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<document id="2C5380DAEAD247DD3880D5777736C89F" ID-CLB-Dataset="3777" ID-DOI="10.11646/zootaxa.5060.2.1" ID-GBIF-Dataset="ed98a754-c3ac-49ff-9973-b8d9839993ff" ID-ISSN="1175-5326" ID-Zenodo-Dep="5627213" ID-ZooBank="F6BFFABC-7F88-473C-9517-D031485140DD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1635492351830" checkinUser="plazi" docAuthor="Moravec, Jiří &amp; Trýzna, Miloš" docDate="2021" docId="03B477221138FFE3DCF8DB452E7EFF7F" docLanguage="en" docName="zootaxa.5060.2.1.pdf" docOrigin="Zootaxa 5060 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Physodeutera (Toxoma) conturbata Moravec 2002" docType="treatment" docVersion="9" lastPageNumber="160" masterDocId="FF8D0F5A113CFFEADC6FD91A2A22FFAA" masterDocTitle="New or rare Madagascar tiger beetles- 23. Physodeutera (Toxoma) lokobensis sp nov., a new species close to Ph. (T.) conturbata Moravec, Ph. (T.) sulcoprothoracica (W. Horn) and Ph. (T.) dubia (Mařan), with revised type designation of the latter (Coleoptera: Cicindelidae)" masterLastPageNumber="182" masterPageNumber="151" pageNumber="155" updateTime="1699467157693" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="413FE3DEBCA8B6EEA468928154BECCDD" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="C8025FDF48EC145EB535DC18A5FC0AFF">New or rare Madagascar tiger beetles- 23. Physodeutera (Toxoma) lokobensis sp nov., a new species close to Ph. (T.) conturbata Moravec, Ph. (T.) sulcoprothoracica (W. Horn) and Ph. (T.) dubia (Mařan), with revised type designation of the latter (Coleoptera: Cicindelidae)</mods:title>
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<mods:name id="AF8695F9B1CF9A8E7A60F8BA70D828CF" type="personal">
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<mods:namePart id="58A7CE7049B49C273AD14467966E0021">Moravec, Jiří</mods:namePart>
</mods:name>
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<mods:namePart id="DDF12AC80253EEF3F7DF3F6AAC27DC8B">Trýzna, Miloš</mods:namePart>
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<mods:typeOfResource id="BF302C1B3B511EB9086A6C87150C6B32">text</mods:typeOfResource>
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<mods:title id="F269B5FE63AB5AD97144C43DC072430E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="836958D213D86EA18DD6DF98DD954ACE">
<mods:date id="B0FA16558682CDD3EE70B2BA7AB45C8D">2021</mods:date>
<mods:detail id="CC24F8185BE61F0344EF9D3F51BE2F22" type="pubDate">
<mods:number id="EF5472CB6B6268FE90730BA4F318C82E">2021-10-29</mods:number>
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<mods:number id="D8956232D76690D578FAC6B474A92166">5060</mods:number>
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<mods:start id="CB6C184D050A15F3B10DC051CD070945">151</mods:start>
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<mods:classification id="2B4B434F9B3FE251DB8A525B8CC80F1F">journal article</mods:classification>
<mods:identifier id="45C412849A62C207FFC30B5A3FD01C0B" type="CLB-Dataset">3777</mods:identifier>
<mods:identifier id="5E61976BA4D3F0E30611A24CFA5BE31C" type="DOI">10.11646/zootaxa.5060.2.1</mods:identifier>
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<mods:identifier id="251AC4C8EC67DC4836CE443E3DE7E15D" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03B477221138FFE3DCF8DB452E7EFF7F" ID-DOI="http://doi.org/10.5281/zenodo.5633545" ID-GBIF-Taxon="189613067" ID-Zenodo-Dep="5633545" LSID="urn:lsid:plazi:treatment:03B477221138FFE3DCF8DB452E7EFF7F" httpUri="http://treatment.plazi.org/id/03B477221138FFE3DCF8DB452E7EFF7F" lastPageId="9" lastPageNumber="160" pageId="4" pageNumber="155">
<subSubSection id="C30795BF1138FFEEDCF8DB452920FDD0" box="[151,770,607,634]" pageId="4" pageNumber="155" type="nomenclature">
<paragraph id="8BA2C6341138FFEEDCF8DB452920FDD0" blockId="4.[151,770,607,671]" box="[151,770,607,634]" pageId="4" pageNumber="155">
<heading id="D0EA71581138FFEEDCF8DB452920FDD0" bold="true" box="[151,770,607,634]" fontSize="11" level="1" pageId="4" pageNumber="155" reason="1">
<taxonomicName id="4C1DBDB71138FFEEDCF8DB452920FDD0" ID-CoL="6VHNB" authority="Moravec, 2002" authorityName="Moravec" authorityYear="2002" box="[151,770,607,634]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="conturbata" subGenus="Toxoma">
<emphasis id="B9691A261138FFEEDCF8DB452920FDD0" bold="true" box="[151,770,607,634]" pageId="4" pageNumber="155">
<emphasis id="B9691A261138FFEEDCF8DB452B1EFDD0" bold="true" box="[151,316,607,634]" italics="true" pageId="4" pageNumber="155">Physodeutera</emphasis>
(
<emphasis id="B9691A261138FFEEDD22DB7A2B8CFDD0" bold="true" box="[333,430,608,634]" italics="true" pageId="4" pageNumber="155">Toxoma</emphasis>
)
<emphasis id="B9691A261138FFEEDDD0DB452861FDD0" bold="true" box="[447,579,607,634]" italics="true" pageId="4" pageNumber="155">conturbata</emphasis>
Moravec, 2002
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30795BF1138FFEEDCF8DB9E2B73FD35" box="[151,337,644,671]" pageId="4" pageNumber="155" type="description">
<paragraph id="8BA2C6341138FFEEDCF8DB9E2B73FD35" blockId="4.[151,770,607,671]" box="[151,337,644,671]" pageId="4" pageNumber="155">
(
<figureCitation id="1326DAB11138FFEEDCF0DB9E2B30FD34" box="[159,274,644,670]" captionStart-0="FIGURES 1–9" captionStart-1="FIGURES 10–18" captionStart-2="FIGURES 19–23" captionStartId-0="5.[151,264,1937,1961]" captionStartId-1="6.[151,264,1867,1891]" captionStartId-2="7.[151,264,1614,1638]" captionTargetBox-0="[177,1410,288,1906]" captionTargetBox-1="[159,1430,191,1829]" captionTargetBox-2="[166,1425,195,1578]" captionTargetId-0="figure-61@5.[160,1427,273,1914]" captionTargetId-1="figure-31@6.[151,1435,181,1844]" captionTargetId-2="figure-17@7.[151,1435,181,1589]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIGURES 1–9. Physodeutera (Toxoma) conturbata Moravec, from type locality, all in SDEI. 1—habitus ♂, 9.2 mm, HT; 2– 3—Antennomeres 1–5: 2—♂, HT; 3—♀, PT; 4–5—♀, maxillary palpus: 4—dorsal view; 5—ventral view; 6–7—labial palpus: 6—female, PT; 7—male, HT (adopted from Moravec 2002a); 8–9— male aedeagus showing internal sac, HT. Bars = 1 mm." captionText-1="FIGURES 10–18. Physodeutera (Toxoma) conturbata Moravec, from type locality, all in SDEI. 10–12—head: 10—♂, HT; 11–12—♀, PT; 13–15—labrum: 13—♂, HT; 14–15—♀, PT; 16–18—pronotum: 16—♂, HT; 17–18—♀, PT. Bars = 1 mm." captionText-2="FIGURES 19–23. Physodeutera (Toxoma) conturbata Moravec, from type locality, all in SDEI. 19–22—elytron: 19—♂, HT; 20—ditto, detail of elytral disc; 21—♀, PT; 22— ditto, detail of posterior area of elytral disc; 23—labels of the holotype. Bars = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5627215" figureDoi-1="http://doi.org/10.5281/zenodo.5627217" figureDoi-2="http://doi.org/10.5281/zenodo.5627219" httpUri-0="https://zenodo.org/record/5627215/files/figure.png" httpUri-1="https://zenodo.org/record/5627217/files/figure.png" httpUri-2="https://zenodo.org/record/5627219/files/figure.png" pageId="4" pageNumber="155">Figs 1–23</figureCitation>
,
<figureCitation id="1326DAB11138FFEEDD71DB9E2B6BFD35" box="[286,329,644,671]" captionStart="FIGURE 106" captionStartId="27.[151,250,1561,1585]" captionTargetBox="[271,1319,572,1531]" captionTargetId="figure-167@27.[262,1326,561,1538]" captionTargetPageId="27" captionText="FIGURE 106. Distribution of four Physodeutera (Toxoma) species in northern part of Madagascar. Adopted from the map of preserved areas in Madagascar according to Moat &amp; Smith (2007), adapted and published previously (Moravec 2010) under permission of the trustee of the Royal Botanic Gardens, Kew, GB." figureDoi="http://doi.org/10.5281/zenodo.5627247" httpUri="https://zenodo.org/record/5627247/files/figure.png" pageId="4" pageNumber="155">106</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30795BF1138FFEEDCF8DBD229F8FD4A" box="[151,986,712,736]" pageId="4" pageNumber="155" type="reference_group">
<paragraph id="8BA2C6341138FFEEDCF8DBD229F8FD4A" blockId="4.[151,986,712,736]" box="[151,986,712,736]" pageId="4" pageNumber="155">
<treatmentCitationGroup id="AB0DE11A1138FFEEDCF8DBD229F8FD4A" box="[151,986,712,736]" pageId="4" pageNumber="155">
<treatmentCitation id="0ABCE0251138FFEEDCF8DBD228FCFD4A" author="Moravec, J." box="[151,734,712,736]" page="118" pageId="4" pageNumber="155" year="2002">
<taxonomicName id="4C1DBDB71138FFEEDCF8DBD228FCFD4A" ID-CoL="6VHNB" authority="Moravec, 2002 a: 118" authorityName="Moravec" authorityPageNumber="118" authorityYear="2002" box="[151,734,712,736]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="conturbata" subGenus="Toxoma">
<emphasis id="B9691A261138FFEEDCF8DBD22B00FD4A" box="[151,290,712,736]" italics="true" pageId="4" pageNumber="155">Physodeutera</emphasis>
(
<emphasis id="B9691A261138FFEEDD5EDBD32BA2FD4A" box="[305,384,713,736]" italics="true" pageId="4" pageNumber="155">Toxoma</emphasis>
)
<emphasis id="B9691A261138FFEEDDE0DBD22BDCFD4A" box="[399,510,712,736]" italics="true" pageId="4" pageNumber="155">conturbata</emphasis>
<bibRefCitation id="EF8CBBC51138FFEEDE6ADBD328FCFD4A" author="Moravec, J." box="[517,734,712,736]" pageId="4" pageNumber="155" refId="ref15451" refString="Moravec, J. (2002 a) Tiger beetles of Madagascar 2. A monograph of the genus Physodeutera (Coleoptera: Cicindelidae). Nakladatelstvi Kabourek, Zlin, Czech Republic, 290 pp." type="book" year="2002">Moravec, 2002a: 118</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 309–316, 724–725.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30795BF1138FFEEDCF8DA152E9DFBAB" pageId="4" pageNumber="155" type="materials_examined">
<paragraph id="8BA2C6341138FFEEDCF8DA152EA6FC83" blockId="4.[151,1437,782,2033]" box="[151,1156,782,809]" pageId="4" pageNumber="155">
<materialsCitation id="3B75CC691138FFEEDCF8DA152EA2FC83" ID-GBIF-Occurrence="3395513318" box="[151,1152,782,809]" country="Madagascar" location="Antseva" municipality="Sambirano" pageId="4" pageNumber="155" specimenCount="1" typeStatus="holotype">
<emphasis id="B9691A261138FFEEDCF8DA152B12FC82" bold="true" box="[151,304,782,809]" pageId="4" pageNumber="155">
<typeStatus id="54A678961138FFEEDCF8DA152AF0FC83" box="[151,210,783,809]" pageId="4" pageNumber="155">Type</typeStatus>
locality
</emphasis>
.
