zenodo.5305430" ID-Zenodo-Dep="5305430" httpUri="https://zenodo.org/record/5305430/files/figure.png" pageId="6" pageNumber="209" startId="6.[151,188,1514,1533]" targetBox="[152,1099,222,1451]" targetPageId="6">
<paragraph id="8BA9EE0CFFEBFFD2CAFCFA1FBC61C90D" blockId="6.[151,1128,1514,1587]" pageId="6" pageNumber="209">
Figs 5–13. 
<taxonomicName id="4C16958FFFEBFFD2CA91FA1FBED5CAC3" authority="Janák &amp; Makranczy, 2016" authorityName="Janák &amp; Makranczy" authorityYear="2016" box="[250,429,1514,1533]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="209" phylum="Arthropoda" rank="species" species="crypticola" status="sp. nov.">
<emphasis id="B962321EFFEBFFD2CA91FA1FBED5CAC3" box="[250,429,1514,1533]" italics="true" pageId="6" pageNumber="209">Thinobius crypticola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2518F65FFEBFFD2CBDFFA1FBE81CAC3" box="[436,505,1514,1533]" pageId="6" pageNumber="209" rank="species">sp. nov.</taxonomicNameLabel>
5 – antenna; 6 – maxillary palp; 7 – scutellum; 8 – male sternite VIII; 9 – male tergite IX; 10 – male sternite IX; 11 – aedeagus, frontal; 12 – female genital appendage; 13 – spermatheca. Scale bars: 0.075 mm for 11, 13; 0.09 mm for 5–7, 9–10, 12; 0.1 mm for 8.
</paragraph>
</caption>
<paragraph id="8BA9EE0CFFEAFFD3CAFBFEC9BD3FCECD" blockId="7.[113,1089,220,499]" pageId="7" pageNumber="210">
The total length (including urogomphi) is 
<quantity id="4CEE43E9FFEAFFD3C83FFEC9BDCFCE6D" box="[596,695,316,339]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="7" pageNumber="210" unit="mm" value="1.25">1.25 mm</quantity>
. The rather slender shaped specimen has the head and urogomphi more sclerotized than the rest of the body, the thoracic segments and the middle part appear soft and at least partly dissolved. A possible explanation is that the cuticle of this delicate specimen does not withstand extended storage in ethanol. The same observation was made by the second author with a larva of 
<taxonomicName id="4C16958FFFEAFFD3C8A5FE49BC48CEED" authorityName="Lobl &amp; Rychlik" authorityYear="1994" box="[718,816,444,467]" class="Insecta" family="Staphylinidae" genus="Thinobius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="210" phylum="Arthropoda" rank="species" species="korbeli">
<emphasis id="B962321EFFEAFFD3C8A5FE49BC48CEED" box="[718,816,444,467]" italics="true" pageId="7" pageNumber="210">T. korbeli</emphasis>
</taxonomicName>
. This state of preservation (
<figureCitation id="132DF289FFEAFFD3CA12FE29BFC1CECD" box="[121,185,476,499]" captionStart="Figs 1–2" captionStartId="2.[841,879,1480,1499]" captionTargetBox="[151,830,690,1602]" captionTargetId="figure-190@2.[151,830,687,1605]" captionTargetPageId="2" captionText="Figs 1–2. Thinobius crypticola sp. nov.: 1 – male holotype, dorsal habitus; 2 – suspected larva (maximum width of head capsule 0.16 mm)." figureDoi="http://doi.org/10.5281/zenodo.5305426" httpUri="https://zenodo.org/record/5305426/files/figure.png" pageId="7" pageNumber="210">Fig. 2</figureCitation>
) does not allow a proper description.
</paragraph>
</subSubSection>
</treatment>
</document>