<location id="8EC290EF1138FFEEDD53DA152BB9FC83" LSID="urn:lsid:plazi:treatment:03B477221138FFE3DCF8DB452E7EFF7F:8EC290EF1138FFEEDD53DA152BB9FC83" box="[316,411,783,809]" country="Madagascar" municipality="Sambirano" name="Antseva" pageId="4" pageNumber="155">Antseva</location>
, northwestern
<collectingCountry id="F30A86A41138FFEEDE50DA1428EAFC82" box="[575,712,782,808]" name="Madagascar" pageId="4" pageNumber="155">Madagascar</collectingCountry>
,
<collectingMunicipality id="6BC65C4E1138FFEEDEBCDA14296EFC82" box="[723,844,782,808]" pageId="4" pageNumber="155">Sambirano</collectingMunicipality>
(see “Distribution” below)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA2C6341138FFEEDCA8DA292E9DFBAB" blockId="4.[151,1437,782,2033]" pageId="4" pageNumber="155">
<emphasis id="B9691A261138FFEEDCA8DA292B4FFCE6" bold="true" box="[199,365,818,845]" pageId="4" pageNumber="155">Type material</emphasis>
.
<materialsCitation id="3B75CC691138FFEEDD15DA282B55FC3F" ID-GBIF-Occurrence="3395513313" collectionCode="SDEI" collectorName="J. Mellis &amp; Coll. W. Horn" determinerName="Jiri" pageId="4" pageNumber="155" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54A678961138FFEEDD15DA282BC0FCE6" box="[378,482,818,844]" pageId="4" pageNumber="155" type="holotype">Holotype</typeStatus>
<specimenCount id="9D1B0DBD1138FFEEDD86DA292BDCFCE7" box="[489,510,819,845]" pageId="4" pageNumber="155" type="male">♂</specimenCount>
in
<collectionCode id="ED0C5EF11138FFEEDE4FDA292843FCE7" box="[544,609,819,845]" pageId="4" pageNumber="155">SDEI</collectionCode>
, labelled: “Antseva / XI. 33” // “NW-Madag. /
<collectorName id="26E8A3E21138FFEED806DA292EE9FCE6" box="[1129,1227,818,845]" pageId="4" pageNumber="155">J. Mellis</collectorName>
” [printed] // “
<collectorName id="26E8A3E21138FFEED90CDA282ADAFCDB" pageId="4" pageNumber="155">Coll. W. Horn</collectorName>
/ DEI Eberswalde” [printed // “
<typeStatus id="54A678961138FFEEDE32DA4C28EAFCDA" box="[605,712,854,880]" pageId="4" pageNumber="155" type="holotype">Holotype</typeStatus>
/
<taxonomicName id="4C1DBDB71138FFEEDEB0DA4C29CEFCDB" authority="" authorityName="Rivalier" authorityYear="1967" box="[735,1004,854,881]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="subGenus" subGenus="Toxoma">Physodeutera (Toxoma)</taxonomicName>
/ conturbata
<taxonomicNameLabel id="A25AA75D1138FFEED8EEDA4D2E9CFCDB" box="[1153,1214,855,881]" pageId="4" pageNumber="155" rank="species">sp. n.</taxonomicNameLabel>
/ det.
<determinerName id="7E7E2AF01138FFEED969DA4C2F08FCDA" box="[1286,1322,854,880]" pageId="4" pageNumber="155">Jiří</determinerName>
<bibRefCitation id="EF8CBBC51138FFEED95DDA4D2AF0FC3F" author="Moravec, J." pageId="4" pageNumber="155" pagination="197 - 216" refId="ref15414" refString="Moravec, J. (2000) New or rare Madagascar tiger beetles - 6. New taxa in the genus Physodeutera (Coleoptera: Cicindelidae). Folia Heyrovskyana, 8, 197 - 216." type="journal article" year="2000">Moravec, 2000</bibRefCitation>
” [red, printed]
</materialsCitation>
.
<materialsCitation id="3B75CC691138FFEEDDEEDA612E1BFC13" ID-GBIF-Occurrence="3395513305" collectionCode="SDEI" collectorName="W. Horn" country="Madagascar" location="Madagaskar" pageId="4" pageNumber="155" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<typeStatus id="54A678961138FFEEDDEEDA612BD0FC3F" box="[385,498,891,917]" pageId="4" pageNumber="155" type="paratype">Paratypes</typeStatus>
.
<specimenCount id="9D1B0DBD1138FFEEDD95DA61281AFC3F" box="[506,568,891,917]" pageId="4" pageNumber="155" type="female">2 ♀♀</specimenCount>
in
<collectionCode id="ED0C5EF11138FFEEDE35DA6128BFFC3F" box="[602,669,891,917]" pageId="4" pageNumber="155">SDEI</collectionCode>
: “Antseva-Ammbobako / N.W.
<collectingCountry id="F30A86A41138FFEEDF93DA602EAAFC3E" box="[1020,1160,890,916]" name="Madagascar" pageId="4" pageNumber="155">Madagaskar</collectingCountry>
” [printed] // “
<collectorName id="26E8A3E21138FFEED94FDA612B0DFC12" pageId="4" pageNumber="155">R. Ramena N.W. Madag.</collectorName>
/
<collectorName id="26E8A3E21138FFEEDD28DA852B89FC12" box="[327,427,926,953]" pageId="4" pageNumber="155">J. Mellis</collectorName>
” [printed] // “
<collectorName id="26E8A3E21138FFEEDE25DA8428CFFC13" box="[586,749,926,953]" pageId="4" pageNumber="155">Coll. W. Horn</collectorName>
/ DEI Eberswalde” [printed]
</materialsCitation>
.
<materialsCitation id="3B75CC691138FFEED82ADA852E99FBAB" ID-GBIF-Occurrence="3395513315" collectionCode="SDEI" country="Madagascar" determinerName="Jiri" pageId="4" pageNumber="155" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D1B0DBD1138FFEED82ADA852E52FC13" box="[1093,1136,927,953]" pageId="4" pageNumber="155" type="female">1 ♀</specimenCount>
in
<collectionCode id="ED0C5EF11138FFEED8F8DA852EF7FC13" box="[1175,1237,927,953]" pageId="4" pageNumber="155">SDEI</collectionCode>
with partly illegible locality label: “Antsabetsienne / XI. 33” [handwritten]; other labels identical. All
<typeStatus id="54A678961138FFEED836DAD92E9BFC77" box="[1113,1209,963,989]" pageId="4" pageNumber="155" type="paratype">paratype</typeStatus>
specimens labelled: “
<typeStatus id="54A678961138FFEEDCCEDAFD2B24FBAB" box="[161,262,999,1025]" pageId="4" pageNumber="155" type="paratype">Paratype</typeStatus>
/
<taxonomicName id="4C1DBDB71138FFEEDD74DAFC2805FBAB" authority="" authorityName="Rivalier" authorityYear="1967" box="[283,551,998,1025]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="subGenus" subGenus="Toxoma">Physodeutera (Toxoma)</taxonomicName>
/ conturbata
<taxonomicNameLabel id="A25AA75D1138FFEEDED5DAFD28D4FBAB" box="[698,758,999,1025]" pageId="4" pageNumber="155" rank="species">sp. n.</taxonomicNameLabel>
/ det.
<determinerName id="7E7E2AF01138FFEEDF53DAFC2942FBAA" box="[828,864,998,1024]" pageId="4" pageNumber="155">Jiří</determinerName>
<bibRefCitation id="EF8CBBC51138FFEEDF08DAFD2E31FBAB" author="Moravec, J." box="[871,1043,999,1025]" pageId="4" pageNumber="155" pagination="197 - 216" refId="ref15414" refString="Moravec, J. (2000) New or rare Madagascar tiger beetles - 6. New taxa in the genus Physodeutera (Coleoptera: Cicindelidae). Folia Heyrovskyana, 8, 197 - 216." type="journal article" year="2000">Moravec, 2000</bibRefCitation>
” [red, printed]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30795BF1138FFEEDCA8DD102B97FA52" pageId="4" pageNumber="155" type="diagnosis">
<paragraph id="8BA2C6341138FFEEDCA8DD102B30FAC3" blockId="4.[151,1437,782,2033]" pageId="4" pageNumber="155">
<emphasis id="B9691A261138FFEEDCA8DD102BE7FB8E" bold="true" box="[199,453,1034,1060]" pageId="4" pageNumber="155">Differential diagnosis</emphasis>
.
<taxonomicName id="4C1DBDB71138FFEEDDBDDD10297FFB8E" box="[466,861,1034,1061]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="conturbata" subGenus="Toxoma">
<emphasis id="B9691A261138FFEEDDBDDD10284EFB8E" box="[466,620,1034,1060]" italics="true" pageId="4" pageNumber="155">Physodeutera</emphasis>
(
<emphasis id="B9691A261138FFEEDE14DD1128F1FB8E" box="[635,723,1035,1060]" italics="true" pageId="4" pageNumber="155">Toxoma</emphasis>
)
<emphasis id="B9691A261138FFEEDE8DDD10297FFB8E" box="[738,861,1034,1060]" italics="true" pageId="4" pageNumber="155">conturbata</emphasis>
</taxonomicName>
and the three other species treated below are clearly distinguished from
<taxonomicName id="4C1DBDB71138FFEEDD1BDD3429B9FBE3" authority="(W. Horn, 1896)" baseAuthorityName="W. Horn" baseAuthorityYear="1896" box="[372,923,1070,1097]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="sikorai" subGenus="Toxoma">
<emphasis id="B9691A261138FFEEDD1BDD34282CFBE2" box="[372,526,1070,1096]" italics="true" pageId="4" pageNumber="155">Physodeutera</emphasis>
(
<emphasis id="B9691A261138FFEEDE4FDD35285AFBE2" box="[544,632,1071,1096]" italics="true" pageId="4" pageNumber="155">Toxoma</emphasis>
)
<emphasis id="B9691A261138FFEEDEE6DD3428F5FBE2" box="[649,727,1070,1096]" italics="true" pageId="4" pageNumber="155">sikorai</emphasis>
(W. Horn, 1896)
</taxonomicName>
and
<emphasis id="B9691A261138FFEEDFBBDD3429D1FBE2" box="[980,1011,1070,1096]" italics="true" pageId="4" pageNumber="155">Ph</emphasis>
. (
<emphasis id="B9691A261138FFEED863DD352E3EFBE2" box="[1036,1052,1071,1096]" italics="true" pageId="4" pageNumber="155">T</emphasis>
.)
<emphasis id="B9691A261138FFEED85BDD342E96FBE2" box="[1076,1204,1070,1096]" italics="true" pageId="4" pageNumber="155">breviformis</emphasis>
(W. Horn, 1904) by their white elytral maculation reduced to only humeral macula, somewhat thinner penultimate palpomeres of labial palpi, and shorter, much stouter aedeagus.
<emphasis id="B9691A261138FFEEDE1FDD6C28ADFB3A" box="[624,655,1142,1168]" italics="true" pageId="4" pageNumber="155">Ph</emphasis>
.
<taxonomicName id="4C1DBDB71138FFEEDEF3DD6D2965FB3A" box="[668,839,1142,1169]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="conturbata" subGenus="Toxoma">
(
<emphasis id="B9691A261138FFEEDEC9DD6D2894FB3A" box="[678,694,1143,1168]" italics="true" pageId="4" pageNumber="155">T</emphasis>
.)
<emphasis id="B9691A261138FFEEDEA3DD6C2965FB3A" box="[716,839,1142,1168]" italics="true" pageId="4" pageNumber="155">conturbata</emphasis>
</taxonomicName>
immediately differs from
<taxonomicName id="4C1DBDB71138FFEED804DD6C2F1FFB3A" authority="Moravec, 2021" authorityName="Moravec" authorityYear="2021" box="[1131,1341,1142,1169]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="164" phylum="Arthropoda" rank="species" species="lokobensis" status="sp. nov." subGenus="Toxoma">
<emphasis id="B9691A261138FFEED804DD6C2EA8FB3A" box="[1131,1162,1142,1168]" italics="true" pageId="4" pageNumber="155">Ph</emphasis>
. (
<emphasis id="B9691A261138FFEED8CEDD6D2E93FB3A" box="[1185,1201,1143,1168]" italics="true" pageId="4" pageNumber="155">T</emphasis>
.)
<emphasis id="B9691A261138FFEED8A8DD6C2F1FFB3A" box="[1223,1341,1142,1168]" italics="true" pageId="4" pageNumber="155">lokobensis</emphasis>
</taxonomicName>
<emphasis id="B9691A261138FFEED92CDD6D2FBEFB3B" bold="true" box="[1347,1436,1143,1169]" pageId="4" pageNumber="155">
<taxonomicNameLabel id="A25AA75D1138FFEED92CDD6D2FBEFB3B" box="[1347,1436,1143,1169]" pageId="4" pageNumber="155" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C1DBDB71138FFEEDCA8DD802B47FB1E" baseAuthorityName="Maran" baseAuthorityYear="1942" box="[199,357,1178,1205]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="164" phylum="Arthropoda" rank="species" species="dubia" subGenus="Toxoma">
<emphasis id="B9691A261138FFEEDCA8DD802AC4FB1E" box="[199,230,1178,1204]" italics="true" pageId="4" pageNumber="155">Ph</emphasis>
. (
<emphasis id="B9691A261138FFEEDC91DD812B2CFB1E" box="[254,270,1179,1204]" italics="true" pageId="4" pageNumber="155">T</emphasis>
.)
<emphasis id="B9691A261138FFEEDD4ADD802B47FB1E" box="[293,357,1178,1204]" italics="true" pageId="4" pageNumber="155">dubia</emphasis>
</taxonomicName>
in having much darker, prevailingly deep cyaneous-violaceous dorsal coloration particularly on elytra, with indistinct shadowy tinge on elytral disc and much finer punctures on the posterior declivity (
<figureCitation id="1326DAB11138FFEED923DDA42FB7FB73" box="[1356,1429,1214,1241]" captionStart="FIGURES 1–9" captionStartId="5.[151,264,1937,1961]" captionTargetBox="[177,1410,288,1906]" captionTargetId="figure-61@5.[160,1427,273,1914]" captionTargetPageId="5" captionText="FIGURES 1–9. Physodeutera (Toxoma) conturbata Moravec, from type locality, all in SDEI. 1—habitus ♂, 9.2 mm, HT; 2– 3—Antennomeres 1–5: 2—♂, HT; 3—♀, PT; 4–5—♀, maxillary palpus: 4—dorsal view; 5—ventral view; 6–7—labial palpus: 6—female, PT; 7—male, HT (adopted from Moravec 2002a); 8–9— male aedeagus showing internal sac, HT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627215" httpUri="https://zenodo.org/record/5627215/files/figure.png" pageId="4" pageNumber="155">Figs 1</figureCitation>
,
<figureCitation id="1326DAB11138FFEEDCF8DDF92AFCFB57" box="[151,222,1251,1277]" captionStart="FIGURES 19–23" captionStartId="7.[151,264,1614,1638]" captionTargetBox="[166,1425,195,1578]" captionTargetId="figure-17@7.[151,1435,181,1589]" captionTargetPageId="7" captionText="FIGURES 19–23. Physodeutera (Toxoma) conturbata Moravec, from type locality, all in SDEI. 19–22—elytron: 19—♂, HT; 20—ditto, detail of elytral disc; 21—♀, PT; 22— ditto, detail of posterior area of elytral disc; 23—labels of the holotype. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627219" httpUri="https://zenodo.org/record/5627219/files/figure.png" pageId="4" pageNumber="155">19–22</figureCitation>
) than in
<taxonomicName id="4C1DBDB71138FFEEDD50DDF82BF9FB56" baseAuthorityName="Maran" baseAuthorityYear="1942" box="[319,475,1250,1277]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="164" phylum="Arthropoda" rank="species" species="dubia" subGenus="Toxoma">
<emphasis id="B9691A261138FFEEDD50DDF82B7CFB56" box="[319,350,1250,1276]" italics="true" pageId="4" pageNumber="155">Ph</emphasis>
. (
<emphasis id="B9691A261138FFEEDD1ADDF92BA7FB56" box="[373,389,1251,1276]" italics="true" pageId="4" pageNumber="155">T</emphasis>
.)
<emphasis id="B9691A261138FFEEDDF4DDF82BF9FB56" box="[411,475,1250,1276]" italics="true" pageId="4" pageNumber="155">dubia</emphasis>
</taxonomicName>
, distinctly anteriad-attenuated lateral margins of its pronotal disc (
<figureCitation id="1326DAB11138FFEED8D3DDF82F19FB57" box="[1212,1339,1250,1277]" captionStart="FIGURES 10–18" captionStartId="6.[151,264,1867,1891]" captionTargetBox="[159,1430,191,1829]" captionTargetId="figure-31@6.[151,1435,181,1844]" captionTargetPageId="6" captionText="FIGURES 10–18. Physodeutera (Toxoma) conturbata Moravec, from type locality, all in SDEI. 10–12—head: 10—♂, HT; 11–12—♀, PT; 13–15—labrum: 13—♂, HT; 14–15—♀, PT; 16–18—pronotum: 16—♂, HT; 17–18—♀, PT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627217" httpUri="https://zenodo.org/record/5627217/files/figure.png" pageId="4" pageNumber="155">Figs 16–17</figureCitation>
) (except for one large aberrant female (
<figureCitation id="1326DAB11138FFEEDD9FDC1C2866FA8B" box="[496,580,1286,1313]" captionStart="FIGURES 10–18" captionStartId="6.[151,264,1867,1891]" captionTargetBox="[159,1430,191,1829]" captionTargetId="figure-31@6.[151,1435,181,1844]" captionTargetPageId="6" captionText="FIGURES 10–18. Physodeutera (Toxoma) conturbata Moravec, from type locality, all in SDEI. 10–12—head: 10—♂, HT; 11–12—♀, PT; 13–15—labrum: 13—♂, HT; 14–15—♀, PT; 16–18—pronotum: 16—♂, HT; 17–18—♀, PT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627217" httpUri="https://zenodo.org/record/5627217/files/figure.png" pageId="4" pageNumber="155">Fig. 18</figureCitation>
), entirely metallic black antennomeres
<quantity id="4CE56BD11138FFEED86DDC1D2E68FA8A" box="[1026,1098,1286,1313]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" metricValueMax="10.16" metricValueMin="5.08" pageId="4" pageNumber="155" unit="in" value="3.0" valueMax="4.0" valueMin="2.0">2–4 in</quantity>
both sexes (
<figureCitation id="1326DAB11138FFEED8B7DC1C2F1DFA8B" box="[1240,1343,1286,1313]" captionStart="FIGURES 1–9" captionStartId="5.[151,264,1937,1961]" captionTargetBox="[177,1410,288,1906]" captionTargetId="figure-61@5.[160,1427,273,1914]" captionTargetPageId="5" captionText="FIGURES 1–9. Physodeutera (Toxoma) conturbata Moravec, from type locality, all in SDEI. 1—habitus ♂, 9.2 mm, HT; 2– 3—Antennomeres 1–5: 2—♂, HT; 3—♀, PT; 4–5—♀, maxillary palpus: 4—dorsal view; 5—ventral view; 6–7—labial palpus: 6—female, PT; 7—male, HT (adopted from Moravec 2002a); 8–9— male aedeagus showing internal sac, HT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627215" httpUri="https://zenodo.org/record/5627215/files/figure.png" pageId="4" pageNumber="155">Figs 2–3</figureCitation>
), darker penultimate palpomere of maxillary palpus in female (
<figureCitation id="1326DAB11138FFEEDE80DC302913FAEF" box="[751,817,1322,1349]" captionStart="FIGURES 1–9" captionStartId="5.[151,264,1937,1961]" captionTargetBox="[177,1410,288,1906]" captionTargetId="figure-61@5.[160,1427,273,1914]" captionTargetPageId="5" captionText="FIGURES 1–9. Physodeutera (Toxoma) conturbata Moravec, from type locality, all in SDEI. 1—habitus ♂, 9.2 mm, HT; 2– 3—Antennomeres 1–5: 2—♂, HT; 3—♀, PT; 4–5—♀, maxillary palpus: 4—dorsal view; 5—ventral view; 6–7—labial palpus: 6—female, PT; 7—male, HT (adopted from Moravec 2002a); 8–9— male aedeagus showing internal sac, HT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627215" httpUri="https://zenodo.org/record/5627215/files/figure.png" pageId="4" pageNumber="155">Fig. 4</figureCitation>
) and wider aedeagus with short, regularly rounded apex (
<figureCitation id="1326DAB11138FFEEDCF0DC542B24FAC3" box="[159,262,1358,1385]" captionStart="FIGURES 1–9" captionStartId="5.[151,264,1937,1961]" captionTargetBox="[177,1410,288,1906]" captionTargetId="figure-61@5.[160,1427,273,1914]" captionTargetPageId="5" captionText="FIGURES 1–9. Physodeutera (Toxoma) conturbata Moravec, from type locality, all in SDEI. 1—habitus ♂, 9.2 mm, HT; 2– 3—Antennomeres 1–5: 2—♂, HT; 3—♀, PT; 4–5—♀, maxillary palpus: 4—dorsal view; 5—ventral view; 6–7—labial palpus: 6—female, PT; 7—male, HT (adopted from Moravec 2002a); 8–9— male aedeagus showing internal sac, HT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627215" httpUri="https://zenodo.org/record/5627215/files/figure.png" pageId="4" pageNumber="155">Figs 8–9</figureCitation>
).
</paragraph>
<paragraph id="8BA2C6341138FFEEDCA8DC682B97FA52" blockId="4.[151,1437,782,2033]" pageId="4" pageNumber="155">
<taxonomicName id="4C1DBDB71138FFEEDCA8DC68288EFA26" baseAuthorityName="W. Horn" baseAuthorityYear="1913" box="[199,684,1394,1421]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="sulcoprothoracica" subGenus="Toxoma">
<emphasis id="B9691A261138FFEEDCA8DC682B43FA26" box="[199,353,1394,1420]" italics="true" pageId="4" pageNumber="155">Physodeutera</emphasis>
(
<emphasis id="B9691A261138FFEEDD1ADC692BEFFA26" box="[373,461,1395,1420]" italics="true" pageId="4" pageNumber="155">Toxoma</emphasis>
)
<emphasis id="B9691A261138FFEEDD8FDC68288EFA26" box="[480,684,1394,1420]" italics="true" pageId="4" pageNumber="155">sulcoprothoracica</emphasis>
</taxonomicName>
immediately differs in having rather matt green-blue elytra with notably distinct, velvety-black shadowy zone on the elytral disc and almost effaced punctures within the area and on posterior declivity (
<figureCitation id="1326DAB11138FFEEDDF8DCA02BCDFA7F" box="[407,495,1466,1493]" captionStart="FIGURES 90–92" captionStartId="23.[151,264,1924,1948]" captionTargetBox="[164,1425,324,1881]" captionTargetId="figure-76@23.[151,1435,309,1901]" captionTargetPageId="23" captionText="FIGURES 90–92. Physodeutera (Toxoma) sulcoprothoracica W. Horn. ♂, “Ambre” (CCJM ex APCA). 90— habitus, 9.9 mm; 91— antennomeres 1–5; 92—antennomeres 1–6. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627241" httpUri="https://zenodo.org/record/5627241/files/figure.png" pageId="4" pageNumber="155">Figs 90</figureCitation>
,
<figureCitation id="1326DAB11138FFEEDD95DCA12843FA7F" box="[506,609,1467,1493]" captionStart="FIGURES 102–105" captionStartId="25.[151,264,1867,1891]" captionTargetBox="[155,1428,191,1827]" captionTargetId="figure-17@25.[152,1436,181,1844]" captionTargetPageId="25" captionText="FIGURES 102–105. Physodeutera (Toxoma) sulcoprothoracica W. Horn. 102–104—elytra: 102—♂, “Ambre” (CCJM ex APCA); 103—♀, LT (SDEI), adopted from SDEI database; 104—apical area, ♂, “Ambre” (CCJM ex APCA); 105—labels of HT (SDEI). Bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627245" httpUri="https://zenodo.org/record/5627245/files/figure.png" pageId="4" pageNumber="155">102–104</figureCitation>
). For other differences see under
<taxonomicName id="4C1DBDB71138FFEEDFB6DCA02E8FFA7E" authority="Moravec, 2021" authorityName="Moravec" authorityYear="2021" box="[985,1197,1466,1493]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="164" phylum="Arthropoda" rank="species" species="lokobensis" status="sp. nov." subGenus="Toxoma">
<emphasis id="B9691A261138FFEEDFB6DCA029DAFA7E" box="[985,1016,1466,1492]" italics="true" pageId="4" pageNumber="155">Ph</emphasis>
. (
<emphasis id="B9691A261138FFEED87FDCA12E02FA7E" box="[1040,1056,1467,1492]" italics="true" pageId="4" pageNumber="155">T</emphasis>
.)
<emphasis id="B9691A261138FFEED858DCA02E8FFA7E" box="[1079,1197,1466,1492]" italics="true" pageId="4" pageNumber="155">lokobensis</emphasis>
</taxonomicName>
<emphasis id="B9691A261138FFEED8DBDCA12F2CFA7F" bold="true" box="[1204,1294,1467,1493]" pageId="4" pageNumber="155">
<taxonomicNameLabel id="A25AA75D1138FFEED8DBDCA12F2CFA7F" box="[1204,1294,1467,1493]" pageId="4" pageNumber="155" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<emphasis id="B9691A261138FFEED929DCA02F47FA7E" box="[1350,1381,1466,1492]" italics="true" pageId="4" pageNumber="155">Ph</emphasis>
.
<taxonomicName id="4C1DBDB71138FFEED91CDCA12B41FA52" baseAuthorityName="W. Horn" baseAuthorityYear="1913" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="sulcoprothoracica" subGenus="Toxoma">
(
<emphasis id="B9691A261138FFEED913DCA12FAEFA7E" box="[1404,1420,1467,1492]" italics="true" pageId="4" pageNumber="155">T</emphasis>
.)
<emphasis id="B9691A261138FFEEDCF8DCC42B41FA52" box="[151,355,1502,1528]" italics="true" pageId="4" pageNumber="155">sulcoprothoracica</emphasis>
</taxonomicName>
below.
</paragraph>
</subSubSection>
<subSubSection id="C30795BF1138FFE2DCA8DF182850FA4F" lastPageId="8" lastPageNumber="159" pageId="4" pageNumber="155" type="description">
<paragraph id="8BA2C6341138FFEEDCA8DF182856F9EA" blockId="4.[151,1437,782,2033]" pageId="4" pageNumber="155">
<emphasis id="B9691A261138FFEEDCA8DF182B4FF9B6" bold="true" box="[199,365,1538,1564]" pageId="4" pageNumber="155">Redescription</emphasis>
. Body (
<figureCitation id="1326DAB11138FFEEDDA6DF182828F9B7" box="[457,522,1538,1565]" captionStart="FIGURES 1–9" captionStartId="5.[151,264,1937,1961]" captionTargetBox="[177,1410,288,1906]" captionTargetId="figure-61@5.[160,1427,273,1914]" captionTargetPageId="5" captionText="FIGURES 1–9. Physodeutera (Toxoma) conturbata Moravec, from type locality, all in SDEI. 1—habitus ♂, 9.2 mm, HT; 2– 3—Antennomeres 1–5: 2—♂, HT; 3—♀, PT; 4–5—♀, maxillary palpus: 4—dorsal view; 5—ventral view; 6–7—labial palpus: 6—female, PT; 7—male, HT (adopted from Moravec 2002a); 8–9— male aedeagus showing internal sac, HT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627215" httpUri="https://zenodo.org/record/5627215/files/figure.png" pageId="4" pageNumber="155">Fig. 1</figureCitation>
) medium-sized to large, length 9.25–10.35 (HT 9.25) mm long, width 2.90–3.20 (HT 2.90) mm, deep cyaneous-violaceous.
</paragraph>
<paragraph id="8BA2C6341138FFEEDCA8DF502EE1F9CF" blockId="4.[151,1437,782,2033]" box="[199,1219,1610,1637]" pageId="4" pageNumber="155">
Head (
<figureCitation id="1326DAB11138FFEEDD7EDF502BB3F9CF" box="[273,401,1610,1637]" captionStart="FIGURES 10–18" captionStartId="6.[151,264,1867,1891]" captionTargetBox="[159,1430,191,1829]" captionTargetId="figure-31@6.[151,1435,181,1844]" captionTargetPageId="6" captionText="FIGURES 10–18. Physodeutera (Toxoma) conturbata Moravec, from type locality, all in SDEI. 10–12—head: 10—♂, HT; 11–12—♀, PT; 13–15—labrum: 13—♂, HT; 14–15—♀, PT; 16–18—pronotum: 16—♂, HT; 17–18—♀, PT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627217" httpUri="https://zenodo.org/record/5627217/files/figure.png" pageId="4" pageNumber="155">Figs 10–12</figureCitation>
) with large eyes, only slightly narrower than body, width
<quantity id="4CE56BD11138FFEED876DF502E9DF9CF" box="[1049,1215,1610,1637]" metricMagnitude="-3" metricUnit="m" metricValue="2.875" metricValueMax="3.1" metricValueMin="2.65" pageId="4" pageNumber="155" unit="mm" value="2.875" valueMax="3.1" valueMin="2.65">2.65–3.10 mm</quantity>
.
</paragraph>
<paragraph id="8BA2C6341138FFEEDCA8DF752E37F906" blockId="4.[151,1437,782,2033]" pageId="4" pageNumber="155">Frons metallic deep green-blue, convex in middle, finely longitudinally striate-wrinkled, wrinkles passing through frons-vertex blunt fold onto vertex; supraantennal plates flat, indistinct.</paragraph>
<paragraph id="8BA2C6341138FFEEDCA8DFAD2E28F896" blockId="4.[151,1437,782,2033]" pageId="4" pageNumber="155">Vertex black-cyaneous with deep, transverse anterior impression reaching eyes, surface coarsely and irregularly (mostly longitudinally) striate-rugulose, striae wavy, passing to irregularly vermicular rugae on posterior and occipital areas; large juxtaorbital areas with denser parallel striae deformed by anterior and two posterior impressions (on either side); sublateral parallel rugae divergent when passing onto temples.</paragraph>
<paragraph id="8BA2C6341138FFEEDCA8DE5D2962F8CA" blockId="4.[151,1437,782,2033]" box="[199,832,1862,1889]" pageId="4" pageNumber="155">Genae deep blue with violaceous lustre, finely wrinkled.</paragraph>
<paragraph id="8BA2C6341138FFEEDCA8DE702E23F82E" blockId="4.[151,1437,782,2033]" box="[199,1025,1898,1925]" pageId="4" pageNumber="155">Clypeus metallic black-blue with green lustre, lateral upper areas reddish.</paragraph>
<paragraph id="8BA2C6341138FFEFDCA8DE942E7BFF53" blockId="4.[151,1437,782,2033]" lastBlockId="5.[151,1436,150,249]" lastPageId="5" lastPageNumber="156" pageId="4" pageNumber="155">
Labrum ivory to ochre-testaceous with metallic black-blue basolateral areas of central convexity (more expanded in female); male labrum (
<figureCitation id="1326DAB11138FFEEDE7ADEA82845F867" box="[533,615,1970,1997]" captionStart="FIGURES 10–18" captionStartId="6.[151,264,1867,1891]" captionTargetBox="[159,1430,191,1829]" captionTargetId="figure-31@6.[151,1435,181,1844]" captionTargetPageId="6" captionText="FIGURES 10–18. Physodeutera (Toxoma) conturbata Moravec, from type locality, all in SDEI. 10–12—head: 10—♂, HT; 11–12—♀, PT; 13–15—labrum: 13—♂, HT; 14–15—♀, PT; 16–18—pronotum: 16—♂, HT; 17–18—♀, PT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627217" httpUri="https://zenodo.org/record/5627217/files/figure.png" pageId="4" pageNumber="155">Fig. 13</figureCitation>
) rather short, length
<quantity id="4CE56BD11138FFEEDF36DEA8299DF867" box="[857,959,1970,1997]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" pageId="4" pageNumber="155" unit="mm" value="0.95">0.95 mm</quantity>
width
<quantity id="4CE56BD11138FFEED87FDEA92E55F867" box="[1040,1143,1971,1997]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="155" unit="mm" value="1.2">1.20 mm</quantity>
, lateral margins conically attenuated towards indistinct lateral indentations, and prominent, acute anterolateral teeth; tridentate anteromedian lobe short and subtruncate with right-angled teeth and bluntly indicated median tooth; female labrum (
<figureCitation id="1326DAB11139FFEFD97BD98D2FB6FF1B" box="[1300,1428,150,177]" captionStart="FIGURES 10–18" captionStartId="6.[151,264,1867,1891]" captionTargetBox="[159,1430,191,1829]" captionTargetId="figure-31@6.[151,1435,181,1844]" captionTargetPageId="6" captionText="FIGURES 10–18. Physodeutera (Toxoma) conturbata Moravec, from type locality, all in SDEI. 10–12—head: 10—♂, HT; 11–12—♀, PT; 13–15—labrum: 13—♂, HT; 14–15—♀, PT; 16–18—pronotum: 16—♂, HT; 17–18—♀, PT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627217" httpUri="https://zenodo.org/record/5627217/files/figure.png" pageId="5" pageNumber="156">Figs 14–15</figureCitation>
) almost as long as wide, length
<quantity id="4CE56BD11139FFEFDD98D9A028BFFF7F" box="[503,669,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="1.275" metricValueMax="1.35" metricValueMin="1.2" pageId="5" pageNumber="156" unit="mm" value="1.275" valueMax="1.35" valueMin="1.2">1.20–1.35 mm</quantity>
, width
<quantity id="4CE56BD11139FFEFDE9ED9A029BAFF7F" box="[753,920,186,213]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.45" metricValueMin="1.25" pageId="5" pageNumber="156" unit="mm" value="1.35" valueMax="1.45" valueMin="1.25">1.25–1.45 mm</quantity>
, anteromedian lobe tridentate with prominent acute or subacute teeth and longer, protruding, almost cylindrical, blunt median tooth.
</paragraph>
<caption id="DF6296BC1139FFEFDCF8DE8B2FA1F85B" ID-DOI="http://doi.org/10.5281/zenodo.5627215" ID-Zenodo-Dep="5627215" httpUri="https://zenodo.org/record/5627215/files/figure.png" pageId="5" pageNumber="156" startId="5.[151,264,1937,1961]" targetBox="[177,1410,288,1906]" targetPageId="5">
<paragraph id="8BA2C6341139FFEFDCF8DE8B2FA1F85B" blockId="5.[151,1437,1937,2033]" pageId="5" pageNumber="156">
<emphasis id="B9691A261139FFEFDCF8DE8B2B19F803" bold="true" box="[151,315,1937,1961]" pageId="5" pageNumber="156">FIGURES 1–9.</emphasis>
<taxonomicName id="4C1DBDB71139FFEFDD2CDE8B2933F803" authority="Moravec" authorityName="Moravec" box="[323,785,1937,1961]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="conturbata" subGenus="Toxoma">
<emphasis id="B9691A261139FFEFDD2CDE8B2BECF803" box="[323,462,1937,1961]" italics="true" pageId="5" pageNumber="156">Physodeutera</emphasis>
(
<emphasis id="B9691A261139FFEFDDB1DE88280FF803" box="[478,557,1938,1961]" italics="true" pageId="5" pageNumber="156">Toxoma</emphasis>
)
<emphasis id="B9691A261139FFEFDE52DE8B288EF803" box="[573,684,1937,1961]" italics="true" pageId="5" pageNumber="156">conturbata</emphasis>
Moravec
</taxonomicName>
, from type locality, all in SDEI. 1—habitus ♂, 9.2 mm, HT; 2– 3—Antennomeres 1–5: 2—♂, HT; 3—♀, PT; 4–5—♀, maxillary palpus: 4—dorsal view; 5—ventral view; 6–7—labial palpus: 6—female, PT; 7—male, HT (adopted from
<bibRefCitation id="EF8CBBC51139FFEFDE35DEC028DFF85B" author="Moravec, J." box="[602,765,2009,2033]" pageId="5" pageNumber="156" refId="ref15451" refString="Moravec, J. (2002 a) Tiger beetles of Madagascar 2. A monograph of the genus Physodeutera (Coleoptera: Cicindelidae). Nakladatelstvi Kabourek, Zlin, Czech Republic, 290 pp." type="book" year="2002">Moravec 2002a</bibRefCitation>
); 8–9— male aedeagus showing internal sac, HT. Bars = 1 mm.
</paragraph>
</caption>
<caption id="DF6296BC113AFFECDCF8DE512F5EF82D" ID-DOI="http://doi.org/10.5281/zenodo.5627217" ID-Zenodo-Dep="5627217" httpUri="https://zenodo.org/record/5627217/files/figure.png" pageId="6" pageNumber="157" startId="6.[151,264,1867,1891]" targetBox="[159,1430,191,1829]" targetPageId="6">
<paragraph id="8BA2C634113AFFECDCF8DE512F5EF82D" blockId="6.[151,1436,1867,1928]" pageId="6" pageNumber="157">
<emphasis id="B9691A26113AFFECDCF8DE512B74F8CE" bold="true" box="[151,342,1867,1892]" pageId="6" pageNumber="157">FIGURES 10–18.</emphasis>
<taxonomicName id="4C1DBDB7113AFFECDD30DE562912F8C9" authority="Moravec" authorityName="Moravec" box="[351,816,1868,1892]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="157" phylum="Arthropoda" rank="species" species="conturbata" subGenus="Toxoma">
<emphasis id="B9691A26113AFFECDD30DE562BC8F8CE" box="[351,490,1868,1892]" italics="true" pageId="6" pageNumber="157">Physodeutera</emphasis>
(
<emphasis id="B9691A26113AFFECDD94DE562868F8C9" box="[507,586,1868,1891]" italics="true" pageId="6" pageNumber="157">Toxoma</emphasis>
)
<emphasis id="B9691A26113AFFECDE34DE5628E8F8CE" box="[603,714,1868,1892]" italics="true" pageId="6" pageNumber="157">conturbata</emphasis>
Moravec
</taxonomicName>
, from type locality, all in SDEI. 10–12—head: 10—♂, HT; 11–12—♀, PT; 13–15—labrum: 13—♂, HT; 14–15—♀, PT; 16–18—pronotum: 16—♂, HT; 17–18—♀, PT. Bars = 1 mm.
</paragraph>
</caption>
<caption id="DF6296BC113BFFEDDCF8DF542ACFF907" ID-DOI="http://doi.org/10.5281/zenodo.5627219" ID-Zenodo-Dep="5627219" httpUri="https://zenodo.org/record/5627219/files/figure.png" pageId="7" pageNumber="158" startId="7.[151,264,1614,1638]" targetBox="[166,1425,195,1578]" targetPageId="7">
<paragraph id="8BA2C634113BFFEDDCF8DF542ACFF907" blockId="7.[151,1436,1614,1710]" pageId="7" pageNumber="158">
<emphasis id="B9691A26113BFFEDDCF8DF542B77F9CC" bold="true" box="[151,341,1614,1638]" pageId="7" pageNumber="158">FIGURES 19–23.</emphasis>
<taxonomicName id="4C1DBDB7113BFFEDDD34DF542905F9CF" authority="Moravec" authorityName="Moravec" box="[347,807,1614,1638]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="158" phylum="Arthropoda" rank="species" species="conturbata" subGenus="Toxoma">
<emphasis id="B9691A26113BFFEDDD34DF542BC4F9CC" box="[347,486,1614,1638]" italics="true" pageId="7" pageNumber="158">Physodeutera</emphasis>
(
<emphasis id="B9691A26113BFFEDDD99DF552867F9CC" box="[502,581,1615,1638]" italics="true" pageId="7" pageNumber="158">Toxoma</emphasis>
)
<emphasis id="B9691A26113BFFEDDE3BDF5428E1F9CC" box="[596,707,1614,1638]" italics="true" pageId="7" pageNumber="158">conturbata</emphasis>
Moravec
</taxonomicName>
, from type locality, all in SDEI. 19–22—elytron: 19—♂, HT; 20—ditto, detail of elytral disc; 21—♀, PT; 22— ditto, detail of posterior area of elytral disc; 23—labels of the holotype. Bars = 1 mm.
</paragraph>
</caption>
<paragraph id="8BA2C634113BFFEDDCA8DFC629F6F895" blockId="7.[151,1437,1756,2035]" pageId="7" pageNumber="158">
Mandibles (
<figureCitation id="1326DAB1113BFFEDDD24DFC62BB9F95C" box="[331,411,1756,1782]" captionStart="FIGURES 10–18" captionStartId="6.[151,264,1867,1891]" captionTargetBox="[159,1430,191,1829]" captionTargetId="figure-31@6.[151,1435,181,1844]" captionTargetPageId="6" captionText="FIGURES 10–18. Physodeutera (Toxoma) conturbata Moravec, from type locality, all in SDEI. 10–12—head: 10—♂, HT; 11–12—♀, PT; 13–15—labrum: 13—♂, HT; 14–15—♀, PT; 16–18—pronotum: 16—♂, HT; 17–18—♀, PT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627217" httpUri="https://zenodo.org/record/5627217/files/figure.png" pageId="7" pageNumber="158">Fig. 10</figureCitation>
) subsymmetrical, brown with testaceous basolateral areas, teeth mahogany-brown with dark margins, shaped as in other species treated below (the exact shape of opened mandibles is not illustrated here as the mandibles in the
<typeStatus id="54A67896113BFFEDDD3CDE3F2BA0F895" box="[339,386,1829,1855]" pageId="7" pageNumber="158">type</typeStatus>
specimens are firmly closed, resistant to rehydration).
</paragraph>
<paragraph id="8BA2C634113BFFEDDCA8DE522F54F801" blockId="7.[151,1437,1756,2035]" pageId="7" pageNumber="158">
Palpi. Maxillary palpi (
<figureCitation id="1326DAB1113BFFEDDDA7DE522809F8C9" box="[456,555,1864,1891]" captionStart="FIGURES 1–9" captionStartId="5.[151,264,1937,1961]" captionTargetBox="[177,1410,288,1906]" captionTargetId="figure-61@5.[160,1427,273,1914]" captionTargetPageId="5" captionText="FIGURES 1–9. Physodeutera (Toxoma) conturbata Moravec, from type locality, all in SDEI. 1—habitus ♂, 9.2 mm, HT; 2– 3—Antennomeres 1–5: 2—♂, HT; 3—♀, PT; 4–5—♀, maxillary palpus: 4—dorsal view; 5—ventral view; 6–7—labial palpus: 6—female, PT; 7—male, HT (adopted from Moravec 2002a); 8–9— male aedeagus showing internal sac, HT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627215" httpUri="https://zenodo.org/record/5627215/files/figure.png" pageId="7" pageNumber="158">Figs 4–5</figureCitation>
) ivory-testaceous with terminal palpomere brownish in male, with last two palpomeres brownish in female; labial palpi in male with brown terminal palpomere, their penultimate (longest) palpomere moderately dilated with subparallel lateral margins, ochre-testaceous in male (
<figureCitation id="1326DAB1113BFFEDDF82DE8A2E0DF801" box="[1005,1071,1936,1963]" captionStart="FIGURES 1–9" captionStartId="5.[151,264,1937,1961]" captionTargetBox="[177,1410,288,1906]" captionTargetId="figure-61@5.[160,1427,273,1914]" captionTargetPageId="5" captionText="FIGURES 1–9. Physodeutera (Toxoma) conturbata Moravec, from type locality, all in SDEI. 1—habitus ♂, 9.2 mm, HT; 2– 3—Antennomeres 1–5: 2—♂, HT; 3—♀, PT; 4–5—♀, maxillary palpus: 4—dorsal view; 5—ventral view; 6–7—labial palpus: 6—female, PT; 7—male, HT (adopted from Moravec 2002a); 8–9— male aedeagus showing internal sac, HT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627215" httpUri="https://zenodo.org/record/5627215/files/figure.png" pageId="7" pageNumber="158">Fig. 7</figureCitation>
), brownish in female (
<figureCitation id="1326DAB1113BFFEDD949DE8A2F4BF801" box="[1318,1385,1936,1963]" captionStart="FIGURES 1–9" captionStartId="5.[151,264,1937,1961]" captionTargetBox="[177,1410,288,1906]" captionTargetId="figure-61@5.[160,1427,273,1914]" captionTargetPageId="5" captionText="FIGURES 1–9. Physodeutera (Toxoma) conturbata Moravec, from type locality, all in SDEI. 1—habitus ♂, 9.2 mm, HT; 2– 3—Antennomeres 1–5: 2—♂, HT; 3—♀, PT; 4–5—♀, maxillary palpus: 4—dorsal view; 5—ventral view; 6–7—labial palpus: 6—female, PT; 7—male, HT (adopted from Moravec 2002a); 8–9— male aedeagus showing internal sac, HT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627215" httpUri="https://zenodo.org/record/5627215/files/figure.png" pageId="7" pageNumber="158">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="8BA2C634113BFFE2DCA8DEAE2910FF7F" blockId="7.[151,1437,1756,2035]" lastBlockId="8.[151,1437,151,2013]" lastPageId="8" lastPageNumber="159" pageId="7" pageNumber="158">
Antennae (
<figureCitation id="1326DAB1113BFFEDDD2CDEAE2BE5F864" box="[323,455,1972,1998]" captionStart="FIGURES 1–9" captionStartId="5.[151,264,1937,1961]" captionTargetBox="[177,1410,288,1906]" captionTargetId="figure-61@5.[160,1427,273,1914]" captionTargetPageId="5" captionText="FIGURES 1–9. Physodeutera (Toxoma) conturbata Moravec, from type locality, all in SDEI. 1—habitus ♂, 9.2 mm, HT; 2– 3—Antennomeres 1–5: 2—♂, HT; 3—♀, PT; 4–5—♀, maxillary palpus: 4—dorsal view; 5—ventral view; 6–7—labial palpus: 6—female, PT; 7—male, HT (adopted from Moravec 2002a); 8–9— male aedeagus showing internal sac, HT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627215" httpUri="https://zenodo.org/record/5627215/files/figure.png" pageId="7" pageNumber="158">Figs 1, 2–3</figureCitation>
) slightly surpassing elytral half; scape whitish-testaceous in male (
<figureCitation id="1326DAB1113BFFEDD8A9DEAE2F2EF864" box="[1222,1292,1972,1998]" captionStart="FIGURES 1–9" captionStartId="5.[151,264,1937,1961]" captionTargetBox="[177,1410,288,1906]" captionTargetId="figure-61@5.[160,1427,273,1914]" captionTargetPageId="5" captionText="FIGURES 1–9. Physodeutera (Toxoma) conturbata Moravec, from type locality, all in SDEI. 1—habitus ♂, 9.2 mm, HT; 2– 3—Antennomeres 1–5: 2—♂, HT; 3—♀, PT; 4–5—♀, maxillary palpus: 4—dorsal view; 5—ventral view; 6–7—labial palpus: 6—female, PT; 7—male, HT (adopted from Moravec 2002a); 8–9— male aedeagus showing internal sac, HT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627215" httpUri="https://zenodo.org/record/5627215/files/figure.png" pageId="7" pageNumber="158">Fig. 2</figureCitation>
), while dark mahogany-brown in female (
<figureCitation id="1326DAB1113BFFEDDDB1DEC22800F858" box="[478,546,2008,2034]" captionStart="FIGURES 1–9" captionStartId="5.[151,264,1937,1961]" captionTargetBox="[177,1410,288,1906]" captionTargetId="figure-61@5.[160,1427,273,1914]" captionTargetPageId="5" captionText="FIGURES 1–9. Physodeutera (Toxoma) conturbata Moravec, from type locality, all in SDEI. 1—habitus ♂, 9.2 mm, HT; 2– 3—Antennomeres 1–5: 2—♂, HT; 3—♀, PT; 4–5—♀, maxillary palpus: 4—dorsal view; 5—ventral view; 6–7—labial palpus: 6—female, PT; 7—male, HT (adopted from Moravec 2002a); 8–9— male aedeagus showing internal sac, HT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627215" httpUri="https://zenodo.org/record/5627215/files/figure.png" pageId="7" pageNumber="158">Fig. 3</figureCitation>
), antennomeres 2–4 metallic black-brown in both sexes, antennomere 5 ochretestaceous or brownish-testaceous, antennomere 6 variably testaceous or brownish, 7–10 brownish, gradually darkened towards terminal antennomere that is smoky-blackish.
</paragraph>
<paragraph id="8BA2C6341134FFE2DCA8D9C52F10FDCB" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="159">
Thorax. Pronotum (
<figureCitation id="1326DAB11134FFE2DDCCD9C52806FF50" box="[419,548,223,250]" captionStart="FIGURES 10–18" captionStartId="6.[151,264,1867,1891]" captionTargetBox="[159,1430,191,1829]" captionTargetId="figure-31@6.[151,1435,181,1844]" captionTargetPageId="6" captionText="FIGURES 10–18. Physodeutera (Toxoma) conturbata Moravec, from type locality, all in SDEI. 10–12—head: 10—♂, HT; 11–12—♀, PT; 13–15—labrum: 13—♂, HT; 14–15—♀, PT; 16–18—pronotum: 16—♂, HT; 17–18—♀, PT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627217" httpUri="https://zenodo.org/record/5627217/files/figure.png" pageId="8" pageNumber="159">Figs 16–18</figureCitation>
), elongate, length
<quantity id="4CE56BD11134FFE2DE9ED9C429B4FF53" box="[753,918,222,249]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" metricValueMax="2.25" metricValueMin="1.95" pageId="8" pageNumber="159" unit="mm" value="2.1" valueMax="2.25" valueMin="1.95">1.95–2.25 mm</quantity>
, width
<quantity id="4CE56BD11134FFE2DF87D9C42EAEFF53" box="[1000,1164,222,249]" metricMagnitude="-3" metricUnit="m" metricValue="1.675" metricValueMax="1.8" metricValueMin="1.55" pageId="8" pageNumber="159" unit="mm" value="1.675" valueMax="1.8" valueMin="1.55">1.55–1.80 mm</quantity>
, dorsally deep cyaneous with violaceous lustre within sulci and on lateral areas of disc and faint bronze lustre in middle in male, greenish lustre in female; anterior and posterior sulci well pronounced, anterior lobe narrower than posterior lobe and disc; lateral margins of disc distinctly attenuated anteriad, except for subparallel margins in one of the females (
<figureCitation id="1326DAB11134FFE2D954D8512FACFECF" box="[1339,1422,331,357]" captionStart="FIGURES 10–18" captionStartId="6.[151,264,1867,1891]" captionTargetBox="[159,1430,191,1829]" captionTargetId="figure-31@6.[151,1435,181,1844]" captionTargetPageId="6" captionText="FIGURES 10–18. Physodeutera (Toxoma) conturbata Moravec, from type locality, all in SDEI. 10–12—head: 10—♂, HT; 11–12—♀, PT; 13–15—labrum: 13—♂, HT; 14–15—♀, PT; 16–18—pronotum: 16—♂, HT; 17–18—♀, PT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627217" httpUri="https://zenodo.org/record/5627217/files/figure.png" pageId="8" pageNumber="159">Fig. 18</figureCitation>
); notopleural sutures almost invisible from above; median line narrow, partly merging with surface sculpture anteriorly; surface of disc distinctly and coarsely transversely rugose on entire discal surface, rugae rather sharp, wavy and occasionally anastomosing, passing to transverse-parallel rugae on lateral areas; posterior lobe bordered with sharp basal rim, dorsolateral bulges rather distinct, pulvinate, fluently passing to irregularly and shallowly rugulose median area; prosternum and mesosternum deep blue, finely wrinkled; proepisterna shiny black-blue with violaceous lustre, nearly smooth; mesepisterna black blue with purple lustre, mesepisternal coupling sulci variable (also between left and right mesepisternum), entirely absent, or present in form of indistinct, rarely distinct pit.
</paragraph>
<paragraph id="8BA2C6341134FFE2DCA8DB712E2CFC47" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="159">
Elytra (
<figureCitation id="1326DAB11134FFE2DD72DB712B83FD2F" box="[285,417,619,645]" captionStart="FIGURES 19–23" captionStartId="7.[151,264,1614,1638]" captionTargetBox="[166,1425,195,1578]" captionTargetId="figure-17@7.[151,1435,181,1589]" captionTargetPageId="7" captionText="FIGURES 19–23. Physodeutera (Toxoma) conturbata Moravec, from type locality, all in SDEI. 19–22—elytron: 19—♂, HT; 20—ditto, detail of elytral disc; 21—♀, PT; 22— ditto, detail of posterior area of elytral disc; 23—labels of the holotype. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627219" httpUri="https://zenodo.org/record/5627219/files/figure.png" pageId="8" pageNumber="159">Figs 19–22</figureCitation>
), elongate,
<quantity id="4CE56BD11134FFE2DE48DB7028EEFD2F" box="[551,716,618,645]" metricMagnitude="-3" metricUnit="m" metricValue="5.875" metricValueMax="6.2" metricValueMin="5.55" pageId="8" pageNumber="159" unit="mm" value="5.875" valueMax="6.2" valueMin="5.55">5.55–6.20 mm</quantity>
long; lateral margins subparallel, only slightly dilated towards arcuate anteapical angles, apices almost subacute in male, rounded in female, indistinctly emarginate towards indistinct sutural spine; juxtaepipleural area with sparse white microsetae; humeral impressions very short, basodiscal convexity and discal impression rather distinct, apical impressions moderate; surface punctate throughout (except for smooth basohumeral limited area), punctures much coarser on anterior elytral half and lateral area, occasionally anastomosing in chains, their elevated anterior margins optically forming subreticulate ornament (in different light angle the intervals appear as if forming crests giving rasp-like appearance of the surface sculpture); posterior half of elytral disc with much smaller and more spaced punctures, particularly within indistinctly shadowed area; elytral coloration deep cyaneous with brighter greenish lustre within discal impression and deep violaceous on lateral and apical areas, and with indistinct, feebly diffusing shade on elytral disc; whitish maculation consisting of distinct humeral macula in male, which is brownish-darkened, barely visible in female.
</paragraph>
<paragraph id="8BA2C6341134FFE2DCA8DAED2906FB9F" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="159">Abdomen. Abdominal ventrites black-blue with violaceous reflections, smooth and glabrous (except for usual sublateral sensory seta (on either side) at ventrite margins.</paragraph>
<paragraph id="8BA2C6341134FFE2DCA8DD25286BFB43" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="159">Legs. Male procoxae and mesocoxae pale testaceous, metacoxae and all female coxae metallic blue with testaceous apices, trochanters in both sexes testaceous, male femora distinctly bicoloured: profemora dorsally brown, while their whole ventral and apical area ochre-yellow; mesofemora brown, their ochre ventral area less expanded; metafemora almost entirely brown; tibiae and tarsi testaceous to brownish, protarsi darkened; female legs much darker, entirely brown to black-brown.</paragraph>
<paragraph id="8BA2C6341134FFE2DCA8DDE92850FA4F" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="159">
Aedeagus (
<figureCitation id="1326DAB11134FFE2DD28DDE92B8DFAA7" box="[327,431,1267,1293]" captionStart="FIGURES 1–9" captionStartId="5.[151,264,1937,1961]" captionTargetBox="[177,1410,288,1906]" captionTargetId="figure-61@5.[160,1427,273,1914]" captionTargetPageId="5" captionText="FIGURES 1–9. Physodeutera (Toxoma) conturbata Moravec, from type locality, all in SDEI. 1—habitus ♂, 9.2 mm, HT; 2– 3—Antennomeres 1–5: 2—♂, HT; 3—♀, PT; 4–5—♀, maxillary palpus: 4—dorsal view; 5—ventral view; 6–7—labial palpus: 6—female, PT; 7—male, HT (adopted from Moravec 2002a); 8–9— male aedeagus showing internal sac, HT. Bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5627215" httpUri="https://zenodo.org/record/5627215/files/figure.png" pageId="8" pageNumber="159">Figs 8–9</figureCitation>
), stout, length
<quantity id="4CE56BD11134FFE2DE32DDE828E7FAA7" box="[605,709,1266,1293]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" pageId="8" pageNumber="159" unit="mm" value="2.25">2.25 mm</quantity>
, width
<quantity id="4CE56BD11134FFE2DF73DDE829A7FAA7" box="[796,901,1266,1293]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="8" pageNumber="159" unit="mm" value="0.45">0.45 mm</quantity>
, dorsally dilated, apical portion conically constricted, while ventral outline almost straight, only slightly ventrad-bent apically and fluently arcuately passing into short, regularly rounded apex; structure of internal sac consisting of distinctly arcuate-bent arciform piece accompanied with shorter, apically bilobed satellite piece, basal plates, and large, longitudinal, apically clavate-rounded, membranous piece. Only cleared aedeagus was illustrated by the first author (
<bibRefCitation id="EF8CBBC51134FFE2D867DC992E9EFA37" author="Moravec, J." box="[1032,1212,1411,1437]" pageId="8" pageNumber="159" refId="ref15451" refString="Moravec, J. (2002 a) Tiger beetles of Madagascar 2. A monograph of the genus Physodeutera (Coleoptera: Cicindelidae). Nakladatelstvi Kabourek, Zlin, Czech Republic, 290 pp." type="book" year="2002">Moravec 2002a</bibRefCitation>
, fig. 316) and since the same aedeagus is photographed here, it is impossible to compare the shape of untreated aedeagus to that of
<taxonomicName id="4C1DBDB71134FFE2D919DCBD2B25FA4F" baseAuthorityName="Maran" baseAuthorityYear="1942" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="164" phylum="Arthropoda" rank="species" species="dubia" subGenus="Toxoma">
<emphasis id="B9691A261134FFE2D919DCBD2FB7FA6B" box="[1398,1429,1447,1473]" italics="true" pageId="8" pageNumber="159">Ph</emphasis>
. (
<emphasis id="B9691A261134FFE2DCCEDCD12A93FA4E" box="[161,177,1483,1508]" italics="true" pageId="8" pageNumber="159">T</emphasis>
.)
<emphasis id="B9691A261134FFE2DCA8DCD12B25FA4F" box="[199,263,1483,1509]" italics="true" pageId="8" pageNumber="159">dubia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1DBDB71134FFE2DD51DCD12833FA4F" authority="Moravec, 2021" authorityName="Moravec" authorityYear="2021" box="[318,529,1483,1509]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="164" phylum="Arthropoda" rank="species" species="lokobensis" status="sp. nov." subGenus="Toxoma">
<emphasis id="B9691A261134FFE2DD51DCD12B7FFA4F" box="[318,349,1483,1509]" italics="true" pageId="8" pageNumber="159">Ph</emphasis>
. (
<emphasis id="B9691A261134FFE2DD1BDCD12BA6FA4E" box="[372,388,1483,1508]" italics="true" pageId="8" pageNumber="159">T</emphasis>
.)
<emphasis id="B9691A261134FFE2DDF4DCD12833FA4F" box="[411,529,1483,1509]" italics="true" pageId="8" pageNumber="159">lokobensis</emphasis>
</taxonomicName>
<emphasis id="B9691A261134FFE2DE77DCD12850FA4F" bold="true" box="[536,626,1483,1509]" pageId="8" pageNumber="159">
<taxonomicNameLabel id="A25AA75D1134FFE2DE77DCD12850FA4F" box="[536,626,1483,1509]" pageId="8" pageNumber="159" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C30795BF1134FFE2DCA8DCF429E7F94B" pageId="8" pageNumber="159" type="distribution">
<paragraph id="8BA2C6341134FFE2DCA8DCF429E7F94B" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="159">
<emphasis id="B9691A261134FFE2DCA8DCF42B7BF9A2" bold="true" box="[199,345,1518,1544]" pageId="8" pageNumber="159">Distribution</emphasis>
. (
<figureCitation id="1326DAB11134FFE2DD1FDCF52BF6F9A0" box="[368,468,1519,1546]" captionStart="FIGURE 106" captionStartId="27.[151,250,1561,1585]" captionTargetBox="[271,1319,572,1531]" captionTargetId="figure-167@27.[262,1326,561,1538]" captionTargetPageId="27" captionText="FIGURE 106. Distribution of four Physodeutera (Toxoma) species in northern part of Madagascar. Adopted from the map of preserved areas in Madagascar according to Moat &amp; Smith (2007), adapted and published previously (Moravec 2010) under permission of the trustee of the Royal Botanic Gardens, Kew, GB." figureDoi="http://doi.org/10.5281/zenodo.5627247" httpUri="https://zenodo.org/record/5627247/files/figure.png" pageId="8" pageNumber="159">Fig. 106</figureCitation>
). Only the
<specimenCount id="9D1B0DBD1134FFE2DE3EDCF5289AF9A3" box="[593,696,1519,1545]" pageId="8" pageNumber="159" type="generic" typeStatus="type">four type</specimenCount>
specimens from the
<typeStatus id="54A678961134FFE2DFCBDCF529F6F9A3" box="[932,980,1519,1545]" pageId="8" pageNumber="159">type</typeStatus>
locality of
<taxonomicName id="4C1DBDB71134FFE2D838DCF52F40F9A3" authority="" authorityName="Rivalier" authorityYear="1967" box="[1111,1378,1519,1545]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="159" phylum="Arthropoda" rank="subGenus" subGenus="Toxoma">
<emphasis id="B9691A261134FFE2D838DCF52ED3F9A3" box="[1111,1265,1519,1545]" italics="true" pageId="8" pageNumber="159">Physodeutera</emphasis>
(
<emphasis id="B9691A261134FFE2D96DDCF52F78F9A2" box="[1282,1370,1519,1544]" italics="true" pageId="8" pageNumber="159">Toxoma</emphasis>
)
</taxonomicName>
<emphasis id="B9691A261134FFE2D905DCF52ACBF987" italics="true" pageId="8" pageNumber="159">conturbata</emphasis>
are known. Antseva (Antseva-Ambobako) lies
<quantity id="4CE56BD11134FFE2DE99DF082944F987" box="[758,870,1554,1581]" metricMagnitude="4" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="8" pageNumber="159" unit="km" value="17.5" valueMax="20.0" valueMin="15.0">15–20 km</quantity>
southeast of the town of Ambanja at the base of the Tsaratanana Massif and the Ramena River, geographically Sambirano (see
<bibRefCitation id="EF8CBBC51134FFE2DFB5DF2D2E42F9FB" author="Viette, P." box="[986,1120,1591,1617]" pageId="8" pageNumber="159" refId="ref17111" refString="Viette, P. (1991) Principales localites ou des Insectes ont ete recueillis a Madagascar. Chief field stations where Insects were collected in Madagascar. Faune de Madagascar. Supplement 2. Privately published, Paris, 88 pp." type="book" year="1991">Viette 1991</bibRefCitation>
). The locality name Antseva must not be confused with homonymous places in other areas of
<collectingCountry id="F30A86A41134FFE2DF10DF412E24F9DF" box="[895,1030,1627,1653]" name="Madagascar" pageId="8" pageNumber="159">Madagascar</collectingCountry>
named as such not only recently (a case of homonymous names and recent changes in names in a number of places throughout
<collectingCountry id="F30A86A41134FFE2D8FCDF652F02F930" box="[1171,1312,1663,1690]" name="Madagascar" pageId="8" pageNumber="159">Madagascar</collectingCountry>
). The same is true of the name of the Sambiranense Ramena River, which must not be confused with the town of Ramena in the northernmost promontory of
<collectingCountry id="F30A86A41134FFE2DDB2DFDD2844F94B" box="[477,614,1735,1761]" name="Madagascar" pageId="8" pageNumber="159">Madagascar</collectingCountry>
. Behaviour of adults unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30795BF1134FFE3DCA8DFF12E7EFF7F" lastPageId="9" lastPageNumber="160" pageId="8" pageNumber="159" type="discussion">
<paragraph id="8BA2C6341134FFE2DCA8DFF1283FF877" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="159">
<emphasis id="B9691A261134FFE2DCA8DFF12B16F8AF" bold="true" box="[199,308,1771,1797]" pageId="8" pageNumber="159">Remarks</emphasis>
. The
<specimenCount id="9D1B0DBD1134FFE2DD17DFF12BC2F8AF" box="[376,480,1771,1797]" pageId="8" pageNumber="159" type="generic">four type</specimenCount>
specimens examined were deposited in a series standing under the name
<taxonomicName id="4C1DBDB71134FFE2D940DFF12AD6F883" authorityName="W. Horn" authorityYear="1892" class="Insecta" family="Carabidae" genus="Prothyma" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="schaumi">
<emphasis id="B9691A261134FFE2D940DFF12AD6F883" italics="true" pageId="8" pageNumber="159">Prothyma schaumi</emphasis>
</taxonomicName>
in the collection of Walther Horn (SDEI). Because of the complex of characters, particularly the shape of its penultimate palpomeres of the labial palpi and the structure of the internal sac of the aedeagus, characteristic of the subgenus
<taxonomicName id="4C1DBDB71134FFE2DD40DE4D2BA5F8DA" authorityName="Rivalier" authorityYear="1967" box="[303,391,1879,1904]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="159" phylum="Arthropoda" rank="subGenus" subGenus="Toxoma">
<emphasis id="B9691A261134FFE2DD40DE4D2BA5F8DA" box="[303,391,1879,1904]" italics="true" pageId="8" pageNumber="159">Toxoma</emphasis>
</taxonomicName>
, this species was described by the first author (
<bibRefCitation id="EF8CBBC51134FFE2DFF9DE4D2E6AF8DB" author="Moravec, J." box="[918,1096,1879,1905]" pageId="8" pageNumber="159" refId="ref15451" refString="Moravec, J. (2002 a) Tiger beetles of Madagascar 2. A monograph of the genus Physodeutera (Coleoptera: Cicindelidae). Nakladatelstvi Kabourek, Zlin, Czech Republic, 290 pp." type="book" year="2002">Moravec 2002a</bibRefCitation>
) in the subgenus
<taxonomicName id="4C1DBDB71134FFE2D963DE4D2F46F8DA" authorityName="Rivalier" authorityYear="1967" box="[1292,1380,1879,1904]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="159" phylum="Arthropoda" rank="subGenus" subGenus="Toxoma">
<emphasis id="B9691A261134FFE2D963DE4D2F46F8DA" box="[1292,1380,1879,1904]" italics="true" pageId="8" pageNumber="159">Toxoma</emphasis>
</taxonomicName>
. The characters clearly differentiate
<emphasis id="B9691A261134FFE2DD80DE61282CF83F" box="[495,526,1915,1941]" italics="true" pageId="8" pageNumber="159">Ph</emphasis>
.
<taxonomicName id="4C1DBDB71134FFE2DE74DE6128E7F83F" box="[539,709,1915,1941]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="conturbata" subGenus="Toxoma">
(
<emphasis id="B9691A261134FFE2DE4BDE612816F83E" box="[548,564,1915,1940]" italics="true" pageId="8" pageNumber="159">T</emphasis>
.)
<emphasis id="B9691A261134FFE2DE25DE6128E7F83F" box="[586,709,1915,1941]" italics="true" pageId="8" pageNumber="159">conturbata</emphasis>
</taxonomicName>
from the
<typeStatus id="54A678961134FFE2DF40DE6129B3F83F" box="[815,913,1915,1941]" pageId="8" pageNumber="159" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C1DBDB71134FFE2DFDCDE612F6CF83F" authority="W. Horn, 1872" authorityName="W. Horn" authorityYear="1872" box="[947,1358,1915,1941]" class="Insecta" family="Carabidae" genus="Megalomma" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="schaumi">
<emphasis id="B9691A261134FFE2DFDCDE612E80F83F" box="[947,1186,1915,1941]" italics="true" pageId="8" pageNumber="159">Megalomma schaumi</emphasis>
W. Horn, 1872
</taxonomicName>
, which is a junior synonym of
<taxonomicName id="4C1DBDB71134FFE2DDFADE8529DEF813" authority="(Audouin &amp; Brulle, 1839)" baseAuthorityName="Audouin &amp; Brulle" baseAuthorityYear="1839" box="[405,1020,1951,1977]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="159" phylum="Arthropoda" rank="species" species="viridicyanea" subGenus="Diarrhiza">
<emphasis id="B9691A261134FFE2DDFADE852B96F813" box="[405,436,1951,1977]" italics="true" pageId="8" pageNumber="159">Ph</emphasis>
. (
<emphasis id="B9691A261134FFE2DDA5DE85281AF813" box="[458,568,1951,1977]" italics="true" pageId="8" pageNumber="159">Diarrhiza</emphasis>
)
<emphasis id="B9691A261134FFE2DE28DE8528F1F813" box="[583,723,1951,1977]" italics="true" pageId="8" pageNumber="159">viridicyanea</emphasis>
(Audouin &amp; Brullé, 1839)
</taxonomicName>
, and also from all other species of the subgenus
<taxonomicName id="4C1DBDB71134FFE2DD69DED9283AF877" authority="Jeannel, 1946" authorityName="Jeannel" authorityYear="1946" box="[262,536,1987,2013]" class="Insecta" family="Carabidae" genus="Physodeutera" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="159" phylum="Arthropoda" rank="subGenus" subGenus="Diarrhiza">
<emphasis id="B9691A261134FFE2DD69DED92B56F877" box="[262,372,1987,2013]" italics="true" pageId="8" pageNumber="159">Diarrhiza</emphasis>
<bibRefCitation id="EF8CBBC51134FFE2DD14DED9283AF877" author="Jeannel, R." box="[379,536,1987,2013]" pageId="8" pageNumber="159" refId="ref15178" refString="Jeannel, R. = (1946) Coleopteres Carabiques de la region Malgache (premiere partie). Faune de l'empire francais. VI. Libraire Larosse, Paris, 372 pp." type="book" year="1946">Jeannel, 1946</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA2C6341135FFE3DCA8D98D2E7EFF7F" blockId="9.[151,1436,151,213]" pageId="9" pageNumber="160">
<materialsCitation id="3B75CC691135FFE3DCA8D98D2E7AFF7F" ID-GBIF-Occurrence="3395513310" collectionCode="SDEI" pageId="9" pageNumber="160" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
It must be noted here that there are in fact
<specimenCount id="9D1B0DBD1135FFE3DECDD98D291AFF1B" box="[674,824,151,177]" pageId="9" pageNumber="160" type="female">three females</specimenCount>
in
<collectionCode id="ED0C5EF11135FFE3DF33D98D29B8FF1B" box="[860,922,151,177]" pageId="9" pageNumber="160">SDEI</collectionCode>
originally labelled as
<typeStatus id="54A678961135FFE3D8FCD98D2EDCFF1B" box="[1171,1278,151,177]" pageId="9" pageNumber="160" type="paratype">paratypes</typeStatus>
instead of the
<specimenCount id="9D1B0DBD1135FFE3DCF8D9A12B3DFF7F" box="[151,287,187,213]" pageId="9" pageNumber="160" type="female">two females</specimenCount>
mentioned in typo error in the monograph of the genus (
<bibRefCitation id="EF8CBBC51135FFE3DFF5D9A12E72FF7F" author="Moravec, J." box="[922,1104,187,213]" pageId="9" pageNumber="160" refId="ref15451" refString="Moravec, J. (2002 a) Tiger beetles of Madagascar 2. A monograph of the genus Physodeutera (Coleoptera: Cicindelidae). Nakladatelstvi Kabourek, Zlin, Czech Republic, 290 pp." type="book" year="2002">Moravec 2002a</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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